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German Pages 105 [112] Year 1980
DAS TIERREICH Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen Herausgeber HEINZ WERMUTH
EDWIN M Ö H N
(Wirbeltiere)
(Wirbellose)
n ä v t a £>ei
Sine systemate chaos Lieferung 101
Liste der rezenten Amphibien und Reptilien
Anomalepididae, Leptotyphlopidae, Typhlopidae Bearbeitet von Donald E. Hahn (Cottonwood, Arizona)
w DE
G Walter de Gruyter • Berlin • New York 1980
Das Tierreich
Berlin
Lieferung 101
Seite I - X I I , 1 - 9 3
1980
ISBN 3 11 006860 6 © 1980 by Walter de Gruyter & Co., vormals G. J. Göschen'sc he Verlagshandlung, J . Guttentag, Verlagsbuchhandlung Georg Reimer, Karl J. Trübner, Veit & Comp., Berlin 30. Alle Rechte, insbesondere das Recht der Vervielfältigung und Verbreitung, sowie der Ubersetzung, vorbehalten. Kein Teil des Werkes darf in irgendeiner Form (durch Photokopie, Mikrofilm oder ein anderes Verfahren) ohne schriftliche Genehmigung des Verlages reproduziert oder unter Verwendung elektronischer Systeme verarbeitet, vervielfältigt oder verbreitet werden. Printed in Germany. Satz: IBM-Composer Walter de Gruyter & Co., Berlin. Druck: Karl Gerike, Berlin. Bindearbeiten: Dieter Mikolai, Berlin.
Liste der rezenten Amphibien und Reptilien
Anomalepididae, Leptotyphlopidae, Typhlopidae
Bearbeitet von
Donald E. Hahn
Die Liste der rezenten Amphibien und Reptilien ist H E R R N P R O F . D R . R O B E R T M E R T E N S T,
Direktor emeritus des Natur-Museums und Forschungs-Institutes Senckenberg zu Frankfurt am Main, als Dank der Fachkollegen für seine überragenden Verdienste um die Herpetologie gewidmet.
Als Beiträge zur Liste der rezenten Amphibien und Reptilien sind bisher erschienen: Amphibia: G y m n o p h i o n a (GORHAM, Ottawa: Lieferung 78, 1962). - S a 1 i e n t i a Ascaphidae, Leiopelmatidae, Pipidae, Discoglossidae, Pelobatidae, Leptodactylidae, Rhinophrynidae (GORHAM, St. John: Lieferung 85, 1966); Hylidae (DUELLMAN, Lawrence: Lieferung 95, 1977). Reptilia: T e s t u d i n e s , C r o c o d y l i a , R h y n c h o c e p h a l i a (WERMUTH, Ludwigsburg, & MERTENS, Frankfurt am Main: Lieferung 100, 1977). - S a u r i a : Gekkonidae, Pygopodidae, Xantusiidae (WERMUTH, Ludwigsburg: Lieferung 80, 1965); Agamidae (WERMUTH, Ludwigsburg: Lieferung 86, 1967); Chamaeleonidae (MERTENS, Frankfurt am Main: Lieferung 83, 1966); Cordylidae - Cordylinae, Chaniaesaurinae - (WERMUTH, Ludwigsburg: Lieferung 87, 1968); Anguidae, Anniellidae, Xenosauridae (WERMUTH, Ludwigsburg: Lieferung 90, 1969); Helodermatidae, Varanidae, Lanthanotidae (MERTENS, Frankfurt am Main: Lieferung 79, 1963). - S e r p e n t e s : Anomalepididae, Leptotyphlopidae, Typhlopidae (HAHN, Cottonwood: Lieferung 101, 1980); Boidae - Boinae, Bolyeriinae, Loxoceminae, Pythoninae - (STIMSON, London: Lieferung 89, 1969); Uropeltidae (GANS, Buffalo: Lieferung 84, 1966); Colubridae - Dipsadinae - (PETERS, Washington: Lieferung 81, 1965).
Als weitere Beiträge erscheinen demnächst: Amphibia: C a u d a t a (BRAME, Pasadena, & GORHAM, St. John). - S a 1 i e n t i a : Bufonidae (COOK, Ottawa). Reptilia: S a u r i a : Iguanidae (ETHERIDGE, San Diego); Lacertidae (BÖHME, Bonn, & WERMUTH, Ludwigsburg); Teiidae (HOTZ, Zürich, & WERMUTH, Ludwigsburg); Amphisbaenidae (GANS, Ann Arbor, & WERMUTH, Ludwigsburg); Scincidae (FUHN, Bukarest). S e r p e n t e s : Aniliidae, Uropeltidae, Xenopeltidae, Acrochordidae (RIEPPEL, Basel); Elapidae (KLEMMER, Frankfurt am Main, & MARX, Chicago); Viperidae, Crotalidae (HAHN, Cottonwood/HOGE, Säo Paulo [?]).
Preface In the period of 1893—96, Boulenger published the first comprehensive checklist of the Scolecophidia. Werner updated these checklists in 1917 (Leptotyphlopidae) and 1921 (Typhlopidae and what is now Anomalepididae). Since this time no fewer than 179 new forms have been described, and revisions of the species of many areas have also appeared. These checklists have here been brought up to date and set out in the DAS TIERREICH format. Literature citation abbreviations are consistant with those used in the series. All primary synonyms and major spelling variations have been listed for each form, as has the first use of the current combination. The present location and museum number of each extant primary type specimen have been listed, where known. Under each taxon I have listed the most recent publication(s) dealing in some detail with that form. Where no publication is listed, it may be assumed that nothing significant has appeared since the original description, or Boulenger and Werner's checklists. Many taxonomic problems still exist within the Scolecophidia. The systematics of most Asian forms especially is still in a primitive state. Numerous species of these secretive burrowing snakes are known only from the holotype, and little is known of the variation that exists in most species. An accurate drawing of the head scutulation of the worm snakes is worth more than a thousand words of description, yet many species have never been figured, and many of those that have, are not accurately drawn. This is mainly due to the small size of most species, and the difficulty in distinguishing the shiny, close-fitting head scales, even under adequate magnification. Scale counts now considered standard are also lacking in most descriptions for the same reasons. I have consistently followed the zoological nomenclature committee's decision to consider all generic names ending in -ops as masculine (1972. Bull. zool. Nomencl. 29 (4): 177). Many new name combinations were formed in McDowell's (1974) important work on the systematics of the Typhlopidae of the New Guinea area. For consistency, it has been necessary to assign the remaining species occurring in the Australasian and Asian areas not mentioned in his work to the most probable generic placement — Typhlops or Typhlina. Museums housing type material are referred to by the following abbreviations. My sincere thanks goes to the listed curators for providing information on their collections. A.M.N.H. A.N.S.P. B.M.N.H. C.A.S. C.M. E.H.T.-H.M.S. F.M.N.H.
American Museum of Natural History, New York. Academy of Natural Sciences of Philadelphia, Pennsylvania British Museum (Natural History), London. (Andrew F. Stimson). California Academy of Sciences, San Francisco. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. (C. J. McCoy). Edward H. Taylor-Hobart M. Smith Collection. Field Museum Natural History, Chicago. (Hymnen Marx).
Vili F.S.M. I.R.S.N. K.U. M.B.L. M.C.Z. M.H.N.L.S. M.H.N.P. M.S.N.G. M.S.N.M. N.M.B. N.M.W. R.G.M.C. R.M.N.H. S.D.S.N.H. S.M.F. T.M. U.M. U.M.M.Z. U.S.N.M. W.A.M. Z.I.L. Z.M.B.
Florida State Museum, Gainesville. Insitut Royal des Sciences Naturelles de Belgique, Brussels. (Gaston F. de Witte). University of Kansas Museum of Natural History, Lawrence. Museu-Bocage, Lisbon. (J. de A. Fernandes). Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts. (Ernest E. Williams). Museo de Historia Natural LaSalle, Caracas. Museum National d'Histoire Naturelle, Paris. (J. Guibé and Rolande RouxEstève). Museo Civico di Storia Naturale "Giacomo Doria", Genova. (Lilia Capocaccia). Museo Civico di Storia Naturale, Milan. Naturhistorisches Museum, Basel. Naturhistorisches Museum, Vienna. (Franz Tiedemann). Musee Royal de l'Afrique Centrale, Tervuren, Belgium. (D. Thys Van Den Audenaerde). Rijksmuseum van Natuurlijke Historie, Leiden. (M. S. Hoogmoed). San Diego Society Natural History, San Diego. (Thomas Fritts). Natur-Museums und Forschungsinstitutes Senckenberg, Frankfurt a. M. Transvaal Museum, Pretoria. Umtali Museum, Umtali, Rhodesia. (Donald G. Broadley). University of Michigan, Museum of Zoology, Ann Arbor. United States National Museum, Washington, D. C. (Ronald I. Crombie). Western Australian Museum, Perth. (G. M. Storr). Zoological Institute, Academy of Sciences, Leningrad. (I. S. Darevsky) Zoologisches Museum, Berlin.
Other persons to whom I am also endebted for information are: Eugene C. Beckham III, F. W. Braestrup, William Branch, R. K. Brooke, Jeanette Covacevich, A. J. Coventry, Richard Forissier, W. Ladiges, R. Laurent, Volker Mahnert, Sven Mathiasson, Per Pethon, Oliver Rieppel, Douglas C. Robinson, Jay M. Savage, Hobart M. Smith, Edward H. Taylor, Richard Thomas, Paul Webber, Heinz Wermuth, Kenneth L. Williams, and Larry D. Wilson. Cottonwood, Arizona, September, 1976. Donald E. Hahn Marcus J. Lawrence Memorial Hospital and Department o f Biology/Verde Valley Branch Yavapai College Cottonwood, Arizona 8 6 3 2 6 / U . S . A .
Systematic Index
Class Reptilia Order Serpentes Family
Anomalepididae Anomalepis A. aspinosus A. columbius A. flavipices A. mexicanus Helminthophis H. flavoterminatus H. frontalis H. praeocularis
2 2 2 2
Liotyphlops L. albirostris L. anops L. caracasensis L. cucutae L. guentheri L. incertus L. metae L. petersii L. rowani L. schubarti L. ternetzii L. wilderi
2 3 3 3 3 3 4 4 4 4 4 4 5
Typhlophis T. squamosus
5 5
Family
Leptotyphlopidae
Leptotyphlops L. affinis L. albifrons L. albipunctus L. amazonicus L. anthracinus L. australis L. bicolor L. bilineatus L. blanfordi L. borrichianus L. boulengeri L. braccianii L. brasiliensis L. bressoni L. brevicaudus L. brevissimus L. burli L. cairi L. collaris L. columbi L. conjunctus L. c. conjunctus L. c. incognitus L. c. latirostris L. c. lepezi L. cupinensis L. debilis L. diaplocius L. dimidiatus L. dissimilis L. distanti L. dugandi L. dulcis L. d. dulcis L. d. dissectus L. d. myopicus L. erythraeus . L. filiformis L. fitzingeri
8 9 9 9 9 9 9 10 10 10 10 10
11 1 1
1 11
12 12 12 12 12 13 13 13 13 14 14 14
X Leptotyphlops gestri L. goudotii L. g. goudotii L. g. ater L. g. magnamaculatus L. g. phenops L. gracilior L. guayaquilensis L. hamulirostris L. humilis L. h. humilis L. h. boettgeri L. h. cahuilae L. h. dugesii L. h. levitoni L. h. lindsayi L. h. segregus L. h. tenuiculus L. h. utahensis L. joshuai L. koppesi L. labialis L. longicaudus L. macrolepis L. macrorhynchus L. m. macrorhynchus L. m. bilmaensis L. macrurus L. maximus L. melanotermus L. melanurus L. munaoi L. narirostris L. n. narirostris L. n. boueti L. nasalis L. natatrix L. nicefori L. nigricans L. n. nigricans L. n. pembae L. nursii L. occiden talis L. peruvianas L. phillipsi L. pyrites L. reticulatus L. rostratus L. rubrolineatus
14 14 15 15 15 15 16 16 16 16 17 17 17 17 18 18 18 18 19 19 19 19 19 20 20 20 20 20 21 21 21 21 22 22 22 22 22 22 23 23 23 23 24 24 24 24 24 24 25
Leptotyphlops rufidorsus L. salgueiroi L. scutifrons L. s. scutifrons L. s. merkeri L. septemstriatus L. signatus L. sundewalli L. teaguei L. telloi L. tenellus L. tesselatus L. tricolor L. undecimstriatus L. unguirostris L. variabilis L. weyrauchi
25 25 25 25 26 26 26 27 27 27 27 27 28 28 28 28 28
Rhinoleptus R. koniagui
28 29
Family
29
Typhlopidae
Rhinotyphlops R. anomalus R. caecus R. crossii R. feae R. gracilis R. graueri R. kibarae R. lalandei R. leucocephalus R. lumbriciformis R. newtonii R. pallidas R. praeocularis R. rufescens R. schinzi R. schlegelii R. s. schlegelii R. s. brevis R. s. mucruso R. s. petersii R. scorteccii R. somalicus R. stejnegeri R. sudanensis
29 29 29 30 30 30 31 31 31 32 32 32 32 33 33 33 33 34 34 34 35 35 35 36 36
XI
Rhinotyphlops unitaeniatus R. wittei
36 36
Typhlina T. acuticauda T. affinis T. albiceps T. angusticeps T. australis T. badila T. bituberculata T. bramina T. broomi T. cumingii T. diversa T. endo tera T. erycina T. exocoeti T. flaviventer T. grypha T. guentheri T. leptosoma T. leucoprocta T. ligata T. lineata T. minima T. multilineata T. nigroterminata T. olivacea T. pinguis T. polygrammica T. p. polygrammica T. p. brongersmai T. p. elberti T. p. florensis T. p. nigrescens T. p. undecimlineata T. próxima T. subocularis T. tovelli T. unguirostris T. wiedii T. yirrikalae
36 37 37 37 38 38 38 39 39 40 40 40 41 41 41 41 42 42 42 43 43 43 43 44 44 44 45 45 45 45 46 46 46 46 47 47 47 48 48 48
Typhlops T. acutus T. andamanensis T. angolensis T. arenarius
49 49 50 50 51
Typhlops ater T. bibronii T. biminiensis T. b. biminiensis T. b. epactia T. b. paradoxus T. bipartitus T. bisubocularis T. boylei T. brongersmianus T. coecatus T. canlaonensis T. capitulatus T. c. capitulatus T. c. gonavensis T. caymanensis T. ceylonicus T. comorensis T. conradi T. costaricensis T. cuneirostris T. decorosus T. decorsei T. depressiceps T. diardi T. d. diardi T, d. muelleri T. dominicanus T. elegans T. exiguus T. floweri T. fornasinii T. giadinhensis T. gierrai T. grandidieri T. granti T. guadeloupensis T. hectus T. hypogius T. hypsobothrius T. inorna tus T. jamaicensis T. jerdoni T. khoratensis T. klemmeri T. koekkoeki T. koshunnensis T. kraali T. lankaensis
51 51 52 52 52 52 52 52 53 53 53 53 53 54 54 54 54 54 54 55 55 55 55 55 56 56 56 57 57 57 57 58 58 58 58 59 59 59 59 59 59 60 60 60 60 60 61 61 61
XII Typhlops lehnen T. leucomelas T. leucostictus T. lineolatus T. 1. lineolatus T. 1. tanganicanus T. lorenzi T. loveridgei T. lumbricalis T. luzonensis T. madagascariensis T. malcolmi T. manilae T. manni T. microstomas T. mints T. monastus T. m. monastus T. m. geotomus T. monensis T. mucronatus T. oatesi T. obtusus T. ocularis T. oligolepis T. pammeces T. platyrhynchus T. porrectus T. pseudosaurus T. punctatus T. p. punctatus T. p. congestus T. p. liberiensis T. pusillus T. reticulatus T. reuteri T. richardi T. r. richardi T. r. capapontus T. r. naugus T. r. platycephalus T. rondoensis T. rostellatus
61 61 61 62 62 62 63 63 63 63 63 64 64 64 64 64 65 65 65 65 65 65 65 66 66 66 67 67 67 67 67 68 68 69 69 69 70 70 70 70 70 70 71
Typhlops ruber T. ruficaudus T. siamensis T. simoni T. socotranus T. stadelmani T. steinhausi T. sulcatus T. syntherus T. tasymicris T. tenebrarum T. tenuicollis T. tenuis T. thurstoni T. tindalli T. trangensis T. trinitatus T. uluguruensis T. unilineatus T. veddae T. vermicularis T. violaceus T. wilsoni T. zenkeri
71 71 71 72 72 72 72 72 73 73 73 73 73 74 74 74 74 75 75 75 75 76 76 76
Incertae sedis Typhlops disparilis T. filiformis T. fuscus T. longissimus T. macrurus T. melanocephalus T. mutilatus T. psittacus T. textilis T. verticalis
76 76 76 77 77 77 77 77 77 78
Fossil Species Typhlops carieri T. grivensis
78 78
Addendum Leptotyphlops
82
wilsoni
Family A n o m a l e p i d i d a e
TAYLOR
Anomalepis JAN 1860 Anomalepis JAN, Iconogr. gén. Ophid., 1 (1. livr.): 1; pls. 5, 6, fig. 1. - S p e c i e s t y p i c a (by monotypy): Anomalepis mexicanus JAN. 1893 Anomalolepis GÜNTHER, Biol. Centr. Amer., Rept. Batr.: 87. - Substitute name for Anomalepis JAN.
R a n g e : Mexico (?), Panama, Colombia, Ecuador, and Peru. Anomalepis aspinosus TAYLOR 1939 Anomalepis aspinosus TAYLOR, Proc. New England Zool. Club. Boston, 17: 92, pi. 5, figs. 5 - 7 . - T e r r a t y p i c a : Perico, Peru. - H o 1 o t y p e : M.C.Z. 14782.
R a n g e : Peru. Known only from the type locality and 28 km N Santa Cruz, Depto. Cajamarca. Anomalepis colombius MARX 1953 Anomalepis Colombia MARX, Fieldiana, Zool., Chicago, 34:197. — T e r r a t y p i c a : La Selva, Pueblo Rico, Department of Caldas, Colombia, 1700 m. - H o l o t y p e : F.M.N.H. 54986.
R a n g e : Known only from the type locality. Anomalepis flavapices PETERS 1957 Anomalepis flavapices PETERS, Amer. Mus. Novit., New York, 1851: 3. - T e r r a t y p i c a : Esmeraldas, Esmeralda Province, Ecuador. - H o l o t y p e : U.S.N.M. 196349.
R a n g e : Lowlands of northwestern Ecuador. Anomalepis mexicanus JAN 1860 Anomalepis mexicanus J AN, Iconogr. gen. Ophid., 1 (1. livr.):l; pis. 5, 6, fig. 1. T e r r a t y p i c a : Mexico. - H o l o t y p e : M.S.N.M. 1939 Anomalepis dentatus TAYLOR, Proc. New England Zool. Club, Boston, 17:90; pi. 5, figs. 1 - 3 - T e r r a t y p i c a : Barro Colorado Island, Canal Zone. H o l o t y p e : M.C.Z. 29220. Discussion:
AMARAL (1927:88); DUNN (1941:511).
R a n g e : Panama and (?) Mexico.
2 Helminthophis P E T E R S 1860 Helminthophis PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1860: 518. - S p e c i e s t y p i c a (by monotypy): Typhlops (Helminthophis) frontalis PETERS = Helminthophis frontalis (PETERS). 1861 Idiotyphlops JAN, Arch. Naturgesch., Berlin, 27:6. - S p e c i e s t y p i c a (by monotypy): Typhlops flavoterminatus PETERS = Helminthophis flavoterminatus (PETERS).
R a n g e : Costa Rica; Colombia and Venezuela. Helminthophis flavoterminatus ( P E T E R S ) 1857 Typhlops flavoterminatus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin 1857:402. T e r r a t y p i c a : Caracas, Venezuela. - H o l o t y p e : formerly in Hamburg Museum; destroyed during World War II. 1860 Idiotyphlops flavoterminatus — JAN. Iconogr. gén. Ophid., 1 (1. livr.) :1; pis. 5, 6, fig. 10. 1873 Helminthophis flavoterminatus - BOCAGE, Jörn. Sci. math. phys. nat., Lisboa, 4:252. D i s c u s s i o n : AMARAL (1924:25); AMARAL (1926:123). R a n g e : Venezuela and Colombia; may have been introduced into Mauritius. Helminthophis frontalis ( P E T E R S ) 1860 Typhlops (Helminthophis) frontalis PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1860:517; pL, figs. 1 - 1 a - T e r r a t y p i c a : Costa Rica. — H o l o t y p e : Z.M.B. 3925. 1881 Helminthophis frontalis - PETERS, S.Ber. Ges. naturi. Freunde Berlin, 1881:69. R a n g e : Costa Rica. Helminthophis praeocularis A M A R A L 1924 Helminthophis praeocularis AMARAL, Proc. New England zool. Club, Boston, 9: 28. - T e r r a t y p i c a : Honda, upper valley of Rio Magdalena, Colombia. H o l o t y p e : M.C.Z. 17960. D i s c u s s i o n : DUNN (1944:47). R a n g e : Inter-andean area of northern Colombia, in Tolima, Santander and Norte de Santander, from 200 to 1200 m. Liotyphlops P E T E R S 1857 Rhinotyphlops PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1857:402. - S p e c i e s t y p i c a (by monotypy): Rhinotyphlops albirostris PETERS = Liotyphlops albirostris (PETERS). - Preoccupied by Rhinotyphlops FITZINGER, 1843. 1881 Liotyphlops PETERS, S.Ber. Ges. naturforsch. Freunde Berlin, 1881:69. - S p e c i e s t y p i c a (by monotypy): Rhinotyphlops albirostris PETERS = Liotyphlops albirostris (PETERS). R a n g e : Costa Rica to Paraguay.
3 Liotyphlops albirostris ( P E T E R S ) 1857 Rhinotyphlopsalbirostris PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin 1857:402. T e r r a t y p i c a : "Veragua", Panama. - S y n t y p e s : Z.M.B. 9529 (2). 1881 Liotyphlops albirostris - PETERS, S.Ber. Ges. naturforsch. Freunde Berlin, 1881: 60. 1883 Typhlops CIdiotyphlops) emunctus GARMAN, Mem. Mus. comp. Zool., Cambridge (Massachusetts), 8(3): 3. - T e r r a t y p i c a : Panama. - H o l o t y p e : M.C.Z. 3971. 1887 Heiminthophis emunctus - COPE, Bull. U.S. nation. Mus., Washington, 32:91. 1903 Heiminthophis Canellei MOCQUARD, Bull. Mus. nation. Hist, nat., Paris, 1903:211. T e r r a t y p i c a : Isthmus of Panama. - H o l o t y p e : M.H.N.P. 3189A. 1916 Heiminthophis bondensis GRIFFIN, Mem. Carnegie Mus., Pittsburgh, 7 (1915): 165. T e r r a t y p i c a : Bonda, Colombia. - H o l o t y p e : C.M. 216. 1932 Liotyphlops albirostris - DUNN, Proc. biol. Soc. Washington, 45:176. D i s c u s s i o n : DUNN (1944:50); SMITH & GRANT (1958:207). R a n g e : Southern Central America and northern South America, including Costa Rica, Panama, Colombia, Venezuela, and Curaçao. Liotyphlops anops (COPE) 1899 Heiminthophis anops COPE, Philadelphia Mus. Sei. Bull., 1:10, pl. 4, figs. l a - f . T e r r a t y p i c a : "New Grenada" (= Colombia): DUNN (1944:8) says "near Bogota". - S y n t y p e s : two formerly in "The Philadelphia Museums"; specimens no longer extant. 1944 Liotyphlops anops - DUNN, Caldasia, Bogotá, 3:48. R a n g e : Provincias Santander and Cundinamarca, Colombia. Liotyphlops caracasensis R O Z E 1952 Liotyphlops caracasensis ROZE, Mem. Soc. Cien. nat. LaSalle, Caracas, 12 (32): 150: figs. 1 - 2 . - T e r r a t y p i c a : Cuartel Urdaneta, Caracas, Venezuela. - H o l o t y p e : Museo de Historia Natural LaSalle, Caracas 4327. D I S C U S S I O N : ROZE ( 1 9 6 6 : 32).
R a n g e : North central Venezuela. Liotyphlops cucutae DUNN 1944 Liotyphlops cucutae DUNN, Caldasia, Bogotá, 3:49; figs. 5 - 6 . - T e r r a t y p i c a : Cucuta, Norte de Santander, Colombia, 215 m. - H o l o t y p e : an un-numbered 163 mm specimen in the Instituto de LaSalle, Bogota. R a n g e : Colombia. Known only from the type locality. Liotyphlops guentheri ( B O U L E N G E R ) 1889 Heiminthophis guentheri BOULENGER, Ann. Mag. nat. Hist., London, (6) 4: 361. T e r r a t y p i c a : Porto Real, Rio de Janeiro, Brazil. - H o l o t y p e : B.M.N.H. 1946.1.11.3. 1970 Liotyphlops guentheri - PETERS & OREJAS-MIRANDA, Bull. U.S. nation. Mus., Washington, 297:182. Range:
Estado do Rio de Janeiro, Brazil.
4 L i o t y p h l o p s incertus (AM AR AL) 1924 Helminthophis incertus AMARAL, Proc. New England Zool. Club, Boston, 9 : 2 9 . T e r r a t y p i c a : Surinam. - H o l o t y p e : M.C.Z. 17846. 1948 Liotyphlops incertus - VANZOLINI, Rev. Brazil Biol., Rio de Janeiro, 8 : 380. Range:
Surinam.
L i o t y p h l o p s m e t a e DUNN 1944 Liotyphlops metae DUNN, Caldasia, Bogotá, 3:49; figs. 3 - 4 . - T e r r a t y p i c a : Villavicencio, Meta, Colombia, 498 m. - H o l o t y p e : Instituto de La Salle, Bogota, no. 8. Range:
Villavicencio, C o l o m b i a . K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
L i o t y p h l o p s petersii (BOULENGER) 1889 Helminthophis petersii BOULENGER, Ann. Mag. nat. Hist., London (6) 4: 360. T e r r a t y p i c a : Guayaquil, Ecuador. - H o l o t y p e : B.M.N.H. 1946.1.11.26. 1944 Liotyphlops petersii - DUNN, Proc. biol. Soc. Washington, 4 5 : 1 7 5 . Range:
Northwestern Ecuador.
L i o t y p h l o p s rowani SMITH & GRANT 1958 Liotyphlops rowanii SMITH & GRANT, Herpetologica, Escondido, 14:207. T e r r a t y p i c a : Pacific shoreline, Ft. Clayton Reservation, Canal Zone. H o l o t y p e : University of Illinois Museum Natural History 41731. Range:
P a c i f i c shoreline, C a n a l Z o n e . K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
L i o t y p h l o p s schubarti VANZOLINI 1948 Liotyphlops schubarti VANZOLINI, Rev. brasil. Biol., Rio de Janeiro, 8 : 379; figs. 1 - 2 . - T e r r a t y p i c a : Cachoeira de Ernas, Säo Paulo, Brazil. - H o l o t y p e : Universidade de Sao Paulo, Departamento de Zoología Collection 425. Range:
Säo P a u l o , Brazil. K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
L i o t y p h l o p s ternetzii (BOULENGER) 1896 Helminthophis ternetzii BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 3:584. T e r r a t y p i c a : Paraguay. - H o l o t y p e : B.M.N.H. 1946.1.11.77. 1924 Helminthophis beui AMARAL, Proc. New England zool. Club, Boston, 9: 29. T e r r a t y p i c a : Säo Paulo, Brazil. - H o l o t y p e : Instituto Butantan 1806. 1928 Helminthophis collenettei PARKER, Ann. Mag. nat. Hist., London, (10) 2 : 9 7 ; fig. - T e r r a t y p i c a : Burity, 2250 ft., 30 miles northeast of Coyaba, Mato Grosso, Brazil. - H o l o t y p e : B.M.N.H. 1946.1.10.73. 1958 Liotyphlops ternetzi - SMITH & GRANT, Herpetologica, Escondido, 14:207. Discussion: Range:
AMARAL (1954:191).
Säo P a u l o a n d M a t o G r o s s o , Brazil; n o r t h e r n A r g e n t i n a ; P a r a g u a y ( ? ) .
5 Liotyphlops wilden (GARMAN) 1883 Typhlops Wilden GARMAN, Sci. Observer, Boston, 4 (5 - 6):48. - T e r r a t y p i c a : Säo Cyriaco, Brazil; noted as Cyriaco, near Sena Providencia, Minas Gerais, Brazil, by HAMMAR (1908: 334); corrected by MARX (1958:496) to Säo Cypriao, Minas Gerais, Brazil. - S y n t y p e s : formerly two in Cornell University, Ithaca, New York (now apparently lost) and M.C.Z. 5126, 18138 and 19585. 1908 Helminthophis wilderi - HAMMAR, Ann. Mag. nat. Hist., London, (8) 1: 334, figs, a - c. 1948 Liotyphlops wilderi - VANZOLINI, Rev. brasil, Biol. Rio de Janeiro 8 : 380.
R a n g e : Minas Gerais, Brazil; Paraguay.
T y p h l o p h i s FITZINGER 1843 Typhlophis FITZINGER, Syst. Rept., 1:24. - S p e c i e s t y p i c a (by monotypy): Typhlops squamosus SCHLEGEL = Typhlophis squamosus (SCHLEGEL). 1844 Cephalolepis DUMERIL & BIBRON, Erpetol. gen., 6 : 314. - S p e c i e s t y p i c a (by monotypy): Cephalolepis leucocephalus DUMERIL & BIBRON = Typhlophis squamosus (SCHLEGEL). - Preoccupied by Cephalolepis RAFINESQUE, 1810.
R a n g e : Trinidad; Atlantic coast of South America from Guianas to Gräo Para, Brazil. Typhlophis squamosus (SCHLEGEL) 1839 Typhlops squamosus SCHLEGEL, Abbild. Amphib., 0 : 36; pl. 32, figs. 9 - 12. T e r r a t y p i c a : Cayenne, French Guiana. - H o l o t y p e : M.H.N.P.; now lost. 1843 Typhlophis squamosus - FITZINGER, Syst. Rept., 1:24. 1844 Cephalolepis leucocephalus DUMERIL & BIBRON, Erpetol. gen., 6 : 314. - T e r r a t y p i c a : French Guiana. - H o l o t y p e : M.H.N.P., lost. 1845 Anilios (?) squamosus - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 136. 1860 Cephalolepis squamosus - JAN, Iconogr. gen. Ophid. 1 (1. livr.): 1; pis. 5, 6, fig. 11. Discussion:
MYERS (1967:75).
R a n g e : Trinidad: Atlantic coast of South America from Guianas to Gräo Para, Brazil.
Family L e p t o t y p h l o p i d a e
STEJNEGER
L e p t o t y p h l o p s FITZINGER 1824 Stenostoma WAGLER, in Spix, Spec. nov. Serp. Brasil: 68; pl. 25, fig. 3. - S p e c i e s t y p i c a (by monotypy): Stenostoma albifrons WAGLER = Leptotyphlops albifrons (WAGLER). - Preoccupied by Stenostoma LATREILLE, 1810. 1843 Leptotyphlops FITZINGER, Syst. Rept., 1 : 2 4 - S p e c i e s t y p i c a (by original designation): Typhlops nigricans SCHLEGEL = Leptotyphlops nigricans (SCHLEGEL). 1843 Eucephalus FITZINGER, Syst. Rept. 1:24. - S p e c i e s t y p i c a (by original designation): Typhlops bilineatus SCHLEGEL = Leptotyphlops bilineatus (SCHLEGEL).
6 1844 Catodon DUMERIL & BIBRON, Erpetol. gen., 6 : 3 1 8 . - S p e c i e s t y p i c a (by monotypy): Typhlops septem-striatus SCHNEIDER = Leptotyphlops septemstriatus (SCHNEIDER). - Preoccupied by Catodon LINNAEUS, 1761. 1845 Epictia GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 139. — S p e c i e s t y p i c a (by subsequent designation: LOVERIDGE 1957: 246): Typhlops undecimstriatus SCHLEGEL = Leptotyphlops undecimstriatus (SCHLEGEL). 1845 Glauconia GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 139. - S p e c i e s t y p i c a (by monotypy): Typhlops nigricans SCHLEGEL = Leptotyphlops nigricans (SCHLEGEL). 1853 Rena BAIRD & GIRARD, Cat. N. Amer. Rept., 1:142. - S p e c i e s t y p i c a (by subsequent designation: STEJNEGER 1891: 501): Rena humilis BAIRD & GIRARD = Leptotyphlops humilis (BAIRD & GIRARD). 1857 Sabrina GIRARD, Proc. Acad. nat. Sei. Philadelphia 1857:181. - S p e c i e s t y p i c a (by monotypy): Typhlops tesselatum TSCHUDI = Leptotyphlops tesselatus (TSCHUDI). 1861 Rhamphostoma JAN, Arch. Zool. Anat. Fisiol, Genova, 1:190. - S p e c i e s t y p i c a (by monotypy): Stenostoma macrorhynchum JAN = Leptotyphlops macrorhynchus (JAN). - Preoccupied by Rhamphostoma WAGLER, 1830. 1861 Tricheilostoma JAN, Arch. Zool. Anat. Fisiol., Genova, 1:190. - S p e c i e s t y p i c a (by subsequent designation: LOVERIDGE 1957: 246): Stenostoma macrolepis PETERS = Leptotyphlops macrolepis (PETERS). 1861 Tetracheilostoma JAN, Arch. Zool. Anat. Fisiol., Genova, 1:190. - S p e c i e s t y p i c a (by monotypy): Typhlops bilineatus SCHLEGEL = Leptotyphlops bilineatus (SCHLEGEL). 1863 Ramphostoma JAN, Elenco sist. Ofidi: 16 (emendation). 1881 Siagonodon PETERS, S.Ber. Ges. naturforsch. Freunde Berlin 1881:71. - S p e c i e s t y p i c a (by original designation): Typhlops septemstriatus SCHNEIDER = Leptotyphlops septemstriatus (SCHNEIDER). 1885 Stenostomophis ROCHEBRUNE, Faune Senegambie, Rept.: 141. - Substitute name for Stenostoma WAGLER, 1824.
R a n g e : Southwestern United States southward throughout most of Central and South America to Uruguay; Bahamas, Dominican Republic, Lesser Antilles; Africa; southwestern Asia; Socotra Island. Leptotyphlops affinis (BOULENGER) 1884 Stenostoma affine BOULENGER, Ann. Mag. nat. Hist., London, (5) 13: 396. T e r r a t y p i c a : Tachira,Venezuela. - H o l o t y p e : B.M.N.H. no. 1946.1.11.16. 1929 Leptotyphlops affinis - AMARAL, Mem. Inst. Butantan, Säo Paulo, 4 : 1 3 8 . Discussion:
ROZE ( 1 9 6 6 : 4 0 - 4 1 ) .
R a n g e : States of Merida and Tachira, Venezuela. Leptotyphlops albifrons (WAGLER) 1824 Stenostoma albifrons WAGLER in Spix, Spec. nov. Serp. Brasil: 68; pi. 25, fig. 3. T e r r a t y p i c a : Vicinity of Belem, Para, Brasil. - H o l o t y p e : Formerly in Zoologische Sammlung des Bayerischen Staates, Munich; destroyed during World War II. 1893 Glauconia albifrons - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:63. 1929 Leptotyphlops albifrons - AMARAL, Mem. Inst. Butantan, Säo Paulo, 4 : 76. D i s c u s s i o n : SMITH & LIST (1958: 271); OREJAS-MIRANDA ( 1 9 6 7 : 4 3 8 - 4 3 9 ) ; and WILSON & HAHN ( 1 9 7 4 : 1 2 0 - 1 2 3 ) .
R a n g e : Known only from the type locality and Rio Grande do Norte, Brasil.
7 L e p t o t y p h l o p s albipunctus ( J A N ) 1861 Stenostoma albifrons var. albipuncta JAN, Iconogr. gen. Ophid., 1 (2. livr.): 2; pi. 5, fig. 1 - T e r r a t y p i c a : Tucuman, Argentina. - H o l o t y p e : in Museum Halle, Germany (?). 1863 Stenostoma (Typhlops) undecimstriatus var. albipuncta - JAN, Elenco sist. Ofidi: 15. 1970 Leptotyphlops albipuncta - PETERS & OREJAS-MIRANDA, Bull. U.S. nation. Mus., Washington, 297:168. Range:
Tucuman, Argentina. K n o w n only f r o m the type locality.
L e p t o t y p h l o p s amazonicus OREJAS-MIRANDA 1969 Leptotyphlops amazonicus OREJAS-MIRANDA, Común, zool. Mus. hist. nat. Montevideo, 10 (124): 1, fig. 1 . - T e r r a t y p i c a : Esmeralda, Territorio Federal Amazonas, Venezuela. - H o l o t y p e : A.M.N.H. 36664. R a n g e : E s t a d o Bolivar t o T e r r i t o r i o ' F e d e r a l A m a z o n a s , V e n e z u e l a ; possibly in A m a z o n i a n l o w l a n d s of E c u a d o r .
L e p t o t y p h l o p s anthracinus BAILEY 1946 Leptotyphlops anthracinus BAILEY, Occ. Pap. Mus. Zool. Univ. Michigan, Ann Arbor, 4 9 4 : 1 . - T e r r a t y p i c a : near Banos, 1800 m elevation, Provincia Pastaza, Ecuador. - H o l o t y p e : U.M.M.Z. 90816. D I S C U S S I O N : OREJAS-MIRANDA (1967:421 - 442). R a n g e : B a n o s a n d A b i t u a g u a in e a s t e r n l o w l a n d s o f E c u a d o r , a n d o n e r e c o r d f r o m B a l z a p a m b a , in w e s t e r n E c u a d o r i a n l o w l a n d s .
L e p t o t y p h l o p s australis FREIBERG & OREJAS-MIRANDA 1968 Leptotyphlops australis FREIBERG & OREJAS-MIRANDA, Physis, Soc. Argent. Cien. Nat., Buenos Aires, 28:145. figs. 1 - 2 - T e r r a t y p i c a : Valcheta, Rio Negro, Argentina. - H o l o t y p e : Museo Argentino de Ciencias Naturales 12525. Range:
Rio Negro to Cordoba, Argentina.
L e p t o t y p h l o p s bicolor ( J A N ) 1860 Stenostoma (Tricheilostoma) bicolor JAN, Iconogr. gén. Ophid., 1 (1 livr.): 1; pi. 5, fig. 15. - T e r r a t y p i c a : Boutry, Gold Coast (= Butre, Ghana). - H o l o t y p e : R.M.N.H. 3683, 100 mm specimen. 1860 Stenostoma (Tricheilostoma) gracile JAN, Iconogr. gén. Ophid., 1 (1. livr.): 1; pls. 5, 6, fig. 14 - T e r r a t y p i c a : None given. - H o l o t y p e : R.M.N.H. 3683. 1890 Glaucoma bicolor - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 92. 1919 Glauconia gruveli - BOULENGER, Proc. zool. Soc. London, 1919: 267. 1916 Glaucoma bicolor gruveli CHABANAUD, Bull. Mus. nation. Hist, nat., Paris, 1916: 366, figs. - T e r r a t y p i c a : Dahomey. - H o l o t y p e : M.H.N.P. 1916-196. 1950 Leptotyphlops bicolor - LEESON, Ident. Snakes Gold Coast: 21. Range:
T o g o , D a h o m e y , s o u t h w e s t C h a d , G h a n a , Nigeria, U p p e r V o l t a ,
Niger, Ivory C o a s t , a n d Mali. F o r e s t a n d savanna.
8 Leptotyphlops bilineatus (SCHLEGEL) 1839 Typhlops bilineatus SCHLEGEL, Abbild. Amphib.: 36; pl. 32, figs. 5 - 8 . - T e r r a t y p i c a : Martinique - S y n t y p e s : R.M.N.H. 3704 (2) and M.H.N.P. 3234. 1843 Typhlops (Eucephalus) bilineatus - FITZINGER, Syst. Rept., 1:24. 1844 Stenostoma bilineatum - DUMÉRIL & BIBRON, Erpétol. gén., 6 : 331. 1845 Epictia bilineatus - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 140. 1861 Stenostoma (Tetracheilostoma) bilineatum - JAN, Iconogr. gén. Ophid., 1 (2. livr.): 2; pis: 5, 6, fig. 12. 1914 Leptotyphlops bilineata - BARBOUR, Mem. Mus. comp. Zool., Cambridge (Massachusetts), 44 (2): 324. Discussion:
THOMAS (1965:1).
R a n g e : Martinique, St. Lucia, Barbados, and questionably Guadeloupe.
Leptotyphlops blanfordi (BOULENGER) 1890 Glauconia blanfordii BOULENGER, Fauna Brit. India, Rept. Batr.: 243, fig. T e r r a t y p i c a : Sind, Punjab, India. - S y n t y p e s : B.M.N.H. 1946.1.16. 8 5 - 8 9 . (5). 1908 Glauconia carltoni BARBOUR, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 5 1 : 3 1 6 . - T e r r a t y p i c a : 70 miles southwest of Amballa, Punjab, India. — H o 1 o t y p e : M.C.Z. 3749. 1943 Leptotyphlops blanfordi - SMITH, Fauna Brit. India, Rept., Amph., 3: 61. 1960 Glauconia blanfondi SUNDERSINGH (error typographicus), Proc. Rajasthan Acad. Sei., 7:53. 1960 Leptotypheops beanyordi SUNDERSINGH (error typographicus), Proc. Rajasthan Acad. Sei., 7:58. D i s c u s s i o n : MINTON (1966:117).
R a n g e : West Pakistan; Baluchistan, North West Frontier Provinces, and Afghanistan; Iran(?).
Leptotyphlops borrichianus (DEGERBQL) 1923 Glauconia borrichiana DEGERB0L, Vidensk. Meddel. dansk naturhist. Foren., Stockholm, 76:113. - T e r r a t y p i c a : Santa Rosa, Mendoza, Argentina. - H o l o t y p e : Universitetets Zoologiske Museum, Denmark, no. R539. 1929 Leptotyphlops borrichianus - AMARAL, Mem. Inst. Butantan, Sào Paulo, 4 : 1 3 9 . D i s c u s s i o n : FREIBERG (1951:259).
R a n g e : Mendoza to Rio Negro, western Argentina, and north to Chamical, La Rioja Province. Leptotyphlops boulengeri (BOETTGER) 1913 Glauconia boulengeri BOETTGER in VOELTZKOW, Reise in Ost-Afrika, 3: 354; pi. 25, fig. 1. - T e r r a t y p i c a : Manda Island, Kenya. - H o l o t y p e : S.M.F. 16700. 1957 Leptotyphlops boulengeri - LOVERIDGE, Bull. Mus. comp. Zool. Cambridge (Massachusetts), 117 (2): 246.
R a n g e : Manda and Lamu Islands, Kenya.
9 L e p t o t y p h l o p s braccianii (SCORTECCI) 1929 Glauconia braccianii SCORTECCI, Atti Soc. ital. Sci. nat., Milano, 67 (3 - 4): 294, fig. - T e r r a t y p i c a : Adi Ugri, Eritrea, Ethiopia. - H o l o t y p e : M.S.N.M. 1917. 1949 Leptotyphlops braccianii - PARKER, Zool. Verh., Leiden, 6 : 2 1 . Range:
E r i t r e a area of E t h i o p i a .
L e p t o t y p h l o p s brasiliensis LAURENT 1949 Leptotyphlops brasiliensis LAURENT, Bull. Inst. roy. Sci. nat. Belgique, Bruxelles, 25 (9): 4, figs. 7 - 9 . - T e r r a t y p i c a : Brasil. - H o l o t y p e : I.R.S.N. Reg. 2049.1.G. 12594. Range: Brasil.
K n o w n f r o m t y p e s p e c i m e n a n d a n o t h e r f r o m Barrieras, Bahia,
L e p t o t y p h l o p s bressoni TAYLOR 1939 Leptotyphlops bressoni TAYLOR, Copeia, New York & c., 1939 (1): 5; pi. 1, figs. 7 - 8 . — T e r r a t y p i c a : Hacienda El Sabino, Urupuan, Michoacan, Mexico. - H o l o t y p e : University of Illinois Museum of Natural History 25076. Discussion: Range:
KLAUBER (1940:122).
Michoacan, Mexico.
L e p t o t y p h l o p s brevicaudus (BOCAGE) 1887 Stenostoma brevicauda BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, 11:194. T e r r a t y p i c a : Dahomey. - H o l o t y p e : M.B.L. - destroyed. 1890 Glauconia brevicauda - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1950 Leptotyphlops brevicauda - LEESON, Identif. Snakes Gold Coast Prov.: 20. Range: savanna.
D a h o m e y , G h a n a , I v o r y C o a s t a n d Mali t o Nigeria. F o r e s t a n d
L e p t o t y p h l o p s brevissimus SHREVE 1964 Leptotyphlops brevissimus SHREVE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 2 1 1 : 1 . - T e r r a t y p i c a : Florencia, ¿aqueta, Colombia. - H o l o t y p e : Museo del Instituto La Salle, Bogota 1311. Discussion: Range:
ORE J AS-MIRAN DA (1967: 433).
Colombia. K n o w n only f r o m the type locality.
L e p t o t y p h l o p s burii (BOULENGER) 1905 Glauconia Burii BOULENGER, Ann. Mag. nat. Hist., London, (7) 16:178. T e r r a t y p i c a : El Kubar (= Al Kubar), Upper Hushabi, near Yemen frontier, Saudi Arabia. - H o l o t y p e : B.M.N.H. 1946.1.16.56. 1965 Leptotyphlops burii - CORKILL & COCHRANE, J. Bombay nat. Hist. Soc., 62 (3): 496. R a n g e : Saudi Arabia. K n o w n o n l y f r o m t h e h o l o t y p e , a n d f r o m D h a l a , 75 mi. N N E Aden.
10 L e p t o t y p h l o p s cairi (DUMÉRIL & BIBRON) 1844 Stenostoma Cairi DUMÉRIL & BIBRON, Erpétol. gén., 6 : 323. - T e r r a t y p i c a : Cairo, Êgypt. - H o l o t y p e : M.H.N.P. 3231. 1890 Glauconia cairi - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1932 Leptotyphlops cairi - PARKER, Proc. zool. Soc. London, 1932: 362. Range:
E g y p t , S u d a n , E t h i o p i a a n d Somalia. I s o l a t e d p o p u l a t i o n s in Ni-
ger(?) a n d M a u r i t a n i a ( ? ) . L e p t o t y p h l o p s collaris HOOGMOED 1977 Leptotyphlops collaris HOOGMOED, Zool. Meded., Leiden, 51 (7): 100, Fig. 1. T e r r a t y p i c a : Base Camp Nassau Mountains, District Marowijne, Surinam. — H o l o t y p e : R.M.N.H. 13468. Range:
N o r t h e r n part of Surinam and F r e n c h Guiana.
L e p t o t y p h l o p s columbi KLAUBER 1939 Leptotyphlops columbi KLAUBER, Trans. San Diego Soc. nat. Hist., 9 : 6 2 ; fig. 3 a - b . - T e r r a t y p i c a : Watlings Island (= San Salvador), Bahamas. H o l o t y p e : C.M. 1364. 1946 Leptotyphlops colombi - BARBOUR & LOVE RIDGE (error typographicus), Bull. Mus. comp. Zool., Cambridge (Massachusetts), 96 (2): 142. Discussion: Range: locality.
LEGLER (1959:112); THOMAS (1965:6).
San S a l v a d o r I s l a n d , B a h a m a s . K n o w n o n l y f r o m t h e t y p e
L e p t o t y p h l o p s conjunctus ( J A N ) 1861 Stenostoma conjunctum JAN, Arch. Zool. Anat. Fisiol., Genova, 1:189. - T e r r a t y p i c a : "Cape of Good Hope", South Africa. Restricted to Eastern Cape Province by BROADLEY & WATSON (1976:493). - H o l o t y p e : Staatliches Museum für Naturkunde, Stuttgart 2519A. Discussion: Range:
BROADLEY & WATSON (1976:493).
Zaire a n d U g a n d a s o u t h t o E a s t e r n C a p e P r o v i n c e , S o u t h A f r i c a .
L e p t o t y p h l o p s
c o n j u n c t u s
c o n j u n c t u s
(JAN)
1876 Stenostomagroutii COPE, J. Acad. nat. Sci. Philadelphia (2) 8:128. - T e r r a t y p i c a : Umvati Mission, Natal, South Africa. - H o l o t y p e : A.N.S.P. 3293. 1890 Glauconia conjuncta - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1933 Leptotyphlops conjunctus - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 74:224. 1956 Leptotyphlops conjunctus conjunctus - LAURENT, Ann. Mus. r. Congo Beige, Bruxelles, (8°, Zool.) 4 8 : 8 2 , 368; fig. 13. D I S C U S S I O N : FITZSIMONS (1962:84); BROADLEY & WATSON (1976:493). R a n g e : E a s t e r n C a p e P r o v i n c e n o r t h t o t h e s o u t h e r n T r a n s v a a l , Swaziland a n d N a t a l . Isolated p o p u l a t i o n at s o u t h e r n end o f L a k e T a n g a n y i k a .
11 L e p t o t y p h l o p s Watson
conjunctus
incognitus
Broad ley &
1976 Leptotyphlops conjunctus incognitus BROADLEY & WATSON, Occ. Pap. nation. Mus. Rhodesia Bulawayo, (B5) 1976 : (8): 494. - T e r r a t y p i c a : Umtali, Rhodesia. - H o l o t y p e : U.M. 6121. R a n g e : Southern Zambia, southern Malawi, northern and eastern Rhodesia, northern and eastern Transvaal, central and southern Mozambique, eastern Swaziland and KwaZulu. A n isolated population in central S o u t h West Africa.
Leptotyphlops
conjunctus
latirostris
(Sternfeld)
1912 Glaucoma latirostris STERNFELD, Wiss Ergebn. dtsch. Zentral-Afrika Exped., 4:264. - T e r r a t y p i c a : northwest of Lake Tanganika, Zaire. - H o l o t y p e : not traced. 1956 Leptotyphlops conjunctus latirostris - LAURENT, Ann. Mus. r. Congo Beige, Bruxelles, (8°, Zool.) 48:82, 368; fig. 14. Discussion: Range:
LAURENT (1956: 368).
Kivu Province, Zaire.
Leptotyphlops
conjunctus
lepezi
(Boulenger)
1901 Glauconia lepezi BOULENGER, Ann. Mus. Congo Belge, Bruxelles, Zool., ( 1 ) 2 : 8 , pi. 3, fig. 2. - T e r r a t y p i c a : Zambi, Zaire. - H o l o t y p e : R.G.M.C. 618.
1956 Leptotyphlops conjunctus lepezi - LAURENT, Ann. Mus. r. Congo Belge, Bruxelles, (8°, Zool.) 48:84. Discussion: Range:
LAURENT (1956:84).
Southern Zaire and Zambia.
Leptotyphlops cupinensis B a i l e y
& Carvalho
1946 Leptotyphlops cupinensis BAILEY & CARVALHO, Boi. Mus. nac. Brazil, Rio de Janeiro, (Nova Ser., Zool.) 5 2 : 1 , figs. 1 - 3 . - T e r r a t y p i c a : Rio Tapirape, tributary of Rio Araguaia, Mato Grosso, Brasil. - H o l o t y p e : Museu Nacional, Rio de Janeiro, 387. 1954 Leptotyphlops eupinensis BATTERSBY & SWINTON (error typographicus), Zool. Record (1952), Reptilia: 130. D i s c u s s i o n : OREJAS-MIRANDA (1967:428). R a n g e : Brasil. Known from the type locality, Serra do Navio, Territorio Amapa, Brasil, and Lely Mountains, Surinam. L e p t o t y p h l o p s debilis ( C h a b a n a u d ) 1918 Glauconia debilis CHABANAUD, Bull. Mus. nation. Mist. nat. Paris 24: 111. T e r r a t y p i c a : Kousri (= Koussri), Chad. - H o l o t y p e : M.H.N.P. 1904-180. Range:
West Africa.
12 Leptotyphlops diaplocius OREJAS-MIRANDA 1969 Leptotyphlops diaplocius OREJAS-MIRANDA, Común. Zool. Mus. hist. nat. Montevideo, 10 (124): 5, fig. 2. - T e r r a t y p i c a : Requena, Montecarmelo, Peru. - H o l o t y p e : A.M.N.H. 55654.
R a n g e : Lower parts of valleys of Rios Ucayali and Huallaga, northeastern Peru. Leptotyphlops dimidiatus (JAN) 1861 Stenostoma dimidiatum JAN, Arch. Zool. Anat. Fisiol., Genova, 1:188. - T e r r a t y p i c a : Brazil, restricted by PETERS & OREJAS-MIRANDA (1970:169) to San Marcos, near confluence of Rios Uriracuera and Tacutu, both tributaries of Rio Branco, Territorio de Roraima, Brazil. - H o l o t y p e : M.S.N.M. - destroyed. 1929 Leptotyphlops dimidiata - AMARAL, Mem. Inst. Butantan, Sao Paulo, 4 : 7 6 . 1967 Leptotyphlops dimidiatus - OREJAS-MIRANDA, Atas Simp. Biota Amaz., 5 : 4 3 3 . Discussion:
OREJAS-MIRANDA ( 1 9 6 7 : 4 3 3 - 4 3 5 ) .
R a n g e : Guianas, northern Brazil, and southeastern Venezuela.
Leptotyphlops dissimflis (BOCAGE) 1886 Stenostoma dissimile BOCAGE, Jörn. Sci. math. phys. nat., Lisboa, 11:174. T e r r a t y p i c a : White Nile. - H o l o t y p e : M.B.L. - destroyed. 1890 Glauconia dissimilis - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1933 Leptotyphlops dissimile - FLOWER, Proc. zool. Soc. London, 1933 (3): 803.
R a n g e : Sudan. Leptotyphlops distanti (BOULENGER) 1892 Glauconia distanti BOULENGER in DISTANT, A Naturalist in the Transvaal: 175, fig. 1. — T e r r a t y p i c a : Pretoria, South Africa. - H o l o t y p e : B.M.N.H. 1946.1.10.82. 1936 Leptotyphlops distanti - PARKER, Novitates Zool., New York, 4 0 : 1 2 1 . 1940 Leptotyphlops conjuncta distanti - BOGERT, Bull. Amer. Mus. nat. Hist., New York, 7 7 ( 1 ) : 13. 1955 Leptotyphlops conjunctus distanti - MERTENS, Abh. senckenb. naturf. Ges., Frankfurt am Main, 490:87. Discussion:
FITZSIMONS (1962:88); BROADLEY & WATSON (1976:501).
R a n g e : Central, northern and eastern Transvaal, and southern Mozambique; relict populations in Natal and KwaZulu. Leptotyphlops dugandi DUNN 1944 Leptotyphlops dugandi DUNN, Caldasia, Bogota 3 ( 1 1 ) : 52. - T e r r a t y p i c a : Juanamina, 20 m, about 11 km southwest of Barranquilla, Departamento Atlantico, Colombia. - H o l o t y p e : an un-numbered specimen in the Colegio Biffi, Barranquilla, Colombia.
R a n g e : Colombia. Known only from the type locality.
13 L e p t o t y p h l o p s dulcis (BAIRD & GIRARD) 1853 Rena dulcis BAIRD & GIRARD, Cat. N. Amer. Rept., 1:142. - T e r r a t y p i c a : between San Pedro and Comanche Springs, Texas. - H o l o t y p e : U.S.N.M. 7296. Discussion:
KLAUBER (1940:91).
R a n g e : South central Kansas, central Oklahoma and the panhandle, c e n t r a l T e x a s w e s t w a r d t h r o u g h s o u t h e r n N e w M e x i c o t o s o u t h e a s t A r i z o n a ; in M e x i c o , r e c o r d e d f r o m t h e s t a t e s of T a m a u l i p a s , N u e v o L e o n , V e r a c r u z , San Luis Potosi, Puebla, Queretaro, Hidalgo, Coahuila, and Chihuahua.
L e p t o t y p h l o p s
dulcis
dulcis
(BAIRD & GIRARD)
1861 Stenostoma dulce - COPE, Proc. Acad. nat. Sci. Philadelphia, 13: 305. 1863 Stenostoma (Rena) dulce - JAN, Elenco sist. Ofidi: 15. 1883 Stenostoma rubellum GARMAN, Mem. Mus. comp. Zool, Cambridge (Massachusetts), 8 (3): 130. - T e r r a t y p i c a : Uvalde, Uvalde County, Texas. - H o l o t y p e : M.C.Z. 4584. 1891 Leptotyphlops dulcis - STEJNEGER, Proc. U.S. nation. Mus., Washington, 14:501. 1892 Glaucoma dulcis - COPE, Proc. U.S. nation. Mus. Washington, 14:590. 1940 Leptotyphlops dulcis dulcis - KLAUBER, Trans, oan Diego Soc. nat. Hist., 9 : 1 0 8 ; map 1. R a n g e : Southern Oklahoma and the panhandle, south through central Texas t o northern Tamaulipas and central Nuevo Leon, Mexico.
L e p t o t y p h l o p s
dulcis
dissectus
(COPE)
1896 Glauconia dissecta COPE, Amer. Natural., Lancaster, 30:753. - T e r r a t y p i c a : Lake Valley, Sierra County, New Mexico. - H o l o t y p e : A.N.S.P. 10752. 1940 Leptotyphlops dulcis dissectus - KLAUBER, Trans. San Diego Soc. nat. Hist., 9: 112; map 1. 1952 Leptotyphlops myopicus dissectus - SMITH & SANDERS, Texas J. Sei., San Marcos, 4 ( 2 ) : 216. Discussion:
KLAUBER (1940:112).
R a n g e : F r o m southern Coahuila and Chihuahua, Mexico n o r t h and northw e s t t h r o u g h t r a n s - P e c o s T e x a s a n d s o u t h e r n N e w M e x i c o t o s o u t h e a s t e r n Ariz o n a ; also s o u t h e r n Kansas, a n d c e n t r a l a n d n o r t h e a s t e r n O k l a h o m a .
L e p t o t y p h l o p s
dulcis
m y o p i c u s
(GARMAN)
1883 Stenostoma myopicum GARMAN, Mem. Mus. comp. Zool., Cambridge (Massachusetts), 8 (3): 6. - T e r r a t y p i c a : Savineto near Tampico, Tamaulipas, Mexico. S y n t y p e s: M.C.Z. 4526, and F.M.N.H. 73386 (2). 1893 Glauconia myopica - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:69. 1929 Leptotyphlops myopica - BARBOUR & LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 69 (10): 345. 1940 Leptotyphlops dulcis myopicus - KLAUBER, Trans. San Diego Soc. nat. Hist., 9 : 1 1 7 ; map. 1.
14 1944 Leptotyphlops myopicus myopicus - SMITH, Publ. Field Mus. nat. Hist., Chicago, (Zool. Ser.), 29:146. D i s c u s s i o n : SMITH & SANDERS (1952: 204); WEBB (1970: 251). R a n g e : F r o m n o r t h e r n V e r a c r u z , P u e b l a , Q u e r e t a r o , and Hidalgo n o r t h ward across s o u t h e r n T a m a u l i p a s t o central N u e v o L e o n , Mexico. Leptotyphlops erythraeus (SCORTECCI) 1928 Glauconia erythraea SCORTECCI, Atti Soc. ital. Sci. nat., Milano, 67 (3 - 4): 293, fig. - T e r r a t y p i c a : Massaua, Eritrea, Ethiopia. - H o l o t y p e : M.S.N.M. 1916. 1949 Leptotyphlops Range:
erythraea - PARKER, Zool. Verh., Leiden, 1949 (6): 21.
Eritrea area of E t h i o p i a .
Leptotyphlops filiformis ( B O U L E N G E R ) 1899 Glauconia filiformis BOULENGER, Bull. Liverpool Mus., 2 : 7 . - T e r r a t y p i c a : Dahamis, 350 ft.; Jenna-agahan, 1200 - 2500 ft.; and Homhil, 1500 - 2500 ft., Socotra. - S y n t y p e s: B.M.N.H. 1946.1.11. 1 - 2, and two others in Liverpool Museum. 1949 Leptotyphlops Range:
filiformis - PARKER, Zool. Verh., Leiden, 1949 (6): 20.
S o c o t r a Island, n o r t h w e s t e r n I n d i a n Ocean.
Leptotyphlops fitzingeri (JAN) 1861 Stenostoma Fitzingeri JAN, Arch. Zool. Anat. Fisiol., Genova. 1:189. - T e r r a t y p i c a : Isle of Rhodes, in Nile near Cairo, Egypt. - H o l o t y p e : M.S.N.M. destroyed. 1893 Glauconia fitzingeri - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:66. Range:
Isle of R h o d e s near Cairo. K n o w n o n l y f r o m the h o l o t y p e .
Leptotyphlops gestri ( B O U L E N G E R ) 1906 Glauconia gestri BOULENGER, Annali Mus. civ. Stor. nat. Giacomo Doria, Genova, 2:210. - T e r r a t y p i c a : Musola and Basile, Fernando Po Island; Buea, Cameroon. - S y n t y p e s : B.M.N.H. 1946.1.17.3 and M.S.N.G. 28877 (2). 1961 Leptotyphlops gestri - PERRET, Bull. Soc. neuchatel. Sci. nat., 84:133. Range:
C a m e r o o n , F e r n a n d o P o Island, and Central A f r i c a n R e p u b l i c .
Leptotyphlops goudotii ( D U M E R I L & B I B R O N ) 1844 Stenostoma Goudotii DUMERIL & BIBRON, Krpetol. gen., 6 : 330. - T e r r a t y p i c a : Valley of Rio Magdalena, Colombia. - H o l o t y p e : M.H.N.P. 1068. R a n g e : Coastal and foothill regions o n Pacific slopes of Mexico f r o m Colima and Michoacan, and V e r a c r u z on t h e Caribbean, s o u t h t h r o u g h o u t Middle America t o C o l o m b i a and Caribbean coast of V e n e z u e l a ; T r i n i d a d ; Bonaire; Margarita Islands; Bay Islands o f H o n d u r a s , San A n d r e s and Providence Islands, S u m a Islands, Cozumel Island, O a x a c a , and Swan Islands, H o n d u r a s .
15 L e p t o t y p h l o p s
goudotii
goudotii
(DUMÉRIL & BIBRON)
1857 Stenostoma fallax PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1857:402. T e r r a t y p i c a : LaGuaira, Venezuela. - H o l o t y p e : Z.M.B. 1876 Stenostoma goudottii - COPE (error typographicus), J. Acad. nat. Sei. Philadelphia (2)8:129. 1929 Leptotyphlops goudotii - AMARAL, Mem. Inst. Butantan, Säo Paulo, 4 : 1 3 9 . 1952 Leptotyphlops albifrons margariate ROZE, Mem. Soc. Cien. nat. La Salle, Venezuela, 12:154; figs. 6 - 7 . — T e r r a t y p i c a : San Francisco de Macanao, Isla Margarita, Venezuela. - H o l o t y p e : M.H.N.L.S. 4515. 1970 Leptotyphlops goudotii goudotii - PETERS & ORE JAS-MI RAN DA, Bull. U.S. nation. Mus., Washington, 2 9 7 : 1 6 9 .
R a n g e : Panama and Colombia to Caribbean coast of Venezuela; Trinidad, Bonaire, and Margarita Islands. L e p t o t y p h l o p s
goudotii
ater
TAYLOR
1939 Leptotyphlops ater TAYLOR, Univ. Kansas Sci. Bull., Lawrence 26 (15): 536, fig. 4. - T e r r a t y p i c a : Managua, Nicaragua. - H o l o t y p e : U.S.N.M. 79957. 1961 Leptotyphlops albifrons ater - COCHRAN, Bull. U.S. nation. Mus., Washington, 220:194. 1970 Leptotyphlops goudotii ater - PETERS & OREJAS-MIRANDA, Bull. U.S. nation. Mus., Washington, 297 (1): 170
R a n g e : Nicaragua and Costa Rica. L e p t o t y p h l o p s
goudotii
m a g n a m a c u l a t u s
TAYLOR
1939 Leptotyphlops magnamaculata TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 26 (15): 532, fig. 1. - T e r r a t y p i c a : Isla Utila, Isias de la Bahia, Honduras. H o l o t y p e : U.S.N.M. 54760. 1950 Leptotyphlops albifrons magnamaculata - DUNN & SAXE, Proc. Acad. nat. Sci. Philadelphia, 102:141. 1970. Leptotyphlops goudotii magnamaculatus - PETERS & ORE JAS-MI RANDA, Bull. U.S. nation. Mus., Washington, 297 1: 170. 1973 Leptotyphlopsphenops - WILSON & HAHN, Bull. Florida State Mus., Gainesville, (biol. Sci.) 17 (2): 120. Discussion:
WILSON & HAHN (1973:120).
R a n g e : Bay Islands of Honduras; San Andres and Providence Islands. L e p t o t y p h l o p s
goudotii
p h e n o p s
(COPE)
1876 Stenostoma phenops COPE, J. Acad. nat. Sei. Philadelphia, 8 (2): 128. - T e r r a t y p i c a : Tehuantepec, Mexico and Coban, Guatemala. - S y n t y p e s : U.S.N.M. 12444, 30289-94. 1937 Leptotyphlops bakewelli OLIVER, Occ. Pap. Mus. Zool. Univ. Michigan, Ann Arbor, 3 6 0 : 1 ; fig. 1, pi. 1, map. - T e r r a t y p i c a : Paso del Rio, Colima, Mexico. H o l o t y p e : U.M.M.Z. 80228. 1939 Leptotyphlops phenops - SMITH, Zool. Ser. Field Mus. nat. Hist., Chicago, 2 4 : 2 8 . 1943 Leptotyphlops phenops phenops - SMITH, Proc. U.S. nation. Mus. Washington, 93:444. 1943 Leptotyphlops phenops bakewelli - SMITH, Proc. U.S. nation. Mus. Washington, 93:445.
16 1956 Leptotyphlopsgadowi DUELLMAN, Copeia New York & c., 1956 (2): 93. - T e r r a t y p i c a: above Apatzingan, 2600 ft., Michoacan, Mexico. - H o l o t y p e : B.M.N.H. 1946.9.7.55. 1970 Leptotyphlops goudotii bakewelli - PETERS & OREJAS-MIRANDA, Bull. U.S. nation. Mus., Washington, 297 (1): 169. 1970 Leptotyphlops goudotii phenops - PETERS & OREJAS-MIRANDA, Bull. U.S. nation. Mus. Washington, 297 (1): 170. D i s c u s s i o n : WILSON &HAHN (1973:120). R a n g e : Coastal and foothill regions on Pacific slopes of Mexico from Colima and Michoacan, and from Veracruz and Yucatan on the Caribbean south through Guatemala, Honduras, El Salvador, Nicaragua, and Suma Islands. Leptotyphlops gracilior ( B O U L E N G E R ) 1910 Glauconia gracilior BOULENGER, Ann. S. African Mus., Cape Town, 5:524. T e r r a t y p i c a : Van Rhynsdorp, Robertson, Clanwilliam, and Matjesfontein, South Africa. - S y n t y p e s: B.M.N.H. 1946.1.16.51; 1946.1.16.93; 10 specimens in South African Museum, Capetown. 1946 Leptotyphlops gracilior - FITZSIMONS, Ann. Transvaal Mus., Pretoria, 20: 355. 1962 Leptotyphlopsgraciliar ISEMONGER (error typographies), Snakes of Africa Southern, Central and East: 93. D i s c u s s i o n : BROADLEY & WATSON (1976:485). Range:
The western Cape Province, South Africa, inland from the coast.
Leptotyphlops guayaquilensis OREJAS-MIRANDA & PETERS 1970 Leptotyphlops guayaquilensis OREJAS-MIRANDA & PETERS, Mitt. zool. Mus. Berlin, 46:439. - T e r r a t y p i c a : Guayaquil, Ecuador. - H o l o t y p e : Z.M.B. 4508. Range:
Guayaquil and Pacific coast of Ecuador.
Leptotyphlops hamulirostris (NIKOLSKI) 1907 Glauconia hamulirostris NIKOLSKI, Ezheg. zool. Muz., St. Petersburg, 10:260. T e r r a t y p i c a : Aguljaschker, 49° 40' E, 31° 40' N, Iran. - L e c t o t y p e : Z.I.L. N10299, 305 (KRAMER & SCHNURRENBERGER, 1963). 1907 Glauconia laticeps NIKOLSKI, Ezheg. zool. Muz., St. Petersburg, 10:288, fig. 6 - 7 . - T e r r a t y p i c a : Kaletolin, Sarchum, and Malamiz, Iran. - S y n t y p e s : Z.I.L. N10294, N10301, and N10340. 1956 Leptotyphlops hamulirostris - MERTENS, Jh. Ver. vaterl. Naturk. Wiirttemb., Stuttgart, 111 (1): 90. D i s c u s s i o n : KRAMER & SCHNURRENBERGER (1963:453). R a n g e : Known only from Iranshar and Aguljaschker, Iran. Leptotyphlops humilis (BAIRD & G I R A R D ) 1853 Rena humilis BAIRD & GIRARD, Cat. N. Amer. Rept., 1:143. - T e r r a t y p i c a : "Valliecitas", California, U.S.A. - H o l o t y p e : U.S.N.M. 2101. D i s c u s s i o n : KLAUBER (1940: 123).
17 R a n g e : S o u t h w e s t and trans-Pecos T e x a s westward through southern and central Arizona, southern Nevada, south-western U t a h , southern California, Baja California, Coahuila, Sonora, Sinaloa, Jalisco, Colima, Chihuahua, Durango, Tamaulipas, and San Luis P o t o s i , M e x i c o .
L e p t o t y p h l o p s
humilis
humilis
(BAIRD&GIRARD)
1857 Stenostoma humUe - PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1857:402. 1893 Glauconia humilis - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:70. 1897 Siagonodon humilis - VAN DENBURGH, Occ. Pap. California Acad. Sci., San Francisco, 5 : 1 5 0 . 1907 Leptotyphlops humilis - RUTHVEN, Bull. Amer. Mus. nat. Hist., New York, 23 (10): 573. 1966 Leptotyphlops chumilis RHODES (error typographies), Zool. Record (1964), Rept.: 81. 1931 Leptotyphlops humilis humilis — KLAUBER, Trans. San Diego Soc. nat. Hist., 6 (23): 340. R a n g e : Coastal and c i s m o n t a n e area o f southern California, and northern and central Baja California; also the Mojave Desert and eastward o f t h e Sierras, f r o m the D e a t h V a l l e y region, through the southern tip o f Nevada, t o central and southeastern Arizona; also Cedros Island.
L e p t o t y p h l o p s
humilis
boettgeri
(WERNER)
1899 Glauconia boettgeri WERNER, Zool. Anz., Leipzig, 22: 116. - T e r r a t y p i c a : p i c a : Unknown. - H o l o t y p e : N.M.W. 15455. 1931 Leptotyphlops humilis slevini KLAUBER, Trans. San Diego Soc. nat. Hist., 6 (23): 338. - T e r r a t y p i c a : LaPaz, Baja California Sur, Mexico. - H o l o t y p e : C.A.S. 53721. 1974 Leptotyphlops humilis boettgeri - SMITH & LARSEN (error typographies), Great Basin Natural., Provo, 34 (2): 94. 1974 Leptotyphlops humilis boettgeri - SMITH & LARSEN, Great Basin Natural., Provo, 34 (2): 93. - Type locality restricted to LaPaz, Mexico. Discussion: Range:
KLAUBER (1940:132). Cape region o f Baja California Sur and Cerralvo Island.
L e p t o t y p h l o p s
humilis
cahuilae
KLAUBER
1931 Leptotyphlops humilis cahuilae KLAUBER, Trans. San Diego Soc. nat. Hist., 6 : 339. - T e r r a t y p i c a : Yaqui Well, San Diego County, California, U.S.A. H o l o t y p e : S.D.S.N.H. 2637. Discussion:
KLAUBER (1940:134).
R a n g e : Colorado and Y u m a deserts o f southeastern California and southwestern Arizona; also V i z c a i n o desert o f central Baja California.
L e p t o t y p h l o p s
humilis
dugesii
(BOCOURT)
1881 Catodon dugesii BOCOURT, Bull. Soc. philomath., Paris, (7) 6 : 8 1 . p i c a : Colima, Mexico. - S y n t y p e s : M.H.N.P. 1651 (2). 1882 Siagonodon dugesii - BOCOURT, Miss. Sci. Mexique, Rept.: 507.
Terra
ty-
18 1887 Rena dugesii - COPE, Bull. U.S. nation. Mus., Washington, 32:64. 1940 Leptotyphlops dugesii - TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 26:538, fig. 5. 1940 Leptotyphlops humilis dugesi - KLAUBER, Trans. San Diego Soc. nat. Hist., 9: 129, map 2. D i s c u s s i o n : KLAUBER (1940:129). Range:
Sonora, Sinaloa, Jalisco and Colima, Mexico.
Leptotyphlops
humilis
levitoni
MURPHY
1975 Leptotyphlops humilis levitoni MURPHY, Proc. California Acad. Sci., San Francisco, 40 (5): 93 - 107, 4 figs. - T e r r a t y p i c a : Isla Santa Catalina, Gulf of California, Mexico (26° 40' N., 110° 47' W.). - H o l o t y p e : C.A.S.: 135146. Range:
Known only from Isla Santa Catalina, Gulf of California.
Leptotyphlops
humilis
lindsayi
MURPHY
1975 Leptotyphlops humilis lindsayi MURPHY, Proc. California Acad. Sci., San Francisco, (4) 40 (5): 93 - 107, 4 figs. - T e r r a t y p i c a : Isla Carmen, Gulf of California, Mexico (25° 57' N., 111° 12' W.). - H o l o t y p e : S.D.S.N.H. 44386. Range:
Known only from Isla Carmen, Gulf of California.
Leptotyphlops
humilis
segregus
KLAUBER
1939 Leptotyphlops humilis segregus KLAUBER, Trans. San Diego Soc. nat. Hist., 9 (14a): 67. - T e r r a t y p i c a : Chalk Draw, Brewster County, Texas, U.S.A. H o l o t y p e : U.S.N.M. 103670. D i s c u s s i o n : KLAUBER (1940:140). R a n g e : Trans-Pecos Texas, southwestern New Mexico, southeastern Arizona; Coahuila, Chihuahua and Durango, Mexico.
Leptotyphlops
humilis
tenuiculus
(GARMAN)
1883 Stenostoma tenuiculum GARMAN, Mem. Mus. comp. Zool., Cambridge (Massachusetts), 8 (3): 5. - T e r r a t y p i c a : San Luis Potosi (city?), Mexico. - H o l o t y p e : M.C.Z. 4519. 1887 Rena tenuicula - COPE, Bull U.S. nation. Mus., Washington, 32:91. 1940 Leptotyphlops humilis tenuiculus - KLAUBER, Trans. San Diego Soc. nat. Hist., 9:143; map 2. D i s c u s s i o n : KLAUBER (1940:143). Range:
San Luis Potosi and Tamaulipas, Mexico.
19 L e p t o t y p h l o p s
humilis
u t a h e n s i s
TANNER
1938 Leptotyphlops humilis utahensis TANNER, Proc. Utah Acad. Sci. Arts Letters, Provo, 15:149. - T e r r a t y p i c a : Saint George, Washington County, Utah, U.S.A. - H o l o t y p e : Brigham Young University 662. Discussion:
KLAUBER (1940:137).
R a n g e : W a s h i n g t o n C o u n t y in e x t r e m e s o u t h w e s t e r n U t a h a n d s o u t h eastern Nevada. L e p t o t y p h l o p s joshuai DUNN 1944 Leptotyphlops joshuai DUNN, Caldasia, Bogotá, 3 (11): 53; figs. 9 - 1 0 . - T e r r a t y p i c a : Jerico, Antioquia, Colombia, 1967 m - H o l o t y p e : an un-numbered 270 mm. specimen in Instituto LaSalle, Bogotá. Range: Colombia.
C e n t r a l a n d w e s t e r n A n d e s , in Provincias A n t i o q u i a a n d Caldas,
L e p t o t y p h l o p s koppesi AMARAL 1955 Leptotyphlops koppesi AMARAL, Mem. Inst. Butantan, Sao Paulo, 26 (1954): 203, figs. 4 - 6 . - T e r r a t y p i c a : Terenos, Mato Grosso, Brazil. - H o l o t y p e : Instituto Butantan 8883. Range:
M a t o G r o s s o , Brazil. K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
L e p t o t y p h l o p s labialis (STERNFELD) 1908 Glaucoma labialis STERNFELD, S.Ber. Ges. naturforsch. Freunde Berlin, 1908:92. T e r r a t y p i c a : Outjo, South West Africa. - H o l o t y p e : Z.M.B. 1962 Leptotyphlops labialis - FITZSIMONS, Snakes Southern Africa: 79; fig. 12. Discussion: Range: ward.
BROADLEY & WATSON (1976:484).
N o r t h e r n p o r t i o n o f S o u t h West A f r i c a , f r o m D a m a r a l a n d n o r t h -
L e p t o t y p h l o p s longicaudus (PETERS) 1854 Stenostoma longicaudum PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1854:621. T e r r a t y p i c a : Tete, Mozambique. - H o l o t y p e : Z.M.B. 4827. 1890 Gluconia longicauda - BOULENGER, Ann. Mag. Nat. Hist., London, (6) 6 : 9 2 . 1929 Glauconia fiechteri SCORTECCI, Atti Soc. ital. nat., Milano, 6 8 : 2 6 6 , figs. T e r r a t y p i c a : Villaggio Duca degli Abruzzi, Somalia. - H o l o t y p e : M.S.N.M. 1930 Glauconia brevirostralis FITZSIMONS, Ann. Transvaal Mus., Pretoria, 14: 38, fig. 2 1 . - T e r r a t y p i c a : Hectorspruit, eastern Transvaal, South Africa. H o l o t y p e : T.M. 4994. 1953 Leptotyphlops longicauda - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 110:247. Discussion:
FITZSIMONS (1962:80): BROADLEY & WATSON (1976:487).
R a n g e : East A f r i c a n l o w l a n d s f r o m s o u t h e r n coastal S o m a l i s o u t h t o t h e eastern Transvaal a n d S w a z i l a n d , r e a c h i n g its w e s t e r n limit in R h o d e s i a a n d eastern Botswana.
20 L e p t o t y p h l o p s macrolepis (PETERS) 1857 Stenostoma macrolepis PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1857:402. T e r r a t y p i c a : Caracas and Puerto Cabello, Venezuela; restricted to Puerto Cabello, Venezuela, by OREJAS-MIRANDA (1967:430). - H o 1 o t y p e : Z.M.B. 1434 1861 Stenostoma (TricheUostoma) macrolepis - JAN, Iconogr. gen. Ophid., 1 (2. livr.): 2; pis. 5, 6, fig. 10. 1922 Leptotyphlops macrolepis - RUTHVEN, Misc. Publ. Mus. Zool. Univ. Michigan, Ann Arbor, 8 : 6 4 . 1933 Leptotyphlops ihlei BRONGERSMA, Zool. Mededel., Leiden, 15:175; figs. 1 - 2. T e r r a t y p i c a : Toegoemoetoe, Surinam. - H o l o t y p e : R.M.N.H. 4466. Discussion: Range:
OREJAS-MIRANDA, (1967:430).
P a n a m a t o C o l o m b i a , V e n e z u e l a , G u i a n a s a n d n o r t h e r n Brazil.
L e p t o t y p h l o p s macrorhynchus ( J A N ) 1861 Stenostoma (Ramphostoma) macrorhynchum JAN, Arch. Zool. Anat. Fisiol., Genova, 1:190. - . T e r r a t y p i c a : Sennaar, Sudan. - H o l o t y p e : formerly M.S.N.M.; now destroyed. D i s c u s s i o n : MINTON (1966:116). R a n g e : Niger, Algeria, M o r o c c o , L i b y a , M a u r i t a n i a , Senegal, T u n i s i a , G u i n e a , G h a n a , Mali, E g y p t , A d e n , T u r k e y , I r a n , Iraq, P a k i s t a n , a n d S i n d , I n d i a .
L e p t o t y p h l o p s
m a c r o r h y n c h u s
m a c r o r h y n c h u s
(JAN)
1890 Glauconia macrorhynchus - BOULENGER, Ann. Mag. nat. Hist., London, 6 (6): 92. 1895 Glauconia algeriensis JAQUET, Bibliogr. Anat., Paris & Nancy, 4 : 7 9 , 7 figs. T e r r a t y p i c a : near Biskra, Algeria. - H o l o t y p e : not traced. 1932 Leptotyphlops macrorhynchus - CORKILL, Snakes and Snake bite in Iraq: 8. 1936 Leptotyphlops macrorhynchus macrorhynchus - ANGEL, Bull. Soc. Zool. France, Paris, 6 1 : 2 7 4 . R a n g e : West a n d N o r t h A f r i c a ; s o u t h w e s t e r n Asia, n o r t h t o T u r k e y a n d east t o S i n d , I n d i a .
L e p t o t y p h l o p s
m a c r o r h y n c h u s
bilmaensis
ANGEL
1936 Leptotyphlops macrorhynchus bilmaensis ANGEL, Bull. Soc. Zool. France, Paris, 6 1 : 2 7 4 . - T e r r a t y p i c a : Oasis of Bilma, Niger. - S y n t y p e s : M.H.N.P. 1936-99-102. Range:
Niger.
L e p t o t y p h l o p s macrurus (BOULENGER) 1899 Glauconia longicauda BOULENGER, Bull. Liverpool Mus., 2 : 7 . - T e r r a t y p i c a : Jena-agahan, 1 2 0 0 - 2 5 0 0 ft.; Dahamis, 350 ft.; and Homhil, 1 5 0 0 - 2 5 0 0 ft., S o c o t r a . - S y n t y p e s : B.M.N.H. 1946.1.11.75-76, 1946.1.11.88-91, 1946.1.10.84-85. - Name preoccupied by Stenostoma longicaudum PETERS, 1854.
21 1903 Glauconia macrura BOULENGER, Bull. Liverpool Mus., 1903:89 (nomen novum pro Glauconia longicauda BOULENGER). 1949 Leptotyphlops macrura - PARKER, Zool. Verh., Leiden, 1949 (6): 20. R a n g e : Socotra Island, northwestern Indian Ocean.
Leptotyphlops maximus LOVERIDGE 1932 Leptotyphlops maximus LOVERIDGE, Proc. biol. Soc. Washington, 45:151. T e r r a t y p i c a : Chilpancingo, Guerrero, Mexico, altitute 4000 - 6000 ft. H o 1 o t y p e : M.C.Z. 33604. D i s c u s s i o n : TAYLOR (1939:4); KLAUBER (1940:120). Range:
Guerrero, Mexico, Morelos and Puebla, Mexico.
Leptotyphlops melanotermus (COPE) 1862 Stenostoma melanoterma COPE, Proc. Acad. nat. Sei. Philadelphia; 1862: 350. T e r r a t y p i c a : Corrientes, Argentina. - H o l o t y p e : A.N.S.P. 5406. 1876 Stenostoma flavifrons WEYENBERGH in NAPP, Die Argentinische Republik (Buenos Aires): 1 6 4 . - T e r r a t y p i c a : Argentina. - T y p e : Not traced. 1893 Stenostoma melanostoma GÜNTHER (in error for Stenostoma melanoterma COPE), BioL Centr. Amer., Rept.: 85. 1893 Stenostoma melanosterna BOULENGER (in error for Stenostoma melanoterma COPE), Cat. Snakes Brit. Mus. (Nat. Hist.), 1:63. 1945 Leptotyphlops striatula SMITH & LAUFE, Proc. biol. Soc. Washington, 58:29; figs. A, B; pi. 5, figs. A, B. - T e r r a t y p i c a : "Yamachi" = Yanacachi, Sur de Yungas, Bolivia. - H o l o t y p e : U.S.N.M. 98889. 1964 Leptotyphlops melanotermus - OREJAS-MI RANDA, Común. Zool. Mus. Hist. Nat. Montevideo, 8 (103): 4. R a n g e : Extreme southern Peru, through Bolivia and northern Argentina to Santa Fe; possibly western and southern Paraguay and southwestern Brazil.
Leptotyphlops melanurus SCHMIDT & WALKER 1943 Leptotyphlops melanurus SCHMIDT & WALKER, Field Mus. nat. Hist., Chicago, (Zool. Ser.) 24: 303. - T e r r a t y p i c a : Chiclin, Libertad, Peru. - H o l o t y p e : F.M.N.H. 34269. R a n g e : Peru. Known only from the type locality.
Leptotyphlops munaoi OREJAS-MIRANDA 1961 Leptotyphlops munaoi OREJAS-MIRANDA, Act. Biol. Venezuelica, Caracas, 3: 85, figs, la - c. - T e r r a t y p i c a : Pozo Hondo, Tambores, Departamento de Tacuarembó, Uruguay, - H o l o t y p e : Museo de Biologia de la Universidad Central de Venezuela 4547. R a n g e : Northern Argentina, Uruguay, and Rio Grande do Sul, Brazil.
22 L e p t o t y p h l o p s narirostris (PETERS) 1867 Stenostoma narirostre PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1867:708. T e r r a t y p i c a : Lagos, Nigeria. - H o l o t y p e : Z.M.B. 5915. R a n g e : Nigeria, C a m e r o o n , D a h o m e y , G h a n a , T o g o , P o r t u g e s e G u i n e a , s o u t h e r n Niger, I v o r y Coast, Senegal, M a u r i t a n i a , a n d Mali.
L e p t o t y p h l o p s
narirostris
narirostris
(PETERS)
1890 Glaucoma narirostris - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1906 Glaucoma narirostre - BOULENGER, Ann. Mag. nat. Hist., London, (7) 18:441. 1950 Leptotyphlops narirostris narirostris - VILLIERS, Les Serpents de l'Ouest Africain: 55. R a n g e : Nigeria, C a m e r o o n , D a h o m e y , G h a n a , T o g o , P o r t u g e s e G u i n e a , Ivory C o a s t , a n d s o u t h e r n Niger.
L e p t o t y p h l o p s
narirostris
b o u e t i
(CHABANAUD)
1917 Glaucoma boueti CHABANAUD, Bull. Mus. nation. Hist, nat., Paris, 1917:9, figs. T e r r a t y p i c a : "Sudan". - H o l o t y p e : M.H.N.P. 1917-24. 1950 Leptotyphlops narirostris boueti - VILLIER, Les Serpents de l'Ouest Africain: 55. Range:
Senegal, M a u r i t a n i a , Mali a n d C h a d .
L e p t o t y p h l o p s nasalis TAYLOR 1939 Leptotyphlops nasalis TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 26 (15): 535, fig. 3. - T e r r a t y p i c a : Managua, Nicaragua. - H o l o t y p e : U.S.N.M. 16134. Range:
Nicaragua. K n o w n only f r o m the type locality.
L e p t o t y p h l o p s natatrix ANDERSSON 1937 Glauconia natatrix ANDERSSON, Ark. Zool., Stockhom, 29A (16): 7. - T e r r a t y p i c a : Tobo swamp, Gambia. - H o l o t y p e : In Naturhistoriska Rijksmuseum, Stockholm. Range:
Gambia. K n o w n only f r o m the type locality.
L e p t o t y p h l o p s nicefori DUNN 1946 Leptotyphlops nicefori DUNN, Caldasia, Bogotá, 4 (17): 121. - T e r r a t y p i c a : Mogotes, Santander, Colombia, 1746 m. - H o l o t y p e : An un-numbered juvenile in the Instituto de LaSalle, Bogotá. Range:
C o l o m b i a . K n o w n o n l y f r o m t h e t y p e locality.
23 L e p t o t y p h l o p s nigricans (SCHLEGEL) 1839 Typhlops nigricans SCHLEGEL, Abbild. Amphib.: 38; pi. 32, figs. 21 - 24. T e r r a t y p i c a : Cape of Good Hope, Africa. - S y n t y p e s : M.H.N.P. 3232 and R.M.N.H. 3705. Range:
Southern Africa.
L e p t o t y p h l o p s 1843 1844 1845 1890 1916 1919 1923 1933 1941 1976
nigricans
nigricans
(SCHLEGEL)
Leptotyphlops nigricans - FITZINGER, Syst. Rept., 1:24. Stenostoma nigricans - DUMÉRIL & BIBRON, Erpétol. gén., 6 : 3 2 6 . Glauconia nigricans - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.,): 139. Glauconia emini BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 1 . - T e r r a t y p i c a : Karagwe, Tanzania. - S y n t y p e s : B.M.N.H. 1946.1.10.86, & 1946.1.10.87 (marked as the lectotype). Glauconia monticola CHABANAUD, Bull. Mus. nation. Hist. nat. Paris 1916: 366; figs. 7 - 9 . - T e r r a t y p i c a : Volcanos of Kivori (Kivu), Zaire. - H o l o t y p e : M.H.N.P. 1916-213. Glauconia kafubi BOULENGER, Rev. zool. afr., Bruxelles, 7 : 1 8 6 . - T e r r a t y p i c a : Kafubu River, Elizabethville (= Lubumbashi), Zaire. - S y n t y p e s : B.M.N.H. 1946.1.11.7-8 (2) and R.G.M.C. 2085. Leptotyphlops nigricans - SCHMIDT, Bull. Amer. Mus. nat. Hist., Washington, 49:53. Leptotyphlops emini - LO VE RIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 74:223. Leptotyphlops emini emini - LOVERIDGE, Proc. biol. Soc. Washington, 5 4 : 1 1 7 . Leptotyphlops nigricans nigricans - BROADLEY & WATSON, Occ. Pap. nation. Mus. Rhodesia, Bulawayo 5 (8): 490.
Discussion:
FITZSIMONS (1962: 81); BROADLEY & WATSON, (1976:490).
R a n g e : D i s c o n t i n u o u s . S o u t h e r n C a p e Province f r o m Calvinia t o l o w e r T r a n s k e i ; c e n t r a l Transvaal; c e n t r a l Z a m b i a t h r o u g h e a s t e r n A n g o l a , e a s t e r n Zaire, R w a n d a , Tanzania; Kenya and Uganda t o the southern Sudan.
L e p t o t y p h l o p s
nigricans
p e m b a e
LOVERIDGE
1941 Leptotyphlops emini pembae LOVERIDGE, Proc. biol. Soc. Washington, 5 4 : 1 7 7 . T e r r a t y p i c a : Wingwi Pwana, Pemba Island, Tanzania. - H o l o t y p e : M.C.Z. 46116. 1976 Leptotyphlops nigricans pembae - BROADLEY & WATSON, Occ. Pap. natn. Mus. Rhodesia, Bulawayo, (B) 5 (8): 485. Range:
P e m b a Island, T a n z a n i a .
L e p t o t y p h l o p s nursii (BOULENGER) 1896 Glauconia nursii BOULENGER, Cat. Snakes Brit. Mus. (nat. Hist.), 3:591. - T e r r a t y p i c a : Aden. - S y n t y p e s : B.M.N.H. 1946.1.16.91-92. 1933 Leptotyphlopsyemenicus SCORTECCI, Atti Soc. ital. Sci. nat., Milano, 72:165. T e r r a t y p i c a : Yemen. - T y p e : M.S.N.M. 1938 Leptotyphlops nursii - PARKER, Ann. Mag. nat. Hist., London, (11) 1:481. Discussion: Range:
SCHMIDT (1953: 353); CORKILL & COCHRANE (1966:494).
Southern Arabian peninsula.
24 L e p t o t y p h l o p s occidentalis FITZSIMONS 1962 Leptotyphlops occidentalis FITZSIMONS, Ann. Transvaal Mus., Pretoria, 24: 239. - T e r r a t y p i c a : Keetmanshoop, Great Namaqualand, South West Africa. - H o l o t y p e : T.M. 5017. Discussion:
BROADLEY & WATSON (1976:485).
R a n g e : F r o m the Richtersveld, Little Namaqualand, northwards through S o u t h West A f r i c a t o t h e K a o k o v e l d , w e s t t o G o b a b e b .
L e p t o t y p h l o p s peruvianus OREJAS-MIRANDA 1969 Leptotyphlops peruvianus OREJAS-MIRANDA, Común, zool. Mus. Hist. nat. Montevideo, 10 (124): 9; pi. 2, fig. 1. - T e r r a t y p i c a : Chanchamayo, Departamento Junin, Peru. - H o l o t y p e : A.M.N.H. 52437. Range:
Peru. K n o w n only f r o m the t y p e locality.
L e p t o t y p h l o p s phillipsi BARBOUR 1914 Leptotyphlops phillipsi BARBOUR, Proc. New England Zool. Club, Boston, 5 : 8 7 , fig. - T e r r a t y p i c a : Petra, Arabia (= Jordan). - H o l o t y p e : M.C.Z. 9650. Range:
N o r t h e r n A r a b i a , J o r d a n , a n d Israel.
L e p t o t y p h l o p s pyrites THOMAS 1965 Leptotyphlops pyrites THOMAS, Breviora, Mus. comp. Zool., Cambridge (Massachusetts), 2 2 2 : 2 ; figs. 1 - 3 . - T e r r a t y p i c a : Pedernales, Pedernales Province, Dominican Republic, Hispaniola. - H o l o t y p e : M.C.Z. 77239. Range:
H i s p a n i o l a . K n o w n o n l y f r o m t h e vicinity o f t h e t y p e l o c a l i t y .
L e p t o t y p h l o p s reticulatus (BOULENGER) 1906 Glauconia reticulata BOULENGER, Ann. Mag. nat. Hist., London, (7) 18:441. T e r r a t y p i c a : Wagga, Goolis Mountains, near Berbera, altitute 3000 - 4000 ft., Somalia. - H o l o t y p e : B.M.N.H. 1946.1.16.53. 1949 Leptotyphlops reticulata - PARKER, Zool. Verh., Leiden, 6 : 1 9 . Range: Somalia.
Somalia. K n o w n o n l y f r o m t h e t y p e l o c a l i t y , a n d Hargeisha,
L e p t o t y p h l o p s rostratus (BOCAGE) 1886 Stenostoma rostratum BOCAGE, Jörn. Sei. math. phys. nat., Lisboa, 11:173. T e r r a t y p i c a : Humbe, Cunene River, Angola. - H o l o t y p e : M.B.L. destroyed. 1890 Glauconia rostrata - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . Range:
Angola.
25 L e p t o t y p h l o p s rubrolineatus (WERNER) 1901 Glauconia albifrons rubrolineata WERNER, Abh. Ber. zool.-anthropol.-ethnol. Mus. Dresden, 9 (2): 6. - T e r r a t y p i c a : Lima, Peru. - H o l o t y p e : Dresden Museum D1752. 1970 Leptotyphlops rubrolineatus - PETERS & OREJAS-MIRANDA, Bull. U.S. nation. Mus., Washington, 2 9 7 : 1 7 1 . Range:
Peru. K n o w n only f r o m the t y p e locality.
L e p t o t y p h l o p s rufidorsus TAYLOR 1940 Leptotyphlops rufldorsum TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 26 (15): 533, fig. 2. - T e r r a t y p i c a : Lima, Peru. - H o l o t y p e : U.S.N.M. 49993. 1943 Leptotyphlops rufidorsus - SCHMIDT & WALKER, Zool. Ser., Field Mus. nat. Hist., Chicago, 24: 302. Range:
K n o w n only f r o m L i m a and Chiclin, Libertad, Peru.
L e p t o t y p h l o p s salgueiroi AMARAL 1955 Leptotyphlops salgueiroi AMARAL, Mem. Inst. Butantan, Sao Paulo, 26 (1954): 203; figs. 1 - 3 . - T e r r a t y p i c a : Ita, Espirito Santo, Brazil. - H o l o t y p e : Instituto Butantan 8876. Range:
Brazil. K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
L e p t o t y p h l o p s scutifrons (PETERS) 1854 Stenostoma scutifrons PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1854:621. T e r r a t y p i c a : Sena, Mozambique. - H o l o t y p e : Z.M.B. - now lost. Discussion:
FITZSIMONS (1962:86); BROADLEY & WATSON (1976:497).
R a n g e : Keyna and Tanzania south to Natal, westward to Angola and S o u t h West A f r i c a .
L e p t o t y p h l o p s
s c u t i f r o n s
s c u t i f r o n s
(PETERS)
1890 Glauconia scutifrons - BOULENGER, Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1908 Glauconia latifrons STERNFELD, S.Ber. Ges. naturf. Freunde Berlin, 1908:94. T e r r a t y p i c a : Damaraland, Windhuk, Outjo, and Gobabis, South West Africa. - S y n t y p e s : Z.M.B.; Lectotype designated B.M.N.H. 64.6.14.5 from Benguela, Angola by Broadley and Watson (1976:483). 1924 Glaucoma okahandjana AHL, Arch. Naturgesch., Berlin, 90 A: 247. - T e r r a t y p i c a : South West Africa. - H o l o t y p e : Z.M.B. 24949. 1935 Leptotyphlops scutifrons - FITZSIMONS, Ann. Transvaal Mus., Pretoria, 16: 308. 1976 Leptotyphlops scutifrons scutifrons - BROADLEY & WATSON, Occ. Pap. nation. Mus. Rhodesia, Bulawayo, B5 (8): 497. Discussion:
FITZSIMONS (1962:86); BROADLEY & WATSON (1976:497).
R a n g e : Southern Tanzania, south t o Natal, westwards to Angola and S o u t h West A f r i c a .
26 L e p t o t y p h l o p s
s c u t i f r o n s
merkeri
(WERNER)
1909 Glauconia Merkeri WERNER, Jahresh. Ver. vaterl. Naturk. Württemberg, Stuttgart, 6 5 : 6 1 . - T e r r a t y p i c a : Mo shi, Tanzania. - S y n t y p e s : M.C.Z. 31627, and Staatliches Museum für Naturkunde, Stuttgart 2517A (designated as lectotype by BROADLEY & WATSON, 1976). 1976 Leptotyphlops scutifrons merkeri - BROADLEY & WATSON, Occ. Pap. nation. Mus. Rhodesia, Bulawayo, B5 (8): 498.
R a n g e : Known from Longido West, Tanzania, Teita, Kenya, and the type locality.
Leptotyphlops septemstriatus (SCHNEIDER) 1801 Anguis septem-striatus SCHNEIDER, Hist. Amphib., 2:341. - T e r r a t y p i c a : none stated. - H o l o t y p e : R.M.N.H. 1830 Typhlops senptemstriatus WAGLER (lapsus), Nat. Syst. Amphib.: 196. 1844 Catodon septemstriatus - DUMERIL & BIBRON, Erpetol. gen., 6 : 3 1 9 . 1860 Stenostoma (Catodon) septemstriatum - JAN, Iconogr. gen. Ophid., 1 (1. livr.): 1; pis. 5, 6, fig. 13. 1881 Siagonodon septemstriatus - PETERS, S.Ber. Ges. naturf. Freunde Berlin, 1881:71. 1893 Glauconia septemstriata - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:71. 1925 Leptotyphlops septemstriatus - MERTENS, Senckenbergiana, Frankfurt am Main, 7:78. ?1934 Leptotyphlops tatacua BRICENO-ROSSI, Bol. Min. Salubr. Agric. Cria, Venezuela, Caracas, 1:1133. — T e r r a t y p i c a : Rio de Oro, Distrito Colon, Estado Zulia, Venezuela. - T y p e : not traced. 1969 Leptotyphlops septemlineata HOFFSTETTER & GASC (lapsus), Biol. Rept., 1 (Chapt. 5): 290. Discussion:
OREJAS-MIRANDA (1967:426).
R a n g e : Northern Brazil, Guianas and southeastern Venezuela.
Leptotyphlops signatus (JAN) 1862 Stenostoma signatum JAN, Arch. Zool. Anat. Fisiol., Genova, 1:188. - T e r r a t y p i c a : none given. - H o l o t y p e : M.H.N.P. 3235. 1893 Glauconia signata - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1: 64.
R a n g e : Known only from Tabora, Tanzania and Kibwezi, Kenya.
Leptotyphlops subcrotillus KLAUBER 1939 Leptotyphlops subcrotilla KLAUBER, Trans. San Diego Soc. nat. Hist., 9: 61; figs. 2a - b. - T e r r a t y p i c a : Grau Tombes, northern Peru; in error, according to SCHMIDT & WALKER (1943:303), who correct it to Grau, Tumbex, Peru. H o l o t y p e : C.A.S. 14554. 1943 Leptotyphlops subcrotillus - SCHMIDT & WALKER, Zool. Ser. Field Mus. nat. Hist., Chicago, 24:303.
R a n g e : Chiclin, Libertad, Peru to Pacific lowlands of southwestern Ecuador.
27 Leptotyphlops sundewalli (JAN) 1862 Stenostoma (Tricheilostoma) Sundewalli JAN, Arch. Zool. A n a t Fisiol., Genova, 1 : 1 9 1 . - T e r r a t y p i c a : "West Africa". - H o 1 o t y p e : M.S.N.M. destroyed. 1890 Glauconia sundevalli BOULENGER (emendation), Ann. Mag. nat. Hist., London, (6) 6 : 9 2 . 1950 Leptotyphlops
sundevalli
- VILLIERS, Les Serpents de l'Ouest Africain: 55.
R a n g e : Forests of Ghana and Togo. Leptotyphlops teaguei
OREJAS-MIRANDA
1964 Leptotyphlops teaguei OREJAS-MI RANDA, Común. Zool. Mus. Hist. Nat. Montevideo, 8 (103): 4; pis. 2 - 3 . - T e r r a t y p i c a : Rio Chotano, between Chota and Cutervo, northern Peru, 2350 m. - H o l o t y p e : Museu Nacional de Historia Natural, Montevideo CH 910.
R a n g e : Northern Peru. Known only from the type locality.
Leptotyphlops telloi
BROADLEY & WATSON
1976 Leptotyphlops telloi BROADLEY & WATSON, Occ. Pap. nation. Mus. Rhodesia, Bulawayo, B5 (8): 500. - T e r r a t y p i c a : Estatuane, Mozambique. - H o l o t y p e : U.M. 30635.
R a n g e : Lebombo Range on the Mozambique-Swaziland border.
Leptotyphlops tenellus
KLAUBER
1939 Leptotyphlops tenella KLAUBER, Trans. San Diego Soc. nat. Hist., 9 (14): 59; fig. l a - b. - T e r r a t y p i c a : Kartabo, Guyana. - H o l o t y p e : A.M.N.H. 14269. tenella - R O Z E , Mem. Soc. Cien. nat. La Salle, Caracas, 1952 Leptotyphlopsalbifrons 12 (32): 143. D i s c u s s i o n : BAILEY & CARVALHO ( 1 9 4 6 : 1 ) ; THOMAS (1965 : 1 ) ; OREJAS-MIRANDA ( 1 9 6 7 : 4 3 5 ) ; and THOMAS (1975 : 250).
R a n g e : Guianas, southeastern Venezuela, Amazonian Brazil, Trinidad, and Amazonian Peru; possibly Ecuador and Antigua.
Leptotyphlops tesselatus
(TSCHUDI)
1845
Typhlops (Stenostoma) tesselatus TSCHUDI, Arch. Naturgesch., Berlin, 1 1 : 1 6 2 . T e r r a t y p i c a : Lima, Peru. - H o l o t y p e : in Neuchatel Museum, Switzerland. 1861 Stenostoma albifrons var. tessellata JAN, Iconogr. gen. Ophid., 1 (2. livr.): 2; pi. 5, fig. 1**. - (Emendation). 1943 Leptotyphlops tessellatus - SCHMIDT & WALKER. Zool. Ser. Field Mus. nat. Hist., Chicago, 2 4 : 3 0 4 .
R a n g e : Known only from the vicinity of Lima, Peru.
28 L e p t o t y p h l o p s tricolor OREJAS-MIRANDA & ZUG 1974 Leptotyphlops tricolor OREJAS-MIRANDA & ZüG, Proc. biol. Soc. Washington, 87 (16): 167 - 174, 3 figs. - T e r r a t y p i c a : Yunca Pampa, 2700 m, Huaylas District, Huaylas Province, Ancash Deapartment, Perú. - H o l o t y p e : Museo de Historia Natural "Javier Prado", Lima 0669. R a n g e : Perú. K n o w n f r o m the Departamentos Cajamarca and Ancash.
L e p t o t y p h l o p s undecimstriatus (SCHLEGEL) 1839 Typhlops undecimstriatus SCHLEGEL, Abbild. Amphib.: 36. - T e r r a t y p i c a : Santa Cruz de la Sierra, Bolivia. - H o l o t y p e : M.H.N.P.; now lost. 1845 Epictia undecimstriata - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 140. 1970 Leptotyphlops undecimstriatus - PETERS & OREJ AS-MI RANDA, Bull. U.S. nation. Mus. Washington, 297:173. Range:
Bolivia. K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
L e p t o t y p h l o p s unguirostris (BOULENGER) 1902 Glauconia unguirostris BOULENGER, Ann. Mag. nat. Hist., London, (7) 9: 338. T e r r a t y p i c a : Cruz del Eje, Cordoba, Argentina. - H o l o t y p e : B.M.N.H. 1946.1.11.52. 1921 Leptotyphlops 148. Range:
unguirostris - SERIE, An. Soc. Cient. Argentina, Buenos Aires, 92:
San J u a n t o S a n t i a g o del E s t e r o , A r g e n t i n a ; s o u t h e r n P a r a g u a y .
L e p t o t y p h l o p s variabilis (SCORTECCI) 1928 Glauconia variabilis SCORTECCI, Atti Soc. ital. Sci. nat., Milano, 67 (3 - 4): 295, fig. 4. - T e r r a t y p i c a : Adi Ugri, Eritrea, Ethiopia. - S y n t y p e s : M.S.N.M. 1915 (2). 1949 Leptotyphlops Range:
variabilis - PARKER, Zool. Verh., Leiden, 1949 (6): 21.
E r i t r e a area o f E t h i o p i a .
L e p t o t y p h l o p s weyrauchi OREJAS-MIRANDA 1964 Leptotyphlops weyrauchi OREJAS-MIRANDA, Común. Zool. Mus. Hist. nat. Montevideo, 9 (103): 1, pi. 1. - T e r r a t y p i c a : Ciudad de Tucuman, Provincia de Tucuman, Argentina. - H o l o t y p e : Museo Nacional de Historia Natural de Montevideo CH 911. Range: Argentina.
Provincias T u c u m a n , S a n t i a g o del E s t e r o , C h a c o , a n d C o r d o b a ,
R h i n o l e p t u s OREJAS-MIRANDA, ROUX-ESTEVE & GUIBÉ 1970 Rhinoleptus OREJAS-MIRANDA, ROUX-ESTÉVE & GUIBÉ, Común. Zool. Mus. Hist. nat. Montevideo, (127): 1 - 4, 3 figs. - S p e c i e s t y p i c a (by monotypy): Typhlops koniagui VILLIERS = Rhinoleptus koniagui (VILLIERS). Range:
Senegal a n d G u i n e a .
29 R h i n o l e p t u s koniagui (VILLIERS) 1956 Typhlops koniagui VILLIERS, Mém. Inst. fran?. Afr. noire, Dakar, 1956 (48): 151, figs. 1 - 3 . - T e r r a t y p i c a : Youkounkoun, Guinea. - H o l o t y p e : M.H.N.P. 8730. 1967 Leptotyphlops koniagui - GUIBÉ, ROUX-ESTÉVE & VILLIERS, Bull. Mus. nation. Hist, nat., Paris, (2) 3 9 : 4 5 2 - 453, figs. 1 - 3. 1970 Rhinoleptus koniagui - OREJAS-MIRANDA, ROUX-ESTÉVE & GUIBÉ, Común. Zool. Mus. Hist. nat. Montevideo, 1970 (127): 1 - 4, 3 figs.
R a n g e : Known only from Bonghari, Casamance, Senegal; Kouroussa, Guinea; and the type locality.
Family T y p h l o p i d a e
MERREM
R h i n o t y p h l o p s FITZINGER 1843 Rhinotyphlops FITZINGER, Syst. Rept., 1:24. - S p e c i e s t y p i c a (by original designation): Typhlops lalandei SCHLEGEL = Rhinotyphlops lalandei (SCHLEGEL). 1844 Onychocephalus DUMÉRIL & BIBRON, Erpétol. gén., 6 : 2 7 2 . - S p e c i e s t y p i c a (by subsequent designation: PETERS 1881:70). Typhlops lalandei SCHLEGEL = Rhinotyphlops lalandei (SCHLEGEL). 1845 Onychophis GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 132. - S p e c i e s t y p i c a (by monotypy): Onychophis franklinii GRAY = Rhinotyphlops lalandei (SCHLEGEL). 1868 Letheobia COPE, Proc. Acad. nat. Sci. Philadelphia, 1868: 322. - S p e c i e s t y p i c a (by subsequent designation: DUNN & DUNN 1940:75): Letheobia pallida COPE = Rhinotyphlops pallidus (COPE).
R a n g e : Africa, south of the Sahara Desert. R h i n o t y p h l o p s a n o m a l u s (BOCAGE) 1873 Onychocephalus anomalus BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, 4 : 2 4 8 ; pi. 1, fig. 3. - T e r r a t y p i c a : Huila, Angola. - H o l o t y p e : M.B.L. destroyed. 1886 typhlops (Onchocephalus) anchietae BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, 1 1 : 1 7 2 . - T e r r a t y p i c a : Huila, South Angola. - H o l o t y p e : M.B.L. 1871; now destroyed by recent fire. 1890 Onychocephalus anchietae - MATSCHIE, Zool. Jahrb., Jena, 5 : 6 0 8 . 1893 Typhlops anchietae - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:40. 1893 Typhlops anomalus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:47. 1950 Typhlopsanchiate ROSE (error typographies), Rept. Amph. South Africa: 252. 1974 Rhinotyphlops anomalus - ROUX-ESTEVE, Mem. nation. Hist, nat., Paris, 87:190.
R a n g e : Southern Angola and northern South West Africa. R h i n o t y p h l o p s caecus ( A . DUM£;RIL) 1856 Onychocephalus caecus A. DUMERIL, Rev. Mag. Zool., Paris, 1856:462; pi. 21, fig. 4. - T e r r a t y p i c a : Gabon. - S y n t y p e s: M.H.N.P. 1063, 1063A. 1864 Typhlops caecus - JAN, Iconogr. gen. Ophid., 1 (4. livr.): 3; pis. 4, 5, fig. 7. 1868 Letheobia caeca - COPE, Proc. Acad. nat. Sci. Philadelphia, 1868:322. 1916 Typhlops acutirostratus ANDERSSON, Meddel. Göteborgs Mus., Zool. Afd., 9 : 2 3 , fig. 3. - T e r r a t y p i c a : Irebu, Zaire. - H o l o t y p e : Naturhistoriska Museet, Göteborg 1375.
30 1923 Typhlops avakubae SCHMIDT, Bull. Amer. Mus. nat. Hist., New York, 49:51, fig. 1. — T e r r a t y p i c a : Avakubi, Medje, Uele, Zaire. - H o l o t y p e : A.M.N.H. 11674. 1961 Typhlops caecus pitmani WITTE, Rev. Zool. Bot. afr., Bruxelles, 64:183. - T e r r a t y p i c a : Bokuma, Bas-Congo, Zaire. - H o l o t y p e : R.G.M.C. 16298. 1961 Typhlops caecus caecus - WITTE, Rev. Zool. Bot. afr., Bruxelles 64:183. 1974 Rhinotyphlops caecus - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:210. R a n g e : Gabon, Sierra Leone, Guinea, Togo, Dahomey, Ghana; Equateur and Leopoldville Provinces, Zaire.
Rhinotyphlops crossii (BOULENGER) 1893 Typhlops crossii BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:52; pi. 3, fig. 5. - T e r r a t y p i c a : Asaba, 180 miles up the Nigeria. - H o l o t y p e : B.M.N.H. 1946.1.11.55. 1974 Rhinotyphlops crossii - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris, 87:199. Range:
Known only from the type locality and Agberi, southern Nigeria.
Rhinotyphlops feae (BOULENGER) 1906 Typhlops feae BOULENGER, Annali Mus. civ. Stor. nat. Giacomo Doria, Genova, (3a) 42:209, fig. 5. - T e r r a t y p i c a : Vista Alegre, 200 - 300 m, San Thome Island, Gulf of Guinea. - H o l o t y p e : M.S.N.G. 28871. 1906 Typhlops principis BOULENGER, Annali Mus. civ. Stor. nat. Giacomo Doria, Genova, (3a) 42:209, fig. 6. - T e r r a t y p i c a : Roco Infante Don Herrique, 100 - 300 m, Principe Island, Gulf of Guinea. - S y n t y p e s: B.M.N.H. 1946.1. 11.56 and M.S.N.G. 28876. 1974 Rhinotyphlops feae - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris, 87:202. Range:
San Thome and Principe Islands, Gulf of Guinea.
Rhinotyphlops gracilis (STERNFELD) 1910 Typhlops gracilis STERNFELD, Mitt. zool. Mus., Berlin, 5 :70. - T e r r a t y p i c a : Kitungulu, Urungu, Rhodesia. - H o l o t y p e : Z.M.B. 1933 Typhlops leptosoma WITTE, Rev. Zool. Bot. afr., Bruxelles, 23:189. - T e r r a t y p i c a : Lukafu, Kundelungu, Katanga, Zaire. - S y n t y p e s : R.G.M.C. 7251 and 7252. 1933 Typhlops katangensis WITTE, Rev. Zool. Bot. afr., Bruxelles, 23 (2): 190. - T e r r a t y p i c a : Lukonzolwa, District Lupuala-Moero, Katanga, Zaire. - H o l o t y p e : R.G.M.C. 7375. 1953 Typhlops kibarae WITTE (part), Inst. Pares nation. Congo beige, Explor. Pare nation. Upemba, Miss. G. F. de Witte . . . (1946 - 1949), 6:150. - T e r r a t y p i c a (of Paratype I.R.S.N. 2203): Kaswabilenga, Upemba, Zaire. 1960 Typhlops gracilis leptosoma - LAURENT, Ann. Mus. r. Congo beige, (8°), Zool., 84:20. 1974 Rhinotyphlops gracilis - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87: 224. Range:
Zambia; Katanga Province, Zaire; southwestern Tanzania.
31 Rhinotyphlops grauen (STERNFELD) 1912 Typhlops grauen STERNFELD, Wiss. Ergebn. deutsch. Zentr.-Afr. Exped., 1907 - 1 9 0 8 , 4 : 264. - T e r r a t y p i c a : northwest of Lake Tanganyika, Zaire. H o 1 o t y p e : Z.M.B. 195 3 Typhlops kibarae WITTE (part), Inst. Pares nation. Congo beige, Explor. Pare nation. Upemba, Miss. G. F. de Witte . . . ( 1 9 4 6 - 1 9 4 9 ) , 6 : 1 5 0 . - T e r r a t y p i c a (of Paratype R.G.M.C. 15353): Kabalo, District Tanganika, Katanga, Zaire. 1956 Typhlops leptosoma polli LAURENT, Ann. Mus. r. Congo beige, (8°), Zool., 8 4 : 1 8 . - T e r r a t y p i c a : junction of Niemba and Lukuga Rivers, Territory Albertville, Zaire. - H o 1 o t y p e : R.G.M.C. 1985. 1960 Typhlops gracilis polli - LAURENT, Ann. Mus. r. Congo beige, (8°), Zool., 8 4 : 1 8 . 1966 Typhlops graven ROBB (error typographicus), Ann. Mag. nat. Hist., London, (13) 9:677. 1974 Rhinotyphlops graueri - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:227. Discussion:
LAURENT (1956:18).
R a n g e : Zaire Provinces of Leopoldville, Equateur, Orientale, Kivu and Katanga; Rwanda; Burundi; and Ujuji, Tanzania. Rhinotyphlops kibarae (WITTE) 1953 Typhlops kibarae WITTE (part), Inst. Pares nation. Congo beige, Explor. Pare nation. Upemba, Miss. G. F. de Witte . . . (1946 - 1949), 6 : 1 5 0 , fig. 35. - T e r r a t y p i c a : Pelenge Gorges, stream below Lupiala, directly even with Lufira, Kaswabilenga, Upemba National Park, Zaire. - H o l o t y p e : I.R.S.N. 2201. 1974 Rhinotyphlops kibarae - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:236.
R a n g e : Katanga Province, Zaire. Rhinotyphlops lalandei (SCHLEGEL) 1839 Typhlops Lalandei SCHLEGEL, Abbild. Amphib.,: 38; pi. 32, figs. 17 - 20. - T e r r a t y p i c a : Cape of Good Hope, South Africa. - S y n t y p e s : M.H.N.P. 1492, 3214, 3214A, 6992, 6992A, R.M.N.H. 3715 (4 specimens). 1843 Rhinotyphlops lalandei - FITZINGER, Syst. Rept., 1:24. 1844 Onychocephalus delalandii DUMERIL & BIBRON, Erpetol. gen., 6 : 2 7 3 . - (Emendation). 1845 Onychophis franklinii GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 132. - T e r r a t y p i c a : Australia (error). - H o l o t y p e : B.M.N.H. 1946.1.10.81. 1845 Onychophis fordii GRAY, Cat. Lizards Brit. Mus. (Nat. Hist): 134. - T e r r a t y p i c a : Cape of Good Hope, South Africa. - H o l o t y p e : Museum Fort Pitt, Chatham; probably now lost. 1845 Onychophis lalandeii - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 132. 1860 Typhlops Smithii JAN, Iconogr. gen. Ophid., 1 (1 livr.): 1; pis. 5, 6, fig. 5. T e r r a t y p i c a : Cape of Good Hope, South Africa. - H o l o t y p e : M.S.N.M. destroyed. 1883 Typhlops (Onychocephalus) lalandei var. paucisquamosa BOETTGER, Ber. Offenbach. Ver. Naturkunde, 2 2 / 2 3 : 1 5 5 . — T e r r a t y p i c a : Smithfield, Transvaal (= Orange Free State). - L e c t o t y p e : S.M.F. 16658 (MERTENS, 1967:88). 1887 Typhlops (Onychocephalus) delalandei - BOETTGER, Ber. senckenb. naturf. Ges., Frankfurt am Main, 1887:154, pi. 5. 1887 Typhlops delalandei - BOULENGER, The Zoologist, London, (3) 11:173. 1898 Typhlops delalandei var. paucisquamata BOETTGER (error typographicus), Kat. Rept.-Samml. Mus. senckenb. naturf. Ges., 2 : 5 .
32 1974 Rhinotyphlops 87:158. Discussion:
lalandei - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris,
FITZSIMONS (1962:71).
R a n g e : Cape Province, S o u t h Africa, west of the A l b a n y district, e x t e n d i n g n o r t h w a r d s t h r o u g h t h e Orange Free State, Transvaal, and n o r t h e r n Cape, and into the southwestern part of southern Rhodesia. R h i n o t y p h l o p s l e u c o c e p h a l u s (PARKER) 1930 Typhlops leucocephalus PARKER, Ann. Mag. nat. Hist., London, (10) 6 : 6 0 5 . T e r r a t y p i c a : Las Anod, 2400 ft. elevation, 8° 28' N, 47° 21' E, Somalia. H o l o t y p e : B.M.N.H. 1946.1.11.96. 1974 Rhinotyphlops leucocephalus - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris, 8 7 : 1 8 3 . Range:
Somalia. K n o w n only f r o m the t y p e locality.
R h i n o t y p h l o p s lumbriciformis (PETERS) 1874 Onychocephalus (Letheobia) lumbriciformis PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1874: 377. - T e r r a t y p i c a : "Zanzibar coast", probably in error. S y n t y p e s: Z.M.B. 8045 & 8045A. 1893 Typhlops lumbriciformis - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1 : 5 4 . 1904 Typhlops kleebergi WERNER, Zool. Anz., Leipzig, 2 7 : 4 6 4 . - T e r r a t y p i c a : Usambara, Tanzania. - H o 1 o t y p e : Z.M.B. 18088. 1974 Rhinotyphlops lumbriciformis - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris, 87:221. Range:
Coastal Kenya and Tanzania.
R h i n o t y p h l o p s n e w t o n » (BOCAGE) 1890 Typhlops (Onychocephalus) Newtonii BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, ( 2 ) 2 : 6 1 . - T e r r a t y p i c a : Rolas Island, south of San Thome Island, Gulf of Guinea, West Africa. - H o 1 o t y p e : M.B.L. 1890. 1920 Typhlops naveli ANGEL, Bull. Mus. nation. Hist, nat., Paris, 2 6 : 1 9 7 , figs. 1 - 3. T e r r a t y p i c a : Principe Island, Gulf of Guinea, West Africa. - H o l o t y p e : M.H.N.P. 1920-9. 1974 Rhinotyphlops newtoni - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:195. Range: Africa.
S a n T h o m e , R o l a s , a n d P r i n c i p e Islands, G u l f of G u i n e a , West
R h i n o t y p h l o p s pallidus (COPE) 1868 Letheobia pallida COPE, Proc. Acad. nat. Sci., Philadelphia, 1886:322. - T e r r a t y p i c a : Zanzibar. - H o l o t y p e : formerly in Essex Institute, Salem, Massachusetts; now M.C.Z. 5723. 1893 Typhlops pallidus - BOULENGER, Cat. Snakes Brit. Mus. (Nat Hist.) 1 : 5 4 . 1974 Rhinotyphlops pallidus - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87 : 217. R a n g e : Z a n z i b a r ; s o u t h e r n S u d a n ; P e m b a Island a n d c o a s t a l b e l t o f K e n y a and Tanzania.
33 R h i n o t y p h l o p s praeocularis (STEJNEGER) 1894 Typhlopspraeocularis STEJNEGER, Proc. U.S. nation. Mus., Washington, 16:709. T e r r a t y p i c a : Stanley Pool, Zaire. - H o 1 o t y p e : U.S.N.M. 20799. 1964 Typhlops praeocularis lundensis LAURENT, Publ. Cult. Cia Diamant. Angola, 6 7 : 9 0 , fig. - T e r r a t y p i c a : Dundo, Angola. - H o l o t y p e : Dundo Museum of the Diamang Lunda Province, Angola 5929. 1964 Typhlops praeocularis praeocularis - LAURENT, Publ. Cult. Cia Diamant. Angola, 67:90. 1974 Rhinotyphlops praeocularis - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:221. Range:
Leopoldville Province, Zaire.
R h i n o t y p h l o p s rufescens (CHABANAUD) 1916 Typhlops rufescens CHABANAUD, Bull. Mus. nation. Hist, nat., Paris, 2 2 : 365, figs. 4 - 6 . — T e r r a t y p i c a : Haut-Oubangui, Central African Republic. H o l o t y p e : M.H.N.P. 1916-262. 1956 Typhlops vryjdaghi LAURENT, Rev. Zool. Bot. afr., Bruxelles, 5 3 : 249. - T e r r a t y p i c a : Bambesa, Zaire. - H o l o t y p e : R.G.M.C. 11949. 1974 Typhlops vrijdaghi ROUX-ESTEVE (spelling variation), Mem. Mus. nation. Hist, nat., Paris, 8 7 : 2 3 0 . 1974 Rhinotyphlops rufescens - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris, 87:230. R a n g e : K n o w n only f r o m Bambesa-Takpwo, lower Uele, Zaire and HautOubangui, Central African Republic.
R h i n o t y p h l o p s schinzi (BOETTGER) 1887
Typhlops (Onychocephalus) schinzi BOETTGER, Ber. senckenb. naturf. Ges., Frankfurt am Main, 1887:154; pi. 5, figs, l a - e, 2. - T e r r a t y p i c a : Noi Xas, Ghansi District, western Kalahari, South West Africa. - H o l o t y p e : S.M.F. 16665. 1893 Typhlops schinzi - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1 : 4 7 . 1974 Rhinotyphlops schinzi - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:186.
R a n g e : F r o m the northwestern Cape Province and Little Namaqualand into the western Kalahari and Great N a m a q u a l a n d , southern Africa.
R h i n o t y p h l o p s schlegelii (BIANCONI) 1847
Typhlops schlegelii BIANCONI, Spec. Zool. Mosambique: 183; pi. 6, fig. 2. T e r r a t y p i c a : Inhambane, Mozambique. - H o l o t y p e : Bologne Museum (?).
Discussion:
FITZSIMONS (1962:73); ROUX-ESTÈVE (1974:164).
R a n g e : Ethiopia; Somilia; southern Sudan; coastal Kenya; Tanzania; southern Zaire; Angola; Rhodesia; Mozambique; Swaziland; n o r t h e r n Zululand, n o r t h e r n and eastern Transvaal.
34 R h i n o t y p h l o p s
s c h l e g e l i i
s c h l e g e l i i
(BIANCONI)
1860 Onychocephalus schlegelii - PETERS, M.Ber. k. preufl. Akad. Wiss. Berlin, 1 8 6 0 : 8 2 . 1933 Typhlops schlegelii schlegelii - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 7 4 : 2 1 4 . 1974 Rhinotyphlops schlegelii schlegelii - ROUX-ESTEVE, Mem. Mus. nation. Hist, nat., Paris, 8 7 : 1 6 4 . Discussion:
FITZSIMONS ( 1 9 6 2 : 7 3 ) .
R a n g e : Southern Mozambique, northern Zululand, Zwaziland to eastern and northern Transvaal south of the Limpopo River.
R h i n t o t y p h l o p s
s c h l e g e l i i
brevis
(SCORTECCI)
1929
Typhlops brevis SCORTECCI, A t t i Soc. ital. Sci. nat., Milano,: 6 8 (3 - 4): 267. T e r r a t y p i c a : Chisimayo (Kismayu), Somalia. - H o l o t y p e : M.S.N.M. 2033. 1949 Typhlops schlegelii brevis - P A R K E R , Zool. Verb., Leiden, 6 : 2 1 . 1974 Rhinotyphlops schlegelii brevis - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 8 7 : 1 8 1 .
Range:
Southern Sudan, Somalia, central Ethiopia, and northern Uganda.
R h i n o t y p h l o p s
s c h l e g e l i i
m u c r u s o
(PETERS)
1854 Onychocephalus dinga PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1 8 5 4 : 6 2 0 . T e r r a t y p i c a : Tete, Sena and Chupanga, Mozambique. Syntypes: Z.M.B. 3942, 3943, 3918; M.C.Z. 32344. - nomen oblitum. 1854 Onychocephalus mucruso PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1854: 6 2 1 . - T e r r a t y p i c a : Macanga (= Makanga), Tete, and Sena, Mozambique. S y n t y p e s : Z.M.B. 3945, 3963, 3946, & 3947. 1860 Onychocephalus varius PETERS, M.Ber. k . preuß. Akad. Wiss. Berlin, 1 8 6 0 : 8 2 . T e r r a t y p i c a : Mozambique. - H o l o t y p e : Z.M.B. 3943. 1881 Typhlops (Onychocephalus) riparius PETERS, S.Ber. Ges. naturf. F r u n d e Berlin, 1 8 8 1 : 5 0 . - T e r r a t y p i c a : Chupanga, Mozambique Holotype: Z.M.B. 3943. 1882 Typhlops dinga - PETERS, Reise n. Mossambique, Zool., 3 : 9 8 ; pi. 14, fig. 2; pi. 14A, fig. 1. 1882 Typhlops varius - PETERS, Reise n. Mossambique, Zool., 3 : 9 7 ; pi. 14, fig. 2; pi. 14A, fig. 1 - l a . 1882 Typhlops mucruso - PETERS, Reise n. Mossambique, Zool. 3 : 9 5 ; pi. 13, fig. 3. 1882 Typhlops riparius - PETERS, Reise n. Mossambique, Zool., 3 : 9 8 ; pi. 14A, fig. 2. 1893 Typhlops mandensis S T E J N E G E R , Proc. U.S. nation. Mus., Washington,: 1 6 : 7 2 5 . T e r r a t y p i c a : Wange, island of Manda, north of Lamu, Kenya. - H o l o t y p e : U.S.N.M. 20125. 1909 Typhlops mucruso varius - CHUBB, Proc. Zool. Soc. L o n d o n , 1 9 0 9 : 5 9 5 . 1909 Typhlops maeruso CHUBB (error t y p o g r a p h i e s ) , Rep. Rhodesia Mus., Bulawayo, 8:35. 1910 Typhlops latirostris S T E R N F E L D , Mitt. Zool. Mus. Berlin, 5 : 6 9 - 7 0 . - T e r r a t y p i c a : Tabora, Tanzania. - H o l o t y p e : Z.M.B. 23459. 1912 Typhlops viridiflavus PERACCA, Annuario Mus. zool. Univ. Napoli, (2) 3 ( 2 5 ) : 3. T e r r a t y p i c a : Bangwelu Lake, Zambia. - H o l o t y p e : Naples Zoological Museum; now presumed lost. 1922 Typhlops excentricus PROCTER, Ann. Mag. nat. Hist., London, (9) 9 : 6 8 5 686. - T e r r a t y p i c a : near Kiiosa, Tanzania. - H o l o t y p e : B.M.N.H. 1946.1.11.51.
35 1933 Typhlops schlegelii excentricus - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 7:216. 1933 Typhlops schlegelii mucruso - LOVERIDGE, Bull. Mus. comp. Zool. Cambridge (Massachusetts), 74: 216. 1936 Typhlops schlegelii mucroso PARKER (error typographicus), Novitates zoologicae, London, 40:120. 1974 Rhinotyphlops schlegelii dinga - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:166. D i s c u s s i o n : FITZSIMONS (1962:75), HAHN (1977:39). R a n g e : Africa, north of the Zambesi in the east through Mozambique and Tanzania to Kenya, and west through Malawi and Zambia to Angola and into the northern parts of South West Africa, and perhaps northern Botswana. Rhinotyphlops
schlegelii
petersii
(BOCAGE)
1873 Onychocephalus petersii BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, 4:248 2 4 9 . - T e r r a t y p i c a : Biballa, Angola. - S y n t y p e s : M.B.L. 1868; now destroyed. 1886 Typhlops (Onychocephalus) humbo BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, 11:171. - T e r r a t y p i c a : Quissange and Benguella, Angola. - S y n t y p e s : M.B.L. 1887. 1893 Typhlops humbo - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:46. 1893 Typhlops hottentotus BOCAGE, Jorn. Sci. math. phys. nat., Lisboa, (2) 3:117. — T e r r a t y p i c a : Humbe, Angola. - H o l o t y p e : M.B.L. 1867; now destroyed. 1974 Rhinotyphlops schlegelii petersii - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris. 87:166. Range:
Southwestern Angola.
Rhinotyphlops scorteccii (GANS & LAURENT) 1965 Typhlops scorteccii GANS & LAURENT, Ann. Mus. Roy. Afr. Centr., Tervuren, (8°) Zool., 134:56, fig. 12. - T e r r a t y p i c a : 10 km south of Afgoi (40 km south of Mogadiscio), on sand dune road to Merca, Benadir Province, Somali. H o l o t y p e : M.C.Z. 74455. 1974 Rhinotyphlops scorteccii - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris. 87:246. Range:
Benadir Province, Somali Republic.
Rhinotyphlops somalicus (BOULENGER) 1895 Typhlops somalicus BOULENGER, Proc. zool. Soc. London, 1895:536; pi. 30, fig. 1. - T e r r a t y p i c a : Beearso, western Somaliland. - H o l o t y p e : B.M.N.H. 1946.1.11.54. 1905 Typhlops acutirostris MOCQUARD, Bull. Mus. nation. Hist, nat., Paris, 11 (2): 7 7 . - T e r r a t y p i c a : Choa, Ethiopia. - S y n t y p e s : M.H.N.P. 1904-603-604 (2). 1923 Typhlops somsalicu SCHMIDT (error typographicus), Bull. Amer. Mus. nat. Hist., New York, 49 (1): 33. 1928 Typhlos erythraeus SCORTECCI (error typographicus), Atti Soc. ital. Sci. nat., Milano, 67 (2 - 3): 291, fig. 1. - T e r r a t y p i c a : Sanganeiti, Eritrea, Ethiopia. - H o l o t y p e : M.S.N.M. 2015. 1974 Rhinotyphlops somalicus - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:200. Range:
Highlands of Ethiopia and western Somalia.
36 Rhinotyphlops stejnegeri (LOVERIDGE) 1931 Typhlops stejnegeri LOVERIDGE, Copeia, Ann Arbor & c., 1931 (3): 92, figs. 1 - 2 . - T e r r a t y p i c a : Kasai, Luebo District, Zaire. - H o l o t y p e : U.S.N.M. 23979. 1933 Typhlops vanderysti WITTE, Rev. Zool. Bot. afr., Bruxelles, 24 :104, figs. 1 - 3. T e r r a t y p i c a : Kikwit, Zaire. - H o l o t y p e : R.G.M.C. 2307. 1974 Rhinotyphlops stejnegeri - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:207.
R a n g e : Kasai Province, Zaire. Rhinotyphlops sudanensis (SCHMIDT) 1923 Typhlops sudanensis SCHMIDT, Bull. Amer. Mus. nat. Hist., New York, 4 9 : 5 1 , fig. 2. - T e r r a t y p i c a : Faradje, Garamba, Zaire. - H o l o t y p e : A.M.N.H. 11677. 1974 Rhinotyphlops sudanensis - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:233.
R a n g e : Orientale Province, Zaire; Uganda (?). Rhinotyphlops unitaeniatus (PETERS) 1878 Typhlops (Letheobia) unitaeniatus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1878:205; pL 2, fig. 5. - T e r r a t y p i c a : Taita, Keyna. - H o l o t y p e : Z.M.B. 9296. 1893 Typhlops unitaeniatus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:55. 1912 Typhlops unitaeniatus var. ataeniatus BOULENGER, Annali Mus. civ. Stor. nat. Giacomo Doria, Genova, (3) 5 : 3 3 1 . - T e r r a t y p i c a : Dolo, Ethiopia. S y n t y p e s : B.M.N.H. 1946.1.10.78, and M.S.N.G. (2). 1936 Typhlops unitaeniatus unitaeniatus - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 79:227. 1939 Typhlops unitaenitus ataeniatus - SCORTECCI, Annali Mus. civ. Stor. nat. Giacomo Doria, Genova, 5 8 : 2 6 4 . 1974 Rhinotyphlops unitaeniatus - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 87:241.
R a n g e : Somalia; southeastern Ethiopia; Kenya, south to Buhuri, northeastern Tanzania. Rhinotyphlops wittei ROUX-ESTÈVE 1966 Typhlops vryjdaghi - WITTE (part), Inst. Pare nation. Congo Expl. P. N. Garamba, Miss Saeg., 48 (Rept.): 50. 1974 Typhlops vrijdaghi ROUX-ESTÈVE (partim; spelling variation), Mém. Mus. nation. Hist, nat., Paris, 87:239. 1974 Rhinotyphlops wittei ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, 8 7 : 2 3 9 . T e r r a t y p i c a : Vandia-Poshe, Region Liki Bembe River, Territory Libenge, District Ubangi, Province Équateur, Zaire. - H o l o t y p e : I.R.S.N. 2511.
R a n g e : Zaire. Known only from the type locality. Typhlina WAGLER 1830 Typhlina WAGLER, Nat. Syst. Amphib.: 196. - S p e c i e s t y p i c a (by subsequent designation: FITZINGER 1843:24): Acontias lineatus „REINWARDT" (nomen nudum). See STIMSON & al. (1977: 204).
37 1843 Ramphotyphlops FITZINGER, Syst. Rept., 1:24. - S p e c i e s t y p i c a (by original designation): Typhlops multilineatus SCHLEGEL = Typhlina multilineata (SCHLEGEL). 1843 Pseudotyphlops FITZINGER, Syst. Rept., 1 : 2 4 . - S p e c i e s t y p i c a (by original designation): Typhlops polygrammicus SCHLEGEL = Typhlina polygrammica (SCHLEGEL). - Preoccupied by Pseudotyphlops SCHLEGEL 1839, Uropeltidae. 1844 Pilidion DUMERIL & BIBRON, Erpetol. gen., 6 : 2 5 7 . - S p e c i e s t y p i c a (by original designation): Typhlina lineatus "REINWARDT" (= WAGLER) = Typhlina lineata WAGLER. - Substitute name for Typhlina, believed (wrongly) to be preoccupied by Typhline WIEGMANN. 1844 Cathetorhinus DUMERIL & BIBRON, Erpetol. gen., 6 : 2 6 8 . - S p e c i e s t y p i c a (by montypy): Cathetorhinus melanocephalus DUMERIL & BIBRON = ?. 1845 Typhlinalis GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.),: 134. - S p e c i e s t y p i c a (by monotypy): Typhlina lineatum WAGLER; substitute name for Typhlina, believed preoccupied by Typhline.
R a n g e : Burma; Thailand; Malaysia; Indonesia; Philippine Islands; Lesser Sundas, and Moluccas; New Guinea; Bismarck Archipelago; Solomon Islands; Palau Islands; possibly New Caledonia; Australia; Loyalty Islands; Fiji Islands. The uncertain inclusion of the all-female species Typhlina (?) bramina expands the range to eastern and western Africa, Madagascar, India, and Pakistan, many Pacific islands, and western Mexico. Typhlina acuticauda (PETERS), New Combination 1877 Typhlops acuticauda, PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1877:416; pi., fig. 2. - T e r r a t y p i c a : Palaos, Palau Islands, Caroline Group. - H o l o t y p e : Z.M.B.
R a n g e : Palau Islands. Typhlina affinis (BOULENGER) 1889 Typhlops affinis BOULENGER, Ann. Mag. nat. Hist., London, (6) 4 : 363. - T e r r a t y p i c a : Queensland. - H o l o t y p e : B.M.N.H. 1946.1.10.56. 1914 Typhlops kenti BOULENGER, Ann. Mag. nat. Hist., London, (8) 1 4 : 4 8 2 . - T e r r a t y p i c a : northern Queensland. - H o l o t y p e : B.M.N.H. 1946.1.11.86. 1921 Typhlops cumingii mansuetus BARBOUR, Proc. New England Zool. Club, Boston, 7 : 9 1 . - T e r r a t y p i c a : Keri Keri, San Cristobal Island, Solomon Islands. H o l o t y p e : M.C.Z. 14270. 1966 Ramphotyphlops kenti - ROBB, Ann. Mag. nat. Hist., London, (9) 13:676. 1966 Ramphotyphlops affinis - ROBB. Ann. Mag. nat. Hist., London, (9) 13:676. 1974 Typhlina affinis - MCDOWELL, J. Herpetol., Athens, 8 (1): 33. Discussion:
COGGER (1975: 348).
R a n g e : Northern and eastern Australia; western Australia in the interior of the Kimberley region; San Cristobal Island, Solomon Islands. Typhlina albiceps (BOULENGER), New Combination 1898 Typhlops albiceps BOULENGER, Ann. Mag. nat. Hist., London, (7) 1:124. T e r r a t y p i c a : Chanthaburi, southeast Thailand. - H o l o t y p e : B.M.N.H. 1946.1.10.50. 1921 Typhlops albicep - WERNER, Arch. Naturgesch., Berlin, 87A (7): 337.
38 1937 Typhlops malaisei RENDAHL, Ark. Zool., Stockholm, 29A (10): 1 1 . - T e r r a t y p i c a : Dawna Hills, Sukli, southern Burma. - H o l o t y p e : Zootomisches Institut der Hochschule zu Stockholm 2999. D i s c u s s i o n : TAYLOR (1965:639). R a n g e : Malay Peninsula; Thailand; Burma; Kedah and Jarak Islands, Malacca Strait; Hongkong (?).
Typhlina angusticeps (PETERS) 1878 Typhlops (Onychocephalus) angusticeps PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1878:417; figs. 3 - 3c. - T e r r a t y p i c a : New Caledonia. - H o l o t y p e : Z.M.B. 1921 Typhlops olivaceus reduncus BARBOUR, Proc. New England Zool. Club, Boston, 7 : 1 0 7 . - T e r r a t y p i c a : Keri Keri, San Cristobal Island, Solomon Islands. H o l o t y p e : M.C.Z. 14269. 1974 Typhlina angusticeps - MCDOWELL, J. Herpetol., Athens, 8 (1): 47. R a n g e : Southermost Solomon Islands: Guadalcanal, San Cristobal, Rennell, and Malaita.
Typhlina australis (GRAY) 1845 Anilios australis GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 1 3 5 . - T e r r a t y p i c a : Western Australia. - L e c t o t y p e : B.M.N.H. 1946.1.10.61. 1860 Typhlops Preissi JAN, Iconogr. gen. Ophid., 1 (1.) (livr.): 1; pL 5, fig. 2. - T e r r a t y p i c a : Southwest Australia. - H o l o t y p e : R.M.N.H. 3689. 1860 Onychocephalus bicolor PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1860:81. T e r r a t y p i c a : Melbourne, Australia. - H o l o t y p e : Z.M.B. 1864 Typhlops bicolor - JAN, Iconogr. gen. Ophid., 1 (4. livr.): 3; pis. 4, 5, fig. 3. 1865 Typhlops australis - PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1865 :262, fig. 3. 1894 Typhlops waitii BOULENGER, Proc. Linn. Soc. New South Wales, Sydney, 9 : 7 1 8 . T e r r a t y p i c a : Northwest Australia. - H o l o t y p e : B.M.N.H. 1946.1.11.74. 1966 Ramphotyphlopsaustralis - ROBB, Ann. Mag. nat. Hist., London, 13 (9): 675. 1974 Typhlina australis - MCDOWELL, J. Herpetol., Athens, 8(1): 35. D i s c u s s i o n : WAITE (1929:189); KINGHORN (1956:60); COGGER (1975: 348). R a n g e : South Australia, across southern parts to southwest Australia; southern inland New South Wales; northwest Victoria.
Typhlina batilla (WAITE) 1894 Typhlops batillus WAITE, Proc. Linn. Soc. New South Wales, Sydney, (2) 9:9; pi. 1, figs. 1 - 3 . - T e r r a t y p i c a : Wagga Wagga, New South Wales, Australia. H o l o t y p e : Australian Museum, Sydney R669. 1966 Ramphotyphlops Obatillus - ROBB, Ann. Mag. nat. Hist. London, (13) 9:676. 1974 Typhlina batilla - MCDOWELL, J. Herpetol., Athens, 8 (1): 47. D i s c u s s i o n : Kinghorn (1956:61). Range:
New South Wales, Australia. Known only from type locality.
39
Typhlina bituberculata (PETERS) 1863 Onychocephalus bituberculatus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1863: 233. - T e r r a t y p i c a : near Adelaide, South Australia. - H o l o t y p e : Z.M.B. 1893 Typhlops bituberculatus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:48. 1966 Ramphotyphlops bituberculatus - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1975 Typhlina bituberculata - COGGER, Rept. Amph. Australia: 349. D i s c u s s i o n : WAITE (1929:196). Range:
Most of Australia except the far northern areas.
Typhlina bramina (DAUDIN) 1803 Eryx braminus DAUDIN, Hist. Rept., 7:279. - T e r r a t y p i c a : Vizagapatam, India. — T y p e : based on Plate. 43 in RUSSELL, 1796, An account of Indian serpents collected on the coast of Coromandel. . .: 48. 1820 Tortrix russelii MERREM, Syst. Amphib.: 84. - T e r r a t y p i c a : Vizagapatam, India. - T y p e : also based an Russell, 1796. 1839 Typhlops russelii - SCHLEGEL, Abbild. Amphib.: 39.. 1845 Argyrophis truncatus GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 138. - T e r r a t y p i c a : Philippines. - S y n t y p e s : B.M.N.H. 1946.1.11.9-11. (3) 1845 Argyrophis bramicus GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 138 (lapsus for braminus). 1846 Onychocephalus capensis A. SMITH, Illustr. Zool. S. Afr., Rept.: pi. 51, fig. 3; pi. 54, figs. 9 - 1 6 . - T e r r a t y p i c a : Interior (?) Cape of Good Hope. S y n t y p e s : B.M.N.H. 65.5.4.74-76. 1860 Ophthalmidium tenue HALLOWELL, Proc. Acad. nat. Sei. Philadelphia, 1860:497. T e r r a t y p i c a : Hong Kong. - S y n t y p e s (3): not traced. 1863 Typhlops (Typhlops) inconspicuus JAN, Elenco sist. Ofidi.: 11. - Madagascar. M.H.N.P. 928 (nomen nudum). 1863 Typhlops (Typhlops) braminus - JAN, Elenco sist. Ofidi.: 11. 1863 Typhlops (Typhlops) accedens JAN, Elenco sist. Ofidi.: 12. - T e r r a t y p i c a : East Indies. — H o l o t y p e : M.S.N.M. - destroyed. 1882 Typhlops (Typhlops) euproctus BOETTGER, Zool. Anz., Leipzig, 5:479. - T e r r a t y p i c a : Nossi-Be, Madagascar. - H o l o t y p e : S.M.F. 16571. 1890 Typhlops beddomii BOULENGER, Fauna Brit. India, Rept. Batr.: 237. - T e r r a t y p i c a : Polghat and Anamallays, 2500 ft.; Kimedy Hills, Vizagapatam district; and Travancore Hills, 2500 - 3000 ft. - S y n t y p e s : B.M.N.H. 1946.1.10.69-72, 1946.1.11. 27-30, 1946.1.11.93-95, 1946.1.10.48, and M.C.Z. 22372. 1906 Typhlops limbricki ANNAN DALE, Mem. asiatic Soc. Bengal, Calcutta, 1 : 1 9 3 . T e r r a t y p i c a : Ramnad, South India. - H o l o t y p e : Indian Museum, Calcutta. 1906 Typhlops braminus vat. arenicola ANNANDALE, Mem. asiatic Soc. Bengal, Calcutta, 1:192. - T e r r a t y p i c a : Ramnad, Madura, South India. - S y n t y p e s : B.M.N.H. 1946.1.11.64 and 2 in the Indian Museum, Calcutta. 1909 Typhlops braminus var. pallidus WALL, J. Bombay nat. Hist. Soc., 19:609. - T e r r a t y p i c a : Dibrugarh, Upper Assam. - H o l o t y p e : not traced. 1909 Typhlops microcephalus WERNER, J.Ber. vaterl. Ver. Naturk. Württemberg, Stuttgart, 65:60. - T e r r a t y p i c a : Madagascar. - H o l o t y p e : not traced. 1910 Glauconia braueri STERNFELD, Mitt. Zool. Mus. Berlin, 5:69. - T e r r a t y p i c a : Bagamoyo, Tanzanzania. - H o l o t y p e : Z.M.B. 1910 Typhlops braueri - BOULENGER, Zool. Ree. (1910), Rept. Batr.: 29. 1919 Typhlops fletcheri WALL, J. Bombay nat. Hist. Soc., 26:556, figs. - T e r r a t y p i c a : Nilgiris Hills, India. - T y p e : not traced. 1943 Typhlops beddomei - SMITH, Fauna Brit. India, Rept. Amphib., 3:45, 54. 1974 Typhlina (?) bramina - MCDOWELL, J. Herpetol., Athens, 8 (1): 21. D i s c u s s i o n : ROUX-ESTEVE (1974:28).
40 R a n g e : Borneo; Nias; Sumatra; Riouw Archipelago; Bangka; Java; Bali; Madura; Flores; Lomblen; Sumba; Timor; Celebes; Saleyer; Butung; Ambon; Ceram; Ternate; Halmahera; Kei Islands; Arn Islands; Lombok; Maldive and Laccadive Islands; Christmas Island; Andamans; Formosa; Hongkong; Sumbawa; Billiton; Nicobars; Hainan; Okinawa; Riu Kius; New Guinea; Saipan; Postitjon Island; Nila; Kai Island; Saparua; Buru; Salajar; New Britian; Bougainville Island; Singapore; Guam; Pelau; Hawaiian Islands; south China; Ceylon; India; Pakistan; Thailand; Burma; Indochina; Malay Peninsula; Nepal; Straits Settlements; Moluccas; vicinity of Darwin, north Australia; Philippines; Arabia; Seychelles; Madagascar; Comoro Islands; Mauritius; Zanzibar; coastal Tanzania; Mozambique; Somalia; South Africa(?); Dahomey; Togo; Ivory Coast; Cameroon; Iran; western Mexico. Comment:
MCDOWELL ( 1 9 7 4 : 2 2 ) presents evidence that this is an all-
female species. Typhlina broomi (BOULENGER) 1898 Typhlops broomi BOULENGER, Ann. Mag. nat. Hist., London, (7) 2 : 4 1 4 . T e r r a t y p i c a : Maldiva, near Cairns, northern Queensland, Australia. - H o l o t y p e : B.M.N.H. 1946.1.10.64. 1966 Ramphotyphlops broomi - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina broomi - MCDOWELL, J. Herpetol., Athens, 8 (1): 31. Discussion:
WAITE (1918:25); KINGHORN (1956:55); and COGGER (1975: 349).
R a n g e : Northern and eastern Australia. Typhlina cumingii (GRAY) 1845 Onychophis cumingii GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 133. - T e r r a t y p i c a : Philippines and "Indian Ocean". - S y n t y p e s : B.M.N.H. 1946.1.11.19-20, 1946.1.10.83. 1886 Onychocephalus cumingii - BOETTGER, Ber. senckenb. naturf. Ges., Frankfurt am Main, 1886:104. 1893 Typhlops cumingii - BOULENGER, Cat. Snakes Brit. Mus. (nat. Hist.), 1:51, pi. 3, fig. 4. 1919 Typhlops rugosa TAYLOR, Philippine J. Sei., Manila, 14:109. - T e r r a t y p i c a : Bunawan, Agusan, Mindanao, Philippines. - H o 1 o t y p e : C.M. 2665. 1919 Typhlops longicauda TAYLOR, Philippine J. Sei., Manila, 14:108. - T e r r a t y p i c a : Bunawan, Agusan, Mindanao, Philippines. - H o l o t y p e : C.M. 2671. 1922 Typhlops dendrophis TAYLOR, Philippine J. Sei., Manila, 16:60. - T e r r a t y p i c a : Bunawan, Agusan, Mindanao, Philippines. - H o l o t y p e : C.M. 2668. 1922 Typhlops mindanensis TAYLOR, Philippine J. Sei., Manila, 16:65. - T e r r a t y p i c a : Bunawan, Agusan, Mindanao, Philippines. - H o l o t y p e : C.M. 2667. 1974 Typhlina cumingii - MCDOWELL, J. Herpetol., Athens, 8 (1): 7.
R a n g e : Mindanao, Polillo, Negros, Bohol, and Marinduque Islands, Philippine Islands. Typhlina diversa (WAITE) 1894 Typhlops diversus WAITE, Proc. Linn. Soc. New South Wales, Sydney, (2) 9 : 1 0 ; pi. 1, figs. 4 - 6 . - T e r r a t y p i c a : Mowen (= Morven), Central Railway, Queensland, Australia. - H o l o t y p e : Queensland Museum J2943 (formerly D4432).
41 1914 Typhlops ammodytes MONTAGUE, Proc. Zool. Soc. London, 1914:642; figs. 8 - 1 0 . - T e r r a t y p i c a : Hermite Island, Monte Bello Islands, Western Australia. - H o l o t y p e : B.M.N.H. 1946.1.11.23. 1966 Ramphotyphlops diversus - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina diversa - MCDOWELL, J. Herpetol., Athens, 8 (1): 21. Discussion:
WAITE (1918: 31); KINGHORN (1956:63); COGGER (1975: 350).
R a n g e : Northwest Australia through the Northern Territory and the whole of Queensland. Typhlina endotera (WAITE) 1918 Typhlops endoterus WAITE, Ree. S. Austra. Mus., Adelaide, 1: 32, fig. 24. T e r r a t y p i c a : Hermannsburg, central Australia. - H o l o t y p e : South Australian Museum, Adelaide R88. 1919 Typhlops leonhardii STERNFELD, Mitt, senckenb. naturf. Ges., Frankfurt am Main, 1 (3): 77. - T e r r a t y p i c a : Hermannsburg, Finke River, central Australia. — L e c t o t y p e : S.M.F. 16666 (MERTENS, 1967:88). 1966 Ramphotyphlops endoterus - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina endotera - MCDOWELL, J. Herpetol., Athens, 8 (1): 21. Discussion:
KINGHORN (1966:64); COGGER (1975 :350).
R a n g e : Central Australia and Warburton Range, Western Australia. Typhlina erycina (WERNER) 1901 Typhlops erycinus WERNER, Verh. zool.-bot. Ges. Wien, 5 1 : 6 1 1 , fig. - T e r r a t y p i c a : German New Guinea. - H o l o t y p e : N.M.W. 1930 Typhlops iridescens DE JONG, Nova Guinea, 1 5 : 4 0 5 . - T e r r a t y p i c a : Albatrosbivak on the Mamberamo River, North Dutch New Guinea. - H o l o t y p e : Buitenzorg Museum. 1974 Typhlina erycina - MCDOWELL, J. Herpetol., Athens, 8 (1): 38.
R a n g e : Northern New Guinea, from Alablrosbivak on the Mameramo River, West Irian, eastward to Lae and Finschhafen in Morobe District, Papua New Guinea. Typhlina exocoeti (BOULENGER), New Combination 1887 Typhlops exocoeti BOULENGER, Proc. zool. Soc. London, 1887:517. t y p i c a : Christmas Island, Indian Ocean. - S y n t y p e s : B.M.N.H. 1946.1.11.78-79.
Terra
R a n g e : Christmas Island, Indian Ocean. Typhlina flaviventer (PETERS) 1865 Typhlops flaviventer PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1865 :271. T e r r a t y p i c a : Ternate Island, Moluccas. - H o l o t y p e : Z.M.B. 1881 Typhlops depressus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1881:217. T e r r a t y p i c a : Duke of York Island. - H o l o t y p e : Z.M.B. 1887 Typhlops aluensis BOULENGER, Proc. zool. Soc. London, 1887:336. - T e r r a t y p i c a : Alu, Shortland Group, Solomon Islands. - H o l o t y p e : B.M.N.H. 1946.1.11.57.
42 1898 Typhlops philococos WERNER, Zool. Anz., Leipzig, 21:553. - T e r r a t y p i c a : Ralum, Bismarck Archipelago, - H o l o t y p e : not traced. 1918 Typhlopsphilococcus WAITE (lapsus), Ree. S. Austral. Mus., Adelaide, 1: 35. 1933 Typhlops buehleri HEDIGER, Verh. naturf. Ges. Basel, 44 (2): 14. - T e r r a t y p i c a : Lorengau, Manus, Admiralty Islands. - H o l o t y p e : N.M.B. 1934 Typhlops supranasalis BRONGERSMA, Zool. Mededel., Leiden, 17:190. - T e r r a t y p i c a : Salawatti Island. - H o l o t y p e : R.M.N.H. 6291. 1934 Typhlops similis BRONGERSMA, Zool. Mededel., Leiden, 17:190. - T e r r a t y p i c a : Dore (= Manokwari), New Guinea. - H o l o t y p e : R.M.N.H. 6295. 1951 Typhlopsalvensis BRATTSTROM & SCHWENKMEYER (error typographies), Herpetologica, San Diego, 7 (4): 195. 1966 Ramphotyphlopsaluensis - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1966 Ramphotyphlops buehleri - ROBB, Ann. Mag. nat. Hist., London, (13) 9: 676. 1974 Typhlina flaviventer - MCDOWELL, J. Herpetol., Athens, 8 (1): 25. R a n g e : Ternate; Halmahera; Morotai; Batjan; Batanta; Salawati; the Vogelkop and Onin Peninsula of New Guinea; Manus; N e w Britian; Duke o f York Island; Nissan Atoll; perhaps all of the Solomon Islands, and at least Bougainville, Shortland Group (Alu), Santa Isabel, N e w Georgia Group (Kolombangara, Ganongga, Narovo), Florida Group (Tulagi), Malaita, Guadalcanal, and San Cristobal; and Vuna Pi in the Fiji Islands. Typhlina grypha (WAITE) 1918 Typhlops gryphus WAITE, Ree. S. Austral. Mus., Adelaide, 1:17, figs. - T e r r a t y p i c a : Marble Bar, northwestern Australia and Gregory Downs, Queensland, Australia.- H o l o t y p e : National Museum of Victoria D12351 (formerly R7102). 1966 Ramphotyphlops gryphus - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1974 Typhina grypha MCDOWELL (misspelling), J. Herpetol., Athens, 8 (1): 45. D i s c u s s i o n : KINGHORN (1956:47); COGGER (1975:350). Range:
Known from widely scattered localities across northern Australia.
Typhlina guentheri (PETERS) 1865 Typhlops (Onychocephalus) Güntheri PETERS, M.Ber. k. preuii. Akad. Wiss. Berlin, 1865:259; pi., fig. 1. - T e r r a t y p i c a : Benguella, West Africa (error). H o l o t y p e : B.M.N.H. 1946.1.10.88. 1893 Typhlops guentheri - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:20. Type locality corrected to northern Australia. 1895 Typhlops nigricauda BOULENGER, Proc. zool. Soc. London, 1895:867; pi. 49, fig. 1. - T e r r a t y p i c a : Daly River and Union Town, northern Australia. S y n t y p e s: B.M.N.H. 1946.1.11.13 and ZoologiskMuseum, Oslo K2011. 1966 Ramphotyphlops guentheri - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1974 Typhlina guentheri - MCDOWELL, J. Herpetol., Athens, 8 (1): 32. D i s c u s s i o n : KINGHORN (1956:52); COGGER (1975:350). Range:
Northwest and northern Australia.
Typhlina leptosoma (ROBB) 1972 Ramphotyphlops leptosoma ROBB, J. roy. Soc. West. Australia, Perth, 55: 39. T e r r a t y p i c a : The Loop, lower Murchison River, Western Australia. - H o l o t y p e : W.A.M. R29623. 1975 Typhlina leptosoma - COGGER, Amph. Rept. Australia: 351. Range:
Known only from the type locality.
43 Typhlina leucoprocta (BOULENGER) 1889 Typhlops leucoproctus BOULENGER, Ann. Mag. nat. Hist., London. (6) 5 : 361. T e r r a t y p i c a : Fly River, southern New Guinea; Murray Island, Torres Straits; and Queensland. - S y n t y p e s : B.M.N.H. 1946.1.10.58-60, 1946.1.11.97, and 1946.1.11.85. 1974 Typhlina (?) leucoprocta - MCDOWELL, J. Herpetol, Athens, 8 (1): 29. D i s c u s s i o n : COGGER (1975: 351). R a n g e : F l y River ( p r e s u m a b l y t h e region of t h e m o u t h ) a n d M a t a in Western District, P a u p a N e w G u i n e a ; M u r r a y I s l a n d , T o r r e s S t r a i t ; a n d n o r t h e r n C a p e Y o r k P e n i n s u l a , a n d e a s t e r n c o a s t o f Q u e e n s l a n d in A u s t r a l i a .
Typhlina ligata (PETERS) 1879 Typhlops ligatus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1879:775, fig. 3. T e r r a t y p i c a : Port Mackay, Queensland, Australia. - H o l o t y p e : Z.M.B. 6231. 1892 Typhlops curtus DOUGLAS-OGILBY, Ree. Austral. Mus., Sydney, 2 : 2 3 . - T e r r a t y p i c a : Gulf of Carpentaria, northern Australia. - H o l o t y p e : not traced. 1966 Ramphotyphlops ligatus - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina ligata - MCDOWELL, J. Herpetol., Athens, 8 (1): 29. D i s c u s s i o n : WAITE (1918:21); and COGGER (1975: 351). Range:
Q u e e n s l a n d , N e w S o u t h Wales, V i c t o r i a , a n d Western A u s t r a l i a .
Typhlina lineata (SCHLEGEL) 1827 Typhlops lineata BOIE, Isis (OKEN), Leipzig, 1827:563 (nomen nudum). - T e r r a t y p i c a : Java. - S y n t y p e s : R.M.N.H. 3696 (2). 1830 Acontias lineatus - WAGLER, Nat. Syst. Amph.: 196. Nomen nudum. 1831 Typhlops lineatus Acontias GRAY in GRIFFITH, Animal Kingdom, 9 (Synops. Spec.): 77. - Nomen nudum. 1839 Typhlops lineatus - SCHLEGEL, Abbild. Amphib.: 39; pl. 32, figs. 32 - 34. 1844 Pilidion lineatum - DUMÉRIL & BIBRON, Erpétol. gén., 6 : 2 5 9 . 1845 Typhlinalis lineatum - GRAY, Cat. Lizards. Brit. Mus. (nat. Hist.): 134. 1863 Typhlops (Typhlina) lineatus - JAN, Elenco sist. Ofidi: 10. 1864 Typhlina lineata - GÜNTHER, Rept. Amph. Brit. India: 171; pl. 16, fig. B. 1896 Typhlops lineatus var. sumatranus WERNER, Verh. zool.-bot. Ges. Wien, 4 6 : 1 3 . T e r r a t y p i c a : Sumatra. - H o l o t y p e : N.M.W. 1896 Typhlops lineatus var. numatrana BOULENGER (error typographies), Zool. Ree. (1896), Rept. Batr: 24. 1966 Ramphotyphlops lineatus - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1974 Typhlina lineata - MCDOWELL, J. Herpetol., Athens, (8 (1): 20. R a n g e : Nias; S u m a t r a ; B o r n e o ; Java; Malay Peninsula; S i n g a p o r e ; Thail a n d ; H o n g k o n g ; I n d o c h i n a ; s o u t h C h i n a (?).
Typhlina minima (KINGHORN) 1929 Typhlops minimus KINGHORN, Ree. Austral. Mus., Sydney, 17 (4): 190 - 191, fig. 1. - T e r r a t y p i c a : Groote Eylandt, Gulf of Carpentaria, northern Australia. - H o l o t y p e : Australian Museum, Sydney R9692.
44 1966 Ramphotyphiops minimus - ROBB, Ann. Mus. nat. Hist., London, (13) 9 : 6 7 6 . 1975 Typhlina minima - COGGER, Amph. Rept. Australia: 352. Discussion:
KINGHORN (1956:51).
R a n g e : Northern Australia. Known only from the type locality.
Typhlina multilineata (SCHLEGEL) 1839 Typhlops multilineatus SCHLEGEL, Abbild. Amphib.,: 40; pi. 32, figs. 39 - 42. T e r r a t y p i c a : west coast of New Guinea. - H o l o t y p e : M.H.N.P. 1067. 1843 Rhinotyphlops (Ramphotyphiops) multilineatus - FITZINGER, Syst. Rept., 1:24. 1844 Onychocephalus multilineatus - DUMÉRIL & BIBRON, Erpétol. gén., 6 : 2 7 6 . 1845 Onychophis multicarinatus GRAY (lapsus), Cat. Lizards Brit. Mus. (Nat. Hist.): 134. 1966 Ramphotyphiops multilineatus - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina multilineata - MCDOWELL, J. Herpetol., Athens, 8 (1): 41.
R a n g e : West Irian, New Guinea; Kei Islands; Salawati.
Typhlina nigroterminata (PARKER) 1931
Typhlops nigroterminatus PARKER, Ann. Mag. nat. Hist., London, (10 8 : 6 0 4 , text fig. - T e r r a t y p i c a : Roebuck Bay, northwest Australia. - H o l o t y p e : B.M.N.H. 1946.1.11.58. 1966 Ramphotyphiops nigroterminatus - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1974 Typhlina nigroterminata
Discussion:
- MCDOWELL, J. Herpetol., Athens, 8 (1): 31.
KINGHORN (1956:54); and COGGER (1975:352).
R a n g e : Northwestern Australia.
Typhlina olivacea (GRAY) 1845 Onychophis olivaceus GRAY, Cat. Lizards. Brit. Mus. (Nat. Hist.): 133. - T e r r a t y p i c a : Philippines. - H o l o t y p e : B.M.N.H. 1946. 1.10.57. 1861 Onychocephalus (Onychophis) olivaceus - PETERS, M.Ber. k. preufc. Akad. Wiss. Berlin, 1861:684. 1886 Onychocephalus olivaceus - BOETTGER, Ber. senckenb. naturi. Ges., Frankfurt am Main, 1886:105. 1893 Typhlops olivaceus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1 : 1 3 . 1918 Typhlops suluensis TAYLOR, Philippine J. Sci., Manila, 13 (d) (5): 257. - T e r r a t y p i c a : Bubuan Island, Sulu Archipelago, Philippines. - H o l o t y p e : Philippine Bureau of Science, Manila 2001. 1921 Typhlops ligorostris SMITH & PROCTER, Ann. Mag. nat. Hist., London, (9) 7 : 35 3. - T e r r a t y p i c a : Teloeti Bay, southern Ceram. - H o l o t y p e : B.M.N.H. 1946.1.11.21. 1921 Typhlops olivaceus olivaceus - BARBOUR, Proc. New England Zool. Club, Boston, 7:107. 1974 Typhlina olivacea - MCDOWELL, J. Herpetol., Athens, 8 (1): 43.
R a n g e : Samar and Bubuan, Philippine Islands; Borneo and Sangihe Islands; Ceram; Ambon; and Mysool. McDowell (1974:43) considers a record for Australia as "probably erroneous".
45 Typhlina pinguis (WAITE) 1897 Typhlopspinguis WAITE, Trans, roy. Soc. S. Austral., Adelaide, 2 1 : 2 5 , pi. 3. T e r r a t y p i c a : South Australia. - H o l o t y p e : South Australian Museum, Adelaide, R 803. 1917 Typhlops opisthopachys WERNER, Jb. Hamburg, wiss. Anst., 34: 34. - T e r r a t y p i c a : Tanga, Tanzania (error). - H o l o t y p e : Zoologisches Museum, Hamburg; destroyed during World War II. 1966 Ramphotyphlops pinguis - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1975 Typhlina pinguis - COGGER, Amph. Rept. Australia: 353. D i s c u s s i o n : WAITE (1918:24, and 1929:196). Range:
S o u t h e r n V i c t o r i a a n d N e w S o u t h Wales t o s o u t h w e s t e r n A u s t r a l i a .
Typhlina polygrammica (SCHLEGEL) 1839 Typhlops polygrammicus SCHLEGEL, Abbild. Amphib.: 40; pi. 32, figs. 35 - 38. T e r r a t y p i c a : Timor. - H o l o t y p e : R.M.N.H. 3712. Discussion:
MCDOWELL (1974: 35).
R a n g e : S u m b a w a I s l a n d ; Flores; L o m b o k ; S u m b a I s l a n d ; T i m o r ; s o u t h e r n N e w G u i n e a ; D a r u I s l a n d ; M u r r a y I s l a n d ; D u n k I s l a n d ; e a s t e r n N e w S o u t h Wales, Queensland, and Victoria, Australia.
T y p h l i n a 1843 1845 1863 1889 1930 1948 1966 1974
p o l y g r a m m i c a
p o l y g r a m m i c a
(SCHLEGEL)
Typhlops (Pseudotyphlops) polygrammicus - FITZINGER, Syst. Rept., 1:24. Argyophis polygrammicus - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 138. Typhlops (Typhlops) polygrammicus - JAN, Elenco sist. Ofidi: 11. Typhlops torresianus BOULENGER, Ann. Mag. nat. Hist., London, (6) 4 : 362. T e r r a t y p i c a : Murray Island, Torres Straits. - S y n t y p e s : B.M.N.H. 1946.1.11.48-49. Typhlops soensis DE JONG, Treubia, Buitenzorg, 12 (1): 117. - T e r r a t y p i c a : Soe, south Timor. - S y n t y p e s : 9 in Buitzenzorg Museum. Typhlops polygrammicus polygrammicus - LO VE RIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 101: 320. Ramphotyphlops torresianus - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. Typhlina polygrammica - MCDOWELL, J. Herpetol., Athens, 8 (1): 35.
R a n g e : T i m o r ; s o u t h e r n N e w G u i n e a ( t h e u p p e r D i d u l River a n d t h e F l y River regions); D a r u I s l a n d ; M u r r a y I s l a n d , a n d D u n k Island.
T y p h l i n a
p o l y g r a m m i c a
b r o n g e r s m a i
(MERTENS)
1929 Typhlops florensis brongersmai MERTENS, Treubia, Buitenzorg, 11:295. - T e r r a t y p i c a : Mao Marru, 450 m., Sumba Island. - H o l o t y p e : Zoologisch Museum, Amsterdam, 11179. 195 3 Typhlops polygrammicus brongersmai - FORCART, Verh. naturf. Ges. Basel, 6 4 : 356. Range:
K n o w n o n l y f r o m S u m b a Island, I n d o n e s i a .
46 Typhlina
polygrammica
elberti
(Roux)
1911 Typhlops elberti ROUX, Zool. Jahrb., Syst., Jena, 30:499. - T e r r a t y p i c a : Sadjang, Lombok. - H o 1 o t y p e : S.M.F. 16669. 1929 Typhlops florensis elberti - MERTENS, Treubia, Buitenzorg, 11:295. 1953 Typhlopspolygrammicus elberti - FORCART, Verh. naturi. Ges. Basel, 64: 356. Range:
Typhlina
Known only from the island of Lombok, Indonesia.
polygrammica
florensis
(BOULENGER)
1897 Typhlops florensis BOULENGER, Ann. Mag. nat. Hist., London, (6) 19:505. T e r r a t y p i c a : Flores. - H o l o t y p e : B.M.N.H. 1946.1,10.47. 1912 Typhlops floris BARBOUR (lapsus), Mem. Mus. comp. Zool., Cambridge (Massachusetts), 44:189. 1927 Typhlops florensis florensis - MERTENS, Senckenbergiana, Frankfurt am Main 9:240. 1953 Typhlops polygrammicus florensis - FORCART, Verh. naturi. Ges. Basel, 64: 356. Range:
Typhlina
Known only from the island of Flores, Indonesia.
polygrammica
nigrescens
(GRAY)
1845 Anilios nigrescens GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 135. - T e r r a t y p i c a : Parramatta, New South Wales, Australia. - L e c t o t y p e : B.M.N.H. 1946.1.11.17. 1863 Typhlops (Typhlops) nigrescens (Anilios) - JAN, Sist. degli Ofidi: 11. 1863 Typhlops rueppelli JAN, Elenco sist. Ophidi : 11. - T e r r a t y p i c a : Sydney, Australia. - L e c t o t y p e : S.M.F. 16617 (MERTENS, 1967:89). 1864 Typhlops Temminckii JAN, Iconogr. gén. Ophid., 1 (3. livr.): 3; pis. 4, 5, fig. 6. T e r r a t y p i c a : Unknown. - H o l o t y p e : R.M.N.H. 1865 Typhlops nigrescens JAN, Elenco sist. Óphidi: 11. - T e r r a t y p i c a : Sydney, 1865 Typhlops (Typhlops)Rüppelli - JAN, Iconogr. gén. Ophid., 1 (9. livr.): 5; pi. 1, fig. 2. 1889 Typhlops regime BOULENGER, Ann. Mag. nat. Hist., London, (6) 4: 362. T e r r a t y p i c a : Queensland, Australia. - S y n t y p e s : B.M.N.H. 1946.1.11.81-83. 1927 Typhlops polygrammicus nigrescens - SMITH, Proc. zool. Soc. London, 1927:219. 1966 Ramphotyphlops nigrescens - ROBB, Ann. Mag. nat. Hist., London, (13) 9:676. 1974 Typhlina polygrammica - MCDOWELL, J. Herpetol., Athens, 8 (1): 35. Range:
Typhlina
Eastern New South Wales, Queensland, and Victoria, Australia.
polygrammica
undecimlineata
(MERTENS)
1927 Typhlops florensis undecimlineatus MERTENS, Senckcnbergiana, Frankfurt am Main, 9:239. - T e r r a t y p i c a : Semongkat Atas, 400 m., Sumbawa, Sunda Isles. - H o l o t y p e : S.M.F. 22092. 1953 Typhlops polygrammicus undecimlineatus - FORCART, Verh. naturf. Ges. Basel, 64:356. Range:
Sumbawa Island, Indonesia.
47 Typhlina p r o x i m a (WAITE) 1885 Typhlops nigrescens MCCOY, Prodr. Zool. Victoria, Dec. 11, pi. 103. - T e r r a t y p i c a : unknown. - T y p e : not traced. - Preoccupied by Typhlops nigrescens GRAY, 1845. 1893 Typhlops proximus WAITE, Ree. Austral. Mus., Sydney, 2:60; pi. 15, figs. 1 - 4. T e r r a t y p i c a : New South Wales, Australia. - H o l o t y p e : Australian Museum, Sydney 6411. 1966 Ramphotyphlopsproximus - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina proxima - MCDOWELL, J. Herpetol., Athens, 8 (1): 35. Discussion:
KINGHORN (1956:48); WAITE (1918:18); COGGER (1975: 353).
R a n g e : North and central Queensland through New South.Wales to northern Victoria, Australia.
Typhlina subocularis (WAITE) 1897 Typhlops subocularis WAITE, Ree. Austral. Mus., Sydney, 3 (3): 69; figs 1 - 3. T e r r a t y p i c a : Duke of York Island, Bismarck Archipelago. — H o l o t y p e : Australian Museum, Sydney R2202. 1918 Typhlops infralabialis WAITE, Ree. S. Austral. Mus., Adelaide, 1 (1): 35 - 36; fig. 25. - T e r r a t y p i c a : Malaita Island, Solomon Islands. - H o l o t y p e : Australian Museum, Sydney R4609. 1921 Typhlops subucularis WERNER (error typographicus), Arch. Naturgesch., Berlin, 87A: 327. 1939 Typhlops solomonis PARKER, Bull. Mus. roy. Hist. nat. Belgique, Bruxelles, 15 (60): 4 - 5 . - T e r r a t y p i c a : Buin, Bougainville Island. - H o l o t y p e : I.R.S.N. Reg. 2029,1.G. 12375. 1948 Typhlops bergi PETERS, Occ. Pap. Mus. Zool. Univ. Michigan, Ann Arbor, 508: 1 - 3 . - T e r r a t y p i c a : Segi Point, New Georgia Island, Solomon Islands. H o l o t y p e : U.M.M.Z. 95445. 1948 Typhlops keasti KINGHORN, Ree. Austral. Mus., Sydney, 22 (1): 67 - 69, fig. 1. - T e r r a t y p i c a : Jacquinot Bay, New Britain. - H o l o t y p e : Australian Museum, Sydney R12856. 1951 Typhlops adamsi TANNER, Great Basin Naturalist, Provo, 11:64 - 6 5 . - T e r r a t y p i c a : Nalimbiu River, Guadalcanal Island. - H o l o t y p e : University of California, Berkeley, Museum of Vertebrate Zoology 40753. 1966 Ramphotyphlops solominis ROBB (error typographicus), Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1966 Ramphotyphlops keasti - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina subocularis - MCDOWELL, J. Herpetol., Athens, 8 (1): 49.
R a n g e : Umboi (or Rooke) Island, New Britain, Duke of York Island, New Ireland, all in the Bismarck Archipelago; Bougainville, New Georgia, Malaita, and Guadalcanal in the Solomon Islands.
T y p h l i n a tovelli (LOVERIDGE) 1945 Typhlops tovelli LOVERIDGE, Proc. biol. Soc. Washington, 58: 111. - T e r r a t y p i c a : Koonowarra Sports Ground, about 5 miles south of Darwin, Northern Territory, Australia. - H o l o t y p e : M.C.Z. 48845. 1975 Typhlina tovelli - COGGER, Amph. Rept. Australia: 353.
R a n g e : Known only from the vicinity of Darwin, northern Australia.
48 Typhlina unguirostris (PETERS) 1867 1879 1893 1966 1975
Typhlops (Onychocephalus) unguirostris PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1867:708; pi., fig. 3. - T e r r a t y p i c a : Rockhampton, Queensland, Australia. - H o l o t y p e : Z.M.B. Typhlops curvirostris PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1879:776; pi., fig. 3. - T e r r a t y p i c a : Port Bowen, northern Australia. - H o l o t y p e : Z.M.B. 7525. Typhlops unguirostris - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:49. Ramphotyphlops unguirostris - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . Typhlina unguirostris - COGGER, Amph. Rept. Australia: 354.
Discussion:
WAITE (1918:20, 1929:195); KINGHORN (1956:50).
R a n g e : K n o w n f r o m w i d e l y s c a t t e r e d localities t h r o u g h o u t e a s t e r n t w o thirds of Australia.
Typhlina wiedii (PETERS) 1867 Typhlops wiedii PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1867:24. T e r r a t y p i c a : Brisbane, Queensland, Australia. - H o l o t y p e : Z.M.B. 1966 Ramphotyphlops wiedii - ROBB, Ann. Mag. nat. Hist., London, (13) 9 : 6 7 6 . 1974 Typhlina wiedii - MCDOWELL, J. Herpetol., Athens, 8 (1): 31. D I S C U S S I O N : WAITE (1918:26); KINGHORN (1956:57); COGGER (1975: 354). R a n g e : M o s t o f A u s t r a l i a , i n c l u d i n g Q u e e n s l a n d , N e w S o u t h Wales, V i c t o r i a , f r o m C a p e Y o r k across n o r t h A u s t r a l i a t o n o r t h w e s t a n d s o u t h w e s t Australia.
Typhlina willeyi (BOULENGER) 1900 Typhlops willeyi BOULENGER in WILLEY, Zool. Results, 2 : 6 0 3 - 604, figs. T e r r a t y p i c a : Lifu, Loyalty Islands. - H o l o t y p e : B.M.N.H. 1946.1.10.80. 1948 Typhlops becki TANNER, Great Basin Naturalist, Provo, 9 (1 - 2): 12. - T e r r a t y p i c a : Tenaru River, Guadalcanal Island. - H o l o t y p e : Brigham Young University 7448. 1974 Typhlina willeyi - MCDOWELL, J. Herpetol., Athens, 8 (1): 47. R a n g e : G u a d a l c a n a l I s l a n d , S o l o m o n Islands; Mare a n d L i f u in t h e L o y a l t y Islands.
Typhlina yirrikalae (KINGHORN) 1942 Typhlops yirrikalae KINGHORN, Ree. Austral. Mus., Sydney, 21 (2): 118, fig. 1. T e r r a t y p i c a : Yirrkalla Mission, Caledon Bay, Arnhem Land, Northern Territory, Australia. - H o l o t y p e : Australian Museum, Sydney R12381. 1963 Typhlops yirrkalae WORRELL (error typographies), Rept. Australia: 94. 1975 Typhlina yirrikalae - COGGER, Amph. Rept. Australia: 354. Discussion: Range:
KINGHORN (1956:51).
Northern Australia. K n o w n only f r o m the type locality.
49 T y p h l o p s OPPEL 1811 Typhlops OPPEL, Ordn. Fam. Gatt. Rept.: 54 - 5 5 . - S p e c i e s t y p i c a (by subsequent designation: FITZINGER 1843:24): Anguis lumbricalis LINNAEUS = Typhlops lumbricalis (LINNAEUS). 1843 Aspidorhynchus FITZINGER, Syst. Rept., 1:24. - S p e c i e s t y p i c a (by original designation): Typhlops eschrichtii SCHLEGEL = Typhlops punctatus (LEACH). 1843 Gerrhopilus FITZINGER, Syst. Rept., 1:24. - S p e c i e s t y p i c a (by original designation): Typhlops ater SCHLEGEL. 1844 Ophthalmidium DUMERIL & BIBRON, Erpetol. gen., 6 : 2 6 2 . - S p e c i e s t y p i c a (by subsequent designation: PETERS, 1881:70): Ophthalmidium longissiMUM DUMERIL & BIBRON = ?. 1845 Anilios GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.),: 1 3 5 . - S p e c i e s t y p i c a (by subsequent designation: LOVERIDGE, 1957:240): Anilos leachii GRAY = Typhlops lumbricalis (LINNAEUS). 1845 Argyrophis GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 136. - S p e c i e s t y p i c a (by subsequent designation: M. A. SMITH: 1943:43): Argyrophis bicolor GRAY = Typhlops diardi SCHLEGEL. 1845 Meditoria GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 139. - S p e c i e s t y p i c a (by monotypy): Meditoria nasuta GRAY = Typhlops lumbricalis (LINNAEUS). 1861 Diaphorotyphlops JAN, Arch. Zool. Anat. Fisiol., Genova, 1:185. - S p e c i e s t y p i c a (by subsequent designation: PETERS, 1881:70): Diaphorotyphlops disparity JAN = ? 1881 Grypotyphlops PETERS, S.Ber. Ges. naturf. Freunde Berlin, 1 8 8 1 : 7 0 . - S p e c i e s t y p i c a (by original designation): Onychocephalus acutus DUMERIL & BIBRON = Typhlops acutus (DUMERIL & BIBRON). 1893 Gryptotyphlops BOULENGER (lapsus), Cat. Snakes Brit. Mus. (Nat. Hist.), 1:56.
R a n g e : Southeastern Europe, the Middle East, and all of tropical and subtropical Asia; most of Africa, Madagascar, and islands of the Indian Ocean; Philippines, Moluccas, and New Guinea; South America, tropical Central America, and the West Indies.
T y p h l o p s acutus (DUMÉRIL & BIBRON) 1844 Onychocephalus acutus DUMÉRIL & BIBRON, Erpétol. gén., 6 : 3 3 3 . - T e r r a t y p i c a : unknown. - H o l o t y p e : M.H.N.P., now lost. 1863 Onychocephalus westermanni LÜTKEN, Vidensk. Meddel. dansk naturhist. Foren., Stockholm, 14: 306; pl. 1, fig. 5. - T e r r a t y p i c a : Bengal. - H o l o t y p e : Universitetets Zoologiske Museum, Denmark R52183. 1845 Typhlops russellii GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 132. - T e r r a t y p i c a : India. - H o l o t y p e : B.M.N.H. 1946.1.11.70. - not Tortrix russeli MERREM, 1820. 1865 Typhlopsexcipiens JAN, Iconogr. gén. Ophid., 1 (9. livr.): 5; pl. 1, fig. 5. - T e r r a t y p i c a : "Indes Orientales". - H o l o t y p e : Collection de l'École technique (Realschule) de Cologne. 1875 Onychocephalus malabaricus BEDDOME (nomen nudum, manuscript name referred to by GÜNTHER, 1875), Proc. zool. Soc. London, 1875:224. - P a t r i a : Nellumbar, India. - B.M.N.H. 1881 Grypotyphlops acutus - PETERS, S.Ber. Ges. naturf. Freunde Berlin, 1881:70. 1890 Typhlops acutus - BOULENGER, Fauna Brit. India, Rept. Batr.: 242. 1893 Gryptotyphlops acutus - BOULENGER, (lapsus), Cat. Snakes Brit. Mus. (Nat. Hist.) 1:56.
50 1949 Typhlops acuta - CONSTABLE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 103 (2): 113. D i s c u s s i o n : M. SMITH (1943:56). R a n g e : India, south of the Ganges Basin and south of Rajputana, west to Baroda and east to Calcutta.
Typhlops andamanensis
STOLICZKA
1871 Typhlops andamanensis STOLICZKA, J. asiat. Soc. Bengal, Calcutta, 40:428; pi. 25, figs. 9 - 1 2 . - T e r r a t y p i c a : Andaman Islands. - H o l o t y p e : formerly in Indian Museum, Calcutta; now N.M.W. 15427. D i s c u s s i o n : M. A. SMITH (1943:56). Range:
Andaman Islands.
Typhlops angolensis (BOCAGE) 1866 Onychocephalus angolensis BOCAGE, Jörn. Sci. math. phys. nat., Lisboa, 1:46, 65. T e r r a t y p i c a : District of Duque of Braganca, Angola. - H o l o t y p e : M.B.L. T79.1134. 1887 Typhlops (Onychocephalus) congicus BOETTGER, Zool. Anz., Leipzig, 10: 650. T e r r a t y p i c a : Povo Netonna, near Banana, Zaire. - H o l o t y p e : S.M.F. 16638. 1893 Typhlops congicus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1: 40. 1910 Typhlops adolfi STERNFELD, Mitt. zool. Mus. Berlin, 5:70. - T e r r a t y p i c a : Fort Beni, Kivu, eastern Zaire. - H o l o t y p e : Z.M.B. 24897. 1916 Typhlops dubius CHABANAUD, Bull. Mus. nation. Hist, nat., Paris, 22: 364, figs. 1 - 3 . - T e r r a t y p i c a : Kivu volcanos, Zaire. - H o l o t y p e : M.H.N.P. 1916.213. 1933 Typhlops lestradei DE WITTE, Rev. Zool. Bot. afr., Bruxelles, 23:207, figs. 1 - 3. T e r r a t y p i c a : Ruhengeri, Ruanda. - H o l o t y p e : R.G.M.C. 7758. 1935 Typhlops kaimosae LO VE RIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 7 9 : 5 . - T e r r a t y p i c a : Kaimosi Forest, near Friends' Africa Mission, Kakamega district, Nyanza Province, Kenya. - H o l o t y p e : M.C.Z. 40060. 1942 Typhlops blanfordi lestradei - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 91:254, pi. 2. 1952 Typhlops ochraceus LAURENT, Rev. Zool. Bot. afr., Bruxelles, 46:269. - T e r r a t y p i c a : Mulenge, Lulenga, Zaire. - H o l o t y p e : R.G.M.C. 17636. 1956 Typhlops angolensis adolfi - LAURENT, Ann. Mus. Congo, Tervuren, Zool., (8°) 48:63, 349; figs. 5 - 6 . 1956 Typhlops angolensis dubius - LAURENT, Ann. Mus. Congo, Tervuren, Zool., (8°) 48:60. 1956 Typhlops angolensis irsaci LAURENT, Ann. Mus. Congo, Tervuren, Zool., (8°) 48: 57, 349; figs. 3 - 4, 6; pi. 7, fig. 4 . - T e r r a t y p i c a : Lwiro, 1750 m, Territory of Kabare, Kivu, Zaire. - H o l o t y p e : R.G.M.C. 17573. 1956 Typhlops angolensis polylepis LAURENT, Ann. Mus. Congo, Tervuren, Zool., (8°) 48:56, 367; fig. 6; pi. 7, fig. 3. - T e r r a t y p i c a : Kiandjo, 1850-1950 m, Territory of Mwenga, Kivu, Zaire. - H o l o t y p e : R.G.M.C. 17571. 1956 Typhlops angolensis angolensis - LAURENT, Ann. Mus. Congo, Tervuren, Zool., (8°) 48:55. 1960 Typhlops angolensis symoensi. LAURENT, Ann. Mus. Congo, Tervuren, Zool., (8°) 84:10. - T e r r a t y p i c a : Ngovi River, 1500-2000 m, Makobola, Fizi, Zaire. - H o l o t y p e : R.G.M.C. 21619. D i s c u s s i o n : LAURENT (1964:422); ROUX-ESTÈVE (1970:2185, 1974:47).
51 R a n g e : Zaire, Angola, coastal C a m e r o o n , s o u t h e r n Central African Republic, Gabon, Congo, western Kenya, Uganda, northern Tanzania, northern Zambia.
T y p h l o p s arenarius (GRANDIDIER) 1872 Onychocephalusarenarius GRANDIDIER, Ann. Sci. nat. (Zool.), Paris, (5) 15 (20): 9. - T e r r a t y p i c a : Morondava, Malagasy Republic. - H o l o t y p e : M.H.N.P. 9 5 - 2 1 6 . 1893 Typhlops boettgeri BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:39. T e r r a t y p i c a : Southwest Madagascar. - S y n t y p e s : B.M.N.H. 1946.1.11.98-99; 1946.1.12.1-4. 1893 Typhlops arenarius - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1:49. Discussion: Range:
GUIBE (1958:195).
Malagasy R e p u b l i c .
T y p h l o p s ater SCHLEGEL 1839 Typhlops ater SCHLEGEL, Abbild. Amphib.,: 39; pi. 32, figs. 29 - 31. - T e r r a t y p i c a : Java. - H o l o t y p e : R.M.N.H. 3714. 1843 Typhlops (Gerrhopilus) ater - FITZINGER, Syst. Rept., 1:24. 1845 Anilios ater - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 136. 1863 Typhlops (Typhlops) ater - JAN, Elenco sist. Ofidi: 11. 1934 Typhlops ater suturalis BRONGERSMA, Zool. Mededel., Leiden, 17:196, fig. 26. T e r r a t y p i c a : Andai, western New Guinea. - H o l o t y p e : R.M.N.H. 6297. 1934 Typhlops ater ater - BRONGERSMA, Zool. Mededel., Leiden, 17:195. 1950 Typhlops hedraea SAVAGE, Proc. California zool. Club, Palo Alto, 1 : 4 9 , fig. 1. T e r r a t y p i c a : 1500 ft. above Luzuriaga, ca. 6 miles southwest of Dumaguete, Oriental Negros, Philippines. - H o l o t y p e : Stanford University 12346. Discussion:
MCDOWELL (1974:16).
R a n g e : Java; T e r n a t e ; C e l e b e s ; H a l m a h e r a ; Waigeu; S a l a w a t i ; a n d A n d a i in West I r i a n . If Typhlops hedraeus is c o n s p e c i f i c , also N e g r o s I s l a n d s in t h e Philippines. A record for southern S o u t h V i e t n a m requires confirmation.
T y p h l o p s bibronii ( A . SMITH) 1846 Onychocephalus Bibronii A. SMITH, Illustr. Zool. S. Africa, Rept.: pi. 51, fig. 2; pi. 54, figs. 5 - 8 . - T e r r a t y p i c a : north of Latakoo (= Kuruman), South Africa. - S y n t y p e s : B.M.N.H. 1946.1.11.22; others not traced. 1887 Typhlops bibronii - BOULENGER, The Zoologist, London, (3) 11:174. Discussion:
ROUX-ESTEVE (1974:138).
R a n g e : Eastern Cape Province t o Natal, Swaziland and Transvaal, n o r t h e r n Cape Province and the Orange Free State, S o u t h Africa.
52 T y p h l o p s biminiensis RICHMOND 1955 Typhlops biminiensis RICHMOND, Amer. Mus. Novit., New York, 1734:2. T e r r a t y p i c a : South Bimini, near Nixon's Harbour, Bahamas. - H o l o t y p e : C.M. 32604. Discussion: Range:
THOMAS (1968:714).
C u b a , C a y m a n Brae, C a y Sal B a n k , w e s t e r n G r e a t B a h a m a s B a n k ,
and Great Inagua.
T y p h l o p s
b i m i n i e n s i s
b i m i n i e n s i s
RICHMOND
1968 Typhlops biminiensis biminiensis - RICHMOND, Copeia, New York & c., 1968 (4): 714; figs. 1 - 6; tab. 1. R a n g e : T h e B a h a m a s Islands; N o r t h a n d S o u t h Bimini, A n d r o s I s l a n d , N e w P r o v i d e n c e , E l b o w Cay (Cay Sal B a n k ) , a n d L i t t l e R a g g e d I s l a n d ; C u b a : R a n c h o L u n a n e a r C i e n f u e g o s a n d t h e east side o f t h e Bahia d e G u a n t a n a m o .
T y p h l o p s
b i m i n i e n s i s
epactia
THOMAS
1968 Typhlops biminiensis epactia THOMAS, Copeia, New York & c., 1968 (4): 715; 6 figs., tab. 1. - T e r r a t y p i c a : 5.4 miles east of West End, southern coastal platform, Cayman Brae, British West Indies. - H o i o t y p e : M.C.Z. 92048. Range:
T y p h l o p s
C a y m a n Brae; t o b e e x p e c t e d o n L i t t l e C a y m a n .
b i m i n i e n s i s
p a r a d o x u s
THOMAS
1968 Typhlops biminiensis paradoxus THOMAS, Copeia, New York & c., 1968 (4): 715; 6 figs.; tab. 1 . - T e r r a t y p i c a : 7.5 miles north of Mathew Town, Great Inagua, southeastern Bahamas. - H o l o t y p e : M.C.Z. 92993. Range:
Great Inagua Island, southeastern Bahamas.
T y p h l o p s bipartitus SAUVAGE 1879 Typhlops bipartitus SAUVAGE, Bull. Soc. philomath. Paris, (7) 3 : 5 9 . - T e r r a t y p i c a : Tidore Island, Moluccas. - S y n t y p e s : M.H.N.P. 1878-214 and 1878-215. Range:
T i d o r e Island, M o l u c c a s .
T y p h l o p s bisubocularis BOETTGER 1893 Typhlops bisubocularis BOETTGER, Zool. Anz., Leipzig, 16: 336. - TERRA p i c a : West Java. - H o l o t y p e : S.M.F. 16683. Range:
Western Java.
ty-
53 T y p h l o p s b o y l e i FITZSIMONS 1932 Typhlops boylei FITZSIMONS, Ann. Transvaal Mus., Pretoria, 15 (1): 38. - T e r r a t y p i c a : Gemsbok Pan, Ghansi district, West Kalahari. - H o l o t y p e : T.M. 4695. Discussion: Range: West A f r i c a .
ROUX-ESTEVE (1974:151).
Western edge o f t h e K a l a h a r i d e s e r t , n o r t h i n t o D a m a r a l a n d , S o u t h
T y p h l o p s brongersmianus VANZOLINI 1972 Typhlops brortgersmai VANZOLINI, Zool. Mededel., Leiden, 47 (3): 27. - T e r r a t y p i c a : Barra de Itaipe, Ilheus, Bahia, Brasil. - H o l o t y p e : Museu de Zoologia de Universidade de Sao Paulo, Brasil 5218. - Preoccupied by Typhlops florensis brongersmai MERTENS, 1920. 1976 Typhlops brongersmianus VANZOLINI (nomen novum pro Typhlops brongersmai VANZOLINI, 1972), Pap. avuls. Zool., Sào Paulo, 29 (24): 247. Range:
Bahia, Brasil. K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
T y p h l o p s c o e c a t u s JAN 1864 Typhlops (Typhlops) coecatus JAN, Iconogr. gen. Ophid., 1 (3. livr.): 3; pis. 4, 5, fig. 2. - T e r r a t y p i c a : Gold Coast. - H o l o t y p e : N.M.B. 310. 1893 Typhlops caecatus BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1: 32. Substitute name for T. coecatus JAN. Discussion: Range:
ROUX-ESTEVE (1974: 34).
G h a n a a n d Ivory C o a s t .
T y p h l o p s canlaonensis TAYLOR 1917 Typhlops canlaonensis TAYLOR, Philippine J. Sci., Manila, (D 11) 1917: 354. T e r r a t y p i c a : Canlaon volcano, 750 m., Negros Island, Philippines. - H o l o t y p e : C.M. 2666. Discussion:
TAYLOR (1922:55).
R a n g e : K n o w n o n l y f r o m t h e t y p e l o c a l i t y and C u e r n o s d e N e g r o s , s o u t h e r n N e g r o s , Philippines.
T y p h l o p s capitulatus RICHMOND 1964 Typhlops capitulatus RICHMOND, Breviora, Mus. comp. Zool., Cambridge (Massachusetts), 2 0 2 : 2 , figs. - T e r r a t y p i c a : Manneville, Haiti, at the northwest end of Lake Saumatre. - H o l o t y p e : M.C.Z. 62636. Discussion: Range:
THOMAS (1965a: 438).
Haiti and Gonave Island, Haiti.
54 T y p h l o p s
c a p i t u l a t u s
c a p i t u l a t u s
RICHMOND
1965 Typhlops capitulatus capitulatus - THOMAS, Copeia, New York & c., 1965 (4): 438. 1966 Typhlops capitulata capitulata - THOMAS, Proc. biol. Soc. Washington, 79 (4): 264. Range: T y p h l o p s
Haiti. c a p i t u l a t u s
gonavensis
RICHMOND
1964 Typhlops gonavensis RICHMOND, Breviora, Mus. comp. Zool., Cambridge (Massachusetts), 2 0 2 : 3 , fig. 2. - T e r r a t y p i c a : Point a Raquette, on the south shore of Gonave Island, Haiti. — H o l o t y p e : Yale Peabody Museum 3003. 1966 Typhlops capitulatus gonavensis - THOMAS, Proc. biol. Soc. Washington, 79 (4): 265. Range:
G o n a v e Island, H a i t i .
T y p h l o p s caymanensis SACKETT 1940 Typhlops caymanensis SACKETT, Notulae Naturae, Acad. nat. Sei. Philadelphia, 4 8 : 1 . - T e r r a t y p i c a : between Pedro Point and North Sound, Grand Cayman Island, West Indies. - H o l o t y p e : A.N.S.P. 22123. Discussion: Range:
THOMAS (1968:717).
G r a n d C a y m a n Island.
T y p h l o p s c e y l o n i c u s M. A . SMITH 1943 Typhlops ceylonicus M. A. SMITH, Fauna Brit. India, Rept. Amphib. 3 : 5 5 . T e r r a t y p i c a : Peradeniya, Ceylon. - H o l o t y p e :
B.M.N.H. 1946.1.11.42.
D I S C U S S I O N : TAYLOR (1947:285). Range:
Sri L a n k a ( C e y l o n ) . K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
T y p h l o p s c o m o r e n s i s BOULENGER 1889 Typhlops comorensis BOULENGER, Ann. Mag. nat. Hist., London, 4 (6): 361. T e r r a t y p i c a : Comoro Islands, Indian Ocean. - H o l o t y p e : B.M.N.H. 1946.1.11.92. 1918 Typhlops capensis RENDAHL, Ark. Zool, Stockholm, 11 (17): 1; figs. 1 - 3. T e r r a t y p i c a : "Capeland", i. e. South Africa. - H o l o t y p e : Zootomisches Institut der Hochschule zu Stockholm. Range:
C o m o r o Islands, I n d i a n O c e a n ; S o u t h A f r i c a n r e c o r d in e r r o r .
T y p h l o p s conradi PETERS 1874 Typhlops conradi PETERS, M.Ber. k. preufe. Akad. Wiss. Berlin, 1874:162; pi., fig. 1. - T e r r a t y p i c a : North Celebes. - H o l o t y p e : Z.M.B. Discussion: Range:
MCDOWELL (1974: 14). Celebes.
55 Typhlops costaricensis JIMENEZ
& SAVAGE
1962 Typhlops costaricensis JIMENEZ & SAVAGE, Rev. Biol, trop., San José, 10:199. T e r r a t y p i c a : Monteverde, Siena de Tilaran, Provincia de Puntarenas, 1500 m., Costa Rica. - H o l o t y p e : Museo Nacional de Costa Rica; on permanent loan to the University of Southern California, Los Angeles.
Rica and Nicaragua. Typhlops cuneirostris
PETERS
1879 Typhlops cuneirostris PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1879:775; pi., fig. 4, - T e r r a t y p i c a : Barawa, Somalia. - H o l o t y p e : Z.M.B. 9531. 1965 Typhlops cuneirostris calabresii GANS & LAURENT, Ann. Mus. Afr. centr., Tervuren, 134:54, pi. 6. - T e r r a t y p i c a : 56 miles from Hargeisa, on Berbera road, Northern Territory, Somalia. - H o l o t y p e : M.C.Z. 74465. 1965 Typhlops cuneirostris cuneirostris - GANS & LAURENT, Ann. Mus. Afr. centr., Tervuren, 134:54. Discussion:
ROUX-ESTÈVE (1974:153).
R a n g e : Somali Republic; Ethiopia (?). Typhlops decorosus BUCHHOLZ
& PETERS
1875
Typhlops (Ophthalmidion) decorosus BUCHHOLZ & PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1875:197. - T e r r a t y p i c a : Cameroons. - H o l o t y p e : Z.M.B. 8322. 1881 Typhlops (Onychocephalus) Buchholzi PETERS, S.Ber. Ges. naturf. Freunde Berlin, 1 8 8 1 : 7 1 . - T e r r a t y p i c a : Mungo, West Africa. - H o l o t y p e : Z.M.B. 8408. 1893 Typhlops buchholzi - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1 : 4 1 . 1893 Typhlops decorosus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1: 38. Discussion:
ROUX-ESTÉVE (1974:73).
R a n g e : Cameroon. Typhlops decorsei
MOCQUARD
1901 Typhlops decorsei MOCQUARD, Bull. Mus. nation. Hist, nat., Paris, 7 : 2 5 5 . Terra typica:
Ambovombe, Madagascar. -
Holotype:
M.H.N.P. 01-242.
D I S C U S S I O N : GUIBE (1958:196).
R a n g e : Madagascar. Typhlops depressiceps 1913 Typhlops Terra 1932 Typhlops Terra
STERNFELD
depressiceps STERNFELD, S.Ber. Ges. naturf. Freunde Berlin, 1913: 384. t y p i c a : New Guinea. - H o l o t y p e : Z.M.B. monochrus VOGT, S.Ber. Ges. naturf. Freunde Berlin, 1932: (4 - 7): 293. t y p i c a : along Sepik River, New Guinea. - H o l o t y p e : Z.M.B.
56 1948 Typhlops monochrous LOVERIDGE (nomen emendatum), Bull. Mus. comp. Zool., Cambridge (Massachusetts), 77: 320. Discussion:
MCDOWELL (1974:19).
R a n g e : Sepik River through Madang to Lae in Morobe District, New Guinea. Typhlops diardi
SCHLEGEL
1839 Typhlops Diardi SCHLEGEL, Abbild. Amphib.: 39. - T e r r a t y p i c a : Indies". - H o l o t y p e : M.H.N.P. 1065. 1863 F. (Typhlops) Diardi - JAN (error typographies), Elenco sist. Ofidi: 11. Discussion:
"East
MCDOWELL (1974:9).
R a n g e : India, Assam, Burma, Indo-China south of Latitude 14°, the Malay Peninsula, Nia Island, Web Island off the northwest tip of Sumatra, Sumatra, Banka, and Borneo. There is a questionable record for New Guinea. T y p h l o p s
diardi
diardi
SCHLEGEL
1845 Argyrophis horsfleldii GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 137. - T e r r a t y p i c a : Khasi Hüls, Assam. - H o l o t y p e : B.M.N.H. 1946.1.11.50. 1861 Typhlops striolatus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1861:922. T e r r a t y p i c a : banks of Ganges River, Calcutta. - H o l o t y p e : B.M.N.H. 1946.1.11.61. 1864 Typhlops horsfieldi - GÜNTHER, Rept. Brit. India: 174. 1872 Typhlops barmanus STOLICZKA, Proc. asiat. Soc. Bengal, Calcutta, 1872:144. T e r r a t y p i c a : near Moulmein, Burma. - H o l o t y p e : Indian Museum, Calcutta. 1908 Typhlops tephrosoma WALL, J. Bombay nat. Hist. Soc., Calcutta, 18 (2): 314. T e r r a t y p i c a : Shillong, Khasi Hills, Assam, India. - H o l o t y p e : B.M.N.H. 1946.1.11.65. 1909 Typhlops cinereus WALL, J. Bombay nat. Hist. Soc., 19:609. - T e r r a t y p i c a : Upper Assam. - T y p e : not traced. - Preoccupied by Typhlops cinereus GUERIN, 1830. 1934 Typhlops diardi diardi - BRONGERSMA, Zool. Meded., Leiden, 17:192.
R a n g e : India, westward as far as Dun Valley; Assam; Burma; Hainan Tao, China. T y p h l o p s
diardi
muelleri
SCHLEGEL
1839 TyphlopsMulleri SCHLEGEL, Abbild. Amphib.: 39; pi. 32, figs. 25 - 28. - T e r r a t y p i c a : Padang, Sumatra. - H o l o t y p e : R.M.N.H. 3718. 1844 Typhlops nigroalbus DUMERIL & BIBRON, Erpetol. gen., 6 : 2 9 5 . - T e r r a t y p i c a : Sumatra. - H o l o t y p e : M.H.N.P. 6991. 1845 Argyrophis bicolor GRAY, Cat. Lizards Brit. Mus.: 136. - T e r r a t y p i c a : Singapore. - H o l o t y p e : B.M.N.H. 1946.1.10.63. 1863 Typhlops (Typhlops) Mülleri - JAN, Elenco sist. Ofidi: 12. 1863 Typhlops (Typhlops) nigroalbus - JAN, Elenco sist. Ofidi: 12. 1863 Typhlops (Typhlops) Schneiden JAN, Elenco sist. Ofidi: 12. - T e r r a t y p i c a : Bangkok, Thailand. - H o l o t y p e : M.S.N.M. - destroyed. 1864 Typhlops bothriorhynchus GÜNTHER, Rept. Brit. India: 174; pi. 16, fig. G. - T e r r a t y p i c a : Pinang, Malaya. - H o l o t y p e : B.M.N.H. 1946.1.11.59.
57 1893 Typhlops muelleri - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1 : 2 5 . 1906 Typhlops kapaladua ANNANDALE, J. Proc. asiat. Soc. Bengal, Calcutta, 1 (8): 208. T e r r a t y p i c a : "Malay Archipelago, probably from Java". - H o l o t y p e : Indian Museum, Calcutta. 1918 Typhlops labialis WAITE, Ree. S. Austral. Mus., Adelaide, 1: 30, fig. 22. - T e r r a t y p i c a : "Western Australia", in error. - H o l o t y p e : W.A.M. R630. 1923 Typhlops diardi nigroalbus - M. A. SMITH, J. nat. Hist. Soc. Siam, Bangkok, 6 : 5 2 . 1934 Typhlos fusconotus BRONGERSMA, Zool. Meded. Leiden, 17:192, text figs. 1 9 - 2 1 . - T e r r a t y p i c a : Dore (= Manokawari), West New Guinea. - H o l o t y p e : R.M.N.H. 6293. 1934 Typhlops diardi mulleri - BRONGERSMA, Zool. Meded., Leiden, 17:193. Discussion:
TAYLOR (1965: 648); MCDOWELL (1974:9).
R a n g e : Indo-China south of latitude 14°, the Malay Peninsula; Nias Island; Web Island off the northwest tip of Sumatra; Banka; Sumatra; and Borneo. Typhlops dominicanus
STEJNEGER
71830 Typhlops cinereus GUÉRIN, Icon. Règne Anim., Rept.: pi. 18, fig. 2. - T e r r a t y p i c a : Guadeloupe. - H o l o t y p e : M.H.N.P. 3219. 1904 Typhlops dominicana STEJNEGER, Rep. U.S. nation. Mus., Washington, 1902: 687. - T e r r a t y p i c a : Island of Dominica, Lesser Antilles. - S y n t y p e s : B.M.N.H. 65.5.4.177; 89.8.14.1-8; and 91.5.11.2. Discussion:
RICHMOND (1966:131).
R a n g e : Dominica, Lesser Antilles. Typhlops elegans
PETERS
1868 Typhlops (Ophthalmidion) elegans PETERS, M.Ber. k. preuli. Akad. Wiss. Berlin, 1868:450; pi. 2, fig. 1. - T e r r a t y p i c a : Principe Island, Gulf of Guinea, West Africa. - S y n t y p e s : Z.M.B. 6066, and another in Zoologischer Sammlung, Stettin. 1893 Typhlops elegans - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.), 1: 3 7 - 3 8 . Discussion:
ROUX-ESTÈVE (1974:60).
R a n g e : Principe Island, Gulf of Guinea. Typhlops exiguus JAN 1864 Typhlops exiguus JAN, Iconogr. gén. Ophid., 1 (3. livr.): 3; pis. 4, 5, fig. 11.— T e r r a t y p i c a : "East Indies". - H o l o t y p e : Staatliches Museum fiir Naturkunde, Stuttgart, Germany. 1921 Typhlops exignus WERNER (error typographies), Arch. Naturgesch., Berlin, 87A (7): 332.
R a n g e : Known only from Belgaum, India. Typhlops floweri
BOULENGER
1899 Typhlops floweri BOULENGER, Proc. zool. Soc. London, 1899:654; pi. 37, fig. 2 - T e r r a t y p i c a : "Siam". - H o l o t y p e : B.M.N.H. 1946.1.10.55. Discussion:
TAYLOR (1965:639).
R a n g e : Thailand.
58 Typhlops fomasinii BIANCONII 1847 Typhlops Fornasinii BIANCONI, Spec. Zool., Mosamb.: 183; pi. 6, fig. 1. - T e r r a t y p i c a : Mozambique. - H o l o t y p e : Musée de Bologne? 1854 Onychocephalus trilobus PETERS, M.Ber. k. preufc. Akad. Wiss. Berlin, 1854: 621. T e r r a t y p i c a : Inhambane, Mozambique. - S y n t y p e s : Z.M.B. 3915 & 3916 and M.C.Z. 21006. 1854 Onychocephalus mossambicus PETERS, M.Ber. k. pre ufi. Akad. Wiss. Berlin, 1854: 621. - T e r r a t y p i c a : Anjoan and Insulae Mozambique. - S y n t y p e s : Z.M.B. 3949 & 3950 and 21007. 1860 Onychocephalus tettensis PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1860:80. T e r r a t y p i c a : Tete, Mozambique. - H o l o t y p e : Z.M.B. 3944. 1860 Typhlops (Ophthalmidion) Bianconii JAN, Iconogr. gén. Ophid., 1 (1. livr.): 1 ; pis. 5 , 6 , fig. 3. - T e r r a t y p i c a : Lagoa, Mossambique. - H o l o t y p e : M.S.N.M. 1863 Typhlops (Typhlops) Fornasinii - JAN, Elenco sist. Ofidi: 12. 1882 "Typhlops tettensis - PETERS, Reise nach Mossambique, 3:92; pi. 15, fig. 1. 1882 Typhlops mossambicus - PETERS, Reise nach Mossambique, 3:93; pi. 15, fig. 2. 1921 Typhlops mossombicus WERNER (error typographies), Arch. Naturgesch. Berlin, 87A (7): 336. Discussion:
LAURENT (1964:421); ROUX-ESTÈVE (1974:146).
R a n g e : Coastal areas of Mozambique from Inhambane to Lourenco Marques; Mulugwe Pan, Gona-re-Zhou, southeastern Rhodesia; and Mseleni, Zululand, Natal, South Africa. Typhlops giadinhensis BOURRET 1937 Typhlops giadinhensis BOURRET, Bull. Gén. Instr. Pubi. Hanoi, Dec. 1937:68, text figs. - T e r r a t y p i c a : Giadinh, South Vietnam. - H o l o t y p e : Laboratoire des Sciences Naturelles de l'Université de Hanoi, now M.H.N.P. 1938-118.
R a n g e : South Vietnam. Known only from the type locality. Typhlops gierrai MOCQUARD 1897 Typhlops gierrai MOCQUARD, Bull. Mus. nation. Hist, nat., Paris, 1897 (4): 122. T e r r a t y p i c a : Tanga, Tanzania. - H o l o t y p e : M.H.N.P. 1897-24. 1923 Typhlops guirrae SCHMIDT (error typographicus), Bull. Amer. Mus. nat. Hist., New York, 49 (1): 33. 1928 Typhlops punctatus gierrai - BARBOUR & LOVERIDGE, Mem. Mus. comp. Zool., Cambridge (Massachusetts), 5 0 : 1 0 6 . Discussion:
LAURENT (1964:417); ROUX-ESTEVE (1974:133).
R a n g e : Mountain forests, in Usambara and Uluguru Mountains, Tanzania. Typhlops grandidieri MOCQUARD 1905 Typhlops grandidieri MOCQUARD, Bull. Mus. nation. Hist, nat., Paris, 9 : 2 8 6 . T e r r a t y p i c a : Madagascar. - S y n t y p e s : M.H.N.P. 05-271 & 05-272. Discussion:
GUIBÉ (1958:191).
R a n g e : Madagascar.
59 T y p h l o p s grand RUTHVEN & GAIGE 1935 Typhlops granti RUTHVEN & GAIGE, Occ. Pap. Mus. Zool. Univ. Michigan, Ann Arbor, 3 0 7 : 2 . - T e r r a t y p i c a : Caja de Muertos Island, 8 miles off Ponce, Puerto Rico. - H o 1 o t y p e : U.M.M.Z. 76669. Range:
S o u t h w e s t e r n P u e r t o R i c o ; Isla C a j a de M u e r t o s .
T y p h l o p s guadeloupensis RICHMOND 1966 Typhlops guadeloupensis RICHMOND, Herpetologica, Lawrence, 2 2 : 1 2 9 , fig. T e r r a t y p i c a : 2 km southwest of Port-Blanc, Guadeloupe. - H o l o t y p e : C.M. 41216. Range:
Guadeloupe.
T y p h l o p s h e c t u s THOMAS 1974 Typhlops hectus THOMAS, Proc. biol. Soc. Washington, 87 (2): 12. - T e r r a t y p i c a : Martineau, ca. 9 km west of Jeremie, Dept. du Sud, Haiti. - H o l o t y p e : M.C.Z. 81149. R a n g e : S o u t h w e s t e r n H i s p a n i o l a s o u t h o f t h e Cul d e Sac-Valle d e N e i b a Plain ( e x c e p t f o r t h e B a r a h o n a P e n i n s u l a ) a n d t h a t p o r t i o n of H i s p a n i o l a n o r t h of t h e Cul d e Sac-Valle de N e i b a P l a i n ; lie G r a n d e C a y e m i t e . Sea level t o 2600 feet. T y p h l o p s h y p o g i u s SAVAGE 1950 Typhlops hypogia SAVAGE, Proc. California, zool. Club, Palo Alto, l : 5 2 - 53;fig. 2. - T e r r a t y p i c a : Cebu, Cebu Island, Philippines. - H o l o t y p e : Stanford University 12347. Discussion: Range:
MCDOWELL (1974:12).
Cebu Island, Philippines.
T y p h l o p s h y p s o b o t h r i u s WERNER 1917 Typhlops hypsobothrius WERNER, Jb. hamburg. wiss. Anst., 34: 34, 35. - T e r r a t y p i c a : Sumatra. - H o l o t y p e : formerly in Zoologisches Museum, Hamburg; destroyed during World War II. Range:
Sumatra.
T y p h l o p s inornatus BOULENGER 1888 Typhlops inornatus BOULENGER, Ann. Mag. nat. Hist., London, (6) 1: 344. T e r r a t y p i c a : Sogere, 1750 ft., Central District, Papua New Guinea. - H o l o t y p e : B.M.N.H. 1946.1.11.80. Discussion:
MCDOWELL (1974:17).
R a n g e : K n o w n f r o m 3 localities in elevated p a r t s of P a p u a , N e w G u i n e a : Sogere ( C e n t r a l District); A l b e r t E d w a r d R a n g e N o r t h e r n District); a n d n e a r Bulolo ( M o r o b e D i s t r i c t ) .
60 T y p h l o p s jamaicensis (SHAW) 1802 Anguis Jamaicensis SHAW, Gen. Zool. 3 : 5 8 8 . - T e r r a t y p i c a : J a m a i c a . H o l o t y p e : not traced. 1845 Meditoria nasuta GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 139. - T e r r a t y p i c a : Berbice, Guyana (error). - H o l o t y p e : B.M.N.H. 1946.1.10.45. 1845 Anilios leachi GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 135. - T e r r a t y p i c a : none given. - S y n t y p e s : B.M.N.H. 1946.1.12.5 & 1946.1.10.65. 1924 Typhlops jamaicensis - COCHRAN, J. Washington Acad. Sci., 14 (8): 175. Discussion: Range:
RUTHVEN & GAIGE (1935:1); THOMAS (1966:256).
Jamaica below 2000 feet.
T y p h l o p s jerdoni BOULENGER 1890 Typhlops jerdoni BOULENGER, Fauna Brit. India, Rept. Batr.: 238. - T e r r a t y p i c a : Khasi Hills, Assam, India. - S y n t y p e s : B.M.N.H. 1946.1.10.66-68. (3). 1912 Typhlops diversiceps ANNANDALE, Rec. Ind. Mus., Calcutta, 8 : 4 4 , fig. 1. T e r r a t y p i c a : Pasighal, 500 ft., Abor district, Assam, India. - H o l o t y p e : Indian Museum, Calcutta 16864. Discussion:
M. A. SMITH (1943:50).
R a n g e : E a s t e r n H i m a l a y a s ( S i k k i m , Darjeeling, D u a r s d i s t r i c t s ) ; U p p e r B u r m a ( L a s h i o ) ; A s s a m ( A b o r a n d Khasi Hills); a n d N e p a l .
T y p h l o p s khoratensis TAYLOR 1962 Typhlops khoratensis TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 4 0 : 248 - 251, fig. 13a. - T e r r a t y p i c a : Muak Lek, Friendship Highway, Sara Buri Province, Thailand, - H o l o t y p e : F.M.N.H. 178263. Discussion: Range:
TAYLOR (1965:643).
K h o r a t P l a t e a u area o f c e n t r a l ( e a s t e r n ) T h a i l a n d .
T y p h l o p s klemmeri TAYLOR 1962 Typhlops klemmeri TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 4 3 : 2 5 3 - 254. T e r r a t y p i c a : Koh Phai, near Kuala Lumpur, Malaya. - H o l o t y p e : F.M.N.H. 17238 Range:
Kuala L u m p u r , Malaya. K n o w n only f r o m t h e type locality.
T y p h l o p s k o e k k o e k i BRONGERSMA 1934 Typhlops koekkoeki BRONGERSMA, Zool. Meded., Leiden, 1 7 : 1 8 6 - 1 8 7 , figs. T e r r a t y p i c a : Palau Bunju, Boenjoe Island, east coast, North Borneo. H o l o t y p e : Zoologisches Museum, Amsterdam 11077. Range: locality.
B o e n j o e Island, o f f N o r t h B o r n e o . K n o w n o n l y f r o m t h e t y p e
61 Typhlops koshunnensis
OSHIMA
1916 Typhlops koshunnensis OSHIMA, Dobutugaku Zasshi, Zool. Mag., Tokyo, 28 (328): 84 - 86. - T e r r a t y p i c a : Koshun, Formosa. - H o 1 o t y p e : Institute of Science Museum, Formosa, no. 4.
R a n g e : Known from Kuraru, Hengchun, Twannaryo and Pingtung, Formosa. Typhlops kraali Do RIA 1874 Typhlops Kraali DORIA, Annali Mus. civ. Stor. nat. Giacomo Doria, Genova, 6 : 347; pl. 12, fig. f. - T e r r a t y p i c a : Kei Islands. - H o 1 o t y p e : M.S.N.G. 29962. Discussion:
MCDOWELL (1974:12).
R a n g e : Ceram; Kei Islands. Typhlops lankaensis
TAYLOR
1947 Typhlops lankaensis TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 31 pt. 2, (13): 287. - T e r r a t y p i c a : 12 miles north of Trincomalee, Ceylon. - H o l o t y p e : F.M.N.H. 100066. Discussion:
DERANIYAGALA (1955 : 20).
R a n g e : Sri Lanka (Ceylon). Typhlops lehnen Roux 1926 Typhlops lehneri ROUX, Rev. Suisse Zool., Genène, 33 (4): 298. - T e r r a t y p i c a : El Pozon, Estado Falcon, Venezuela, - H o l o t y p e : N.M.B. 9020. Discussion:
ROZE (1966: 35); THOMAS (1974:4).
R a n g e : Estado Falcon, Venezuela. Typhlops leucomelas
BOULENGER
1890 Typhlops leucomelas BOULENGER, Fauna Brit. India, Rept. Batr.: 237 - 238. T e r r a t y p i c a : Haycock Mountain, 2000 ft., 40 miles from Point de Galle, Ceylon. - H o l o t y p e : B.M.N.H. 1946.1.10.46. Discussion:
TAYLOR (1947:286).
R a n g e : Hills of Sri Lanka (Ceylon). Typhlops leucostictus
BOULENGER
1898 Typhlops leucostictus BOULENGER, Ann. Mag. nat. Hist., London, (7) 1:124. — T e r r a t y p i c a : Liberia. - H o l o t y p e : B.M.N.H. 1946.1.11.24. Discussion:
ROUX-ESTÈVE (1974: 37).
R a n g e : Liberia. Known only from the holotype.
62 T y p h l o p s lineolatus JAN 1863 Typhlops (Ophthalmidion) lineolatus JAN, Elenco sist. Ofidi: 13. - T e r r a t y p i c a : Sierra Leone. - H o l o t y p e : formerly in Uppsala Museum, Sweden; now lost. 1864 Typhlops lineolatus - JAN, Iconogr. gén. Ophid., 1 (9. livr.): 5; pi. 1, fig. 4. Discussion:
ROUX-ESTÈVE (1974:90).
R a n g e : Senegal; G a m b i a ; Liberia; G u i n e a ; Sierra L e o n e ; Ivory C o a s t ; G h a n a ; Nigeria; C a m e r o o n ; s o u t h w e s t C h a d ; s o u t h e r n S u d a n ; E t h i o p i a ; U g a n d a ; s o u t h w e s t K e n y a ; T a n z a n i a ; w e s t e r n Z a m b i a ; Z a i r e ; A n g o l a ; Congo-Brazzaville. T y p h l o p s
l i n e o l a t u s
l i n e o l a t u s
JAN
1889 Typhlops blanfordi BOULENGER, Ann. Mag. nat. Hist., London, (6) 4 : 363. T e r r a t y p i c a : Senafe, Ethiopia. - H o l o t y p e : B.M.N.H. 1946.1.11.67. 1893 Typhlops boulengeri BOCAGE, Jörn. Sci. nath. phys. nat., Lisboa, (2) 3 : 1 1 7 . T e r r a t y p i c a : Quindumbo, Congo. - S y n t y p e s : B.M.N.H. 1946.1.11.18; M.B.L. 1888. - destroyed. ? 1904 Typhlops bocagei BETHENCOURT-FERREIRA, Jörn. Sci. nath. phys. nat., Lisboa, (2) 7 : 1 1 4 . - T e r r a t y p i c a : Cabicula, Bom Jesus, near Quanza, Angola. H o l o t y p e : M.B.L. - now destroyed. 1910 Typhlops tornieri STERNFELD, Mitt. zool. Mus. Berlin, 5 : 6 9 . - T e r r a t y p i c a : Kilimandjaro Mountain, Tanzania. - S y n t y p e s : Z.M.B. 20722 & 20723. 1928 "Typhlops intermediate between punctatus and gierrai" - BARBOUR & LOVERIDGE, Mem. Mus. comp. Zool., Cambridge (Massachusetts), 5 0 : 1 0 6 . - L o c a l i t y : Amani, Usambara Mountains, Tanzania. 1956 Typhlops angolensis blanfordi - LAURENT, Ann. Mus. roy. Congo beige, Tervuren, (8°) Zool., 4 8 : 6 3 . 1956 Typhlops schmidti LAURENT, Ann. Mus. Congo, Zool., Tervuren, (8°) 4 8 : 7 1 ; figs. 9 - 1 1 ; pi. 8, fig. 4. - T e r r a t y p i c a : Nyunzu, 700 m, Albertville Territory, Tanzania. - H o l o t y p e : R.G.M.C. 17996. 1964 Typhlops schmidti schmidti - LAURENT, Bull. Mus. comp. Zool. Cambridge (Massachusetts), 130: (6): 417. 1964 Typhlops boulengeri usambaricus LAURENT, Bull. Mus. comp. Zool. Cambridge (Massachusetts), 130 (6): 416. - T e r r a t y p i c a : Amani, Usambara Mountains, Tanzania. - H o l o t y p e : M.C.Z. 38699. 1968 Typhlops schmidti wilsoni LAURENT, Arnoldia, Bulawayo, 3 : 3 1 . - T e r r a t y p i c a : Mkanda, Chipangali, Zambia. - H o l o t y p e : U.M. 9444. - Preoccupied by Typhlops wilsoni WALL, 1908. 1974 Typhlops lineolatus lineolatus - ROUX-ESTÈVE, Mem. Mus. nation. Hist, nat., Paris, (A, Zool.) 87: 80. R a n g e : Senegal; G a m b i a ; L i b e r i a ; G u i n e a ; Sierra L e o n e ; Ivory C o a s t ; G h a n a ; Nigeria; C a m e r o o n ; s o u t h w e s t C h a d ; s o u t h e r n S u d a n ; E t h i o p i a ; U g a n d a ; southwest Kenya; n o r t h and central Tanzania; western Zambia; Zaire; Angola; Congo-Brazzaville. T y p h l o p s
l i n e o l a t u s
t a n g a n i c a n u s
LAURENT
1964 Typhlops schmidti tanganicanus LAURENT, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 130:418. - T e r r a t y p i c a : Liwale, Tanzania. - H o l o t y p e : M.C.Z. 57439. 1974 Typhlops lineolatus tanganicanus - ROUX-ESTÈVE, Mem. Mus. nation. Hist, nat., Paris, (A, Zool.) 87:80. Range:
Southeastern Tanzania.
63 Typhlops lorenzi
WERNER
1909 Typhlops lorenzi WERNER, Mitt, naturhist. Mus. Hamburg, 2 6 : 2 0 9 . - T e r r a t y p i c a : Palau Miang-besar, off east coast of Borneo. - H o l o t y p e : formerly in Zoologisches Museum, Hamburg; destroyed during World War II.
R a n g e : Palau Miang-besar off Borneo. Known only from the type locality. Note: This species may belong in the genus Typhlina.
Typhlops loveridgei
CONSTABLE
1949 Typhlops loveridgei CONSTABLE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 103 (2): 110 - 111. - T e r r a t y p i c a : "probably from North India". - H o l o t y p e : M.C.Z. 2283.
R a n g e : North India? Known only from the type locality.
Typhlops lumbricalis (LINNAEUS) 1758 Artguis lumbricalis LINNAEUS, Syst. Nat., Ed. 10, 1:288. - T e r r a t y p i c a : "America". - H o l o t y p e : not traced. 1811 Typhlops lumbricalis - OPPEL, Ordn., Fam. Gatt. Rept.: 55. 1843 Typhlops Cubae BIBRON, in DE LA SAGRA, Historia fis. pol. nat. de Cuba, 4 Rept.: 204, pi. 22. - T e r r a t y p i c a : Cuba. - S y n t y p e s : M.H.N.P. 3218 & 3218a. 1959 Typhlops silus LEGLER, Herpetologica, Escondido, 15:105, fig. 1. - T e r r a t y p i c a : Banes, Provincia Oriente, Cuba. - H o l o t y p e : K.U. 47469. Discussion:
RICHMOND (1964:7).
R a n g e : Cuba and the Isla de Pinos; the Bahamas Islands.
Typhlops luzonensis
TAYLOR
1919 Typhlops luzonensis TAYLOR, Philippine J. Sci., Manila, 14:105. T e r r a t y p i c a : low on side of Mount Maquiling, Luguna, Luzon, Philippines. - H o l o t y p e : C.M. 2653. Discussion:
TAYLOR (1922:52); MCDOWELL (1974:12).
R a n g e : Luzon, Negros, and Marinduque Islands, Philippines.
Typhlops madagascariensis 1877
BOETTGER
Typhlops madagascariensis BOETTGER, Abh. senckenb. naturi". Ges., Frankfurt/ Main, 11: 3; pl. 1, fig. 1 . - T e r r a t y p i c a : Nossi Bé, Madagascar. - H o l o t y p e : S.M.F. 16604.
Discussion:
GUIBÉ (1958:195).
R a n g e : Madagascar.
64 Typhlops malcolmi T A Y L O R 1947 Typhlops malcolmi TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 31, pt. 2, (13): 291 - 292. - T e r r a t y p i c a : 12 miles north of Trincomalee, Eastern Province, Ceylon. - H o 1 o t y p e : formerly EHT-HMS 30072; now presumed lost. R a n g e : Near Trincomalee, Sri Lanka (Ceylon). Known only from the type locality.
Typhlops manilae T A Y L O R 1919 Typhlops manilae TAYLOR, Philippine J. Sei., Manila, 14:106. - T e r r a t y p i c a : "Filipinas", probably from Luzon, per Taylor. - H o l o t y p e : an unnumbered specimen in the Santo Tomas Museum, Manila. D i s c u s s i o n : TAYLOR (1922:56). R a n g e : Luzon Island, Philippines? Known only from the type specimen.
Typhlops manni LOVERIDGE 1941 Typhlops manni LOVERIDGE, Proc. U.S. nation. Mus., Washington, 91 (3128): 1 1 8 . - T e r r a t y p i c a : Harbel, Liberia. - H o l o t y p e : U.S.N.M. 109634. 1952 Typhlops angeli GUIBÉ, Bull. Mus. nation. Hist, nat., Paris, (2) 24 (1): 79. T e r r a t y p i c a : Nzo, Mont-Nimba, 500 m, French Guinea. - H o l o t y p e : M.H.N.P. 1951-1. D i s c u s s i o n : ROUX-ESTÈVE (1974:41). R a n g e : Liberia and Guinea.
Typhlops microstomus COPE 1866 Typhlops microstomus COPE, Proc. Acad. nat. Sci. Philadelphia, 18:125 - 126. T e r r a t y p i c a : Yucatan, Mexico. - H o l o t y p e : U.S.N.M. 61064. D I S C U S S I O N : TAYLOR ( 1 9 3 9 : 4 4 3 ) .
R a n g e : The northern half of the Yucatan Peninsula.
Typhlops minis JAN 1860 Typhlops (Diaphorotyphlops) mirus JAN, Iconogr. gén. Ophid., 1 (1. livr.): 1; pis. 4, 5, fig. 7 . - T e r r a t y p i c a : Ceylon. - H o l o t y p e : R.M.N.H. 3721. 1864 Typhlops mirus - GÜNTHER, Rept. Brit. India: 176. D I S C U S S I O N : TAYLOR ( 1 9 4 7 : 2 8 4 ) .
R a n g e : Peradeniya, Sri Lanka (Ceylon) only definite locality.
65 Typhlops monastus THOMAS Typhlops 1966
monastus
monastus
THOMAS
Typhlops monastus monastus THOMAS, Proc. biol. Soc. Washington, 7 9 : 2 5 7 - 260. T e r r a t y p i c a : between Lawyers River and Cassava Ghaut, St. Peter's Parish, Montserrat, British West Indies. - H o l o t y p e : M.C.Z.81112.
R a n g e : Island of Montserrat, British West Indies. Typhlops 1966
monastus
geotomus
THOMAS
Typhlops monastus geotomus THOMAS, Proc. biol. Soc. Washington, 7 9 : 260 - 262. T e r r a t y p i c a : about 1 mile north of Carlisle, St. Mary's Parish, Antigua, British West Indies. - H o l o t y p e : M.C.Z. 81115.
R a n g e : St. Cristopher, Nevis, Antigua, Barbuda, and Great Bird Islands, West Indies. Typhlops monensis SCHMIDT 1926 Typhlops monensis SCHMIDT. Publ. Field Mus. nat. Hist., Zool., 1 2 : 1 5 7 , fig. 1. T e r r a t y p i c a : Mona Island, West Indies. - H o l o t y p e s : Naturhistorisches Museum, Hamburg, 1582; destroyed during World War II. D I S C U S S I O N : RUTHVEN & GAIGE (1935 :12).
R a n g e : Mona Island, West Indies. Typhlops mucronatus BOETTGER 1880
Typhlops (Ophthalmidion) mucronatus BOETTGER, Zool. Anz., Leipzig, 1880: 279. - T e r r a t y p i c a : Lukubé, Nossi Be (= Nosy Bé), Madagascar. L e c t o t y p e : S.M.F. 16626.
Discussion:
GUIBÉ ( 1 9 5 8 : 1 9 7 ) .
R a n g e : Madagascar. Typhlops oatesi BOULENGER 1890
Typhlops oatesi BOULENGER, Fauna Brit. India, Rept. Batr.: 238. - T e r r a t y p i c a : Table Island, Cocos Group, Andamans. - S y n t y p e s : B.M.N.H. 1946.1.10.51-52.
Discussion:
M. A. SMITH ( 1 9 4 3 : 5 3 ) .
R a n g e : Andaman Islands. Typhlops obtusus PETERS 1865
Typhlops (Onychocephalus) obtusus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1865: 260; pi. fig. 2. - T e r r a t y p i c a : Shire River, Nyasaland, Malawi. S y n t y p e s : B.M.N.H. 1946.1.11.31-32. 1953 Typhlops tettensis obtusus - LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 1 1 0 : 2 4 3 .
66 1964 Typhlops obtusus - LAURENT, Bull. Mus. comp. Zool, Cambridge (Massachusetts), 130:420. 1968 Typhlops obtusus palgravei LAURENT, Arnoldia, Bulawayo, 3 (31): 1 - 2. T e r r a t y p i c a : Umtali, Rhodesia. - H o l o t y p e : U.M. 12764. 1968 Typhlops obtusus obtusus - LAURENT, Arnoldia, Bulawayo, 3 (31): 1. Discussion: Range:
LAURENT (1964:420); ROUX-ESTÈVE (1974:69). Malawi;Mozambique;Rhodesia.
Typhlops ocularis
PARKER
1927 Typhlops ocularis PARKER, Ann. Mag. nat. Hist., London, (9) 19: 379, fig. T e r r a t y p i c a : Antongil Bay, Maroansetra Forest, Madagascar. - H o l o t y p e : B.M.N.H. 1946.1.11.25. 1918 Typhlops albanalis RENDAHL, Ark. Zool., Stockholm, 11 (17): 3; figs. 4 - 6. T e r r a t y p i c a : Cape Province, South Africa (in error). - H o l o t y p e : Zootomisches Instituts der Hochschule zu Stockholm. DISCUSSION: GUIBÉ (1958:193); ROUX-ESTÈVE (1974:205). Range:
Madagascar.
Typhlops oligolepis
WALL
1909 Typhlops oligolepis WALL, J. Bombay nat. Hist. Soc., 19: 339, fig. - T e r r a t y p i c a : Nagri Valley, Darjeeling District, 5000 ft., India. - H o l o t y p e : B.M.N.H. 1946.1.11.85, formerly 1926.8.18.3, from "E. Himalayas". Stated to be the type although the the original description gives as type a specimen from Nagri Valley in the Darjeeling Museum. D i s c u s s i o n : M. A. SMITH (1943:55). Range:
The eastern Himalayas area of Sikkim, India.
Typhlops pammeces
GÜNTHER
1864 Typhlops tenuis GÜNTHER, Rept. Brit. India: 176; pi. 16, fig. C. - T e r r a t y p i c a : Madras, India. - H o l o t y p e : B.M.N.H. 1946.1.11.34. Preoccupied by Typhlops tenuis SALVIN, 1860. 1864 Typhlops pammeces GÜNTHER, Rept. Brit. India: 444. Substitute name for T. tenuis. 1898 Typhlops braminus pammeces - BOETTGER, Zool. Anz., Leipzig, 1898 (5): 2. 1906 Typhlops psammophilus ANNANDALE, Mem. Asiat. Soc. Bengal, Calcutta, 1:193. T e r r a t y p i c a : Ramnad, Madura, South India. — S y n t y p e s : B.M.N.H. 1946.1.11.60, & 2 in the Indian Museum, Calcutta. 1943 Typhlops psammeces M. A. SMITH (error typographies), Fauna Brit. India, 3 Serpentes: 48. Range:
Southern India.
67 T y p h l o p s platyrhynchus STERNFELD 1910 Typhlops platyrhynchus STERNFELD, Mitt. zool. Mus. Berlin, 1910 (5); 69. T e r r a t y p i c a : Tanga, Tanzania. - S y n t y p e s : M.C.Z. 39798, Z.M B 20729, 20729A & 20729B. D I S C U S S I O N : ROUX-ESTEVE (1974:45). Range:
Tanzania. K n o w n only f r o m the t y p e locality.
T y p h l o p s porrectus STOLICZKA 1871 Typhlops porrectus STOLICZKA, J. asiat. Soc. Bengal, Calcutta, 4 0 : 4 2 6 ; pi. 25, figs. 1 - 4 . - T e r r a t y p i c a : Bengal and Roorkee, India. - S y n t y p e s : N.M.W. 15358 (2) & 15357 (2). 1909 Typhlops mackinnoni WALL, J. Bombay nat, Hist. Soc., 19:805,fig. - T e r r a t y p i c a : Mussorie, 6000 ft., western Himalayas. - H o l o t y p e : B.M.N.H. 1946.1.23.56. 1913 Typhlops venningi WALL, J. Bombay nat. Hist. Soc., 2 2 : 5 1 5 , 3 figs. - T e r r a t y p i c a : Pyawbwe, 700 Ft., Upper Burma. - S y n t y p e s : B.M.N.H. 1946.1.10.53-54. 1949 Typhlops porrecta - CONSTABLE, Bull. Mus. comp. Zool, Cambridge (Massachusetts), 103 (2): 112. Discussion: Range:
MINTON (1966:116).
P a k i s t a n ; I n d i a ; Sri L a n k a ( C e y l o n ) ; a n d U p p e r B u r m a .
T y p h l o p s pseudosaurus DRYDEN & TAYLOR 1969 Typhlops pseudosaurus DRYDEN & TAYLOR, Univ. Kansas Sci. Bull,, Lawrence, 58:270. - T e r r a t y p i c a : Harmon Village, Guam, Mariana Islands. - H o l o t y p e : F.M.N.H. 189357. Range:
K n o w n only f r o m the type locality.
T y p h l o p s punctatus (LEACH) 1819 Acontias punctatus LEACH in BOWDICH, Miss. Cape Coast Cast. Ashantee; 493. T e r r a t y p i c a : Fanti, Ashanti, Ghana. - H o l o t y p e : B.M.N.H. 1946.1.11.47. 1893 Typhlops punctatus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:42. Discussion:
LAURENT (1964:411); ROUX-ESTÈVE (1974:79).
R a n g e : Sierra L e o n e ; Liberia; G u i n e a ; Ivory C o a s t ; G h a n a ; Nigeria; C a m e r o o n ; F e r n a n d o P o Island; C e n t r a l A f r i c a n R e p u b l i c ; G a b o n ; Congo-Brazzaville; Zaire; U g a n d a ; c o a s t a l Senegal; G a m b i a ; P o r t u g e s e G u i n e a ; c e n t r a l Mali; T o g o ; s o u t h w e s t e r n Niger; s o u t h e r n C h a d ; s o u t h w e s t E t h i o p i a ; a n d D a h o m e y .
T y p h l o p s
p u n c t a t u s
p u n c t a t u s
(LEACH)
1839 Typhlops Eschrichtii SCHLEGEL, Abbild. Amphib.: 37; pi. 32, figs. 13, 16. T e r r a t y p i c a : Coast of Guinea, Africa. - H o l o t y p e : R.M.N.H. 3700. 1843 Typhlops (Aspidorhynchus) Eschrichtii - FITZINGER, Sys. Rcpt., 1: 24.
68 1844 Ophthalmidion eschrichtii - DUMÉRIL & BIBRON, Erpétol. gén., 6 : 2 6 5 . 1845 Onychophispunctata - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 133. 1880 Typhlops (Ophthalmidion) Eschrichtii - MÜLLER, 1. Nachtr. Kat. herpetol. Sammlung Basier Mus.: 24. 1881 Aspidorhynchus eschrichtii - PETERS, S.Ber. Ges. naturf. Freunde Berlin, 1881:70. 1888 Typhlops (Aspidorhynchus) Eschrichtii - BOETTGER, Ber. senckenb. naturf. Ges. Frankfurt am Main, 1888:42. 1893 Typhlops punctatus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1 : 4 2 . 1920 Typhlops milleti CHABANUAD, Bull. Mus. nation. Hist, nat., Paris, 2 6 : 4 6 3 . T e r r a t y p i c a : Togo. - H o l o t y p e : M.H.N.P. 1920-21. 1936 Typhlops punctatus punctatus - LO VE RIDGE, Zool. Ser. Field Mus. Nat. Hist., Chicago, 2 2 : 1 9 . Discussion:
LAURENT (1964:411); ROUX-ESTÈVE (1974:79).
R a n g e : Coastal Senegal; Gambia; Portugese Guinea; Guinea; central Mali; Ivory Coast; Ghana; Dahomey; Togo; Nigeria; southwestern Niger; southern Chad; Central African Republic; Cameroon; Uganda; and southwest Ethiopia. T y p h l o p s
punctatus
congestus
(DUMÉRIL&BIBRON)
1844 Onychocephalus congestus DUMÉRIL & BIBRON, ErpétoL gén., 6 : 334. - T e r r a t y p i c a : none given. - H o l o t y p e : Fort Pitt, Chatham; now presumed lost. 1845 Onychophis barrowii GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 133. - T e r r a t y p i c a : "? India" (in error). - H o l o t y p e : 1946.1.11.63. 1881 Typhlops (Onychocephalus) crassatus PETERS, S.Ber. Ges. naturf. Freunde Berlin, 1 8 8 1 : 5 0 . - T e r r a t y p i c a : Chinchoxo, Congo. - H o l o t y p e : Z.M.B. 9149. 1923 Typhlops congestus - SCHMIDT, Bull. amer. Mus. nat. Hist., New York, 4 9 : 4 8 5 . 1974 Typhlops punctatus congestus - ROUX-ESTÈVE, Mém. Mus. nation. Hist, nat., Paris, (A, Zool.) 8 7 : 7 9 .
R a n g e : Humid forests of Nigeria, Cameroon, Fernando Po Island, Central African Republic, Gabon, Zaire, Congo-Brazzaville, and Uganda. T y p h l o p s
p u n c t a t u s
liberiensis
(HALLOWELL)
1848 Onychocephalus liberiensis HALLOWELL, Proc. Acad. nat. Sci., Philadelphia, 1848 : 5 9 . - T e r r a t y p i c a : Liberia. - H o l o t y p e : A.N.S.P. 3243. 1848 Onychocephalus nigro-lineatus HALLOWELL, Proc. Acad. nat. Sci., Philadelphia, 1 8 4 8 : 6 0 . - T e r r a t y p i c a : Liberia. - H o l o t y p e : A.N.S.P. 3302. 1864 Typhlops Kraussi JAN, Iconogr. gén. Ophid.; 1 (3. livr.): 3; pi. 6, fig. 2. - T e r r a t y p i c a : Gold Coast. - S y n t y p e s : N.M.B. 299 and 300. 1864 TyphlopsHallowelli JAN, Iconogr. gén. Ophid., (1 (4. livr.)3; pis. 4, 5, fig. 6. T e r r a t y p i c a : Gold Coast. - H o l o t y p e : N.M.B. 311. 1864 Typhlops liberiensis - JAN, Iconogr. gén. Ophid., 1 (5. livr.): 4; pis. 5, 6, fig. 1. 1864 Typhlops liberiensis var. intermedia JAN, Iconogr. gén Ophid., 1 (livr. 5): 4; pis. 5, 6; fig. 2. - T e r r a t y p i c a : Liberia. - H o l o t y p e : formerly in Zoologisches Museum, Hamburg; destroyed during World War II. 1874 Onychocephalus kraussi REICHENOW, Arch. Naturgesch., Berlin, 1874 (1).: 291. 1877 Typhlops (Ophthalmidion) eschrichtii, var. intermedia - PETERS, M.Ber. k. preuß. Akad. Wiss Berlin, 1877:614. 1887 Typhlops (Ophthalmidion) kraussi - BOCAGE, Jörn. Sci. math. phys. nat., Lisboa, 11:180. 1958 Typhlops leprosus TAYLOR & WEYER, Univ. Kansas Sci. Bull., Lawrence, 38: 1 2 0 4 . - T e r r a t y p i c a : Harbel, Liberia. - H o l o t y p e : F.M.N.H. 120231.
69 1964 Typhlops punctatus liberiensis - LAURENT, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 130 (6): 412. D i s c u s s i o n : ROUX-ESTÈVE (1974:79). R a n g e : Sierra Leone; Liberia; Guinea; Ivory Coast; and Ghana.
Typhlops pusillus
BARBOUR
1914 Typhlops pusillus BARBOUR, Mem. Mus. comp. Zool., Cambridge, (Massachusetts), 44 (2): 323. - T e r r a t y p i c a : Cape Haitien, Dept. du Nord, Haiti. - H o l o t y p e : M.C.Z. 8719. 1966 Typhlops pusilla - THOMAS, Proc. biol. Soc. Washington, 79:264. R a n g e : Hispaniola; lie Grande Cayemite; Ile de la Gonàve; Ile de la Tortue; Isla Catalina; and Isla Saona.
Typhlops reticulatus
(LINNAEUS)
1758 Anguis reticulata LINNAEUS, Syst. Nat., Ed. 10, 1: 228. - T e r r a t y p i c a : "America". - H o l o t y p e : none designated. 1782 Anguis rostralis WEIGEL, S.Ber. Ges. naturf. Freunde Berlin, 1782 (3): 191. T e r r a t y p i c a : Surinam. - H o l o t y p e : Z.M.B.? 1788 Anguis nasutus GMELIN, Syst. Nat., Ed. 13, 1:1120. - T e r r a t y p i c a : unknown. - H o l o t y p e : not traced. 1801 Anguis crocotatus SCHNEIDER, Hist. Amphib. 2: 340. - T e r r a t y p i c a : none given. - H o l o t y p e : not traced. 1802 Anguis rostratus DAUDIN, Hist. Nat. Rept., 7: 316. - In error for Anguis rostralis WEIGEL. 1844 Typhlops reticulatus - DUMÉRIL & BIBRON, Erpétol. gén., 6:282, pi. 60. 1845 Agryrophis reticulatus - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 137. 1851 Ophthalmidion crassum DUMÉRIL, Cat. méthod. Coll. Rept. Mus. Paris: 202. T e r r a t y p i c a : unknown. - H o l o t y p e : M.H.N.P. 208. 1863 Typhlops (Typhlops) lumbricalis var. Troscheli JAN, Elenco sist. Ofidi: 1 1 . T e r r a t y p i c a : unknown. - H o l o t y p e : M.S.N.M. - destroyed. 1863 Typhlops (Typhlops) reticulatus - JAN, Elenco sist. Ofidi: 11. 1864 Typhlops reticulatus nigrolactea JAN, Iconogr. gén. Ophid., 1 (4. livr.) 5: 3; pi. 6, fig. Id. - T e r r a t y p i c a : none given. - H o l o t y p e : M.S.N.M. - destroyed. 1864 Typhlops reticulatus Troscheli - JAN, Iconogr. gén. Ophid., 1 (4. livr.) 5:3; pi. 6, fig. lc. 1864 Typhlops reticulatus var. crassa - JAN, Iconogr. gén. Ophid., 1 (5. livr.): 4; pi. 5, fig. 7. 1938 Typhlops retisculatus NAKAMURA (error typographies), Dobutugaku Zasshi, Zool. Mag., Tokyo, 50 (4): 192. D i s c u s s i o n : ROZE (1966: 37); VANZOLINI (1972:27). R a n g e : Tropical South America east of the Andes.
Typhlops reuteri
BOETTGER
1881 Typhlops (Typhlops) Reuteri BOETTGER, Zool. Anz., Leipzig, 1881:650. T e r r a t y p i c a : Nossi Bé (= Nosy Bé), Madagascar. - H o l o t y p e : S.M.F. 16589. 1882 Typhlops (Typhlops) Lenzi BOETTGER, Zool. Anz., Leipzig, 1882:478. - T e r r a t y p i c a : Nossi Bé (= Nosy Bé), Madagascar. - L e c t o t y p e : S.M.F. 16590 (MERTENS, 1967:89). 1958 Typhlops reuteri - GUIBÉ, Mém. Inst. Sci. Madagascar, Tananarive, (A) 12:192. Range:
Madagascar.
70 Typhlops richardi DUMÉ RIL & BIBRON 1844
Typhlops Richardi DUMÉRIL & BIBRON, Erpétol. gén., 6 : 2 9 0 . - T e r r a t y p i c a : St. Thomas, Virgin Islands. - S y n t y p e s : M.H.N.P. 3220 and R.M.N.H. 3713. 1863 Typhlops (Typhlops) Richardi - JAN, Elenco sist. Ofidi: 12.
Discussion:
THOMAS (1966a: 190).
R a n g e : Virgin Islands; Puerto Rico; Caja de Muertos; Vieques; and Culebra Islands. Typhlops 1966
richardi
richardi
DUMÉRIL & BIBRON
Typhlops richardi richardi - THOMAS, Revista Biol, trop., San José, 1 3 : 1 9 0 .
R a n g e : St. Croix, St. Thomas, St. John, Tortola, and Prickly Pear Islands, Virgin Islands. Typhlops
richardi
c a p a p o n t u s THOMAS
1966 Typhlops richardi capapontus THOMAS, Revista Biol, trop., San José, 1 3 : 1 8 7 201; 6 figs. - T e r r a t y p i c a : Anegada Island, British Virgin Islands, in the vicinity of the the Settlement. - H o 1 o t y p e : M.C.Z. 77220.
R a n g e : Island of Anegada, British Virgin Islands. Typhlops 1966
richardi
naugus
THOMAS
Typhlops richardi naugus THOMAS, Revista Biol, trop., San José, 1 3 : 1 8 7 - 201, figs. - T e r r a t y p i c a : Virgin Island, on the hillside above Pond Bay, British Virgin Islands. - H o l o t y p e : M.C.Z. 77221.
R a n g e : Known only from the Island of Virgin Gorda, British Virgin Islands. Intermediates between T. r. richardii and T. r. naugus occur on Beef Island, Virgin Islands. Typhlops
richardi
platycephalus
DUMÉRIL & BIBRON
1844
Typhlops platycephalus DUMÉRIL & BIBRON, Erpétol. gén., 6 : 2 9 3 . - T e r r a t y p i c a : „Martinique" (error); corrected to Puerto Rico by STEJNEGER ( 1 9 0 4 : 687). - H o 1 o t y p e : M.H.N.P. 1066. 1863 Typhlops (Typhlops) platycephalus - JAN, Elenco sist. Ofidi: 12. 1965 Typhlops richardi platycephala - THOMAS, Revista Biol, trop., San José, 1 3 : 1 9 0 .
R a n g e : Puerto Rico, Caja de Muertos, Vieques and Culebra Islands, Cayo Palominitos, Cayo Diablo, and Cayo de Tierra. Typhlops rondoensis LOVERIDGE 1942
Typhlops tettensis rondoensis LOVERIDGE, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 9 1 : 2 5 6 . - T e r r a t y p i c a : Nchingidi, Rondo Plateau, southeastern Tanzania. - H o l o t y p e : M.C.Z. 48066.
71 1964 Typhlops rondoensis - LAURENT, Bull. Mus. comp. Zool., Cambridge (Massachusetts), 130 (6): 419. Discussion:
ROUX-ESTÈVE (1974:136).
R a n g e : Rondo Plateau and Newala region in southeastern Tanzania. Typhlops rostellatus
STEJNEGER
1904 Typhlops rostellatus STEJNEGER, Rep. U.S. nation. Mus., Washington, 1902:686; fig. 146 - 147. - T e r r a t y p i c a : Lares, Porto Rico. - H o l o t y p e : U.S.N.M. 25463. 1966 Typhlops rostellata - THOMAS, Proc. biol. Soc. Washington, 7 9 : 2 6 4 . Discussion:
RUTHVEN & GAIGE (1935 :11).
R a n g e : Puerto Rico. Typhlops ruber 1897
BOETTGER
Typhlops ruber BOETTGER, Zool. Anz., Leipzig, 2 0 : 1 6 4 . - T e r r a Samar Island, Philippines. - H o l o t y p e : S.M.F. 16616.
Discussion:
typica:
TAYLOR (1922:50);MCDOWELL (1974:12).
R a n g e : Known from Samar and Mindoro Islands, Philippines. Typhlops ruficaudus (GRAY) 1845 Anilios ruflcauda GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 136. - T e r r a t y p i c a : "Philippines". - S y n t y p e s : B.M.N.H. 1946.1.11.4-6. 1861 Typhlops (Anilios) ruflcauda - PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1861:684. 1861 Typhlops Jagorii PETERS, M.Ber. k. preufi. Akad. Wiss. Berlin, 1861:684. T e r r a t y p i c a : Mount Isarog, Camarines, Luzon, Philippines. - H o l o t y p e : N.M.W. 13294. 1863 Typhlops (Typhlops) dichromatus JAN, Elenco sist. Ofidi: 12. - T e r r a t y p i c a : "Monado?", later stated to be "Island of Rhodes" (in error). - H o l o t y p e : M.S.N.M. - now destroyed. 1864 Typhlops dichromatus - JAN, Iconogr. gén. Ophid., 1 (3. livr.): 3; pis. 4, 5, fig. 1. 1867 "Typhlops petersii STEINDACHNER, Verh. zool.-botan. Ges. Wien, 1867:515; pi. 13, figs. 7 - 9 . - T e r r a t y p i c a : "Philippines". - H o l o t y p e : N.M.W. 1886 Typhlops ruflcauda - BOETTGER, Ber. senckenb. naturf. Ges., Frankfurt am Main, 1886:104. Discussion:
MCDOWELL (1974:12).
R a n g e : Luzon, Tablas, and Marinduque Islands, Philippines. Typhlops siamensis
GÜNTHER
1864 Typhlops siamensis GÜNTHER, Rept. Brit. India: 175; pi. 16, fig. D. - T e r r a t y p i c a : "Siam". - H o l o t y p e : B.M.N.H. 1946. 1.11.69. 1943 Typhlops diardi (partim) - M. A. SMITH, Fauna Brit. India, Rept. Amphib., 3, (Serpentes): 5 1 - 5 2 .
72 1965 Typhlops siamensis - TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 45 (9): 654 655.
Range: Typhlops simoni
Thailand. BOETTGER
1879 Onychocephalus simoni BOETTGER, S.Ber. Ges. naturf. Freunde Berlin, 1878/ 1879: 58. - T e r r a t y p i c a : Syria; corrected to Haiffa (Israel) by MERTENS ( 1 9 6 7 : 8 7 ) . - L e c t o t y p e : S.M.F. 16673 (MERTENS, 1967:87). 1893 Typhlops simonii - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:51.
R a n g e : Israel and Syria. Typhlops socotranus
BOULENGER
1889 Typhlops socotranus BOULENGER, Ann. Mag. nat. Hist., London, (6) 4 : 362. T e r r a t y p i c a : Island of Socotra, Indian Ocean. - S y n t y p e s : B.M.N.H. 1946.1.11.72-73. 1903 Typhlops sokotranus STEINDACHNER, (error typographicus), S.Ber. Akad. Wiss. Wien, math.-naturwiss. Kl., 112:13. D I S C U S S I O N : ROUX-ESTEVE (1974:43).
R a n g e : Socotra Island, northwestern Indian Ocean. Typhlops stadelmani
SCHMIDT
1936 Typhlops stadelmani SCHMIDT, Proc. biol. Soc. Washington, 4 9 : 4 8 . - T e r r a t y p i c a : Subirana Valley, 2800 Ft., Yoro, Honduras. - H o l o t y p e : M.C.Z. 38701.
R a n g e : Honduras. Known only from the type locality. Typhlops steinhausi
WERNER
1909 Typhlops steinhausi WERNER, Mitt, naturhist. Mus. Hamburg, 2 6 : 2 0 9 . - T e r r a t y p i c a : Cameroon. - S y n t y p e s : Two, originally in Zoologisches Museum, Hamburg; destroyed during World War II. Neotype: M.H.N.P. 1964-139. 1911 Typhlops batesii BOULENGER, Ann. Mag. nat. Hist., London, (8) 8: 370. - T e r r a t y p i c a : Bitye, Cameroon. - S y n t y p e s : B.M.N.H. 1946.1.23.48-50. 1965 Typhlops steinhousi ROUX-ESTÈVE (error typographicus), Cah La Maboke, 3 (1): 51 - 52. D I S C U S S I O N : ROUX-ESTÈVE (1974:63).
R a n g e : Coastal Nigeria; southern Cameroon; southeastern Central African Republic. Typhlops sulcatus
COPE
1868 Typhlops sulcatus COPE, Proc. Acad. nat. Sci. Philadelphia, 2 0 : 1 2 8 . - T e r r a p i c a : Navassa Island, West Indies. - H o l o t y p e : U.S.N.M. 12371. 1966 Typhlops sulcata - THOMAS, Proc. biol. Soc. Washington, 79: 264.
ty-
73 1964 Typhlops haitiensis RICHMOND, Breviora, Mus. comp. Zool., Cambridge (Massachusetts), 2 0 2 : 5 - 6, fig. 3. - T e r r a t y p i c a : Manneville, D é p t de l'Ouest, Haiti. H o l o t y p e : M.C.Z. 62635. R a n g e : Southwestern Hispaniola, including the Tiburon Peninsula of Haiti w e s t t o t h e N o r m e D u b o i s P e n i n s u l a east o f A q u i n , t h e Cui de Sac-Valle de N e i b a plain, n o r t h t o 10.1 k m S E M o n t r o u i s , a n d t h e P e n i n s u l a de B a r a h o n a ; Isla A l t o V e l o ; Ile d e la G o n a v e ; Ile G r a n d e C a y e m i t e ; a n d Navassa Island.
T y p h l o p s syntherus THOMAS 1965 Typhlops syntherus THOMAS, Copeia, New York & c., 1965 (4): 436. - T e r r a t y p i c a : 5 km. north of Pedernales, Pedernales Province, Dominican Republic. H o l o t y p e : M.C.Z. 77215. R a n g e : K n o w n o n l y f r o m t h e l o w e l e v a t i o n areas o f t h e B a r a h o n a P e n i n sula o f H i s p i n o l a .
T y p h l o p s tasymicris THOMAS 1974 Typhlops tasymicris THOMAS, Occ. Pap. Mus. Zool. Louisiana State Univ., Baton Rouge, 4 6 : 1 - 5, fig. 1 . - T e r r a t y p i c a : 1 mile east of Vincennes, St. David Parish, Grenada, Lesser Antilles. - H o l o t y p e : F.S.M. 21547. Range:
G r e n a d a , Lesser Antilles.
T y p h l o p s tenebrarum TAYLOR 1947 Typhlops tenebrarum TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 31, pt. 2, (13): 292 - 294. - T e r r a t y p i c a : 12 miles north of Trincomalee, Ceylon. H o l o t y p e : formerly EHT-HMS 30063; now presumed lost. R a n g e : N e a r T r i n c o m a l e e , Sri L a n k a ( C e y l o n ) . K n o w n o n l y f r o m t h e type locality.
T y p h l o p s tenuicollis (PETERS) 1864 Onychocephalus (Ophthalmidion) tenuicollis PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1864: 272; pi., fig. 2. - T e r r a t y p i c a : Himalayas. - H o l o t y p e : Z.M.B. 1871 Typhlops theobaldianus STOLICZKA, J. asiat. Soc. Bengal, Calcutta 4 0 : 4 2 9 ; pi. 25, figs. 5 - 8 . - T e r r a t y p i c a : Assam. - H o l o t y p e : Indian Museum, Calcutta. 1876 Typhlops tenuicollis - THEOBALD, Cat. Rept. Brit. India: 123. Discussion: Range:
M. A. SMITH (1943:50).
K n o w n d e f i n i t e l y o n l y f r o m S a m a g u t i n , N a g a Hills, A s s a m , I n d i a .
T y p h l o p s tenuis SALVIN 1860 Typhlops tenuis SALVIN, Proc. zool. Soc. London, 1860:454. - T e r r a t y p i c a : Coban, Vera Paz, Guatemala. - H o l o t y p e : B.M.N.H. 1946.1.11.71.
74 1867
Typhlops basimaculatus COPE, Proc. Acad. nat. Sci. Philadelphia, 1866:320. T e r r a t y p i c a : Cordoba and Orizaba, Veracruz, Mexico. - H o l o t y p e : U.S.N.M. 6602. 1869 Typhlops perditus PETERS, M.Ber. k. preuß. Akad. Wiss. Berlin, 1869:435. T e r r a t y p i c a : Orizaba, Veracruz, Mexico. - H o l o t y p e : Z.M.U. 1885 Typhlops praelongus MÜLLER, Verh. naturf. Ges. Basel, 7:674. - T e r r a t y p i c a : Cordoba, Veracruz, Mexico. - N.M.B. 1939 Typhlops praelongis TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 26 (13): 442; misspelling. Discussion: Range: of M e x i c o .
STUART (1948:61).
M o d e r a t e e l e v a t i o n s in A l t a V e r a p a z , G u a t e m a l a , a n d G u l f area
T y p h l o p s thurstoni BOETTGER 1890 Typhlops thurstoni BOETTGER, Ber. senckenb. naturf. Ges., Frankfurt am Main, 1890:298. - T e r r a t y p i c a : Nilgiris Hills, South India. - H o l o t y p e : not traced. 1924 Typhlops walli PROCTER, Ann. Mag. nat. Hist., London, (9) 13:139, fig. - T e r r a t y p i c a : Wynaad, South India. - H o l o t y p e : B.M.N.H. 1946.1.11.53. 1936 Typhlops hurstoni BOURRET (error typographies), Les Serpents de l'Indochine, 2:11.
Discussion: Range:
M. A. SMITH (1943:49).
S o u t h India.
T y p h l o p s tindalli M. A . SMITH 1943 Typhlops tindalli M. A. SMITH, Fauna Brit. India, Rept. Amphib., 3 : 5 3 . t y p i c a : Nilambur, Malabar District, India. - S y n t y p e s : B.M.N.H. 1946.1.8.92-93 & 1946.1.8.95. Range:
Terra
Nilgiri Hills a n d M a l a b a r D i s t r i c t , I n d i a .
T y p h l o p s trangensis TAYLOR 1962 Typhlops trangensis TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 4 3 : 2 5 1 - 253, fig. 14. - T e r r a t y p i c a : Khao Chong, Forest Experiment Station, Trang Province, Thailand. - H o l o t y p e : F.M.N.H. 178236. Discussion: Range:
TAYLOR (1965:646).
Trang Province, Thailand. K n o w n only f r o m the type locality.
T y p h l o p s trinitatus RICHMOND 1965 Typhlops trinitatus RICHMOND, Proc. biol. Soc. Washington, 7 8 : 1 2 1 , fig. 1. T e r r a t y p i c a : Arima Road, 3 miles above Simla, Trinidad. - H o l o t y p e : A.M.N.H. 89820. Range:
Trinidad and Tobago.
75 Typhlops uluguruensis BARBOUR & LOVERIDGE 1928 Typhlops uluguruensis BARBOUR & LOVERIDGE, Mem. Mus. comp. Zool. Cambridge (Massachusetts), 5 0 (2): 104. - T e r r a t y p i c a : Nyange, 850 m, Uluguru Mountains, Tanzania. - H o l o t y p e : M.C.Z. 23080. Discussion:
ROUX-ESTÈVE ( 1 9 7 4 : 4 0 ) .
R a n g e : Uluguru Mountains, Tanzania.
Typhlops unilineatus DUMERIL & BIBRON 1844 Onychocephalus unilineatus DUMERIL t y p i c a : Cayenne, French Guiana. 1863 Typhlops (Onychocephalus) unilineatus 1893 Typhlops unilineatus - BOULENGER,
& BIBRON, Erpetol. gen., 6 : 2 7 8 . - T e r r a H o l o t y p e : M.H.N.P. 1064. - JAN, Elenco sist. Ofidi: 13. Cat. Snakes Brit. Mus. (Nat. Hist.) 1 : 1 5 .
R a n g e : Surinam and French Guiana.
Typhlops veddae TAYLOR 1947
Typhlops veddae TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 31, pt. 2 (13): 294 295. - T e r r a t y p i c a : 12 miles north of Trincomalee, Ceylon. - H o l o t y p e : F.M.N.H. 100033.
R a n g e : Near Trincomalee, Sri Lanka (Ceylon). Known only from the type locality.
Typhlops vermicularis MERREM 1803 Anguis lumbricalis DAUDIN, Hist. nat. Rept., 7 : 308. — T e r r a t y p i c a : none given. - T y p e : not traced. - Preoccupied by Anguis lumbricalis Linnaeus, 1758. 1820 Typhlops vermicularis MERREM, Tent. Syst. Amphib.: 158. - T e r r a t y p i c a : "Archipelago, Asia". Restricted to "Greek Islands" by MERTENS & MOLLER ( 1 9 2 8 : 4 9 ) . - T y p e : not traced. 1833 Typhlops flavescens BIBRON & BORY, in BORY, Exped. Sci. Moree, Zool.: 72; pi. 13, fig. 3. - T e r r a t y p i c a : unknown. - T y p e : not traced. 1845 Argyrophis vermicularis - GRAY, Cat. Lizards Brit. Mus. (Nat. Hist.): 137. 1863 Typhlops (Typhlops) vermicularis - JAN, Elenco sist. Ofidi: 12. 1864 Typhlops (Typhlops) syriacus JAN, Iconogr. gen. Ophid., 1 (3. livr.): 3; pis. 4, 5, fig. 5. - T e r r a t y p i c a : Beirut, Lebanon. -- H o l o t y p e : M.S.N.M. destroyed. 1869 Typhlops vermicalis BREHM (lapsus), Illustr. Thierleben, Ed. 1, 5 : 1 8 9 . 1874 Typhlopspersicus BLANFORD, Ann. Mag. nat. Hist., London, (4) 14: 34. T e r r a t y p i c a : southwest of Karman, 6000 f t . , southern Iran. - S y n t y p e s : B.M.N.H. 1946.1.11.87; other(?)s not traced. 1875 Typhlops lumbricalis (not of LINNAEUS) - SCHREIBER, Herpetol. Europ.: 315.
R a n g e : Yugoslovia, Albania, southern Bulgaria, Transcaucasian republics of the U.S.S.R. (Dagestan, southern Turkmenia, southern Uzbekistan, and southern and western Tajikistan), Greece, Ionian Islands, Rhodes, Turkey, Syria, Israel, Lebanon, lower Egypt, Iraq, Iran, and Afghanistan.
76 Typhlops violaceus TAYLOR 1947 Typhlops violaceus TAYLOR, Univ. Kansas Sci. Bull., Lawrence, 31, pt. 2, (13): 289 - 90. - T e r r a t y p i c a : 12 miles north of Trincomalee, Ceylon. - H o 1 o t y p e : F.M.N.H. 100068.
R a n g e : Near Trincomalee, Sri Lanka (Ceylon). Known only from the type locality. Typhlops wilsoni WALL 1908 Typhlops wilsoni WALL, Bombay J. nat. Hist. Soc., 18 (4): 796. - T e r r a t y p i c a : Maidan Mihaftan, 30 miles east of Schustar in southwest Iran. — H o 1 o t y p e : not traced.
R a n g e : Southwestern Iran. Known only from the type locality. Typhlops zenkeri STERNFELD 1908 Typhlops zenkeri STERNFELD, S.Ber. Ges. naturf. Freunde Berlin, 1908:92. T e r r a t y p i c a : Kribi, Cameroon. - H o l o t y p e : Z.M.B. 11091. 1914 Typhlops vermis BOULENGER, Ann. Mag. nat. Hist., London, (8) 14:482. T e r r a t y p i c a : Bitye, Cameroon. - S y n t y p e s : B.M.N.H. 1946.1.10.89-98 (10). Discussion:
ROUX-ESTÈVE (1974: 35).
R a n g e : Cameroon. Incertae Sedis Typhlops disparilis JAN 1860 Typhlops (Diaphorotyphlops) disparilis JAN, Iconogr. gén. Ophid., 1 (1. livr.): 1; pis. 5, 6, fig. 6. - T e r r a t y p i c a : none given. - H o l o t y p e : M.S.N.M. destroyed. 1881 Diaphorotyphlops disparilis - PETERS, S.Ber. Bes. naturf. Freunde Berlin, 1881 : 70. 1893 Typhlops disparilis - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:53.
R a n g e : Unknown. T y p h l o p s filiformis DUMÉRIL & BIBRON 1844 Typhlops filiformis DUMÉRIL & BIBRON, Erpétol. gén., 6 : 307. - T e r r a p i c a : none given. - H o l o t y p e : M.H.N.P. 929. 1863 Typhlops (Typhlops) filiformis - JAN, Elenco sist. Ofidi: 11.
ty-
R a n g e : Unknown. Typhlops fuscus (A. DUMÉRIL) 1851 Opthalmidion fuscum A. DUMÉRIL, Cat. method, coll. Rept., Mus. Paris: 203. T e r r a t y p i c a : "Java". - H o l o t y p e : M.H.N.P. 1062. 1863 Typhlops fuscus (Opthalmidion) - JAN, Elenco sist. Ofidi: 13.
R a n g e : Unknown.
77
Typhlops longissimus
DUMERIL & BIBRON
1844 Ophthalmidion longissimum DUMERIL & BIBRON, Erpetol. gen., 6:263. - T e r r a t y p i c a: "North America". - H o 1 o t y p e: M.H.N.P. 1061. 1864 Typhlops longissimus - JAN, Iconogr. gen. ophid., 1 (5. livr.): 4; pis. 5, 6, fig. 6. 1957 Ophthalmidium longissimum - LOVE RIDGE (error typographies), Bull. Mus. comp. Zool., Cambridge (Massachusetts), 117 (2): 240. Range:
Unknown.
Typhlops macrurus
(PETERS)
1860 Onychocephalus macrurus PETERS, M.Ber. k. preufi. Akad. Berlin, 1860:78, pi. - T e r r a t y p i c a : none given. - H o l o t y p e : not traced. D I S C U S S I O N : ROUX-ESTEVE ( 1 9 7 4 : 2 4 9 ) .
Range:
Unknown.
Typhlops melanocephalus
( D U M E R I L & BIBRON)
1844 Cathetorhinus melancephalus DUMERIL & BIBRON, Erpetol. gen., 6:270. T e r r a t y p i c a : none given. - H o l o t y p e : M.H.N.P. 138. 1860 Typhlops (Typhlina) melanocephalus - JAN, Iconogr. gen. Ophid., 1 (1. livr.): 1; pis. 5, 6, fig. 8. 1863 Typhlops (Cathetorhinus) melanocephalus - JAN, Elenco sist. Ofidi: 10. 1893 Typhlops melanocephalus - BOULENGER, Cat. Snakes Brit. Mus. (Nat. Hist.) 1:15. Range:
Unknown.
C o m m e n t : Probably belongs in the genus Typhlina, although R o u x ESTEVE ( 1 9 7 4 : 1 5 7 ) placed C. melanocephalus in die genusRhinotyphlops.
Typhlops mutilatus
WERNER
1900 Typhlops mutilatus WERNER, Zool. Anz., Leipzig, 23:196; fig. 1 - 2. t y p i c a : Malacca, Malaya. - S y n t y p e s : Z.M.B. (3) - now lost. Range:
Terra
Unknown; not from Malaya.
Typhlops psittacus
WERNER
1903 Typhlops psittacus WERNER, Zool. Anz., Leipzig, 26 (693): 248. t y p i c a : "Mexico". - H o l o t y p e : I.R.S.N. 2017.
Terra
D i s c u s s i o n : TAYLOR (1939: 443). Range:
Unknown; probably not Mexican in origin.
Typhlops textilis JAN 1863 Typhlops (Typhlops) textilis JAN, Elenco sist. Ofidi: 1 1 . - T e r r a unknown. - H o l o t y p e : M.S.N.M. - destroyed. Range:
Unknown.
typica:
78 T y p h l o p s verticalis ( A . SMITH) 1846 Onychocephalus verticalis A. SMITH, Illustr. Zool. S. Afr., Rept.: pi. 54, figs. A & 1 8 - 2 0 . - T e r r a t y p i c a : Interior of South Africa. - H o 1 o t y p e : B.M.N.H. 1946.1.10.99. 1887
Typhlops verticalis - BOULENGER, The Zoologist, London, ( 3 ) 1 1 : 1 7 4 .
Discussion: Range:
ROUX-ESTEVE (1974: 249). Unknown. Fossil Species
T y p h l o p s cariei HOFFSTETTER 1946 Typhlops cariei HOFFSTETTER, Bull. Mus. nation. Hist. Nat., Paris, (2) 18: 3: 313 314. - T e r r a t y p i c a : Island of Mauritius, Indian Ocean. - H o l o t y p e : M.H.N.P. Range:
K n o w n only f r o m Mauritius.
T y p h l o p s grivensis HOFFSTETTER 1946 Typhlops grivensis HOFFSTETTER, Bull. Mus. nation. Hist, nat., Paris, (2) 18: 3: 311 - 313, fig. 1. - T e r r a t y p i c a : Eocene of Isére, France. - H o l o t y p e : in Museum de Lyon, France. Range:
E o c e n e o f Isère, F r a n c e . K n o w n o n l y f r o m t h e t y p e l o c a l i t y .
Typhlops Edwardsi (= Omoiotyphlops priscus ROCHEBRUNE, 1 8 8 4 : 1 4 9 ) , Odontomophis atavus ROCHEBRUNE ( 1 8 8 4 : 1 4 9 ) , a n d t h e g e n u s Simoliophis SAUVAGE ( 1 8 8 0 : 6 7 1 ) h a v e b e e n s h o w n n o t t o b e l o n g t o t h e f a m i l y T y p h l o p i d a e (HOFFSTETTER: 1 9 4 6 : 3 0 9 .
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SAUVAGE, H. E. (1880); Sur l'existence d'un Reptile du Type Ophidien dans les couches à Ostrea columba des Charentes. C.R. hebd. Séanc. Acad. Sci. Paris. 9 1 : 6 7 1 - 672. SCHMIDT, K. P. (1953): Amphibians and Reptiles of Yemen. - Fieldiana, Zoology, Chicago, 34 (24): 253 - 261. - & WALKER, W. F. (1943): Snakes of the Peruvian Coastal Region. - Pubi. Field Mus. nat. Hist., Chicago, (Zool. Ser.) 2 4 : 2 9 7 - 338. SMITH, H. M. & GRANT, C. (1958): New and Noteworthy Snakes from Panama. - Herpetologica, Escondido, 14:207 - 215. - & LIST, J. C. (1958): The Type of the Blind Snake Stenostomaalbifrons. Herpetologica, Escondido, 13: 271.
81 - & SANDERS, O. (1952): Distributional Data on Texan Amphibians and Reptiles. Texas J. Sci., San Marcos 1952 (2): 204 - 219. SMITH, M. A. (1943): The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese sub-region, Reptilia and Amphibia. 3, Serpentes, xii + 583 pp. - Taylor and Francis, London. STEJNEGER, L. (1891): Notes on some North American Snakes. - Proc. U.S. nation. Mus., Washington, 1 4 : 5 0 1 - 505. STUART, L. C. (1948): The Amphibians and Reptiles of Alta Verapaz Guatamala. - Misc. Pubi. Mus. Zool. Univ. Michigan, Ann Arbor, 6 9 : 1 - 109. TAYLOR, E. H. (1922): The Snakes of the Philippine Islands. - Philippine Bur. Sci., Manila, Monogr. 16, 312 pp., 37 pis., 32 text figs. - (1939): Mexican Snakes of the Genus Typhlops. - Univ. Kansas Sci. Bull., Lawrence, 26 (13): 441 - 444. - (1939a): Herpetological Miscellany No. 1. - Univ. Kansas Sci. Bull., Lawrence, 26 (15): 489 - 571. - (1947): Comments on Ceylonese Snakes of the Genus Typhlops with Descriptions of New Species. - Univ. Kansas Sci. Bull., Lawrence, 31, pt. 2 (13): 283 - 298. - (1965): The Serpents of Thailand and adjacent Waters. - Univ. Kansas Sci. Bull., Lawrence, 45 (9): 609 - 1096. THOMAS, R. (1965): The Genus Leptotyphlops in the West Indies with Description of a New Species from Hispaniola (Serpentes, Leptotyphlopidae). - Breviora, Mus. comp. Zool., Cambridge (Massachusetts), 2 2 2 : 1 - 12. - (1965a): A New Species of Typhlops from the Barahona Peninsula of Hispaniola. - Copeia, Washington, 1965 (4): 436 - 439. - (1966): Leeward Islands Typhlops (Reptilia, Serpentes). - Proc. biol. Soc. Washington, 7 0 : 2 5 5 - 266. - (1966a): A Reassessment of the Virgin Islands Typhlops with the Description of Two New Subspecies. - Rev. Biol, trop., San José, 13 (2): 187 - 201. - (1968) The Typhlops biminiensis Group of Antillean Blind Snakes. - Copeia, Washington, 1968 (4): 7 1 3 - 722. - (1974): A New Species of Lesser Antillean Typhlops (Serpentes: Typhlopidae). - Occ. Pap. Mus. Zool. Louisiana St. Univ., Baton Rouge, 4 6 : 1 - 5. - (1975): The Hemipenis of Leptotyphlops tenella KLAUBER (Serpentes: Leptotyphlopidae) and a New Distributional Record. - J. Herpetol., Lawrence, 9 (2): 250 - 251. VANZOLINI, P. E. (1972): Typhlops brongersmai spec. nov. from the Coast, of Bahia, Brasil (Serpentes, Typhlopidae). - Zool. Meded., Leiden, 47 (3): 27 - 29. WAITE, E. R. (1918): Review of the Australian Blind Snakes (Family Typhlopidae). - Ree. S. Austral. Mus., Adelaide, 1 : 1 - 34. - (1929): The Reptiles and Amphibians of South Australia. - Gov. Print., Adelaide: 189 - 198. WEBB, R. G. (1970): Reptiles of Oklahoma. - Stovall Mus., Univ. Oklahoma, i x - x i + 370 p. WERNER, F. (1917): Versuch einer Synopsis der Schlangenfamilie der Glauconiiden. - Mitt, zool. Museum, Hamburg, 1917:191 - 208. - (1921): Synopsis der Schlangenfamilie der Typhlopiden auf Grund des Boulenger'sehen Schlangenkatalogs ( 1 8 9 3 - 1 8 9 6 ) . - Arch. Naturg., Berlin 87, A (7): 266 - 338. WILSON, L. D. & D. E. HAHN (1973): The Herpetofauna of the Isias de la Bahia, Honduras. Bull. Florida State Mus., Gainesville, (Biol. Sci.) 17 (2): 93 - 150. WORRELL, E. (1963): Reptiles of Australia. - xv + 207 pp. - Sydney.
Addendum While this paper was being edited, a number of significant works have appeared or have been accepted for publication dealing with the Leptotyphlopidae and Typhlopidae. To make this checklist as up-to-date and as useful as possible, a few comments on these publications appear to be in order. STIMSON, R O B B , and UNDERWOOD ( 1 9 7 7 ) showed that the genus Typhlina was based on two species, Acontias lineatus REINWARDT, a nomen nudum at the time, and Typhlops sentemstriatus SCHNEIDER, a species now belonging in the genus Leptotyphlops. The next available generic name for the Australian area typhlopids is Ramphotyphlops FITZINGER, 1 8 4 3 . The International Commission on Zoological Nomenclature was requested to supress the name Typhlina WAGLER, 1 8 3 0 , and to place the generic names Leptotyphlops FITZINGER, 1 8 4 3 , and Ramphotyphlops FITZINGER, 1 8 4 3 on the Official List of Generic Names in Zoology. SMITH ( 1 9 7 7 ) was in agreement with these findings and recommendations. HAHN ( 1 9 7 8 ) reviewed the Asian and Socotran Island Leptotyphlops forms, and named one new species: Leptotyphlops wilsoni
HAHN
1978 Leptotyphlops wilsoni HAHN, J. Herpet., Lawrence, 12 (4): 4 7 7 - 4 8 9 . - T e r r a t y p i c a : Kirschon, Socotra Island. - H o l o t y p e : B.M.N.H. 1957.1.10.28.
R a n g e : Socotra Island, northwestern Indian Ocean. In the same paper Leptotyphlops erythraeus,L. braccianii, and L. phillipsi were shown to be conspecific with L. macrorhynchus. Leptotyphlops nursi was considered to be a race of L. blanfordi. Glauconia laticeps was removed from the synonymy of L. hamulirostris, and tentatively placed in that of/,. b. blanfordi. THOMAS (1976), in an as yet unpublished dissertation, has made a significant contribution to our understanding of the systematics of the Antillean Typhlops. — HAHN (1977) reviewed ROUX-ESTEVE'S "Revision Systematique des Typhlopidae d'Afrique, Reptilia — Serpentes" and provided a summary taxonomic innovations. The revival of the genus Rhinotyphlops was questioned. — HAHN (1979a) showed that Leptotyphlops amazonicus belongs in the synonomy of L. signatum. — HAHN (1979b, c, d; in press) summarized the literature dealing with the family Leptotyphlopidae, the genus Leptotyphlops, and the species L. humilis, L. dulcis, andZ,. maximus. - ZUG (1977) reported on the distribution and variation of Leptotyphlops tricolor. - HOOGMOED (1977) reviewed the Leptotyphlops species occurring in Surinam.
Literature Cited HAHN, D. E. (1977): Review of Revision Systématique des Typhlopidae d'Afrique, Reptilia Serpentes. - J. herpet Assoc. Africa, Umtali, (15): 34-39. - (1978): A Brief Review of the Genus Leptotyphlops (Serpentes: Leptotyphlopidae) of Asia, with Description of a New Species. - J. Herpet., Lawrence, 12 (4): 477-489. - (1979a): The Identity of the Blind Snake Stenostoma signatum JAN, 1861 (Serpentes: Leptotyphlopidae). - Herpetologica, Lawrence, 35 (1): 57-60. - (in press, 1979b): Leptotyphlopidae; Leptotyphlops. - Cat. amer. Amph. Rept., New York, 230: 1 - 4 . - (in press, 1979c): Leptotyphlops dulcis. - Cat. amer. Amph. Rept., New York, 231: 1-2.
- (in press, 1979d): Leptotyphlops humilis. - Cat. amer. Amph. Rept., New York, 232: 1-4. - (in press): Leptotyphlops maximus. - Cat. amer. Amph. Rept., New York. HOOGMOED, M. S. (1977): On a New Species of Leptotyphlops from Surinam, with Notes on the other Surinam Species of the Genus (Leptotyphlopidae, Serpentes). - Notes on the Herpetofauna of Surinam V. - Zool. Meded., Leiden, 51 (7): 99-123. SMITH, H. M. (1977): Comment on the Proposed Conservation of Leptotyphlops and Ramphotyphlops Z. N. (S.) 2155. - Bull. zool. Nomencl., 34 (3): 141-142. STIMSON, A. F., J. ROBB & G. UNDERWOOD (1977): Leptotyphlops and Ramphotyphlops FITZINGER, 1843 (Reptilia, Serpentes): Proposed Conservation under the Plenary Powers. Z. N. (S.) 2155. - Bull. zool. Nomencl., 33 (3-4): 204-207. THOMAS, R. (1976): Systematics of the Antillean Blind Snakes of the Genus Typhlops (Serpentes: Typhlopidae). Diss. Abstr. Int., Ann Arbor, 37 (6): 1 - 4 . ZUG, G. R. (1977): Distribution and Variation of Leptotyphlops tricolor. - Copeia, Washington, (4): 744-745.
Register accedens, Typhlops 39 acuticauda, Typhlina 37 - , Typhlops 37 acutiiostratus, Typhlops 29 acutirostris, Typhlops 35 acutus, Grypotyphlops 49 - , Gryptotyphlops 49 - , Onychocephalus 49 -.Typhlops 49,50 adamsi, Typhlops 47 adolfi, Typhlops 50 affine, Stenostoma 6 affinis, Leptotyphlops 6 - , Ramphotyphlops 37 - , Typhlina 37 - , Typhlops 37 albanalis, Typhlops 66 albiceps, Typhlina 37 - , Typhlops 37 albifrons albipunctata, Stenostoma 7 - ater, Leptotyphlops 15 - , Glauconia 6 Leptotyphlops 6 - magnamaculata, Leptotyphlops 15 - margaritae, Leptotyphlops 15 - rubrolineata, Glauconia 25 - , Stenostoma 6 - tenella, Leptotyphlops 27 - tesselata, Stenostoma 17 albipuncta, Stenostoma undecimstriatus 7 - , Stenostoma albifrons 7 - , Typhlops undecimstriatus 7 albipunctus, Leptotyphlops 7 albirostris, Liotyphlops 3 Rhino typhlops 3 algeriensis, Glauconia 20 aluensis, Ramphotyphlops 42 Typhlops 41 alvensis, Typhlops 42 amazonicus, Leptotyphlops 7 ammodytes, Typhlops 41 anchiate, Typhlops 29 anchietae, Onychocephalus 29 - , Typhlops 29 andamanensis, Typhlops 50 angeli, Typhlops 64 angolensis angolensis, Typhlops 50 - blanfordi, Typhlops 62 - dubius, Typhlops 50 - irsaci, Typhlops 50
Onychocephalus 50 - polylepis, Typhlops 50 - symoensi, Typhlops 50 - . T y p h l o p s 50 - , Typhlops angolensis 50 angusticeps, Onychocephalus 38 - . T y p h l i n a 38 - . T y p h l o p s 38 Anilios 49 Anomalepididae 1 Anomalepis 1 Anomalolepis 1 anomalus, Onychocephalus 29 Rhino typhlops 29 - . T y p h l o p s 29 anops, Helminthophis 3 - , Liotyphlops 3 anthracinus, Leptotyphlops 7 arenarius, Onychocephalus 51 - . T y p h l o p s 51 arenicola, Typhlops braminus 39 Argyrophis 49 Aspidorhynchus 49 aspinosus, Anomalepis 1 ataeniatus, Typhlops unitaeniatus 36 atavus, Odontomophis 78 ater, Anilios 51 - ater, Typhlops 51 - , Gerrhophilus 51 - , Leptotyphlops 15 - , Leptotyphlops albifrons 15 - , Leptotyphlops goudotii 15 - suturalis, Typhlops 51 - . T y p h l o p s 51 - , Typhlops ater 51 australis, Typhlops 38 - , Leptotyphlops 7 - , Ramphotyphlops 38 - , Typhlina 38 - . T y p h l o p s 38 avakubae, Typhlops 30
bakewelli, Leptotyphlops 15 - , Leptotyphlops phenops 16 barmanus, Typhlops 56 barrowii, Onychophis 68 basimaculatus, Typhlops 74 batesii, Typhlops 72 batilla, Typhlina 38
85 batillus, Ramphotyphlops 38 - , Typhlops 38 beanyordi, Leptotyphlops 8 becki, Typhlops 48 beddomii, Typhlops 39 bergi, Typhlops 47 beui, Helminthophis 4 bianconii, Ophthalmidion 58 - . T y p h l o p s 58 bibronii, Onychocephalus 51 - , Typhlops 51 bicolor, Argyrophis 56 - , Glauconia 7 - gruveli, Glauconia 7 - , Leptotyphlops 7 - , Onychocephalus 38 - , Stenostoma 7 - , Tricheilostoma 7 - . T y p h l o p s 38 bilineatum, Stenostoma 8 - , Tetracheilostoma 8 bilineatus, Epictia 8 - , Eucephalus 8 - , Leptotyphlops 8 -.Typhlops 8 bilmaensis, Leptotyphlops macrorhynchus 20 biminiensis biminiensis, Typhlops 52 - epatica, Typhlops 52 - paradoxus, Typhlops 52 - . T y p h l o p s 52 - , Typhlops biminiensis 52 bipartitus, Typhlops 52 bisubocularis, Typhlops 5 2 bituberculata, Typhlina 39 bituberculatus, Onychocephalus 39 - , Ramphotyphlops 39 - . T y p h l o p s 39 blanfondi, Glauconia 8 blanfordi, Glauconia 8 - , Leptotyphlops 8 - lestradei, Typhlops 50 - . T y p h l o p s 62 - , Typhlops angolensis 62 bocagei, Typhlops 62 boettgeri, Glauconia 17 - , Leptotyphlops humilis 17 - , Typhlops 51 bondensis, Helminthophis 3 borrichiana, Glauconia 8 borrichianus, Leptotyphlops 8 bothriorhynchus, Typhlops 56 boueti, Glauconia 22 - , Leptotyphlops narirostris 22 boulengeri, Leptotyphlops 8 - , Glauconia 62 - , Typhlops 8 - usambaricus, Typhlops 62 boy lei, Typhlops 53 braccianii, Glauconia 9 - , Leptotyphlops 9
bramicus, Argyrophis 39 bramina, Typhlina 39 braminus arenicola, Typhlops 39 - , Eryx 39 - pallidus, Typhlops 39 - pammeces, Typhlops 66 - . T y p h l o p s 39 brasiliensis, Leptotyphlops 9 braueri, Glauconia 39 - . T y p h l o p s 39 brevicauda, Glauconia 9 - , Stenostoma 9 brevicaudus, Leptotyphlops 9 bressoni, Leptotyphlops 9 brevirostralis, Glauconia 19 brevis, Rhinotyphlops schlegelii 34 - . T y p h l o p s 34 - , Typhlops schlegelii 34 brevissimus, Leptotyphlops 9 brongersmai, Typhlina polygrammica 45 - , Typhlops 53 - , Typhlops florensis 45 - , Typhlops polygrammicus 45 brongersmianus, Typhlops 53 broomi, Ramphotyphlops 40 -.Typhlina 40 - , Typhlops 40 buchholzi, Ophthalmidion 55 - . T y p h l o p s 55 buehleri, Ramphotyphlops 42 - , Typhlops 42 burii, Glauconia 9 - , Leptotyphlops 9 caeca, Lethobia 29 caecatus, Typhlops 53 caecus caecus, Typhlops 30 - , Onychocephalus 29 - pitmani, Typhlops 30 - , Rhinotyphlops 29 - , Typhlops 29 - , Typhlops caecus 30 cahuilae, Leptotyphlops humilis 17 cairi, Glauconia 10 - , Leptotyphlops 10 - , Stenostoma 10 calabresi, Typhlops cuneirostris 55 canellei, Helminthophis 3 canlaonensis, Typhlops 53 capapontus, Typhlops richardi 70 capensis, Onychocephalus 39 - , Typhlops 54 capitulata capitulata, Typhlops 54 - , Typhlops capitulata 54 capitulatus capitulatus, Typhlops 54 - gonavensis, Typhlops 54 - , Typhlops 53 - , Typhlops capitulatus 54 caracasensis, Liotyphlops 3 cariei, Typhlops 78
86 carltoni, Glauconia 8 Cathetorhinos 37 Catodon 6 caymanensis, Typhlops 54 Cephalolepis 5 ceylonicus, Typhlops 54 chumilis, Leptotyphlops 17 cinereus, Typhlops 56,57 coecatus, Typhlops 53 collaris, Leptotyphlops 10 collenettei, Helminthophis 4 Colombia, Anomalepis 1 colombius, Anomalepis 1 colombi, Leptotyphlops 10 columbi, Leptotyphlops 10 comorensis, Typhlops 54 congestus, Onychocephalus 68 - , Typhlops 68 - , Typhlops punctatus 68 congicus, Onychocephalus 50 - , Typhlops 50 conjuncta distanti, Leptotyphlops 12 - , Glauconia 10 conjunctum, Stenostoma 10 conjunctus, conjunctus, Leptotyphlops 10 - incognitus, Leptotyphlops 11 - latirostris, Leptotyphlops 11 - lepezi, Leptotyphlops 11 - , Leptotyphlops 10 - , Leptotyphlops conjunctus 10 conradi, Typhlops 54 costaricensis, Typhlops 55 crassa, Typhlops reticulatus 69 crassatus, Onychocephalus 68 - . T y p h l o p s 68 crassum, Ophthalmidion 69 crocotatus, Anguis 69 crossii, Rhinotyphlops 30 - , Typhlops 30 cubae, Typhlops 63 cucutae, Liotyphlops 3 cumingii mansuetus, Typhlops 37 - , Onychocephalus 40 - , Onychophis 40 - , Typhlina 40 - , Typhlops 40 cuneirostris calabresii, Typhlops 55 - , Typhlops 55 - , Typhlops cuneirostris 55 - cuneirostris, Typhlops 55 cupinensis, Leptotyphlops 11 curtus, Typhlops 43 curvirostris, Typhlops 48
delalandei paucisquamata, Typhlops 31 - , Typhlops 31 - , Onychocephalus 31 dentatus, Anomalepis 1 depressiceps, Typhlops 55 depressus, Typhlops 41 Diaphrotyphlops 49 diaplocius, Leptotyphlops 12 diardi diardi, Typhlops 56 - muelleri, Typhlops 56, 57 - nigroalbus, Typhlops 57 -.Typhlops 56,71 - , Typhlops diardi 56 dichromatus, Typhlops 71 dimidiata, Leptotyphlops 12 dimidiatum, Stenostoma 12 dimidiatus, Leptotyphlops 12 dinga, Onychocephalus 34 - , Rhinotyphlops schlegelii 35 - , Typhlops 34 disparilis, Diaphorotyphlops 76 - . T y p h l o p s 76 dissecta, Glauconia 13 dissectus, Leptotyphlops dulcis 13 - , Leptotyphlops myopicus 13 dissimile, Leptotyphlops 12 - , Stenostoma 12 dissimilis, Glauconia 12 -.Leptotyphlops 12 distanti, Glauconia 12 - , Leptotyphlops 12 - , Leptotyphlops conjuncta 12 diversa, Typhlina 40, 41 diversiceps, Typhlops 60 diversus, Ramphotyphlops 41 - . T y p h l o p s 40 dominicana, Typhlops 57 dominicanus, Typhlops 57 dubius, Typhlops 50 - , Typhlops angolensis 50 dugandi, Leptotyphlops 12 dugesii, Catodon 17 - , Leptotyphlops humilis 17,18 - , Rena 18 - , Siagonodon 17 dulce, Rena 13 - , Stenostoma 13 dulcis dulcis, Leptotyphlops 13
decorsei, Typhlops 55 decorosus, Ophthalmidium 55 - , Typhlops 55 dendrophis, Typhlops 40 debilis, Glauconia 11 -.Leptotyphlops 11
edwardsi, Typhlops 78 elberti, Typhlina polygrammica 46 - , Typhlops 46 - , Typhlops florensis 46 - , Typhlops polygrammicus 46 elegans, Ophthalmidion 57
- dissectus, Leptotyphlops 13 -.Glaucoma 13 -.Leptotyphlops 13 - , Leptotyphlops dulcis 13 - . R e n a 13
87 - , Typhlops 57 emini emini, Leptotyphlops 23 - , Glauconia 23 - , Leptotyphlops 23 - , Leptotyphlops emini 23 - pembae, Leptotyphlops 23 emunctus, Helminthophis 3 - , Idiotyphlops 3 - , Typhlops 3 endotera, Typhlina 41 endoterus, Ramphotyphlops 41 - , Typhlops 41 epatica, Typhlops biminiensis 52 Epictia 6 erycina, Typhlina 41 erycinus, Typhlops 41 erythraea, Glauconia 14 erythraeus, Leptotyphlops 14 - . T y p h l o p s 35 eschrichtii, Aspidorhynchus 67 - intermedia, Ophthalmidion 68 - intermedia, Typhlops 68 - , Ophthalmidion 68 - . T y p h l o p s 67 Eucephalus 5 eupinensis, Leptotyphlops 11 euproctus, Typhlops 39 excentricus, Typhlops 34 - , Typhlops schlegelii 35 excipiens, Typhlops 49 exignus, Typhlops 57 exiguus, Typhlops 57 exocoeti, Typhlina 41 - , Typhlops 41 fallax, Stenostoma 15 feae, Rhinotyphlops 30 - . T y p h l o p s 30 fìechteri, Glauconia 19 filiformis, Glauconia 14 - , Leptotyphlops 14 - . T y p h l o p s 76 fitzingeri, Glauconia 14 - , Leptotyphlops 14 - , Stenostoma 14 flavescens, Typhlops 75 flaviceps, Anomalepis 1 flavifrons, Stenostoma 21 flaviventer, Typhlina 4 1 , 4 2 - . T y p h l o p s 41 flavoterminatus, Helminthophis 2 - , Idiotyphlops 2 - , Typhlops 2 fletcheri, Typhlops 39 florensis brongersmai, Typhlops 45 - elberti, Typhlops 46 - florensis, Typhlops 46 - , Typhlina polygrammica 46 - , Typhlops 46 - , Typhlops florensis 46
- , Typhlops polygrammicus 46 - undecimlineatus, Typhlops 46 floris, Typhlops 46 floweri, Typhlops 57 fordii, Onychophis 31 fornasinii, Typhlops 58 franklini, Onychophis 31 frontalis, Helminthophis 2 - , Typhlops 2 fusconotus, Typhlops 57 fuscus, Typhlops 76 gadowi, Leptotyphlops 16 geotomus, Typhlops monastus 65 Gerrhopilus 49 gestri, Glauconia 14 - , Leptotyphlops 14 giadinhensis, Typhlops 58 gierrai, Typhlops 58, 62 - , Typhlops punctatus 58 Glauconia 6 gonavensis, Typhlops 54 - , Typhlops capitulatus 54 goudotii ater, Leptotyphlops 15 - bakewelli, Leptotyphlops 16 - , Leptotyphlops 14, 15 - magnamaculatus, Leptotyphlops 15 - phenops, Leptotyphlops 15,16 - , Stenostoma 14 goudottii, Stenostoma 15 gracile, Stenostoma 7 - , Tricheilostoma 7 graciliar, Leptotyphlops 16 gracilior, Glauconia 16 - , Leptotyphlops 16 gracilis leptosoma, Typhlops 30 - polli, Typhlops 31 - , Rhinotyphlops 30 - , Typhlops 30 grandidieri, Typhlops 58 granti, Typhlops 59 graueri, Rhinotyphlops 31 - , Typhlops 31 graveri, Typhlops 31 grivensis, Typhlops 78 groutii, Stenostoma 10 gruveli, Glauconia 7 - , Glauconia bicolor 7 grypha, Typhlina 42 Grypothyphlops 49 Gryptotyphlops 49 guadeloupensis, Typhlops 59 guayaquilensis, Leptotyphlops 16 guentheri, Helminthophis 3 - , Liothyphlops 3 - , Onychocephalus 42 - , Ramphotyphlops 42 - , Typhlina 42 - , Typhlops 42 guirrae, Typhlops 58
88 haitiensis, Typhlops 73 hallowelli, Typhlops 68 hamulirostris, Glauconia 16 - , Leptotyphlops 16 hectus, Typhlops 59 hediaea, Typhlops 51 Helminthophis 2 horsfieldi, Argyrophis 56 hottentotus, Typhlops 35 humile, Stenostoma 17 humilis boettgeri, Leptotyphlops 17 - cahuilae, Leptotyphlops 17 - dugesii, Leptotyphlops 17,18 - , Glauconia 17 - humilis, Leptotyphlops 17 - , Leptotyphlops 15,16 - , Leptotyphlops humilis 17 - levitoni, Leptotyphlops 18 - lindsayi, Leptotyphlops 18 - , Rena 15,16 - segregus, Leptotyphlops 18 - , Siagonodon 17 - slevini, Leptotyphlops 17 - tenuiculus, Leptotyphlops 18 - utahensis, Leptotyphlops 19 humbo, Onychocephalus 35 - . T y p h l o p s 35 hurstoni, Typhlops 74 hypogius, Typhlops 59 hypsobothrius, Typhlops 59 Idiothyphlops 2 ihlei, Leptotyphlops 20 incertus, Helminthophis 4 - , Liothyphlops 4 incognitus, Leptotyphlops conjunctus 11 inconspicuus, Typhlops 39 infralabialis, Typhlops 47 inornatus, Typhlops 59 intermedia, Ophthalmidion eschrichtii 68 Typhlops eschrichtii 68 Typhlops liberiensis 68 iridescens, Typhlops 41 irsaci, Typhlops angolensis 50 jagorii, Typhlops 71 jamaicensis, Anguis 60 - . T y p h l o p s 60 jerdoni, Typhlops 60 joshuai, Leptotyphlops 19 kafubi, Glauconia 23 kaimosae, Typhlops 50 kapaladua, Typhlops 57 katangensis, Typhlops 30 keasti, Rhamphotyphlops 47 - , Typhlops 47 kenti, Ramphotyphlops 37 - , Typhlops 37 khoratensis, Typhlops 60
kibarae, Rhinotpyhlops 31 - , Typhlops 30, 31 kleebergi, Typhlops 32 klemmeri, TVphlops 60 koekkoeki, Typhlops 60 koniagui, Rhinoleptus 29 - . T y p h l o p s 29 koppesi, Leptotyphlops 19 koshunnensis, Typhlops 61 kraali, Typhlops 61 kraussi, Onychocephalus 68 - , Ophthalmidion 68 - . T y p h l o p s 68 labialis, Glauconia 19 - , Leptotyphlops 19 - , Typhlops 57 lalandei, Onychophis 31 - paucisquamosa, Typhlops 31 - , Rhino typhlops 31, 32 - , Typhlops 31 lankaensis, Typhlops 61 laticeps, Glauconia 16 latifrons, Glauconia 25 latirostris, Leptotyphlops conjunctus 11 - , Typhlops 34 leachi, Anilios 60 lehneri, Typhlops 61 lenzi, Typhlops 69 leonhardii, Typhlops 41 lepezi, Glauconia 11 - , Leptotyphlops conjunctus 11 leprosus, Typhlops 68 Leptatyphlops 17 leptosoma polli, Typhlops 31 - , Ramphotyphlops 42 - , Typhlina 42 - . T y p h l o p s 30 - , Typhlops gracilis 30 Leptotyphlopidae 5 Leptotyphlops 5 lestradei, Typhlops 50 - , Typhlops blanfordi 50 Letheobia 29 leucocephalus, Cephalolepis 5 - , Rhinotyphlops 32 - , Typhlops 32 leucomelas, Typhlops 61 leucoprocta, Typhlina 43 leucoproctus, Typhlops 43 leucostictus, Typhlops 61 levitoni, Leptotyphlops humilis 18 liberiensis intermedia, Typhlops 68 - , Onychocephalus 68 - , Typhlops 68 - , Typhlops punctatus 68, 69 ligata, Typhlina 43 ligatus, Typhlops 43 Ramphotyphlops 43 ligorostris, Typhlops 44
89 limbricki, Typhlops 39 lindsayi, Leptotyphlops humilis 18 lineata, Typhlina 43 - . T y p h l o p s 43 lineatum, Pilidion 43 - , Typhlinalis 43 lineatus, Acontias 43 - numatrana, Typhlops 43 - , Ramphotyphlops 43 - sumatranus, Typhlops 43 - , Typhlops 43 lineolatus lineolatus, Typhlops 62 - , Ophthalmidion 62 - tanganicus, Typhlops 62 - , Typhlops 62 - , Typhlops lineolatus 62 Liotyphlops 2 longicauda, Glauconia 19, 20 - , Typhlops 40 longicaudum, Stenostoma 19 longicaudus, Leptotyphlops 19 longissimum, Ophthalmidion 77 longissimus, Typhlops 77 lorenzi, Typhlops 63 loveridgei, Typhlops 63 lumbricalis, Anguis 63, 75 - troscheli, Typhlops 69 - , Typhlops 63, 75 lumbriciformis, Lethobia 32 - , Onychocephalus 32 - , Rhinotyphlops 32 lundensis, Typhlops praeocularis 33 luzonensis, Typhlops 63 mackinnoni, Typhlops 67 macrolepis, Leptotyphlops 20 - , Stenostoma 20 - , Tricheilostoma 20 macrorhynchum, Ramphostoma 20 —, Stenostoma 20 macrorhynchus bilmaensis, Leptotyphlops 20 - , Glauconia 20 - , Leptotyphlops 20 - , Leptotyphlops macrorhynchus 20 - macrorhynchus, Leptotyphlops 20 macrura, Glauconia 21 - , Leptotyphlops 21 macrurus, Leptotyphlops 20 - , Onychocephalus 77 - , Typhlops 77 madagascariensis, Typhlops 63 macruso, Typhlops 34 magnamaculata, Leptotyphlops 15 magnamaculatus, Leptotyphlops goudotii 15 malabaricus, Onychocephalus 49 malaisei, Typhlops 38 malcolmi, Typhlops 64 mandensis, Typhlops 34 manilae, Typhlops 64
manni, Typhlops 64 mansuetus, Typhlops cumingii 37 margaritae, Leptotyphlops albifrons 15 maximus, Leptotyphlops 21 Meditoria 49 melanocephalus, Cathetorhinus 77 - , Typhlops 77 melanosterna, Stenostoma 21 melanostoma, Stenostoma 21 melanoterma, Stenostoma 21 melanotermus, Leptotyphlops 21 melanurus, Leptotyphlops 21 merkeri, Glauconia 26 - , Leptotyphlops scutifrons 26 metae, Liotyphlops 4 mexicanus, Anomalepis 1 microcephalus, Typhlops 39 microstomus, Typhlops 64 milleti, Typhlops 68 mindanensis, Typhlops 40 minima, Typhlina 43, 44 minimus, Ramphotyphlops 44 - , Typhlops 43 mirus, Diaphorotyphlops 64 - , Typhlops 64 monastus geotomus, Typhlops 65 - monastus, Typhlops 65 - . T y p h l o p s 65 - , Typhlops monastus 65 monensis, Typhlops 65 monochrous, Typhlops 56 monochrus, Typhlops 55 monticola, Glauconia 23 mossambicus, Onychocephalus 58 - . T y p h l o p s 58 mossombicus, Typhlops 58 mucronatus, Ophthalmidion 65 - , Typhlops 65 mucroso, Typhlops schlegelii 35 mucruso, Onychocephalus 34 - , Rhinotyphlops schlegelii 34 - . T y p h l o p s 34 - , Typhlops schlegelii 35 - varius, Typhlops 34 muelleri, Typhlops diardi 56, 57 mulleri, Typhlops 56 multicarinatus, Onychophis 44 multilineata, Typhlina 44 multilineatus, Onychocephalus 44 - , Ramphotyphlops 44 - , Rhinotyphlops 44 - , Typhlops 44 munaoi, Leptotyphlops 21 mutilatus, Typhlops 77 myopicus dissectus, Leptotyphlops 13 - , Leptotyphlops myopicus 14 - myopicus, Leptotyphlops 14 narirostre, Glauconia 22 -.Stenostoma 22
90 narirostris boueti, Leptotyphlops 22 - , Glaucoma 22 - , Leptotyphlops 22 - , Leptotyphlops narirostris 22 - narirostris, Leptotyphlops 22 nasalis, Leptotyphlops 22 nasuta, Meditoria 60 nasutus, Anguis 69 natatrix, Glauconia 22 - , Leptotyphlops 22 naugus, Typhlops richardi 70 naveli, Typhlops 32 newtonii, Onychocephalus 32 - , Rhinotyphlops 32 - . T y p h l o p s 32 nicefori, Leptotyphlops 22 nigrescens, Anilios 46 - , Typhlina polygrammica 46 - , Typhlops 46,47 - , Typhlops polygrammicus 46 nigricans, Glauconia 23 - , Leptotyphlops 23 - , Leptotyphlops nigricans 23 - nigricans, Leptotyphlops 23 - pembae, Leptotyphlops 23 - , Stenostoma 23 - . T y p h l o p s 23 nigricauda, Typhlops 42 nigroalbus, Typhlops 56 - , Typhlops diardi 57 nigrolactea, Typhlops reticulatus 69 nigrolineatus, Onychocephalus 69 nigroterminata, Typhlina 44 nigroterminatus, Ramphotyphlops 44 - . T y p h l o p s 44 numatrana, Typhlops lineatus 43 nursii, Glauconia 23 - , Leptotyphlops 23 oatesi, Typhlops 65 obtusus obtusus, Typhlops 66 -.Onychocephalus 6 5 , 6 6 - palgravei, Typhlops 66 -.Typhlops 65,66 - , Typhlops obtusus 66 occidentalis, Leptotyphlops 24 ochraceus, Typhlops 50 ocularis, Typhlops 66 okahandjana, Glauconia 25 oligolepis, Typhlops 66 olivacea, Typhlina 44 olivaceus olivaceus, Typhlops 44 - , Onychocephalus 44 - , Onychophis 44 - reduncus, Typhlops 38 - , Typhlops 44 Typhlops olivaceus 44 Onychocephalus 29 Onychophis 29
Ophthalmidium 49 opisthopachys, Typhlops 45 palgravei, Typhlops obtusus 66 pallida, Lethobia 32 pallidus, Rhinotyphlops 32 - . T y p h l o p s 32 Typhlops braminus 39 pammeces, Typhlops 66 - , Typhlops braminus 66 paradoxus, Typhlops biminiensis 52 paucisquamata, Typhlops delalandei 31 paucisquamosa, Typhlops lalandei 31 pembae, Leptotyphlops emini 23 - , Leptotyphlops nigricans 23 perditus, Typhlops 74 peruvianus, Leptotyphlops 24 persicus, Typhlops 75 petersii, Helminthophis 4 - , Liotyphlops 4 - , Onychocephalus 35 - , Rhinotyphlops schlegelii 35 - , Typhlops 71 phenops bakewelli, Leptotyphlops 15, 16 - , Leptotyphlops 15 - , Leptotyphlops goudotii 15,16 - , Leptotyphlops phenops 15, 16 - phenops, Leptotyphlops 15 - , Stenostoma 15 phillipsi, Leptotyphlops 24 philococcus, Typhlops 42 philococos, Typhlops 42 Pilidion 37 pinguis, Ramphotyphlops 45 - . T y p h l i n a 45 - . T y p h l o p s 45 pitmani, Typhlops caecus 30 platycephalus, Typhlops 70 - , Typhlops richardi 70 platyrhynchus, Typhlops 67 polli, Typhlops gracilis 31 - , Typhlops leptosoma 31 polygrammica brongersmai, Typhlina 45 - elberti, Typhlina 46 - florensis, Typhlina 46 - nigrescens, Typhlina 46 - polygrammica, Typhlina 45, 46 - , Typhlina polygrammica 45 - undecimlineata, Typhlina 46 polygrammicus, Argyrophis 45 - brongersmai, Typhlops 45 - elberti, Typhlops 46 - florensis, Typhlops 46 - nigrescens, Typhlops 46 - polygrammicus, Typhlops 45 - , Pseudotyphlops 45 - , Typhlops 45 - , Typhlops polygrammicus 45 - undecimlineatus, Typhlops 47 polylepis, Typhlops angolensis 50
91 porrecta, Typhlops 67 porrectus, Typhlops 67 praelongis, Typhlops 74 praelongus, Typhlops 74 praeocularis, Helminthophis 2 - ludensis, Typhlops 33 - praeocularis, Typhlops 33 - , Rhinotyphlops 33 - , Typhlops 33 - , Typhlops praeocularis 33 preissi, Typhlops 38 principis, Typhlops 30 priscus, Omoiotyphlops 78 proxima, Typhlina 47 proximus, Ramphotyphlops 47 Typhlops 47 psammeces, Typhlops 66 psammophilus, Typhlops 66 pseudosaurus, Typhlops 67 Pseudotyphlops 37 psittacus, Typhlops 77 punctata, Onychophis 68 punctatus, Acontias 67 - congestus, Typhlops 68 - gierrai, Typhlops 58 - liberiensis, Typhlops 68, 69 - punctatus, Typhlops 67, 68 - , Typhlops 6 2 , 6 7 , 6 8 - , Typhlops punctatus 67, 68 pusilla, Typhlops 69 pusillus, Typhlops 69 pyrites, Leptotyphlops 24 Ramphostoma 6 Ramphotyphlops 37 reduncus, Typhlops olivaceus 38 reginae, Typhlops 46 Rena 6 reticulata, Anguis 69 - , Glauconia 24 reticulatus, Argyrophis 69 - crassa, Typhlops 69 - , Leptotyphlops 24 - nigrolactea, Typhlops 69 - troscheli, Typhlops 69 - . T y p h l o p s 69 retisculatus, Typhlops 69 reuteri, Typhlops 69 Rhamphostoma 6 Rhinoleptus 28 Rhinotyphlops 2, 29 richardi capapontus, Typhlops 70 - naugus, Typhlops 70 - platycephalus, Typhlops 70 - richardi, Typhlops 70 - . T y p h l o p s 70 - , Typhlops richardi 70 riparius, Onychocephalus 34 - . T y p h l o p s 34 rondoensis, Typhlops 7 0 , 7 1
- , Typhlops tettensis 70 rostellatus, Typhlops 71 rostralis, Anguis 69 rostrata, Glauconia 24 rostratum, Stenostoma 24 rostratus, Anguis 69 - , Leptotyphlops 24 rowani, Liotyphlops 4 rubellum, Stenostoma 13 ruber, Typhlops 71 rubrolineata, Glauconia albifrons 25 rubrolineatus, Leptotyphlops 25 rueppelli, Typhlops 46 rufescens, Rhinotyphlops 33 - , Typhlops 33 ruficauda, Anilios 71 - , Typhlops 71 ruficaudus, Typhlops 71 rufidorsum, Leptotyphlops 25 rufidorsus, Leptotyphlops 25 rugosa, Typhlops 40 russelii, Tortrix 39 russelli, Typhlops 39, 49
Sabrina 6 salgueiroi, Leptotyphlops 25 schinzi, Onychocephalus 33 - , Rhinotyphlops 33 - . T y p h l o p s 33 schlegelii brevis, Rhinotyphlops 34 - brevis, Typhlops 34 - dinga, Rhinotyphlops 35 - excentricus, Typhlops 35 - mucroso, Typhlops 35 - mucruso, Rhinotyphlops 34 - mucruso, Typhlops 35 - , Onychocephalus 34 - petersii, Rhinotyphlops 35 - , Rhinotyphlops 33 - , Rhinotyphlops schlegelii 34 - schlegelii, Rhinotyphlops 34 - schlegelii, Typhlops 34 - . T y p h l o p s 33 - , Typhlops schlegelii 34 schmidti schmidti, Typhlops 62 - tanganicanus, Typhlops 62 - . T y p h l o p s 62 - , Typhlops schmidti 62 - wilsoni, Typhlops 62 schneideri, Typhlops 56 schubarti, Liotyphlops 4 scorteccii, Rhinotyphlops 35 - . T y p h l o p s 35 scutifrons, Glauconia 25 - , Leptotyphlops 25 - , Leptotyphlops scutifrons 25 - merkeri, Leptotyphlops 26 - scutifrons, Leptotyphlops 25 - , Stenostoma 25
92 segregus, Leptotyphlops humilis 18 senptemstriatus, Typhlops 26 septemstriata, Glauconia 26 septemstriatum, Catodon 26 - , Stenostoma 26 septemstriatus, Anguis 26 - , Catodon 26 - , Leptotyphlops 26 - , Siagonodon 26 Siagonodon 6 siamensis, Typhlops 7 1 , 7 2 signata, Glauconia 26 signatum, Stenostoma 26 signatus, Leptotyphlops 26 silus, Typhlops 63 similis, Typhlops 42 Simoliophis 78 simoni, Onychocephalus 72 - . T y p h l o p s 72 slevini, Leptotyphlops humilis 17 smithii, Typhlops 31 socotranus, Typhlops 72 soensis, Typhlops 45 sokotranus, Typhlops 72 solominis, Ramphotyphlops 47 solomonis, Typhlops 47 somalicus, Rhinotyphlops 35 - , Typhlops 35 somsalicu, Typhlops 35 squamosus, Anilios 5 - , Typhlophis 5 - , Typhlops 5 stadelmani, Typhlops 72 steinhausi, Typhlops 72 steinhousi, Typhlops 72 stejnegeri, Rhinotyphlops 36 - , Typhlops 36 Stenostoma 5 Stenostomophis 6 striatula, Leptotyphlops 21 striolatus, Typhlops 56 subcrotilla, Leptotyphlops 26 subcrotillus, Leptotyphlops 26 suboculaiis, Typhlina 47 - , Typhlops 47 subucularis, Typhlops 47 sudanensis, Rhinotyphlops 36 - . T y p h l o p s 36 sulcata, Typhlops 72 sulcatus, Typhlops 72 suluensis, Typhlops 44 sumatranus, Typhlops lineatus 43 sundevalli, Glauconia 27 - , Leptotyphlops 27 sundewalli, Leptotyphlops 27 - , Stenostoma 27 - , Tricheilostoma 27 supranasalis, Typhlops 42 suturalis, Typhlops ater 51 symoensi, Typhlops angolensis 50
syntherus, Typhlops 73 syriacus, Typhlops 75 tanganicanus, Typhlops lineolatus 62 - , Typhlops schmidti 62 tasymicris, Typhlops 73 tatacua, Leptotyphlops 26 teaguei, Leptotyphlops 27 telloi, Leptotyphlops 27 temminckii, Typhlops 46 tenebrarum, Typhlops 73 tenella, Leptotyphlops 27 - , Leptotyphlops albifrons 27 tenellus, Leptotyphlops 27 tenue, Ophthalmidium 39 tenuicula, Rena 18 tenuicollis, Onychocephalus 73 - , Ophthalmidion 73 - , Typhlops 73 tenuiculum, Stenostoma 18 tenuiculus, Leptotyphlops humilis 18 tenuis, Typhlops 6 6 , 7 3 tephrosoma, Typhlops 56 ternetzii, Helminthophis 4 - , Liotyphlops 4 tesselata, Stenostoma albifrons 27 tesselatus, Leptotyphlops 27 - , Stenostoma 27 - . T y p h l o p s 27 Tetracheilostoma 6 tettensis, Onychocephalus 58 - rondoensis, Typhlops 70 - , Typhlops 58 textilis, Typhlops 77 thurstoni, Typhlops 74 tindalli, Typhlops 74 tornieri, Typhlops 62 torresianus, Ramphotyphlops 45 - . T y p h l o p s 45 tovelli, Typhlina 47 - . T y p h l o p s 47 trangensis, Typhlops 74 Tricheilostoma 6 tricolor, Leptotyphlops 28 trilobus, Onychocephalus 58 trinitatus, Typhlops 74 troscheli, Typhlops lumbricalis 69 - , Typhlops reticulatus 69 truncatus, Argyrophis 39 Typhlina 36 Typhlinalis 37 Typhlopidae 29 Typhlophis 5 Typhlops 49 uluguruensis, Typhlops 75 undecimlineata, Typhlina polygrammica 46 undecimstriata, Epictia 28 undecimlineatus, Typhlops florensis 46 - , Typhlops polygrammicus 47
93 undecimstriatus albipunctata, Stenostoma 7 - albipunctata, Typhlops 7 undecimstriatus, Leptotyphlops 28 - , Typhlops 28 unguirostris, Glauconia 28 - , Leptotyphlops 28 - , Onychocephalus 48 - , Ramphotyphlops 48 - , Typhlina 48 - . T y p h l o p s 48 unilineatus, Onychocephalus 75 - . T y p h l o p s 75 unitaeniatus ataeniatus, Typhlops 36 - , Letheobia 36 - , Rhinotyphlops 36 - , Typhlops 36 - , Typhlops unitaeniatus 36 - unitaeniatus, Typhlops 36 usambaricus, Typhlops boulengeri 62 utahensis, Leptotyphlops humilis 19 vanderysti, Typhlops 36 variabilis, Glauconia 28 - , Leptotyphlops 28 varius, Onychocephalus 34 - , Typhlops 34 - , Typhlops mucruso 34 veddae, Typhlops 75 venningi, Typhlops 67 vermicalis, Typhlops 75 vermicularis, Argyrophis 75 - . T y p h l o p s 75
vermis, Typhlops 76 verticalis, Onychocephalus 78 - . T y p h l o p s 78 violaceus, Typhlops 76 viridiflavus, Typhlops 34 vrijdaghi, Typhlops 33, 36 vryjdaghi, Typhlops 33, 36 waitii, Typhlops 38 walli, Typhlops 74 westermanni, Onychocephalus 49 weyrauchi, Leptotyphlops 28 wiedii, Ramphotyphlops 48 - , Typhlina 48 - , Typhlops 48 wilderi, Helminthophis 5 - , Liotyphlops 5 - , Typhlops 5 willeyi, Typhlina 48 - . T y p h l o p s 48 wilsoni, Leptotyphlops 82 - , Typhlops 76 - , Typhlops schmidti 62 wittei, Rhinotyphlops 36 yirrikalae, Typhlina 48 - , Typhlops 48 yirrkalae, Typhlops 48 zenkeri, Typhlops 76
DAS TIERREICH Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen Herausgeber R o b e r t Mertens und Willi Hennig Schriftleiter: Heinz W e r m u t h
Lfg.
Ersch.jahr
Titel
Bearbeiter
Umfang
Preis in DM
1922
VI, 614 Seiten, 34 Abbildungen
120,-
1923
X X X I I , 5 8 4 Seiten, 380 Abbildungen
115,50
1924
XXVIII,478Seiten, 209 Abbildungen
144,-
1926
XXXVI,885 Seiten, 340 Abbildungen
300,-
1926
XVI, 119 Seiten, 55 Abbildungen
42,-
1927
XL, 707 Seiten, 6 Abbildungen
247,50
1928
XXX, 300 Seiten, 197 Abbildungen
126,50
1929
XXIII, 388 Seiten, 307 Abbildungen
180,-
CRUSTACEA COPEPODA: II. Cyclopoida Gnathostoma
1929
XVI, 102 Seiten, 4 2 Abbildungen
C. Graf A t t e m s
MYRIAPODA: II. Scolopendromorpha
1930
XIX, 308 Seiten, 387 Abbildungen
150,-
55
E. Ahl
AMPHIBIA: Anura Polypedatidae
1931
XVI, 477 Seiten, 320 Abbildungen
232,50
56
S. Thor
ARACHNOIDEA ACARIÑA: Bdellidae, Nicoletiellidae, Cryptognathidae
1931
XIII, 87 Seiten, 9 3 Abbildungen
57
M. Beier
PSEUDOSCORPIONIDEA I: Subord: Chthoniinea et Neobisiinea
1932
XX, 258 Seiten, 271 Abbildungen
120,-
58
M. Beier
PSEUDOSCORPIONIDEA II: Subord. Cheliferinea
1932
XXI, 294 Seiten, 300 Abbildungen
146,20
45
A. Schmidt
COLEOPTERA:
Aphodiinae
46
F. Nieden
AMPHIBIA: Anura I
47
W. Kükenthal
COELENTERATA:
48
J. J. Kieffer
HYMENOPTERA: Proctotrupoidea:
49
F . Nieden
AMPHIBIA: Anura Engystomatidae
SO
E. Giglio-Tos
ORTHOPTERA:
51
H. Stichel
LEPIDOPTERA:
52
C. Graf A t t e m s
MYRIAPODA: /.
53
F. Kiefer
54
Gorgonaria
Scelionidae
II
Mantidae
Nemeobiinae
Geophilomorpha
III
56,30
56,20
Lfg.
Bearbeiter
Ersch.jahr
Titel
Umfang
Preis in DM
1933
XIV, 62 Seiten, 57 Abbildungen
37,50
1933
XI, 84 Seiten, 102 Abbildungen
45,-
PISCES: 2. Dipnoi
1933
IX, 20 Seiten, 25 Abbildungen
10-
F. Werner
REPTILIA: Loricata
1933
XIV, 40 Seiten, 33 Abbildungen
26,20
63
M. Holly
PISCES: 4. Crossopterygii
1933
X, 23 Seiten, 30 Abbildungen
10,-
64
F. Bryk
LEPIDOPTERA: Baroniidae, Teinopalpidae, Parnassiidae I
1934
XXIII, 131 Seiten, 87 Abbildungen
75,-
65
F. Bryk
LEPIDOPTERA: Parnassiidae II (Subfam. Parnassiinae)
1935
LI, 790 Seiten, 698 Abbildungen
412,50
66
E. Marcus
ARTHROPODA: Tardigrada
1936
XVI, 329 Seiten, 306 Abbildungen
150,-
67
M. Holly
PISCES: 4. Ganoidei
1936
XVI, 65 Seiten, 56 Abbildungen
68
C. Graf Attems
MYRIAPODA: 3. Polydesmoidea I. Fam. Strongylosomidae
1937
XXII, 300 Seiten, 343 Abbildungen
150,-
69
C. Graf Attems
MYRIAPODA: 3. Polydesmoidea II. 1938 Fam. Leptodesmidae, Platyrhachi1938 dae, Oxydesmidae, Gomphodesmidae
XXVIII,487 Seiten, 509 Abbildungen
243,80
70
C. Graf Attems
MYRIAPODA: 3. Polydesmoidea III
1940
XXXII,568Seiten, 719 Abbildungen
300,-
71
S. Thor & C. Willmann
ACARINA: 3.-71 a: Eupodidae, Penthalodidae, Penthaleidae, Rhagidiidae, Pachygnathidae, Cunaxidae. -72 b Trombidiidae
1941/1947 XXXVI,541 Seiten, 599 Abbildungen
72
H. Wermuth
REPTILIA: Helodermatidae
1958
16 Seiten, 9 Figuren
73
M. Beier
ORTHOPTERA: Tettigoniidae (Pseudophyllinae I)
1962
XII, 468 Seiten, 245 Abbildungen
210,-
74
M. Beier
ORTHOPTERA: Tettigoniidae (Pseudophyllinae II)
1960
396 Seiten, 241 Figuren
262,50
75
K. Odening
TREMATODA (Digenea): Plagiorchiidae III (Haematoloechinae) und Omphalometridae
1960
IV, 77 Seiten, 60 Figuren
59
M. Holly
CYCLOSTOMATA
60
S. Thor
ACARINA: Tydeidae,
61
M. Holly
62
Ereynetidae
37,50
389,20
16,-
64,-
Lfg.
Bearbeiter
Titel
Ersch.jahr
Umfang
Preis in DM
76
K. Sanft
AVES (Upupae): Bucerotidae
1960
IV, 176 Seiten, 106 Figuren
77
R. Kilias
GASTROPODA PROSOBRANCHIA: Tonnidae
1962
63 Seiten, 47 Figuren
28,-
78
St. W. Gorham
GYMNOPHIONA. Liste der rezenten Amphibien und Reptilien
1962
X, 25 Seiten
15,-
79
R. Mertens
HELODERMATIDAE, VARANIDAE, LANTHANOTIDAE. Liste der rezenten Amphibien u. Reptilien
1963
X, 26 Seiten
14,-
80
H. Wermuth
GEKKONIDAE, PYGOPODIDAE, XANTUSIIDAE. Liste der rezenten Amphibien und Reptilien
1965
XXII, 246 Seiten
81
J. A. Peters
COLUBRIDAE (Dipsadinae). Liste der rezenten Amphibien und Reptilien
1965
VIII, 18 Seiten
82
J. Illies
PLECOPTERA, Katalog
1966
XXX, 632 Seiten, 20 Abbildungen
83
R. Mertens
CHAMAELEONIDAE. Liste der rezenten Amphibien und Reptilien
1966
X, 37 Seiten
26,-
84
C. Gans
UROPELTIDAE. Liste der rezenten Amphibien und Reptilien
1966
29 Seiten
14,-
85
St. W. Gorham
ASCAPHIDAE, LEIOPELMATIDAE, PIPIDAE, DISCOGLOSSIDAE, PELOBATIDAE, LEPTODACTYLIDAE, RHINOPHRYNIDAE. Liste der rezenten Amphibien und Reptilien
1966
XVI, 222 Seiten
107,-
86
H. Wermuth
AGAMIDAE. Liste der rezenten Amphibien und Reptilien
1967
XIV, 127 Seiten
72,-
87
H. Wermuth
CORDYLIDAE (Cordylinae + Gerrhosaurinae). Liste der rezenten Amphibien und Reptilien
1968
X, 30 Seiten
24,-
88
F. Haas
BIVALVIA: Unionacea
1969
X, 663 Seiten, 5 Abbildungen
89
A. F. Stimson
BOIDAE, (Boinae + Bolyeriinae + Loxoceminae + Pythoninae). Liste der rezenten Amphibien und Reptilien
1969
XI, 49 Seiten
156,-
117,80
12,50
294,-
380,-
38,-
Lfg-
Bearbeiter
Titel
Ersch.jahr
Umfang
90
H. Wermuth
ANGUIDAE, ANNIELLIDAE, XENOSAURIDAE. Liste der rezenten Amphibien und Reptilien
1969
XII, 41 Seiten
91
M. Fischer
HYMENOPTERA: Braconidae (Opiinae I )
1972
XII, 620 Seiten, 463 Figuren
92
R. Kilias
GASTROPODA PROSOBRANCHIA: Cymatiidae
1973
VIII, 235 Seiten. 149 Figuren
93
P. Banaiescu & T. T. Nalbant
PISCES, TELEOSTEI: Cyprinidae (Gobioninae)
1973
VII, 304 Seiten, 154 Figuren, 19 Karten
94
P. Zwick
PLECOPTERA, Phylogenetisches System und Katalog
1973
XXXII, 465 Seiten, 75 Abbildungen
95
E. Duellmann
HYLIDAE, CENTROLENIDAE, PSEUDIDAE. Liste der rezenten Amphibien und Reptilien
1976
XIX, 225 Seiten
96
M. Fischer
HYMENOPTERA: Braconidae (Opiinae II / Amerika)
1976
etwa 1038 Seiten, 890 Figuren
97
S. G. Kiriakoff
LEPIDOPTERA NOCTUIFORMES: Agaristidae I
1976
IX, 180 Seiten, 39 Figuren
98
S. G. Kiriakoff
LEPIDOPTERA NOCTUIFORMES: Agaristidae II
1976
VIII, 165 Seiten, 52 Figuren
99
S. G. Kiriakoff
LEPIDOPTERA NOCTUIFORMES: Agaristidae III
1976
VI, 86 Seiten, 26 Figuren
100
H. Wermuth & f R . Mertens
TESTUDINES, CROCODYLIA, RHYNCHOCEPHALIA. Liste der rezenten Amphibien und Reptilien
1976
XXVII, 174 Seiten
101
D. E. Hahn
ANOMALEPIDIDAE, LEPTOTYPHLOPIDAE, TYPHLOPIDAE. Liste der rezenten Amphibien und Reptilien
1980
XII, 93 Seiten