Catalog of fossil spores and pollen. Volume 44. Tertiary and Upper Cretaceous Spores and Pollen from Africa, Europe, Canada, Australia and New Zealand

Citation preview

ISSN 0148-642X

CATALOG OF FOSSIL SPORES AND POLLEN Volume 44 Tertiary and Upper Cretaceous Spores and Pollen from Africa, Europe, Canada, Australia and New Zealand

PREPARED BY H.T. AMES and W. SPACKMAN

Coal Research Section THE PENNSYLVANIA STATE UNIVERSITY Published at University Park, Pennsylvania

1985

Copyright © by The Pennsylvania State University (1985)

Copying of material in the Catalog of Fossil Spores and Pollen for commercial exploitation is prohibited.

CATAWG OF FOSSIL SPORES AND POLLEN

Compilers and Editors W. Spackman

H.T. Ames Preparators

R.M. Krebs

D.L. Krebs Editorial Committee

W.G. Chaloner, Bedford College, University of London, London, England A.T. Cross, Michigan State University, East Lansing, Michigan J. Jansonius, Imperial Oil Co., Calgary, Alberta R.M. Kosanke, U.S. Geological Survey, Denver, Colorado G.O.W. Kremp, Palynodata, Inc., Tucson, Arizona E.A. Stanley, Phillips Petroleum Co., Bartlesville, Oklahoma Business Manager Karen Copenhaver

TABLE OF CONTENTS

Compiler's Preface . . Clarke, R.T. and N.O. Fredericksen, 1968 Some new sporomorphs from the Upper Tertiary of Nigeria . . . . . .

ii 1

Clarke, R.T., 1966 PeregrinipoZZis nigericus, a new palynomorph from the Upper Tertiary of Nigeria . . . . . . . . . . . .

14

Martin, H.A. and G.E. Rouse, 1966 Palynology of Late Tertiary sediments from Queen Charlotte Islands, Bri tish Columbia. . . . . . . . . . . . . . . . . . . . . . . . . ,

17

Martin H.A., 1973 The palynology of some Tertiary Pleistocene deposits, Lachlan River Valley, New South Wales. . . . . . . . . . . . . . . . . . . . . .

33

McIntyre, D.J., 1968 Further new pollen species from New Zealand Tertiary and uppermost Cretaceous deposits . . . . . . ................

73

Nagy, E., 1968 Moss spores in Hungarian Neogene strata . . . . . . . . . . . . . . 100 Nagy, E., 1973 Palynological data for the Neogene of Cserehat . . . . . . . . . . . 120 Pacltova, B. and P. Simoncsics, 1970 New types of spores (genera and species) from the Bohemian Miocene

124

Harris, W.K., 1972 New form species of pollen from southern Australian early Tertiary sediments. . . . . . . . . . . . . . . . . . . . . . 145 Appendix Emended Species

169

Index .

170

44:i

COMPILER'S PREFACE Descripti"ons of new taxa provided by Volume 44 are republished from the works of:

Clarke, 1966; Clarke and Frederiksen, 1968; Martin and

Rouse, 1966; Martin, 1973; McIntyre, "1968; Nagy, 1968; Nagy, 1973; Pacltova and Simoncsics, 1970; and Harris, 1972.

A total of 12 new

genera, 117 new species and one new forma comprise the body of this volume, and a single emendation appears in the accompanying appendix. Specimens assigned to new species reported here were recovered principally from sediments of Tertiary or younger age.

Although all Tertiary

epochs are represented, emphasis was placed on occurrences of Miocene and younger age, including sediments reported to be of questionable Pleistocene age. Paper selection for these younger Tertiary taxa has produced a volume with considerable geographic diversification.

Type localities for

these taxa include: African sites (Nigeria), European sites (Hungary and Czechoslovakia), North American sites (Canada), Australian sites (New South Wales), and some New Zealand localities. ａｲ･ｯｚｩｰｳｾ＠

ｇ｡ｭ｢ｩ･ｲｮｾ＠

New genera treated in Volume 44 include; ｍ｡ｲｧｩｮｰｯｚｳｾ＠ ｎｵｭｚｩｰｯｳｾ＠

ｐ･ｲｧｩｮｰｯｚｳｾ＠

ｳｰｯｲｩｴ･ｾｅｐｨｭ＠

ｐｨ｡･ｯ｣ｲｳｰｩｴｾ＠

ｐｯｚｹｲｩｦ｣ｴ･ｳｾ＠

and SvatopZukisporites.

ｒｩ｣｡･ｳｰｯｲｴｾ＠

EncaZyptaePorisporites

With the exception of Gambierina all of the

generic descriptions for the above taxa are republished in conjunction with CFSP treatment of their designated type species. was necessary in the case of ｇ｡ｭ｢ｩ･ｲｮｾ＠

An alternative treatment

because its type was not a new species

name in the publication in which the new generic name was proposed (Harris, 1972), and its type species, G. ･､ｗ｡ｲｳｩｾ＠

had been previously treated in

CFSP 15-112.

44:ii

The editors express their appreciation to the authors who aided in our efforts to republish the systematics presented here.

We thank them

for their permission to republish, supplying additional illustrative material, and additional informative stratigraphic or geographic detail not provided in their original works.

44:iii

Species n.me:

AreoZipoUis aZiquantuZus Clarke & Frederiksen 1968

5

4

2

3

6

8

7

PI. 4. All specimens y 900.- 2. 3. A. aliqllanAreolipollis aliquanlulus, slide 10176,-\-2, 30.9 x 123.8, size 27 x 31l lu/us, holotype. Slide 13521A-2, 28.1/117.8, size 20 x 27 /1,4. A. uliqllunllllus, slide 10194A-1, 28.0 y 116.0, size 23 x 30 ", 5. A. uliquunlulus, paratype. Slide -I694A-1, 2!U x 11G.8, size Hl x ao ", Ii. 11.• aliqllanlllills, slide 13521A-2, 34.3 x 111.5, size 22 x 2M ", 7. A. uliquanlu/us, slide (jM79A-:l, -1,:1.0 x 11-1.5, size 2-1 x ＺｾＨｪ＠ 1', 8. A. aliqllunilllus, slide 6892A-l, 25.0 x 123.5, size 2-1 x 35 "

ii,

Description:

Diagnosis.-Pollen grains 2-colporate, bilateral. Polar axis of 13 specimens 27-38 fl, mean 32 fl. Equatorial diameter of 12 specimens 19-27 fl, mean 23 fl. Colpi narrow, 0.5-1.0 fl wide, extending onehalf to three-fourths length of the grain, margins distinct. Ora lalongate, distinct, 1.0-2.5 x 2.0-5.0 fl. There are several irregular rows of faint polygonal areolae on each side of the col pi. Exine 1-2 fl thick at poles and 2-3 fl thick at equator. Sexine tectate with distinct infratectal bacula at poles and irregular infratectal bacula at equator. Design finely reticulate in polar regions grading to coarser and more irregularly reticulate at equator. X exine about 0.5-1.0 fl thick. Remarks.-Although A. aliquanillius possesses only faint an'olae, it is ne\'ertheless similar to areoloidate pollen of the Acanlhaceae (se e Beloperone cali[ornica Bentham, PI. 4, Fig. 1). AnomofJanfl."t: schlechleri Harms, figured in Erdtman (1966, p. 60), in the family Araliaceae, is morphologically similar to AreolifJollis aliqllrtnillills, CATALOG OF FOSSIL SPORES AND POLLEN

44:1

t:vt:n possessing lalongate ora, hut it does not have areo lac. These fossil forms are believed to have closer affinity with the Aeanthaceae than with the .\raliaceae. A. aliqllflnillllls difTers from A. ordin Ils 1and A. cOIl{erllls1in possessing distinct ora and less pronoll need areolae. The specific name aliquanlllius is Latin for "little" or "small" and refers to the size of the areolae. Holotype.-Slide 13521A-2, coordinates 28.1 x 117.8 (PI. 4, Fig. :i). Paratype.-Slide 4694A-l, coordinates 28.1 x 116.8 (PI. 4, Fig. 5).

Clarke, R.T. and N.O. Frederiksen, 1968, p. 213, pI. 4, figs. 2-8 Some new sporomorphs from the Upper Tertiary of Nigeria: Grana Palynologica, v. 8, no. 1, pp. 210-224, 5 pIs., 1 text-fig. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southeast" coast Type horizon: Late Tertiary Preparation: HF, KCl-HN03, K2C0 3, concentrated by swirling in watchglass, safranin 0 stain 2 Palynological slide colI., Mobil Field Research Laboratories, Repository: Dallas, Texas Holotype: Slide l352lA-2, coord. 28.1 x 117.8, (PI. 4, fig. 3) Author: Reference:

1

[See AreoUpoUis ordinus, CFSP 44: 5" and A. confertus, CFSP 44: 3.]

2[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.]

CATALOG OF FOSSIL SPORES AND POLLEN

44:2

Species n.me:

Areo"lipoUis confertus Clarke & Frederiksen 1968

4

3

5

PI. 3. All specimens y 900. - ' 3. A.reolipo/lis COII/frtllS, holotype. Slide 1-t-l41A-1, -to.7 " 119.3, size 21 x 33 ", 4. A. ｣ｯｮＯ･ｲｴｵＮｾＬ＠ paralypc. Slide 15222A-2, 37.8 x 112.0, size 23 x 33 /" 5. A. confertus, slide 15223A-2, 2:1 . 112.0, size 20 x 2;) "

Description:

Diagnosis.-Pollen grains 2-colporate, bilateral. Polar axis of five specimens 29-34 fl' ,mean 32 fl. Equatorial diameter of five specimens 20-27 fl' mean 24 fl. Colpi about 1 fl wide, extending one-half to two-thirds length of grain, their margins usually indistinct. Ora indistinct, (sometimes only suggested by a widening of the colpi), about 1.5 fl wide. There are two to three irregular rows of polygonal areolae, 1-2 fl diameter on each side of colpi. Sexine finely reticulate, 1.0-1.5 fl thick at poles, 1.5-3.0 fl t;,ick at equator. l\'exine thin. Remarks.-A. confertus is quite similar to the acanthaceous spe('ies Jacobinia candicans Bentham and Hooker (PI. 3, Figs. I, 2) and .!ncobinin carnea (Lindley) l\"ichols illustrated in Raj (HHH, pI. 10, figs. 4-7 &: pI. 16, figs. 4-7) in number and arrangement of the areolae. Hm\,eyer, the fossil species has indistinct ora whereas ｜Ｇ･ｲｾ＠ pronounced ora (or pores) are present on these species of Jacobinia. A.. confertus difTers from A. ordinus1in the greater number and irregular arrangement of the areolae. The specific name confertus is Latin for "crowded" and refers to the number and arrangement of the areolae. Holotype.-Slide Fig. 3). Paratype.-Slide Fig. 4).

1-l-!41A-l, .coordinates

40.7 x 119.3

(PI.

3,

15222A-2,

37.8 x 112.0

(PI.

3,

CATALOG OF FOSSIL SPORES AND POLLEN

coordinates

44:3

Clarke, R.T. and N.O. Frederiksen, 1968, p. 212, pl. 3, figs. 3-5 Some new sporomorphs from the Upper Tertiary of Nigeria: Grana Palynologica, v. 8, no. 1, pp. 210-224, 5 pIs., 1 text-fig. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southeast" coast Type horizon: Late Tertiary Preparation: HF, KCI-HN03 .• K2 C0 3, concentrated by swirling in watchglass, safranin 0 stain 2 Palynological slide colI., Mobil Field Research Laboratories, Repository : Dallas, Texas Holotype: Slide 14441A-l, coord. 40;7 x 119.3, (PI. 3, fig. 3) ａｾｨｯ｣Ｚ＠

Reference :

1

[See Areolipollis ordinus, CFSP 44:5.]

2[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.]

CATALOG OF FOSSIL SPORES AND POLLEN

44:4

Specie. name:

9

AreoZipoUis ordinus Clarke & Frederiksen 1968

6

7

8

10

PI. :3. All specimens x 900.. 6. -t. orciilllls, purulype. Slide XU\J2A-2, ·10.2 x 125.0, size 2ti x 34 It, 7. A. orriinlls, slide 9007A-l, 30.8 x 121.(i, sizlo IX' 2li /', 8 ...I. ordilltls, slide 8002A-2, :31.2' 122.7, size 22 ｾＳＷ＠ /I, 9. A. IIrdintls, ｨｯｬｾﾷｰＧＮ＠ Slide 0007A-l, 38.9 x 118.·1, size 23' :-11 /', 10. II. ordintls, slide 0007A-l, 23.:-1 x 11 1.7, sizl' 2:-1' :n /'

Description:

Diagnosis.-PoIlen grains 2-colporate to 2-colpate, bilateral. Polar axis of seven specimens 26-37 fl, mean 32 fl. Equatorial diameter of eight specimens 18-26 fl, mean 22 fl. Colpi 1-2 fl wide, length varies from two-thirds to almost full length of grain, margins indistinct. Ora, when observed, are indistinct, about 4-5 fl diameter. One distinct ro\\' of 4-7 areolae and an adjacent row of partly formed areolae border each side of colpi. Areolae polygonal in surface view, 1.5-2.5 fl in diameter, separated by grooves about 0.5-1.0 fl wide. Sexine very finely reticulate, about 1.0 fl thick. :\ exine thin, about 0.5 fl. Remarks.-A. ordinus is probably related to the modern family Acanthaceae. Although fossil specimens have indistinct ora, the areolate and finely reticulate ornamentation is similar to that of Beloperone and Justicia pollen. Justicia cayennensis Ｈｾ･ｳ＠ von Esenheck) Lindau, pictured in Bremekamp (1938, tah. 16, fig. H) possesses a similar arrangement of the areolae. The specific name ordinus is Latin for "line" or "row" and refers to the linear arrangement of the areolae. modern genera, but the entire group is included in one fossil genus because positive identification of affinities at the generic level is difficult. Probable botanical affinities are discussed in the remarks section for each species. The generic prefix refers to the areolate ornamentation exhih ited ｨｾＧ＠ the species assigned to this genus. Holotype.-Slide 9007A-l, coordinates 38.9 Y 118.4 (PI. 3, Fig. 9). Paratype.-Slide 8992A-2, coordinates 40.2 x 125.0 (PI. 3, Fig. 6).

CATALOG OF FOSSIL SPORES AND POLLEN

44:5

[])iagnosis.-Pollen grains 2-col po rate to 2-colpate (?), hi lateral. Ora often indistinct. .-\reolae in rows or irregularly arranged along borders of apertures, mesocolpia finely reticulate. Remarks.-Species of Areolipollis are similar to po lIen from the modern genera Beloperone, .Iacobinia and Jus/icia, family Acanthal'eae (see Raj 1961). The species of Areolipollis possibly represent seycral modern genera: but the en6re group is included in one fossil genus because positive identification of affinities at the generic level is diffieull. Probable botanical affinities are discussed in the remarks .section for each species. The generic prefix refers to the areolate ornamentation exhib ited by the species assigned to this genus. Type species.-A.reolipollis ordinus sp. nov.] 1

Clarke, R.T. and N.O. Frederiksen, 1968, p. 212, pI. 3, figs. 6-10 Some new sporomorphs from the Upper Tertiary of Nigeria: Grana Palynologica, v. 8, no. 1, pp. 210-224, 5 pIs., 1 text-fig. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southeast" coast Type horizon: Late Tertiary Preparation: HF, KCI-HN03, K2C0 3 , concentrated by swirling in watchglass, safranin 0 stain 2 Palynological slide colI., Mobil Field Research Laboratories, Repository : Dallas, Texas Holotype: Slide 9007A-I, coord. 38.9. x 118.4, (PI. 3, fig. 9) Author: Reference:

1

[Bracketed text is identified as the generic description of Areolipollis in Clarke &Frederiksen, 1968, p. 211.]

2[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.] TYPE SPECIES OF THE GENERIC NAME Areolipollis Clarke &Frederiksen, 1968

CATALOG OF FOSSIL SPORES AND POLLEN

44:6

Areolipollis vespiformis Clarke &Frederiksen 1968

9

10

11

12

13

1'1. .J. All SP('dlll('IIS . \11111. \). A. vespi/ormi.• , holotype. Slicte 4692A-l, :{5.0 x 112.0, siz(, :W x :12/,,10. ,I. Iwspi/"I'III.is, slide 1;,2:IIA-I, 27.M x 11;).:1, size 2;' x ·11 /', 11. A. llespi/ormis, slide (iMM2A-:I, 2M.7" 112.J, size 21. x 311 /" 12. A. IIcspi/ormis, paralype. Slide ·16U.IA-I, 1U.1 x 115.Ii, size 18 x 28/1, IJ. A.lIespi/ormis, slide 4717A-1, 2J.(i x 121.2, size 21 x :lIi I'.

Description:

Diagnosis.-Pollen grains 2-colporate, bilateral. Polar axis of senn specimens 28-41 fl, mean 33 fl. Equatorial diameter of seven specimens 18-25 fl, mean 22 fl. Colpi 0.5-1.0 fl wide, extending ahout two-thirds length of the grain, margins distinct to indistinct. Ora lalongate to circular, distinct, 2-4 fl in size. Areolae polygonal, distind to indistind, about 1-2 fl diameter, generally occurring in one or two regular rows on each side of col pi. Exine 1-2 fl thick at poles, thickening up to 6 fl at equator. Sexine thickcr than nexine, uut both layers thicken at the equator. Sexine finely reticulate in polar regions, irregularly reticulate at equator. Hl'll1urks.-A. lIespi{ormis is morphologically similar to Jusiici(/ /1011(/ Yahl, but the modern pollen grains are 3-colporate. Justicia secunda Yahl, figured in Bremekamp (1938. tab. 16, fig. 8) is a 2-colporate form with a thickened sexine in the mesocolpial equatorial region. The drawing, however, shows a flaring of the sexine in outline whereas the thickening in the fossil forms coincides \\ ith constriction of the nexine in .the equatorial region, and the outline is more or less straight. A. vespij'ormis differs from A. aliqualliulus 1 in haying thicker sexine and nexine at the equator than at the poles. The specific name vespi{ormis is deriyed from the Latin word IIcspa, meaning "wasp" and refers to the wasp-like outline of the interior cavity. Ho]otype.-Sli(·le 4GU2A-1, coordinates 35.0 x 112.0 (PI. 4, Fig. 9). Paratype.-Slide 4G9·L\-l, coordinates 19.1 x 115.6 (PI. 4, Fig. 12).

CATALOG OF FOSSIL SPORES AND POLLEN

44:7

Clarke, R.T. and N.O. Frederiksen, 1968, p. 214, pI. 4, figs. 9-13 Some new sporomorphs from the yPper Tertiary of Nigeria: Grana Palynologica, v. 8, no. 1, pp. 210-224, 5 pIs., 1 text-fig. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southeast" coast Type horizon: Late Tertiary Preparation: HF, KCl-HN03 .• K2 C0 3, concentrated by swirlinJ in watchglass, safranin 0 stain 2 Palynological slide colI., Mobil Field Research Laboratories, Repository: Dall as , Te xas Holotype: Slide 4692A-l, coord. 35.0 x 112.0, (PI. 4, fig. 9) Author: Reference:

l[See AreoZipoZZis ｡ｚｩｱｵｮｴｳｾ＠

CFSP 44: 1.]

2[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.]

CATALOG OF FOSSIL SPORES AND POLLEN

44:8

Specie. n.me:

MarginipoUis concinnus Clarke & Frederiksen 1968

10

6

4

3

7

8

5

9

PI. 2. All specimens x 900.- 3. Murginipollis concinnus, paratype. Slidr ｾＹＺ＾ＸａＭＲＬ＠ 44.4 x·123.1, size 22 x 40",4. M. concinnus, slide 15236A-1, 3i.5 y 124.6, size 24,3:> 1', 5. JI. concinnus, slide 15236 A-1, 25.6" 119.4, size 25 x 401'-, 6 . .1I. concilllllls, slide 8959A-1, 26.2 x 112.0, size 22 x44 1', i. M. concinnus, slide 15225A-1, 26.;)' 112.2, sizr 24 x 42 1', 8. M. concinnus, slide 15239A-1, 44.0 x 11 i.2, size 32,' 31; ", 9..11. concinnus, slide 15231A-1, 24.3 x 123.3, size 34".391',10. M. concinnus, hololype. Slide 15225A-1, 32.0 x 124.0, size 32 x 40 1'. Description:

Diagnosis.-Pollen grains 3-colpate (syncolpate), prolate-spheroidal to prolate. Polar axis of 23 specimens 36-44 /1:, mean .to fl. Equatorial diameter of 21 specimens 22-3.t fl, mean 2i fl. Colpi margins smooth, greatly thickened, 1.5-2.5 fl thick in equatorial region, becoming thicker, up to 5 fl towards the poles. :\Iargins often cupped toward colpi (see PI. 2, Fig. 6). :\Iesocolpial exine, margins excepted, about 0.5 fl thick, finely granular with irregular pits and thickenings (areoloidate?) near the margins. Apparent thin strip of exine present between main parts of mesocolpia and their margins. Remarks.-.llarginipollis concinnus resembles pollen grains of the lecythidaceous Planchonia-type (Erdtman 1966, pp. 222, 2n); e.g., the pollen of Planchonia speclabilis :\Ierrill (PI. 1, Figs. 1,2), BarringIonia racemosa Roxburgh (PI. 1, Figs. 3, 4), B. aC/llangula Gaertner

CATALOG OF FOSSil SPORES AND POllEN

44:9

(PI. 1, Fig. 5; PI. 2, Figs. 1,2), B. macroslachya Kurz (PI. 1, Fig. 6) and ｂｾ＠ reliculala ｾＱｩｱｵ･Ｎ＠ See also the illustrations of Plnnchonia crenala ｾｉｩ･ｲｳ＠ in Erdtman (I.e., p. 223) and of Bnrringlonin rrtcelllo.m shown in Kuhitzki (1965, pI. 147-149). The fossil forms appear to resemble the genus Barringlonia in the irregular thickenings near the margins. The specific name concinnlls is Latin for "pleasing" or "well put together". Holotype.-Slide 15225A-l, coordinates 32.0 x 124.0 (PI. 2, Fig. 10). Paratype.-Slide 8988A-2, coordinates 44.4 x 123.1 (PI. 2, Fig. 3).

[Remarks.-The generic prefix is Latin for "borders" or "edges" and refers to the thickened areas along the colpi. This genus difTers from Tricolpiles Couper emend.. Potonie in being syncolpate and possessing thickened colpi margins that extend to the poles. Diagnosis.-Pollen grains 3-colpate (syncolpate), prolate-spheroidal to prolate, colpi margins thick, generally smooth, typically becoming thicker and enlarged toward the poles. Type speeies.-J[nrginipollis concinnus sp. nov.] 1

Clarke, R.T. and N.O. Frederiksen, 1968, P. 211, pI. 2, figs. 3-10 Some new sporomorphs from the Upper Tertiary of Nigeria: Grana Palynologica, v. 8. no. I, pp. 210-224, 5 pIs., 1 text-fig. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southeast" coast Type horizon: Late Tertiary Preparation: HF, KCl-HN03. K2 C0 3, concentrated by swirling in watchglass, safranin 0 stain 2 Palynological slide colI., Mobil Field Research Laboratories, HeposilorYI Dallas, Texas Holotype: Slide l5225A-l, coord. 32.0 x 124.0, (PI. 2, fig. 10) Author: Reference:

1

[Bracketed text is identified as the generic description of MarginipoZZis in Clarke &Frederiksen, 1968, p. 210.]

2

[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.] TYPE SPECIES OF THE GENERIC NAME MarginipoZZis Clarke &Frederiksen, 1968 (Monotypic in Clarke &Frederiksen, 1968)

CATALOG OF FOSSIL SPORES AND POLLEN

44:10

Specie. n.me:

NwnmuZipoUis neogenicus Clarke & Frederiksen 1968

3

1

2

6

5

4

PI. 5. All spccimcns x !l00 eXCl'pt ｾｯＮ＠ !l ( x 550).-1. Xwnnlllliflol/is neogeniclls, slide 152;!2A-2, 27.!1 x 118.8, size. ·IX x 52 /" 2. N. Ileogenicus, hololype. Slide 1522·IA-l, 2,1.0 x 11!l.O, size 40 x '48/,,:t N. IlcogrtliCIIS, slide 15222A-1, :H.7 x 116.3, size. 27 x 30 /" ＧｾＮ＠ N. IlcogeniCIIs, slide !J007A-l, :1-1.:1 x 11:1.B, size :i2 x 3!J /" 5. N. neogelliclls, slide l.'i222A-l, 25.5 x 120.0, size 25 x 2X /', 6, N. lleogclliclls, slide 15222A-2, :17.7 x I1B.9, size of ornamented oval 2fi x :iO 1', 7. N. neogelliclIs, slide 15223A-l, 32.0 x 12·1.0, size of ornatlll'nted oval 2B x :i2 /'

A

c

B ｾｉｯ［ＬＭｓｅｘｎ＠

-NEXINE

Text-fig. 1. Xummuliflo/lis nl'ogenicus. A. "Poral" view, cross-section through margins. B. "Poral" view, surface. C. ＧｾｯｮＭｰｲ｡ｬ＠ view, cross-section through center.

CATALOG OF FOSSIL SPORES AND POLLEN

44:11

Description:

Diagnosis.-Pollen grains 2-porate, bilateral (discoidal), circular or nearly so in "poral" view, ellipsoidal in "non-poral" view. O"erall diameter of nine specimens 24-48 x 26-52 p" mean 33 x 38 p,. Pores ragged, 2-4 p, diameter. Sexine ramihaeulate, crassisexinous towards margins, o"erlaps nexine hy one-third to one-half the radius of the grain (see Text-fig. 1). ｾ＠ exine thin, psilate to punctate. Area extending for 7-15 p, radius around pores crassinexinous (sexine pre'sent ?), coveted with . scattered small processes. These areas sometimes tear ofT like opercula (PI. 5, Figs. 6, 7). Hemarks.-S. neogenicus probably has affinities with the family Acanthaceae; see palynogram of Rhytiglossa lactea Lindau ex Engler (syn. ｉＮｾｯｧｬｗＩ＠ in Erdtman (1966, p. 31) and illustrations in Haj (1961, pI. 16, figs. 1-3) and Rossignol (1962, pI. 1, figs. 1-3). The specific name neogeniClls refers to the strata from which the sporomorphs were reco\"ered. Holotype.-Slide 1522·L\-1, coordinates 24.0 x 119.0 (PI. 5, Fig. 2). Paratype.-Slide 9007A-l, coordinates 34.2 x 113.8 (PI. 5, Fig. 4).

[Diagnosis.-Pollen grains 2-porate, bilateral (discoidal), circular or nearly so in "poral" vie\\" , ellipsoidal in "non-poral" view ..-\rea surrounding the pores crassinexinous (sexine present ?), covered with scattered small processes. Sexine ｲ｡ｭｩ｢｣ｵｬｴｾＬ＠ crassisexinous, becoming thinner toward the pores. Remarks.-The generic prefix is derived from the Latin nl!mmulus, meaning a "small coin" and refers to the resemblance of the palynomorph with certain small British \Yest African coins with holes in the center. Type species.-.Yummulipollis neogenicus sp. nov.]1

CATALOG OF FOSSIL SPORES AND POLLEN

44:12

Specie. n.me:

NummulipoUis neogenious Clarke & Frederiksen 1968

Clarke, R.T. and N.O. Frederiksen, 1968, p. 215, pI. 5, figs. 1-7, text-fig. 1 Reference: Some new sporomorphs from the Upper Tertiary of Nigeria: Grana Palynologica, v. 8, no. 1, pp. 210-224, 5 pIs., I text-fig. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southeast" coast Type horizon: Late Tertiary Preparation: HF, KCl-HN03 K2C0 3, concentrated by swirling in watchglass, safranin 0 stain 2 Palynological slide colI., Mobil Field Research Laboratories, Repository: Dallas, Texas Holotypc: Slide l5224A-l, coord. 24.0 x 119.0, (PI. 5, fig. 2) Author:

1

[Bracketed text is identified as the generic description of NummulipoUis in Clarke & Frederiksen, 1968, p. 215.]

2[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.] TYPE SPECIES OF THE GENERIC NAME NummulipoUis Clarke &Frederiksen, 1968 (Monotypic in Clarke &Frederiksen, 1968)

CATALOG OF FOSSIL SPORES AND POLLEN

44:13

Speciel ".me:

PeregrinipoUis nigericus Clarke 1966

4

2

3

6

7

5

8

Figs. 1-8. - Peregrinipollis nigericlls gen. et sp. nov. All specimens ( x 1000) were photographed on Ilford Pan F film with an Exakta camera on a ｌｾｩｴｺ＠ Ortholux microscope using a Zeiss Xeofluar 100/1.30 oil immersion objective, a 10 /. ocular and a blue filter. Fig. 1. Slide 900:1-A-l, coordinates 31.0 x 120.5; dimensions 36 x 44 II. - Fig. 2. Slide 8961-,-\-2, coordinates 27.3 x 120.1; dimensions 36 x 36 II. - Fig. 3. Slide 8961-,-\-1, coordinates 21.5 :< 116.1; dimensions 38 x 44 ft. - Fig. 4. Holotype. Slide 8961-,,\-1, coordinates 37.0 x 124.3; dimensions 40 x 52,1. - Fig. 5. Paratype. Slide 8961-,-\-1, coordinates 22.0 x 110.7: dimensions Ｔｾ＠ x 52 It. - Fig. 6. Slide 11886-,,\-2, coordinates 32.0 x 114.5; dimensions -t7 x 50 ,Il. - Fig. 7. Slide 11886-A-l, coordinates 29.0 x 122.0; dimensions 44 x 50 ft. - Fig. 8. Slide 11886-A-2, coordinates 36.0 /. 113.8; dimensions 40 x 48,ll.

CATALOG OF FOSSIL SPORES AND POLLEN

44:14

Description:

Specific Description: Pollen grains tricolpate, subspherical to prolate. Mesocolpia coarsely reticulate. Colpi usually indistinct and irregular, delimited by two or more sub-parallel muri (see Figs. 6, 7 and 8). Muri rope-like, 2-5 p. wide, frequently forming a reticulum surrounding a central void. Lumina diameter 4-5 p.. A thin "inner layer" (endexine ?), preserved in very few specimens, is apparently connected to the reticulum by short rods 1-2 p. wide and 1-2 p. high. The rod-like structures are generally well preserved (see Fig. 3). Apocolpia of most specimens are thin and covered with grana or small verrucae 0.5-1.5 p. wide and 1 p. high. The rope-like muri grade into the granulate-verrucose sculpture at the juncture of the mesocolpia and apocolpia. Equatorial dimensions of 24 specimens, 32-50 p., mean 41 p.. Polar dimensions of 26 specimens, 36-52 p., mean 46 p.. Occurrence: L"pper Tertiary sediments from the :\Iobil Exploration Nigeria 1m:. offshore concessions. Remarks: The coarsely reticulate rnesocolpia and granulate-verrucose apocolpia distinguish Peregrinipollis nigericlls from previously described palynomorphs. Botanical affinities of P. nigericus are unknown. Two poorly preserved specimens appeared to have psilate apocolpia (due to corrosion ?). Holotype: Slide 8961A-l, coordinates 37.0 x 12-1.3 (Fig. 4). Paratype: Slide 8961A-l, coordinates 22.0 x 110.7 (Fig. 5).

[Generic Description: Pollen grains tricolpate, suhspherical to prolate. l\Iesocolpia coarsely reticulate. Apocolpia granular to finely verrucose or psilate. Type Species: Peregrinipollis nigericus sp. nov.] 1

CATALOG OF FOSSIL SPORES AND POLLEN

44: IS

Specie. name:

PeregrinipoUis nigeriaus Clarke 1966

Author: Clarke, R.T., 1966, p. 546, Reference: PeregrinipoUis nigeriaus,

figs. 1-8 A new palynomorph from the Upper Tertiary of Nigeria: Grana Palynologica, v. 6, no. 3, pp. 544-546, figs. 1-8. Type locality: Nigeria, Mobil Exploration Nigeria Inc. concession off the "southwest" coast Type horizon: Tertiary, Upper Preparation: HF, KCl-HN0 3 , K2C0 3 , swirled in watchglass to concentrate, safranin stain 2 Palynological slide colI., Mobil Field Research Laboratories, Repository: Dallas, Texas Holotype: Fig. 4, slide 896lA-l, coord: 37.0 x 124.3

°

1

[Bracketed text is identified as the generic description of Peregrinipollis in Clarke 1966, p. 546.]

2[This information provided in personal communication from R.T. Clarke to H.T. Ames, dated 3/14/83.] TYPE SPECIES OF THE GENERIC NAME Peregrinipollis Clarke, 1966 (Monotypic in Clarke, 1966)

CATALOG OF FOSSIL SPORES AND POLLEN

44: 16

Specie. n.me:

Cedrus periaZata Martin & Rouse 1966

58

57

60

59

62

63

61

64

FIGs. 57-64. Cedrlls perialata sp. nov. (Fig. 57, ho!otype). Figs. 57, 58, and 64, X 1000; Figs. 59-63, X 500. Description:

Pollen grains circular in outline, 49-i8 f.J, in height by 46--83 f.J, in length, usually about 55-65 f.J, in both dimensioas. The grains appear to be relatively fiat, somewhat like a thick biconvex lens. The exine of the cap is 4-5 f.J, thick, tectatc, with well-developed mcscxinous rods. The cap grades almost imperceptibly into the weakly dcveloped bladder, and the bladdcr has the same tectate structure of the prominently radiating mesexinous rods. The exine of the bladders is often thicker than that of the cap, reaching 12 f.J, in some specimens. The bladder is constricted rather abruptly at the leptoma forming a distinct notch. The margin of the bladder along thc leptoma is perfectly even and smooth. The body is formed by the endexine, and has a much finer reticulate pattern than the exine of the cap or bladders (see Fig. 58). CATALOG OF FOSSIL SPORES AND POLLEN

44 :17

The characteristic features of Cedrus perialata include the circular outline, the even outline of the bladdcr margin ｡ｬｯｮｾ＠ the Icptomu, and the thickness of the exine of the bladders (up to 12 J.L). I t can be distinguished from C. parvisaccata (Zauer 19.54, p. 31) by a much larger size range, a thicker exine, and the circular outline. I t differs from C. crispa (Zauer 1954, pp. 41-42) of the Oligoccne of the lJ .s.S. R. by a slllaller size ｲ｡ｬＱｾ･Ｎ＠ circular outline (as against semicircular), thicker exine, amJ finer ｲ･ｬｩ｣ｵ｡ｯｾＮ＠ The only reports of pollen of Cedrus in North America to date are those by Martin and Gray (1962), Gray (1964, p. 29, Table I), and Gray and Sohma (1964, p. 119) but no descriptions were given. However, Dr. Gray has verified (personal communication) that grains similar to C. peria/ala, and possibly other species, occur in the Tertiary of the northwestern United States. The assignment of the Skonun specimens to Cedrus appears to be justified, although it must be admitted that many of the fossil grains differ in detail from pollen of any of the three extant species. However, Zauer has pointed out that a good deal of variation exists in the extant pollen, and it seems most likely that the Skonun specimens are those of Cedrus rather than another genus of conifers. Howevcr, pollcn of Dacrydiul1! is similar in having a bladder virtually encompassing the body, and notchlike leptoma. holotypc Fig-, 57

Martin, H.A. and G.E. Rouse. 1966. p. 194. pI. VII, figs. 57-64 Palynology of Late Tertiary sediments from Queen Charlotte Islands, British Columbia: Canadian J. Bot .• vol. 44. pp. 171-208. 12 pIs .• 2 tabs. Type locality: Canada. British Columbia. Queen Charlotte Islands. NE Graham Island Type horizon: Late Tertiary. late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Preparlltion: Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter). 20-30% HN03 (1-6 hrs.), 2% KC03sol. (10-60 mins.), saturated sols. of safranin in H20 and alcohol. mounted in Gelva V-15 plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated Holotype: Pl. VII. fig. 57 Author: Reference:

CATALOG OF FOSSIL SPORES AND POLLEN

44:18

Species n.me:

Fagus granuZata Martin & Rouse 1966

93 ilCIIdillg the h;\cllb. The IIIost diagnostic feature is the complete irregularity of the width of the bacula; delicate, slender bacula are randomly mixed with stout, stuIllp-like bacula. and with all grades in between. Size range: 41-57 "". This form is common in the Skonun, and is readily distinguished from the three other species of Osmunda spores. Its affiliation with modern species is unknown. holotype Fig. 29

Martin, H.A. and G.E. Rouse, 1966, p. 189, pI. IV, figs. 29-30 Palynology of Late Tertiary sediments from Queen Charlotte Islands, British Columbia; Canadian J. Bot., vol. 44, pp. 171-208, 12 pIs., 2 tabs. Type locality: Canada, British Columbia, Queen Charlotte Islands, NE Graham Island Type horizon: Late Tertiary, late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Preparlltion: Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter), 20-30% HN0 3 (1-6 hrs.), 2% KC0 3 sol. (10-60 mins.), saturated sols. of safranin in H20 and alcohol, mounted in Gelva V-IS plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated Holotype: Pl. IV, fig. 29 Author: Reference:

CATALOG OF FOSSIL SPORES AND POLLEN

44:26

Species name:

Osmunda regaUtes Martin & Rouse 1966

31 FIG.

31. Osmunda regalites sp. nov. (holotype). )\ 1000

Description:

Trilete spores, spherical in outline. Laesurae simple, usually gaping slightly, and ranging from 18 to 22 p. in length. A thin but distinct margo borders the commissure. The wall is about 1 p. thick, and appears to be rigid, as little folding has been observed. The ornamentation is characteristically rugulate, with the rugulae short aIld thick, and packed close together. I ndividllal rugLIlae range to 3 p. in length but rarely exceed 0.5 p. in height. These spores are very similar to those of the extant Osmunda regalis, and have been named from their similarity in ornamentation. Fig. 31 (holotype)

Martin, H.A. and G.E. Rouse, 1966, p. 189, pI. V, fig. 31 Palynology of Late Tertiary sediments from Queen Charlotte Islands, British Columbia: Canadian J. Bot., vol. 44, pp. 171-208, 12 pIs., 2 tabs. Type locality: Canada, British Columbia, Queen Charlotte Islands, NE Graham Island Type horizon: Late Tertiary, late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Preparlltion: Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter), 20-30% HN0 3 (1-6 hrs.), 2% KC0 3 sol. (10-60 mins.), saturated sols. of safranin in H20 and alcohol, mounted in Gelva V-IS plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated Holotype: PI. V, fig. 31 Author: Reference:

CATALOG OF FOSSIL SPORES AND POLLEN

44:27

Species n.me:

Salix apiculata Martin & Rouse 1966

86

87

FIGs. 86 and 87. Salix apiculata sp. nov. (Fig. 87. holotype). X WOO

Description:

Tricolpate pollen grains, elliptical in outline, 20-25 X 13-17 p. in size, with a wall thickness of less than O.S p.. Colpae simple with thin borders, about 16 p.long. The apiculate ornamentation is regular and even. The individual apiculae are 1 p. long, very slender, and spaced O.S p. apart, and are slightly baculate in some specimens. In size and general outline, this species resembles Salix albaejormis1Agranovskaya (in Pokrovskaya and Stel'mak 1960). However, S. albacformis is described as having a definite reticulate pattern, whereas S. apiculata has no apparent muri linking the apiculae. holotype Fig. 87

Martin, H.A. and G.E. Rouse, 1966, p.19S, pI. VIII, figs. 86-87 Palynology of Late Tertiary sediments from Queen Charlotte Islands. British Columbia; Canadian J. Bot .• vol. 44. pp. 171-208, 12 pIs .• 2 tabs. ｔｾ＠ ｬｾＺ＠ Canada. British Columbia, Queen Charlotte Islands. NE Graham Island ｔｾ＠ horizon: Late Tertiary. late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Preparation: Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter). 20-30% HN03 (1-6 hrs.). 2% KC0 3 sol. (10-60 mins.), saturated sols. of safranin in H20 and alcohol. mounted in Gelva V-IS plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated ｈｯｬｾｰ･Ｚ＠ Pl. VIII, fig. 87

Author: Reference:

1

[See Salix ｡ｬ｢･ｦｯｾｩｳＬ＠

CATALOG OF FOSSIL SPORES AND POLLEN

CFSP 16:104.] 44:28

Species name:

Salix minuta Martin & Rouse 1966

84

85

FIGs. IH alill 85. Salix milll/lit sp. nov. (Fig. 84, holotypc). X 1000

Description:

Tricolpate pollen grains broadly to narrowly elliptical, 11-13 p. by 7-8 p.. Colpae simple, their borders 7-10 p. long. The wall is thin, less than O.S p.. The ornamentation consists 01 tiny apiculae, less than O.S p. apart. These grains are undoubtedly Salix, but their small size does not compare with other fossil or ext alit species. holotypc Fig. 84

Martin, H.A. and G.E. Rouse, 1966, p. 195, pI. VIII, figs. 84-85 Palynology of Late Tertiary sediments from Queen Charlotte Islands, British Columbia; Canadian J. Bot., vol. 44, pp. 171-208, 12 pIs., 2 tabs. Type ｬｯ｣｡ｾＺ＠ Canada, British Columbia, Queen Charlotte Islands, NE Graham Island Type horizon: Late Tertiary, late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Preparlltion: Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter), 20-30% HN0 3 (1-6 hrs.), 2% KC0 3 sol. (10-60 mins.), saturated sols. of safranin in H20 and alcohol, mounted in Gelva V-IS plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated Holotype: Pl. VI I I, fig. 84 Author:

Reference:

CATALOG OF FOSSIL SPORES AND POLLEN

44:29

Species n.me:

Selaginella sinuites Martin & Rouse 1966

7 FIGS.

Description:

8

7 and 8. Selaginelta sinuites sp. nov. (Fig. 7, holotype). X 1000

Synonym: Lusatisporis perinatlls Krutzsch (1963b) Spore outline circular or only slightly triangular. Perispore decidedly coarsely granular, finely wrinkled, and extending- from 2 to 5 J.L he yond the limits of the t·lIdospore. ThickIless of perispore ca. 1 I1linon. Thc Iacsurac of lhc trilctc mark are wavy, and the Illar)!;O is IlllICh wrinkled, thus g-iving I.he appearance of a sinuous ribbon. Small radial folds on the perispore occur in the interrauials. Rangc of diaIllctcr: 2R-4X p. Except for a smaller size range, the SkonllIl sporcs appcar conspccilic with those of Lusatisporis perinatus described hy Krutzsch (1963b). However, Krutzsch distinguishes Lusaiisl)oris from Perotrilites Erdtman ex Couper (1953) by the fact that Lusatisl)oris has a single spore wall divided into inner and outer layers, whereas Pcrotrilites has a pcrispore. It is quite clear that the Skonun spores have a distinct perispore, and hence would be c1assilied with Perotrilites. However, because of ·their close similarity to spores of extant Selaginella, the present authors prefer to apply a new form species of this genus, and one whose spores will not be confused with similar spores from the Mesozoic called Densoisporites perillatus 1by Couper (1958). This species has spores virtually identical with those of the extant species Selaginella wallacei and S. oregona from the west coast of North America. The spores of S. wallacei and S. orcgona cannot be distinguished from each other on either size or other morphological characters. holotype Fig. 7

Martin, H.A. and G.E. Rouse, .1966, p. 185, pI. I, figs. 7-8 Palynology of Late Tertiary sediments· from Queen Charlotte Islands, British Columbia: Canadian J. Bot., vol. 44, pp. 171-208, 12 pIs., 2 tabs. ｔｾ＠ ｬｯ｣｡ｾＺ＠ Canada, British Columbia, Queen Charlotte Islands, NE Graham Island ｔｾ＠ horizon: Late Tertiary. late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Preparl\tion: Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter), 20-30% HN0 3 (1-6 hrs.), 2% KC03so1. (10-60 mins.), saturated sols. of safranin in H20 and alcohol, mounted in Gelva V-IS plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated Holotype: Pl. I, fig. 7 Author: Reference:

1

[See Densoisporites perinatus, CFSP 15:130.]

CATALOG OF FOSSIL SPORES AND POLLEN

44:30

Species n.me:

Tsuga heterophyUites Martin & Rouse 1966

46

47

FIGS. 46 alld 47. TSllga heterophyllites sp. nov. (Fig. 46, holotype). X 1000

Description;

Monosaccate pollen grains, circular to elliptical in outline, with a saucershaped body encompassed by a fringe-like saccus. Range of diameter 37-78 J..L, with most specimens about 60 J..L. The ornamentation of the body is finely verrucate, with the semicircular verrucae arranged somewhat concentrically, and measuring about 1 J..L. The leptoma is barely distinguishable by slightly finer verrucae on the distal surface of the body. The fringe-like velum is from 8 to 10 J..L wide, and consists of radially aligned convolutions. In well-preserved specimens, fine spinae up to 2 J..L long can be found attached to the convolutions of the saccus. Grains assigned to this species are identical with those of the extant and indigenous Tsuga hctcrophylla. ho\otype Fig. 46

Martin, H.A. and G.E. Rouse, 1966, p. 192, pI. VI, figs. 46-47 Palynology of Late Tertiary sediments from Queen Charlotte Islands, British Columbia: Canadian J. Bot., vol. 44, pp. 171-208, 12 pIs., 2 tabs. Type locality; Canada, British Columbia, Queen Charlotte Islands, NE Graham Island Type horizon; Late Tertiary. late Miocene-early Pliocene (channel sampling of suitable beds from top to bottom of each outcrop) Prepar\\tion; Decalcification and desilicification with HCl and HF (adjusted for various concentrations of mineral matter), 20-30% HN03 (1-6 hrs.), 2% KC03sol. (10-60 mins.), saturated sols. of safranin in H20 and alcohol, mounted in Gelva V-IS plastic dissolved in dioxane (refractive index 1.467 at 20°C.) Repository: Not indicated Holotype: Pl. VI, fig. 46 Author; Reference;

CATALOG OF FOSSIL SPORES AND POLLEN

44:31

Species n.me:

Tsuga minisacca Martin & Rouse 1966

49

48

FIGs. 48 and 4s sp., Ｑｦｾ＠ d,'hiseenee mark which runs only to ｴｨＬｾ＠ zone, as in the gcnerotype deserihed hy BHARDwA Y (I. c.).

or

Holotype: HGJ Pollen Laboratory. Bore-hole: no.: Zgv. 59; S''lmple: 22. Slide: 1. Cross-tahle no.: 39.3 to 112.0 Locus ·typicus: Zengoviirkony Stratum typieum: Upper Helvetian, grey fish scale clay marl; hore-hole' no.: Zgv. 59; depth: 51.3 to 56.0 m.

[Diagnosis: Trilete microspore with circular outline and conve!x side. Dehiscence mark straight, reaching the equator and thickened on both sides. Exine scahrate. At the distal pole, ornamentatioJl in a rotund area. Hl'lllark: The recl'll! species PlweoC('ros /1II/bicIl/OSIIS (Brntern) Prosk. is very similar to this gellcrotype both ill the ordcr of magnitudc allli JIIorphologically. According to the Volume "Die Lehermoose" of K. MULLER (pp. 1305 an,l 1313) tllf' ｾｰｬＧ･ｩｳ＠ of the gl'lIera ;(II'/IO(:e/'Os all,] Plta"o("/'ms lIlay I... t1iffl'fI'ntiated by their spores. So it was done by PROSKAUER in separating also the genera above mentioned (I. c.). GCllerotype: Phaeoeel'osporites baranyaensis n. g. n. sp. Locus typicus: see the description of the species Stratum typicum: see the description of the species] 2

CATALOG OF FOSSIL SPORES AND POLLEN

44: IDS

PhaeoceY'ospoY'i tes baY'anyaensis Nagy 1968

Species name:

Nagy, E., 1968, p. 116, pI. I, figs . 7-8, text-fig. 3 Reference: Moss spores in Hungarian Neogene Strata: Acta Bot. Acad. Sci. Hungaricae, v. 14, no. 1-2, pp. 113-132, 8 pIs., 7 text-figs. Type locality : Hungary, Zengovarkony, Hungarian Geo!. Inst. Bore-hole no. Zgv59, depth 51.3 to 56. Om. Type horizon : Miocene, Helvetian (upper) Preparation : Not given Repository : Hungarian Geo!. Inst. Pollen Lab. ColI., Budapest Holotype : PI. I, figs. 7-8, bore-hole no. Zgv. 59, sample 22, slide 1, coord. 39.3 x 112.0 Author :

l[See TY'ilites lachanae, CFSP 2:126.] 2

[Bracketed text republished from the generic description of PhaeoceY'ospoY'ites, Nagy, 1968, p. 115.] TYPE SPECIES OF THE GENERIC NAME PhaeoceY'ospoY'ites Nagy, 1968

CA TALOG OF FOSSIL SPORES AND POLLEN

44: 106

Specie. n.me: Phaeocerosporites transversus Nagy 1968

1 1,111,12. I'hn,-,wao-',II"'j,,-s

2

'''II/Sf''-'',,,,,, ｉＮｾＬ＠

Ilolnly' .. ·.

Description:

Diagnosis: Trilete, spheroid spore, 66 X 58 p in size, with a 4 to 5 p broad zone. Dehiscence mark straight, running up to the equator. On both faces a thickening similar to that of the zone and tapering towards the equator may be seen. On the proximal face, in addition to the fine scabrate ornamentation, there are also separated grana, surrounding roughly the pole. The middle part of the distal face - an area of about 40 p in diameter - is sculptured by meandering elements, processes and depressions reaching the margin here and there. Differential diagnosis: The species in question is separated from Phaeocerosporilcs ))aranyaensis1n. g. n. sp. hy the ornamentation of its distal and hy the granae of the proximal face. Due to the more pronounced ornamentation the border of the spore is more uneven too. Remarks: Two specimens were found, togl'thl'r with an other spure of fairly simple sculpturing, in the same sample of the material from the Mccsek Mountains. Knowing the extremely high intraspecific variability of recent spores, a eloser connection between both forms may be assumed, but for lack of evidence they are separated - for the moment - morphologically.

CATALOG OF FOSSIL SPORES AND POLLEN

44:107

Jlolot ｾ＠ pc': IIG 1 Poll,·" LalHlrat ur)'. ｂｵｮｾＭｨｯｬＨＢ＠ 1111.: Zgv. ｾＹ［＠ Samplco: 22. Slide': ｾＮ＠ ｃｲｯｾｳＭｴ｡｢ｬＨＢ＠ IHI.: 30.0 to 111.0. ｌｏＨＬｉＱｾ＠ t Ypiell!': ｚＢｉＱｾｻ［｜Ｇ｡＠ rkollY Stratum typieum: Upper HeI\'('tian, gr('y, fish scale day marl; ｉｊｏｲＨＧＭｬｷｾ＠ 110.: Zg\". 59; depth; 51.0 to 56.0 m.

Author:

Nagy, E., 1968, p. 125, pl. II, figs. 1-2 Moss spores in Hungarian Neogene Strata: Acta Bot. Acad. Sci. Hungaricae, v. 14, no. 1-2, pp. 113-132, 8 pIs., 7 text-figs. Type locality: Hungary, Zengovarkony, Hungarian Geo!. Inst. Bore-hole no. Zgv59, depth 51.3 to 56. Om. Type horizon: Miocene, Helvetian (upper) Preparation: Not given Repository: Hungarian Geol. Inst. Pollen Lab. ColI., Budapest Holotype: Pl. II, figs. 1-2, bore-hole no. Zgv. 59, sample 22, slide 2, coord. 30.0 x 111.0

Reference:

l[See Phaeocerosporites baranyaensis, CFSP 44:104.]

CA TALOG OF FOSSIL SPORES AND POLLEN

44: 108

Specie. n.me:

Riooiaespori tes hungarious Nagy 1968

1

3

2 Plate III. 1.

Hi('(';/If·': Specimen in PI. UXI, Figs. 5-8; 39 ｾｭＮ＠ Locus typicuB and slide: Svatopluk Mine, Czechoslovakia. - 28/1 (6,0-101,1:1 Z = 31,3-103,1 L). Stratum typicum: Miocene brown-coal seam, overlying complex. Deposition of origin al: Paleontological Institute of the Charles University, Prague, Czechoslovakia.

Pacltova, B. &P. Simoncsics, 1970, p. 606, pl. CXI, figs. 5-14, pl. CXII, figs. 1-15, text-fig. 9 New types of spores (genera and species) from the Bohemian Miocene: Reference: Palaont., Abh. B-Palaobot., III., no. 3/4, pp. 599-617, pIs. CV-CXIII, 9 text-figs., 2 tabs. Type locality: Czechoslovakia, southern Bohemia, near Mydlovary, Svatopl uk mine, sample 28, depth 7.95-8.lSm. Type horizon: Miocene, Helvetian-Tortonian age, Mydlovary beds Not indicated Preparation: Collection of Pa1eont. Inst., Charles Univ., Prague Repository: Holotype: Pl. CXI, figs. 5-8, slide 28/1, coord. 6.0 x 10l.9Z = 31.3 x l03.lL (Coordinates for specimen location are provided for both a Zeiss Nfpk (identified by a "Z") and a Leitz Ortholux (identified by an "L") microscope.)

Author:

CATALOG OF FOSSIL SPORES AND POLLEN

44: 135

Specie. name:

Poroisporoi tes mioaeniaus Pac1tova &Simoncsics 1970

10

11 ｐｌａｔｉｾ＠

12 GlX

J'orisporites miocenicU8 gen. et sp. n. Figs. 10--12:

H o\otYPUB t'iid,,: 21'/1 (20,5-105,:1 JI: = :14,1-117,5 L)

Text-fig. 6. l'urispur'ites miocenicus gell. et tip. n. A - proximal face; B - distal face; C - optical cross-section

Description:

Diagnosis: see the diagnosis of the genus and text-fig. 6. Hoio!,ypIlH: ::ipOCilllOll illl'\' C\X, Fig. 10-12; 34 ,.un. Locus t.ypiclIs and slido: ::lvn.topluk Mino, ｃｾ･｣ｨｯｳｬｶｵＮｫｩＱ＠ - 28/1 (20,5-105,3 Z = 34,1-117,5 L). ::ltra!,um typicum: Miocene brown-coal seam, OVCI'lyillg complex. Deposition of original: Paleontological Institute of the Charles University, Prague, Czechoslovakia.

[Diagnosis: Up to now monotypic trilete spore genus with cingulum and rounded convex equatorial contour. Cingulum gradually widening on sides, in the centrc of sides with an aperture or a hole. Enuoexine equally thin, V = 6-7/1 on one side, 3/1 on the angles. Equatorial l)Ontour of central cell concave, proximal face covered with fine warts, distal face covered wit.h eoanw wart.H. Y -mlLl'k ruuning to cingulum, r = 5/5. Hemarks: Porispuritcs gen. n. differs from other cingulate spore genera. in having perforations of cingululII. Botanically, it can he a Cyatheaceae (llemitelia). '1' y P IIt1 goneris: l'uT'isporites miocenicus sp. n.] 1

CATALOG OF FOSSIL SPORES AND POLLEN

44 :136

Pacltova, B. & P. Simoncsics, 1970, p. 60S, pI. CIX, figs. 10-12, text-fig. 6 New types of spores (genera and species) from the Bohemian Miocene: Reference: Palaont., Abh. B-Palaobot., III., no. 3/4, pp. 599-617, pIs. CV-CXIII, 9 text-figs., 2 tabs. Type locality: Czechoslovakia, southern Bohemia, near Mydlovary, Svatopluk mine, sample 28, depth 7.95-8.15m. Type horizon: Miocene, He1vetian-Tortonian age, Mydlovary beds Not indicated Preparation: Repository: Collection of Paleont. Inst., Charles Univ., Prague Holotype: Pl. CIX, figs. 10-12, slide 28/1, coord. 20.5 x 10S.3Z = 34.1 x 117.SL (Coordinates for specimen location are provided for both a Zeiss Nfpk (identified by a "Z") and a Leitz Ortholux (identified by an "L") microscope.) Author:

l[Bracketed text republished from the generic description of Porisporites, Pacltova &Simoncsics, 1970, p. 60S.] TYPE SPECIES OF THE GENERIC NAME Porisporites Pacltova &Simoncsics, 1970 (Monotypic in Pacltova &Simoncsics, 1970)

CATALOG OF FOSSIL SPORES AND POLLEN

44:137

Specie. n.me:

SvatopZukisporites hymenophorus Pac1tova &Simoncsics 1970

PLATE eVIl SvatopluldspoTitcsllunwfwplwTuB gOIl.

Fig. 3:

Holotypus Slide: 10/1 (14,5-118,4 Z

=

ct Rp. n.

47,2-111,7 L)

-..

PLATE eVllI ｓｉｾ｡ｦｬＬＢ＠

FigI'!. 1,2:

.. isJlfl,·it(·s 11!IIIU'II(IJl/ulI'IIH glJl.

Figs. 1;-8:

CATALOG OF FOSSil SPORES AND POllEN

II.

Slillo: '28/3 (39,8-104,9 L)

44: 142

Species n.me:

ZZivisporis major' Pac1 tova & Simoncsics 1970

9

10

11

PLATE CXIII ｬ＼Ｇｩｧｾ

12

Ｎ＠ !) -11

: Fig:;. 12, 1:1:

8lide: '!.8/:!. (48,S - 104,3 L) Slillo: 28/3 (:Hi,7 - 10!I,:1 L)

13

ｾ＠

. .. . '.

.

ｾＮ＠ .

.

.

A

.\: IIwl'itlhulHl H.. ﾷＨｾｴｩＢｉ［＠ till rlU'(':

n

-

1\: I' - IU'I,xiIIUlI r,W(\; tt - (liM· "p' Ic'n I ｃﾷｉＧｕａ ﾷ ｾｬﾷＨＮｴｮ＠

CATALOG OF FOSSIL SPORES AND POLLEN

44:143

Description:

DiagllosiH: A l'ct.ieuloid speeies of Zlivi8poris with pCl'ispore directed hackwards ill tlw din'('f iOIl of I,llI' rays of Y -mal'k, in fOl'm of a t.llin 1l1l'milralW wit.h ｉｩｴＮＬｨｾ＠ spinae. Proximal pal't oI'lHlIlll'ntl'd with irl't'glllHt' fUI'I'ows 01' \l'Hl'tS. 'I'll!' rays or t,III' Y -lIIal'k HI'I' lIot, 1,11'111'. till'." lin', J'('llI'I'sl'llkd lIy t.ilil,kl'l' IInhl'oken dphiscoIH'" linOR follo\l'illg the dil'l'd.ion of rays lip 1.0 Lill' mal'gill (If pl'l'ispores (I' ｾ］＠ ri/ri). Distal ± nett.ed. J II the cellt.re of Ow dist.:il suri'al'e t.h., IId,lol'd folds HI'C I II 1d,l'I'. 011 t,lH' IIIHI'gill f1aU,m', 'I'\I'o-layl'I,.,d wall, equat.orial diameter ahout 41.tm, illlH'r \l'all lwlo\\' 0,5 !-,-Ill. V = 8-1O/q. Hell text-fig. a.

t: ,'\11 n I' k S: By i I" t hiekl'lIillg of Iwl "'d folds 011 I Ill' dislal I'llI'll Z/':/'':,,/lOri,y mrtjor sp. II, d i fT,'l's 1"1'011\ ZI; 1'; . •'111)1';.' ＢＯｉｬｲＧｦ［ＬＮｾＨｎ＠ Ali Y I !HiH) tH'W eo III h. 11-11(1 fl'l >II I Z. 11'(1 II 8r1(/. II ＯｬＱＩｩＧｉＬｾＨ＠ N ,Iii Y I !HiH) lun\' ('0 III h. Bol all it'lIlly "I}t'nkill!!. il i" I\. "pt'eil'" of' I hI' gl'IlIlH Hil'ci(/.. LII I,h" "It III}! I" :;\'II-I,opluk 2H II-hOIl!, 10 "pl'eillll'm' of' Ihn HiZl' of HO - 120 ｾｭ＠ have been fouud. M"I'I' spcei,'s which could belollg to tho gl'IlIlS Zli'l!ispol'is l' ACLTOYA ('.ollid he eitm/ 011 tilt, ImHis of litt'I'H'·Y dllta. A!'col'dillg to t.h!' presellt-day klloll'lodgc the forms mcntioncd call be Hsclwtailled ill til(' Lowt'1' CI't'!,HC,'OIl" lip to Plioc('Il(, till' of AII",t.l'alil\., Sihpl'ia !llld EIIl'op(', TIll' 7.li/·;sjlol'is !o;: Specimell ill PI. CVJ. Figs. 1,2; 108 ｾＱｉｬＮ＠ Loells Iypiclls nlld ",lido: Svatup/uk Mille, C),;l'ohoslovakia. - 2H/4 (45,8-!J!J,7 L). HI 1'1\.11I1\l t,ypicull1: Mioe('lll' bl'o\\'ll·onal seam, nV('l'lyillg cOllJplex. De p u 8 i t. i 0 II of ol'igillal: Paleolltological lllst.itut e of t.ho Clml'los Unin'I'Hit y, l'rllglll', ('),;I'eh"sl",' .. kiH.

Pacltova, B. &P. Simoncsics, 1970, p. 602, pl. CVI, figs. 1-2, pl. CVII, figs. 1-2, pl. CXIII, figs. 6-13, text-fig. 3 New types of spores (genera and species) from the Bohemian Miocene: Reference: Palaont., Abh. B-Palaobot., III., no. 3/4, pp. 599-617, pIs. CV-CXIII, 9 text-figs., 2 tabs. Type locality: Czechoslovakia, southern Bohemia, near Mydlovary, Svatopluk mine, sample 28, depth 7.95-8.l5m. Miocene, Helvetian-Tortonian age, Mydlovary beds Type horizon: Not indicated Preparation: Collection of Paleont. Inst., Charles Univ., Prague Repository: Pl. CVI, figs. 1-2, slide 28/4, coord. 45.8 x 99.7L (Coordinates Holotype: for specimen location are provided for a Leitz Ortholux microscope.)

Author:

l[See Ricciaesporites hungaricus, CFSP 44:113. These proposed nov. combs. are not validly published for want of required documentation per Art. 33.2, ICBN 1978.]

CATALOG OF FOSSIL SPORES AND POLLEN

44: 144

Specie. n.me:

AmosopoUis dilwynensis Harris 1972

4 Figs. 4,5.

5

Amosopollis dilwYllellsis sp. nov. Py 015, 38.7: 100.1. Fig. 5, X 1250. Notice the granulate margin of the sulcus. Note: Unless otherwise spcdfied the figures are X5()() in normal Iransmilled light.

Description: Synonymy: A mo.l'O[1olli.f crud/ormis sensu Harris 1965a: 97, PI. 29, fig. 26. Pollen grains in rhomboidal tetrads. Individual grains prolate to sub-prolate. Exine 2 pm thick. psilate to scabrate and finely granulate. except near the margins of the aperture where grana 1-1.5 I,m diam. arc present. Aperture is a long gaping sulcus extending the full length of the grain. Margins of sulcus not ragged. Dimensions: (10 specimens) Overall tliam. of tetrad 50 (60) 68 p.m. Individual grains 22 (34) 40 p'm diam. Type locality: Dilwyn Bay. Victoria. Pebble Point Formation. Paleocene.

Distribution: A. dilwynensis is a rare species but has been observed in Paleocene sediments from the Murray and Otway Basins, and a similar form has heen reported (Harris 1965b) from Queensland in sediments of similar age. Compari.l'On and ajJinity: A. dilwynell.\·;s is in ｾ･ｮｲ｡ｬ＠ larger than the ｾ･ｮｯｴｹｰ＠ bllt can also be distinguished hy the psilatc-scahrate sculpture and more importantly by the straight margins of the sulcus. Holotype: Preparation and slide numberST209/2, 39.3: 100.7. Py 015. Figs, 4. 5.

Author: Harris, W. K., 1972, p. Reference: New form species of

54, figs. 4-5 pollen from southern Australian early Tertiary sediments: Trans. Roy. Soc. South Australia, vol. 96, pt. 1, pp. 53-65, 49 figs. 1 tab. Type locality: Australia, Lat. 38°44'00"S, Long. l43°l0 1 30"E, Otway Basin, SE side of Dilwyn Bay, 1.2m. above base of formation Type horizon: Tertiary, Paleocene, Pebble Point fm. Preparation: Crush sample, dilute HCl; 5g. carbonate-free sample, 50cc 60% HF (boil to sludge), 40ml. 10% HCl (boil 1-2 mins.), 60ml. Schultze sol. (heat gently 1 min.), 4Oml. 10% K2C03 (heat to boiling), 30ml. conc. HN03 (10 to 20 secs.), 30ml. 10% K2C03 (20 secs.), glycerin jelly mounts, safran in 0 stain, gold size seal (per Harris, 1965) Repository: Geo!. Surv. South Australia, Palaeont. ColI. (Holotype catalog numbers bear a Py prefix.) Holotype: Figs. 4-5, slide no. ST209/2, coord. 39.3 x 100.7, Cat. no. Py015 (Coordinates apply to Leitz Orthoplan (715494) microscope, Palynological Laboratory, Geol. Surv. South Australia, Adelaide.)

CATALOG OF FOSSIL SPORES AND POLLEN

44:145

Specie. n.me:

Eriaipites arassiexinus Harris 1972

15

16

Figs. 15,16. Ericipites crassiexinus sp. nov. Fig. 15.-S660/1, 52.9: 96.9. Fig. 16.-S560/1, 19.2: 107.2. Note: Un'less otherwise specified the figures arc X500 in normal transmilled ligh!.

Description:

Pollen united in tetrads. Individual grains indistinctly tricolporate, tetrahedral in shape and strongly united in the tetrad. Exine 2.53.5 I!m thick. sex me as thick as nexine psilate. Apertures complex, colpi ahout 14 I!m long and 1.5 I-lm wide. Pores indistinct and dillicult to detect, 2 I-lm diam. Dimensions: Overall diam. 35 (42) 53 I-lm. Individual grains 24 (30) 35 I-lm diam.

Distribution: Often a very common form in middle and upper Eocene sediments. Comparison and affinity: The psilate nature of the exine and the larger size of this species distinguishes it from E. scC/bratus Harris. Pollen or this type chnracterise the Order Ericnles. HolOlype: Preparation nnd slide numberS660/ I. 52.9: 96.9. Py 415. Fig. 15.

Type locality: Bore, Hd. Cummins at 35.743.3 m. Wanilla Formation, Middle Eocene.

Harris, W.K., 1972, p. 54, figs. 15-16 New form species of pollen from southern Australian early Tertiary sediments: Trans. Roy. Soc. South Australia, vol. 96, pt. 1, pp. 53-65, 49 figs. 1 tab. Type locality: Australia, Lat. 34°l5'10"S, Long. l35°40'45"E, Cummins Basin, Hd. Cummins (W. Con. Res. adj. Sec. 16) Bore, 35.7-43.3m. Type horizon: Tertiary, Middle Eocene, Wanilla fm. Preparation: Crush sample, dilute HCl; 5g. carbonate-free sample, 50cc 60% HF (boil to sludge), 40ml. 10% HCl (boil 1-2 mins.), 60ml. Schultze sol. (heat gently 1 min.), 40ml. 10% K2C03 (heat to boiling), 30ml. conc. HN03 (10 to 20 secs.), 30ml. 10% K2C03 (20 secs.), glycerin jelly mounts, safranin 0 stain, gold size seal (per Harris, 1965) Repository: Geol. Surv. South Australia, Palaeont. ColI. (Holotype catalog numbers bear a Py prefix.) Holotype: Fig. 15, slide no. S660/l, coord. 52.9 x 96.9, Cat. no. Py4l5 (Coordinates apply to Leitz Orthoplan (715494) microscope, Palynological Laboratory, Geol. Surv. South Australia, Adelaide.)

Author: Reference:

CATALOG OF FOSSIL SPORES AND POLLEN

44 :146

Specie. n.me: Proteacidites clintonensis Harris 1972

32

33

34

31

30

Figs. 30-34. Pm/i'llcidi/('s clill/oll(,lIsis sp. nov. Figs. 30. 31.-S741/2. 47.11: 96.3. Fig. 31. X 1250, high foclIs on polar region. Figs. 32-34.-S705/1. 31.3: 105.2. Fig. 33, X 1250, high foclIs on polar region; fig. 34. foclIs on apcrtmal region. Note: Unless otherwise sJlecified the figures are X500 in normal transmitted light.

Description: Pollen liub-isopolar. oblate :lIlgulaperturate. triporate. Amh more or less trianglliar with conCave sides. Pores circular 20-35 p'm in diam. Exine 3 ILm thick. sexine slightly thinner than ncxine. Capita of bacula coalesce 10 form groups up to 7 p'm wide and show an LO pattern. Elements rounded in optical section. Nexine in region of pores. alternately thick and thin. The sexine is readily lost by corrosion. Dilll£,I/.\·ioll.\·: Equatorial diam. 62 (75) 98 p.m. Dis/riblllioll: The species is almost ubiqui-

tous in Eocene sediments and is particularly common in the Triori/es IIIaglli/iclIs1Zonule. It ranges from Middle Eocene to at least Lower Miocene. Comparisoll alld affinity: This species is similar to P. rec/omargilllls Cookson but has much

CATALOG OF FOSSIL SPORES AND POLLEN

larger apertures and strongly concave sides. Figure 30 more closely resembles P. rec/oIIIal'gillll.l'2 with its finer ornament. larger size and straighter sides. It is possible that the two forms intergrade. Cookson's figure (1950, fig. 27) of P. rec/olllal'gillll.f appears to show some thickening of the nexine about the apertures. The species is distinguished from P. incurvaflu 2 by the nature of the sculpture and the characteristic aperture. The exine does not thin markedly near the apertures as it does in P. illCllrvatllS. Holo/ype: Preparation and slide numberS7051 I. ;.\,/,.ill/l/iol/: Common in. the "Wilklltllllll Formation" but less common in other Middle Eocene (Proteacidites ｣ｯｮｬｲ｡ｧＬｷｾ＠ Zonule) assemblages. Compari,fOlI alld affillity: The strongly invaginated colpi and psilate exine make this 11 very distinctive species. Its natural affinities are unknown.

T.l'pr Incality: Lake Torrens Bore 3A at 247.8 m. "Wilkatana Formation", Middle

lIolotypl': Preparation and ｳｬｩ､ｾ＠ numherST2411 12.35.1: 9!U. Py 176. Fig. 9.

Eocene.

Harris, W.K., 1972, p. 54, figs. 8-9 New form species of pollen from southern Australian early Tertiary sediments: Trans. Roy. Soc. South Australia, vol. 96, pt. 1, pp. 53-65, 49 figs. 1 tab. Type locality: Australia, Lat. 3l o l4'00"S, Long. l38°0l'45"E, Pirie-Torrens Basin, Lake Torrens Bore 3A, 247.8m. Type horizon: Tertiary, Middle Eocene, "Wilkatana" fm. Preparation: Crush sample, dilute HCl; 5g. carbonate-free sample, 50cc 60% HF (boil to sludge), 40ml. 10% HCl (boil 1-2 mins.), 60ml. Schultze sol. (heat gently 1 min.), 40ml. 10% K2C03 (heat to boiling), 30ml. conc. HN03 (10 to 20 secs.), 30ml. 10% K2C03 (20 secs.), glycerin jelly mounts, safran in 0 stain, gold size seal (per Harris, 1965) Repository: Geol. Surv. South Australia, Palaeont. ColI. (Holotype catalog numbers bear a Py prefix.) Holotypc: Fig. 9, slide no. ST24l/l2, coord. 35.1 x 98.1, Cat. no. Py176 (Coordinates apply to Leitz Orthoplan (715494) microscope, Palynological Laboratory, Geol. Surv. South Australia, Adelaide.) 1 [See Proteaaidites ｡ｯｮｦｲｧｳｵｾ＠ CFSP 44:150.] CATALOG OF FOSSIL SPORES AND POLLEN 44: 163 Author: Reference:

"Triol'ites"lpsilatus Harris 1972

Species n.rne:

7

6 Figs. 6, 7.

"Triorites" ｰｳｩｬｑｉＮｾ＠

sp. nov. X 1250.

Fig. 6.-S564/1, 32.8: 100.6.

Fig. 7.-5562/1,

37.7: 104.9.

Description:

Pollen radiosymmetric, isopolar, oblate, triorate. Amb sub-triangular, sides straight to slightly convex. Ora sunken, 2-4 Jl-m wide, circular. Exine 2J1-m except around 'apertures where it thickens to 3 or 4 Jl-m. Exine psilate. Dimensions: Equatorial diam. 24 (32) 40 Jl-m . TYflt' loculity: PollIa No. I Bore at 55.1 Ill . 1'11\'11'1'1111 Formnt illll, M hhtlt' POrt'Ill'.

/)i.\'trilllltioll : This spccies is a very common

forlll

ｴｨＯＧｏｉｾｯｬｩ＠

the

I ,ower Tertiary

in

southern Australia. It first appears in the Prineetown M einher and range!\ through to

the Lllwer M illcellc. The uJlJlcr limit hn)! not been determined. Comparison and affinity: "T" psilallIs is compantble and may be conspecific with "T." .fcahratuiCouper. The ornament on the latter however is scabntte. This species would more appropriately be placed ina new genus (see section on TrioritC's below). Holotype: Preparation and slide numberS564/ I, n.R: 100.fl. Py 411. Fig. fl.

Author : Harris. W.K •• 1972. p. Reference: New form species of

54. figs. 6-7 pollen from southern Australian early Tertiary sediments: Trans. Roy. Soc. South Australia. vol. 96. pt. 1. pp. 53-65. 49 figs. 1 tab. Type locality: Australia. Lat. 33°33'00"S. Long. l35°20'00"E. Polda Basin. Polda No.1 Bore, 55.lm. Type horizon: Tertiary. Middle Eocene, Poelpena fm. Preparation: Crush sample, dilute HCl; 5g. carbonate . . free sample. 50cc 60% HF (boil to sludge). 40ml. 10% HCl (boil 1-2 mins.). 60ml. Schultze sol. (heat gently 1 min.). 40ml. 10% K2C03 (heat to boiling). 30ml. conc. HN03 (10 to 20 secs.). 30ml. 10% K2C03 (20 sees.), glycerin jelly mounts. safranin 0 stain. gold size seal (per Harris, 1965) Repository: Geol. Surv. South Australia. Palaeont. ColI. (Holotype catalog numbers bear a Py prefix.) Holotype: Fig. 6, slide no. S564/l. coord. 32.8 x 100.6, Cat. no. Py4ll (Coordinates apply to Leitz Orthoplan (715494) microscope. Palynological Laboratory. Geol. Surv. South Australia. Adelaide.) 1

[The quotation marks enclose the generic name following the practice of Harris; see caption to figs. 6 and 7.]

2[See Tl'iorites scabl'atus, CFSP 4:3.] CATALOG OF FOSSIL SPORES AND POLLEN

44: 164

Specie. n.me:

Triporopollenites gemmatus Harris 1972

11

13

10

14

Figs. 10, II. Triporopollenites gemmallls sp. nov. Fig. I 0.-S650/1 , 43.4: 104.5, single grain. Fig. 11.-S547/1, 31.7: 98.4, tetrad. Figs. 13, 14. T. gemmatlls sp. nov. Fig. 13.-S547/1, 31.7: 98.4, X 1250 high focus. Fig. 14.5741/2, 38.6: 106.8. Nole: Unless olherwise specified Ihe figures are X500 in normal transmitted lighl.

Description:

Pollen occasionally free but most commonly united in tetrads. Tetrads 34-40 p'm in overall diam. Individual pollen radiosymmetric, oblate. sub-isopolar. triorate. Amb sub-triangular with straight to convex sides. Exine 4-5 p'm thick (including ornament). Sexine and nexine difficult to separate but nexine appears to be thicker than sexine. Exine covered with verrucae 2-3 p'm wide, sphaerjcal and 2 p'm high. Verrucae separated from each other (by 2-3 ,tm) by granulate ornament. Apertures obscured by ornament, porate or orate opening 1.5-2.5 p'm wide. Dimensions: Individual pollen. equatorial diam. 25 (29) 31 p.m. Type locality: Lake Cootabarlow Bore 2 at

CATALOG OF FOSSIL SPORES AND POLLEN

163.4 m. Great Artesian Basin. Murnpeowie Formation. Upper Eocene. Di.\·trihlllion: Appears to be restricted to Middle and middle-upper Eocene sediments from the Pirie-Torrens and Great Artesian Basins and Eyre Peninsula. Comparison a1ld affinity: T. gemmatus is similar to T. bullis Gruas-Cavagnetto (1966) from the Sparnacian of the Paris Basin but this species is more or less circular and appears to have a more strongly thickened rim to the aperture. Holotype: Preparation and slide number-

S54711. 31.7: 98.4.

Py 720.

Fig. 11.

44: 165

Harris, W. K., 1972, p. 55, figs. 10-11, 13-14 New form species of pollen from southern Australian early Tertiary sediments: Trans. Roy. Soc. South Australia, vol. 96, pt. 1, pp. 53-65, 49 figs. 1 tab. Type locality: Australia, Lat. 30°16' 30"S, Long. 140°08' 30"E, Great Artesian Basin, Lake Cootabarlow Bore 2, l63.4m. Type horizon: Tertiary, Upper Eocene, Murnpeowie fm. Preparation: Crush sample, dilute HCl; 5g. carbonate-free sample, 50cc 60% HF (boil to sludge), 40ml. 10% HCl (boil 1-2 mins.), 60ml. Schultze sol. (heat gently 1 ｭｩｮＮＩｾ＠ 40ml. 10% K2C03 (heat to boiling), 30ml. conc. HN03 (10 to 20 secs.), 30ml. 10% K2C0 3 (20 secs.), glycerin jelly mounts, safran in 0 stain, gold size seal (per Harris, 1965) Repository: Geol. Surv. South Australia, Palaeont. ColI. (Holotype catalog numbers bear a Py prefix.) Holotype: Fig. 11, slide no. S547/l, coord. 31. 7 x 98.4, Cat. no. Py720 (Coordinates apply to Leitz Orthoplan (715494) microscope, Palynological Laboratory, Geol. Surv. South Australia, Adelaide.) Author: Reference:

CATALOG OF FOSSIL SPORES AND POLLEN

44:166

Genus name:

Gambierina Harris 1972

12 Fig. 12.

Gambierina edwardsii (Cookson & Pike) Harris comb. nov. ST 208/2, 32.1: 103.7. Note: Unless otherwise spc..:ilied the figures are X5()() in normal transmitted light.

Description:

Genus GAMBIERINA gen. nov. Type species: ｔｲｩｯｬＨＧＬｾ＠ ＨＧ､ｬＯｲＮｾｩ＠ lCookson & Pike (in part) 1954: 214. pI. 2, figs. 101.2 105, 106. ｄｩｬｧｯｳＬｾＺ＠

Pollen radiosymmetric, oblate. lobate. angulaperturate, triorate. Apertures sunken. Sexine imperforate tectate, thinner than nexine, the two separated by a faintly discernable baculate layer, which forms a "nick" point in the apertural region. Aperture formed by sexine larger than that of the nexine. Nexine thickens more rapidly than sexine about the apertures. Exine psilate.

Figured specimell: Fig. 12. Remarks: The characters of the exine, the apertures and general shape distinguish this genus from Trioriles. As Dettmann & Playford (1968. p. 86) have pointed out. the species figured by Cookson & Pike (1954, particularly figs. 104 and 105) as T. edwardsii is distinct in being unthickened about the apertures. Dettmann & Playford (1968) summarised the present status of the genus Trioriles but

CATALOG OF FOSSIL SPORES AND POLLEN

chose to continue using the diagnosis of Couper ( 1953) pending a review by the present author. Potonie (1960) c1e:lfly indicated that the two species T. maglli!iclI,\·3 Cookson and T. clavllllls 3 Cookson were morphologically comparahle and distinct from other forms allocated to the genus. However, Potonie gave no indication as to where these other forms should be placed. It is clear that T. mllKnificlIs and T. ｣ｬ｡ｶＮｾ＠ are very closely related morphologically and perhaps phylogenetically. Indeed Cookson (1957. p. 49) goes so far as to state that "there is little or no doubt that they were produced by closely related plants. Both spccies have the same shape, type of ora and exine stratification. and structure. . . " Thus these two species form a natural grouping and all other species assigned to the genus are better accomodated elsewhere. Couper's (1953) diagnosis is too broad and suggestive of a suprageneric category. Mildenhall & Harris (1971) have reached similar conclusions.

44:167

Author: Harris, W.K., 1972, p. Reference: New form species of

55, fig. 12 pollen from southern Australian early Tertiary sediments: Trans. Roy. Soc. South Australia, vol. 96, pt. I, pp. 53-65, 49 figs. 1 tab. 1

[See Triorites edwardsii, CFSP 15:112.]

2[Jansonius &Hills 1976 "GFFS" card no. 1085 designated the specimen illustrated in Cookson &Pike, 1954, pl. 2, fig. 101 as a lectotype of Gambierina edWardsii.] 3[See Triorites magnificus, CFSP 15:42, and T. cZavatus, CFSP 15:66.]

GENERIC DESCRIPTICN OF THE NEW GENUS Gambierina Harris, 1972 (Monotypic in Harris, 1972)

CATALOG OF FOSSIL SPORES AND POLLEN

44: 168

Species n.me:

Myrtaaeidites euaaZyptoides Cookson & Pike 1954 f. aonvexus Cookson &Pike 1954 emend. Martin 1973

88 Figs. 88, 89.-·!IIyrtaceidites ellca/yptoides f.

89 COllveXIIS

(Cookson and Pike) emend.

Description:

Emended Diagllo!is.1- The grains are sub triangular in outline, sides curved and angles rounded, syncolpate or parasyncolpate with well-defined colpae. The polar island varies from non-existent to large (6 J,lm). The pattern of the exine (0.5-1 J,lm thick) is faintly scabrate or reticulate, often best defined near the equator, tending to fade out towards the colpae and pores. The pores have a narrow vestibule above which the exine is smooth and very clearly defined, possibly thickened.

Size. -18(22)30 J,lm. Afllnitics {lnd COl/lparisons. - Cookson and Pike (l954b) desaibe the exine as thicker around the pores. On these specimens, sometimes a thickening cannot be detected but the exine is always smooth and clearly defined around the pore. Otherwise, they agree well with the original diagnosis, which has been emended to include grains which are not thickened around the pores. The original authors have suggested a possible affinity with Eucalyptus and all the above characteristics may be seen on living pollen of this genus. Known Range. -Pliocene to Pleistocene (Cookson and Pike 1954b). Distribution. - The Lachlan and Cowra Formations. It is one of the common fossil species of the Myrtaceae.

Author: Reference:

Martin, H.A., 1973, p. 22, figs. 88-89 The palynology of some Tertiary Pleistocene deposits, Lachlan River Valley, New South Wales: Australian J. Bot., sup. ser. no. 6, pp. I-57, 202 figs. 1

[See Myrtaaeidites euaaZyptoides, CFSP 15:106; note that the nomenclatural irregularity denoted in CFSP footnote "1" Cop. cit.) is not corrected by the proposed emendation.]

CATALOG OF FOSSIL SPORES AND POLLEN

44: 169

APPENDIX

INDEX Volume 44 All references to taxa mentioned in Volume 44 are listed in this index, along with all significant citations of geographic and stratigraphic names. For names of taxa covered in this volume, the first page number given is for the page on which reproduction of the information begins, and this number is italicized. Also, for names covered in this volume, the specific and infraspecific epithets are listed independently from the generic names in order to facilitate searches for synonymy.

Aequitriradites ------------------------------------------------ 44:140 AgZaoreidia ---------------------------------------------------- 44:162 aZiquantuZus, AreoZipoZZis --------------------------------------- 44:1 AZnipoZZenites -------------------------------------------------- 44:49 AZnus ----------------------------------------------------------- 44:49 aZsovadasensis, MicrofoveoZatosporis --------------------------- 44:121 A1sovadasz

------------------------------------------------

44:120--123

AmosopoZZis cruciformis ----------------------------------------------- 44:145 diZwynensis ----------------------------------------------- 44:145 anguZocZavatus, IZexpoZZenites ---------------------------------- 44:75 annuZites, MYrica ----------------------------------------------- 44:23 Anomopanax schZechteri ------------------------------------------- 44:1 Anthoceros ------------------------------------------------ 44:105, 118 apicuZata, SaZix ------------------------------------------------ 44:28 AreoZipoZZis ----------------------------------------------------- 44:6 aZiquantuZus ------------------------------------------ 44:1, 7--8 confertus ------------------------------------------------ 44:3, 2 ordinus ----------------------------------------------- 44:5, 2--4 vespiformis ------------------------------------------------- 44:7 Ascarina Zucida ----------------------------------------------------- 44:73 Zucida var. ZanceoZata ------------------------------------- 44:73 ascarinoides, CZavatipoZZenites --------------------------------- 44:73 Asia (western)

-------------------------------------------------

44:144 44:88 44:84 Australia ---------------- 44:25, 33, 35--48, 50--70, 72, 97, 144--146, 148--149, 151, 153--156, 158, 160--164, 166 Austro-Ma1aya --------------------------------------------------- 44:25 Austromyrtus ---------------------------------------------------- 44:47 Aviemore -------------------------------------------- 44:78, 82--83, 94 aviemorensis, LiZiacidites -------------------------------------- 44:78 Bakony Mountains ----------------------------------------------- 44:105 baranyaensis, Phaeocerosporites -------------------------------- 44:104

asperamarginis, TricoZpites ------------------------------------asperatus, Proteacidites ----------------------------------------

44:170

Barringtonia acutangula -------------------------------------------------- 44:9 macrostachya Ｍｾ＠ 44:10 racemosa ------------------------------------------------ 44:9--10 reticulata ------------------------------------------------- 44:10 Barunga -------------------------------------------------------- 44:162 barungensis, Sparganiaceaepollenites --------------------------- 44:162 BeZoperone

----------------------------------------------------

44:5--6

californica ------------------------------------------------- 44:1 bipatterna,Polyporina ------------------------------------------ 44:51 Bogacs ---------------------------------------------------- 44:102--103 Borsod Basin --------------------------------------------------- 44:103 Borsod county --------------------------------------------- 44:102--103 borsodensis, Ephemerisporites --------------------------------- 44:102 Bortonian-Kaiatan --------------------------------------- 44:87, 96--98 Boundary Creek -------------------------------------- 44:80, 85, 88, 92 British Columbia ------------------------------------------.- 44:18--32 Burrungu1e member ---------------------------------------------- 44:150 Calamity Point Ｍｾ 44:74, 76--77, 79, 93, 95 Canacomyrica monticola ------------------------------------------ 44:97 canacomyricoides, Triorites ------------------------------------- 44:97 Canada ---------------------------------------------------.-- 44:18--32 Casuarina --------------------------------------------------- 44:61, 68

Cedrus crispa ----------------------------------------------------- 44:18 parvisaccata ----------------------------------------------. 44:18 perialata -------------------------------------------------- 44:17 Cheb basin ----------------------------------------------------- 44:125 chenopodiaceoides, Polyporina ----------------------------------_ 44:52 Cibotium spp. --------------------------------------------------- 44:33 Cingulatisporites ornatus ---------------------------------------------------- 44:33 problematicus ---------------------------------------------- 44:33 cinnamomites, Osmunda ------------------------------------------- 44:25 Cirratriradites ------------------------------------------------ 44:140 classis, Triorites Ｍｾ 44:69 Clavatipollenites ascarinoides ----------------------------------------------- 44:73 hughesii ----------------------------------------------- 44:73--74 Clerea ---------------------------------------------------- 44:140 C1ifden -------------------.--------- 44:74, 76--77, 79, 89, 93, 95, 99 clifdenensis, Ilexpollenites ------------------------------------ 44:76 clintonensis, Proteacidites -----------------------------------.. 44:147 Clinton Formation ---------------------------------------------- 44:162 concinnus, Marginipollis ----------------------------------------- 44:9 concretus, Proteacidites --------------------------------------- 44:149 confertus, Areolipollis ------------------------------------------ 44:3 co nfrago sus, Proteacidites ------------------------------------- 44:150 convexus, Myrtaceidites eucalyptoides f. ----------------------- 44:169 cooksonii, Tricolporopollenites --------------------------------- 44:64

44:171

Cowra Formation

--------

44:38--39, 44, 46, 51--54, 57--58, 70--71, 169

oowrensis, Retioulatisporites ----------------------------------- 44:57 orassiexinus, Erioipites --------------------------------------- 44:146 Cretaceous, Lower ---------------------------------------------- 44:144 Cserehat -------------------------------------------------- 44:120--123 Cummins Basin --------------------------------------------- 44:146, 156

Cyathea aeneifolia ------------------------------------------------australis Ｍｾ oooperi ---------------------------------------------------leiohhardtiana --------------------------------------------novae-oaledoneae ------------------------------------------paZeospora

porospora

---------------------------------------------

44:36 44:34 44:34 44:34 44:34

44:34, 36

--------------------------------------------------

44:36 44:37 woollsiana ------------------------------------------------- 44:34 Cyathidites ---------------------------------------------------- 44:105 australis ---------------------------------------------- 44:34--35 minor -------------------------------------------------- 44:34--35 Cystopteris fragilis -------------------------------------------- 44:41 Czechoslovakia --------- 44:126, 128, 130--131, 135--137, 139--140, 144 Daorydium ------------------------------------------------------- 44:18 Dannevirke -------------------------------------------------- 44:77, 92 Dannevirke, Lower Ｍｾ＠ 44:84 Dannevirke, Upper ------------------------------------------- 44:87, 97 sp.

--------------------------------------------------------

Deltoidospora diaphana ----------------------------------------------- 44:37--38 granulomargo ----------------------------------------------- 44;3? inoonspioua ------------------------------------------------ 44:38 densifoveatus, Trioolpites -------------------------------------- 44:89 densipunotatus, Trioolpites ------------------------------------- 44:90 Densoisporites perinatus ---------------------------------------- 44:30 Di1wyn Bay

-----------------------------------------------------

dilwynensis, Amosopollis Di1wyn formation

Dodonaea

44:145 44:145 44:154, 156

---------------------------------------

------------------------------------------

aff. visoosa .---------------------------------------------oamfieldii ------------------------------------------------pinnatq ---------------------------------------------------sphaerioa --------------------------------------------------

44:39 44:39 44:39 44:39

Drimys

insipida --------------------------------------------------- 44:40 lanoeolata ------------------------------------------------- 44:40 tetradites ------------------------------------------------- 44:40 Dumortiera hirsuta --------------------------------------------- 44:118 eohinatus, Retioulatisporites ----------------------------------- 44:58 Ediacara

--------------------------------------------------

44:159--160

Embothrium ------------------------------------------------------ 44:86 Enoalyptaesporites --------------------------------------------- 44:101 pliooenious ----------------------------------------------- 44:100

44: 172

EnaaZypta rhabdoaarpa ----------------------------------------------vuZgaris --------------------------------------------------

44:100 44:100 Eocene --------------------------------------------- 44:87, 96, 98, 147 Eocene, Middle --------------- 44:77, 97, 146--147, 149--151, 154--157, 160--161, 163--164 Eocene, Middle-Upper ---------------------------- 44:148, 154, 158--159 Eocene-Oligocene -------------------- 44:48, 50, 55, 60, 62--63, 65, 67 Eocene, Upper -------------------------- 44:97, 146, 157, 162, 165--166 EPhemerisporites Ｍｾ 44:103 borsodensis ----------------------------------------------- 44:102 EPhemerum serratum --------------------------------------------- 44:103

Eriaipites arassiexinus ---------------------------------------------- 44:146 saabratus ------------------------------------------------- 44:146 esobaZteus, PoZyaoZpites ---------------------------------------- 44:83 Ettrick

---------------------------------------------------

44:147, 158 44:169 44:169 Europe --------------------------------------------------------- 44:144 Eyre Peninsula -------------------------------------------- 44:155, 165

euaaZyptoides f. aonvexus, MYrtaaeidites ----------------------EuaaZyptus ----------------------------------------------------Fagus

grandifoZia ------------------------------------------------ 44:19 granuZata -------------------------------------------------- 44:19 syZvatiaa -------------------------------------------------- 44:19 Foraminisporites daiZyi ---------------------------------------- 44:105 Fossombronia --------------------------------------------------- 44:144 f08suZatus, MonoporopoZZenites ---------------------------------- 44:79 foveoZites, Lyaopodium Ｍｾ 44:21 fromensis, Proteaaidites --------------------------------------- 44:152 Gambierina ------------------------------------------------ 44:167--168 edWardsii -------------------------------------------- 44:167--168 gemmatus, TriporopoZlenites ------------------------------------ 44:165 GZeiaheniidites (Toridistalisporis) simpZex ---------------------------------------------- 44:124, 126 toriaonaavus ----------------------------------------- 44:125--126 GOmphrena ------------------------------------------------------ 44:123 Graham Island

-----------------------------------------------

44:18--32

granuZata, Fagus ------------------------------------------------ 44:19 granuZata, PoZyporina ------------------------------------------- 44:53 granuZomargo, DeZtoidospora ------------------------------------- 44:37 granuZomargo, Klukisporites ------------------------------------- 44:42 Gravisporites sphaera ------------------------------------------ 44:104 Great Artesian Basin

----------------------------

44:152--153, 165--166

HeZiaa arguta ----------------------------------------------------aarrii ----------------------------------------------------divariaata ------------------------------------------------insauZpta -------------------------------------------------Helvetian Helvetian, -----------------------------------------------------lower ______________________________________________ _

44:55 44:55 44:55 44:55 44:120 44:121

44:173

Helvetian-Tortonian age ----------- 44:126, 128, 131, 135, 137, 140, 144 Helvetian (upper) ------------ 44:105--106, 108, 111--112, 114--115, 117 Hemitelia ------------------------------------------------------- 44:136 ｨ･ｴｾｰｹｬｩｳＬ＠ Tsuga ------------------------------------------- 44:31 Hicriopteris laevissima ----------------------------------------- 44:125 Hidas ------------------------------------------------------ 44:101, 119 hidasensis, Saxosporites ---------------------------------------- 44:118 hungaricus, Ricciaesporites ------------------------------------- 44:109 Ｎｾｮｧ｡ｲｹ＠ ----------------- 44:101, 103, 106, 108, 112, 115, 117, 119--123 hymenophorus, Svatoplukisporites -------------------------------- 44:138

Hypolepis muelleri -------- - ------------------------------------------punctata ---------------------------------------------------spinysporis ------------------------------------------------tennifolia --------------------------------------------------

44:41 44:41 44:41 44:41 Ikamatua ----------------------------------------------------- 44:87, 96 Ilex ------------------------------------------------------------- 44:75

Ilexpollenites anguloclavatus ------------------------------------------ 44:75, 77 clifdenensis -------------------------------------------- 44:76--77 megagemmatus ------------------------------------------------ 44:77 inargutus, Tricolpites ------------------------------------------- 44:91 inconspicua, Deltoidospora --------------------------------------- 44:38 inspissatus, Tricolpites -------------------------------------- - -- 44:92 ｉｮｴｾｰｵ｣ｯｳｲｩ＠

reticulatus ------------------------------------------------ 44:128 reticuloides ----------------------------------------------- 44:127 Intratriporopollenites magnificans ------------------------------------------------- 44:69 magnificus -------------------------------------------------- 44:69 pilosus ----------------------------------------------------- 44:69 Introlimbatus, Triorites ----------------------------------------- 44:99 irregulites, Osmunda --------------------------------------------- 44:26 Isoglossa -------------------------------------------------------- 44:12 Ivanhoe ------------------ 44:33, 37, 48--50, 55--56, 60, 62--63, 65, 67 Ivanhoe formation -------------------------------- 44 : 35, 38--39, 51, 64 ivanhoensis, Proteacidites --------------------------------------- 44:55 ivanhoensis, Tricotporopollenites -------------------------------- 44:65 Jacobinia --------------------------------------------------------- 44:6 candicans ---------------------------------------------------- 44:3 carnea ------------------------------------------------------- 44:3 Japan ---------------------------------------------------- 44:19, 25, 32

Juglans cinerea ----------------------------------------------------horniana ---------------------------------------------------periporites ------------------------------------------------Juglanspollenites verus -----------------------------------------Justicia ---------------------------------------------------------cay ennens is -------------------------------------------------flava -------------------------------------------------------secunda -----------------------------------------------------Kermadee Islands ________________________________________________ _

44:20 44:20 44:20 44:20 44:6 44:5 44:7 44:7 44:73

44:174

Klukisporites granulomargo ------------------------------------------------ 44:42 lachlanensis ---------------------------------------- ＴＺＳｾ＠ 42, 58 pseudoreticulatus ------------------------------------------- 44:43 Knight formation ------------------------------------------------ 44:150 Knight group ------------------------- 44:48, 50, 55, 60, 62--63, 65, 67 Kopi Anomaly ---------------------------------------------------- 44:149 ｫｯｰｩ･ｮｳｾ＠ Froteacidites ---------------------------------------- 44:154 ｬ｡｣ｨｮ･ｳｩｾ＠ Klukisporites -------------------------------------- 44:43 Lachlan formation ---------------- 44:33--45, 47--48, 51--52, 54, 56, 59, 61--64, 66, 68--69, 71--72, 169 Lachlan River Valley -------------------- 44:33, 35--43, 45--47, 51--54, 56--59, 61, 64, 66, 68--70, 72 Laevigatosporites ovatus ----------------------------------------- 44:41 Lake Cootabar1ow ------------------------------------------- 44:165--166 Lake Eyre -------------------------------------------------- 44:152--153 Lake Torrens ------------------------------------- 44:150--151, 161, 163 Landon Age ----------------------------------------------- 44:75, 86, 91

Liliacidites aviemorensis -----------------------------------------------variegatus -------------------------------------------------waitunaensis ------------------------------------------------

44:?8 44:78 44:78 Long Beach --------------------------------------------------- 44:89, 99 Luma ------------------------------------------------------------- 44:48

Lusatisporis perinatus 44:30 undulosus 44:120 Lycopodium foveolites -------------------------------------------------- 44:21 haleakalae -------------------------------------------------- 44:21 selago ------------------------------------------------------ 44:22 serratum ---------------------------------------------------- 44:22 triZobites ---------------------------------------------- ＴＺＲｾ＠ 21 Macadamia tetraphylla -------------------------------------------- 44:55 ｭ｡ｪｯｲｾ＠ Zlivisporis ---------------------------------------------- 44:141 Malaya

-----------------------------------------------------------

44:25

Marginipollis ---------------------------------------------------- 44:10 concinnus ---------------------------------------------------- 44:9 ｭ･｣ｳｫｮｩｾ＠ Rudolphisporites ----------------------------------- 44:116 Mecsekisporites cerebra lis ------------------------------------------------- 44:130 zengoevarkonyensis f. minor --------------------------- ＴＺＱＲＹｾ＠ 131 zengoevarkonyensis f. minor -------------------------------- 44:129 Mecsek Mountains ------------------------------------------- 44:107, III Mediterranean --------------------------------------------------- 44:144 ｭ･ｧ｡ｴｵｳｾ＠ Ilexpollenites ------------------------------------- 44:77 Mesozoic ----------------------------------------------------- 44:30, 59

Micrantheum ericoides hexandrum spinyspora

44:44 44:44 44:44

44:175

MicrofoveoZatosporis aZsovadasensis --------------------------------------------- 44:121 eZZipsoides ------------------------------------------------ 44:121 minisacca, Tsuga ------------------------------------------------- 44:32 minisporis, ReticuZoidOsporites ---------------------------------- 44:59 minor, Mecsekisporites zengoevarkonyensis f. -------------------- 44:129 minuta, SaZix ---------------------------------------------------- 44:29 minutiscabratus, MonosuZcites ------------------------------------ 44:80 Miocene

----------------------------- 44:73--74, 76--79, 81--83, 89--90, 93--96, 99, 106, 108, 112, 115, 117, 126, 128, 130--131, 135--137, 139--140, 144 44:18--32 Miocene, late Ｍｾ Miocene, Lower Ｍｾ 44:147, 162 miocenicus, Porisporites ---------------------------------------- 44:136 Misko1e ---------------------------------------------------- 44:120--123 "Monoporites" subreticuZata ------------------------------------- 44:162 MonoporopoZZenites fossuZatus ------------------------------------ 44:79

MonosuZcites minutiscabratus --------------------------------------------otagoensis -------------------------------------------------spinosus ---------------------------------------------------waitakiensis ------------------------------------------------

44:80

44:81 44:45 44:81 Mowbray River ---------------------------------------------------- 44:91 Murnpeowie formation ------------------------ 44:152--153, 157, 165--166 Murray basin -------------------------------- 44:48, 50, 55, 60, 62--63, 65, 67, 145, 148, 152, 158 Myd10vary ------------------------- 44:126, 128, 131, 135, 137, 140, 144 Myd10vary beds -------------------- 44:126, 128, 131, 135, 137, 140, 144 Myrica annuZites ------------------------------------------------- 44:23

Myricipites coryphaeus -------------------------------------------------- 44:23 dub ius ------------------------------------------------------ 44:23 Myrtaceidi tes eucaZyptoides f. convexus ---------------------------------- 44:169 protrudiporens ---------------------------------------------- 44:46 rhodamnoides -------------------------------------------- 44:47--48 xanthomyrtoide8 ----------------------------------------- 44:48, 47 Neogene ---------------------------------------------------- 44:120--123 neogenicus, NummuZipoZZis ---------------------------------------- 44:11 New Caledonia

-.-----------------------------------------------

44:75, 97

New South Wales ------------------------------ 44:33, 35--48, 50--70, 72 New Zealand -------------------------- 44:39--40, 73--80, 82--83, 85--99 Nigeria -------------------------------------- 44:2, 4, 6, 8, 10, 13, 16 nigericus, PeregrinipoZZis --------------------------------------- 44:14 North America ------------------------------------------- 44:18, 30, 144 North Maslin sands ---------------------------------------------- 44:150 Nothofagus --------------------------------------------------- 44:56, 62 spp. -------------------------------------------------------- 44:63 NummuZipoZZis ---------------------------------------------------- 44:13 neogenicus ---------------------------------------------- 44:11, 13

44: 176

ｾｰｍｲ＠

44:24 44:24 44:49 ｯ｢ｚ｡ｴｵｳｾ＠ 44:61 ｯ｢ｚ｡ｴｕＸｾ＠ Oligocene ------------------------------------------- 44:18, 75, 86, 91 Oligocene, Middle ----------------------------------------------- 44:40 ｯｲ､ｩｮｵｳｾ＠ AreoZipoZZis -------------------------------------------- 44:5

fructicans ------------------------------------------------spicuZites ------------------------------------------------PoZyorificites ----------------------------------------StephanocoZpites ---------------------------------------

Orites exceZsa ---------------------------------------------------ZancifoZia ------------------------------------------------ｯｲｮ｡ｴｵｳｾ＠ CinguZatisporites -------------------------------------Osmunda cinnamomea ------------------------------------------------cinnamomites ----------------------------------------------irreguZites -----------------------------------------------regaZis ---------------------------------------------------regaZites --------------------------------------------------

44:55 44:55 44:33

44:25 44:25 44:26 44:27 44:27 Otway Basin ---------------------------------------------------- 44:145 Oxymitra ------------------------------------------------------- 44:144 Paleocene -------------------- 44:80, 84--85, 88, 92, 96, 145, 152--153 ｰ｡ｚ･ｯｳｲｾ＠ cyathea --------------------------------------------- 44:34 Pannonian ------------------------------------------------------ 44:123 Pannonian, lower ------------------------------------------ 44:102--103 Pannonian (upper) ----------------------------------------- 44:101, 119 Pareora Age ------------- 44:73--74, 76--79, 81--83, 89--90, 93--96, 99 Paris Basin ---------------------------------------------------- 44:165 Pebble Point formation ----------------------------------------- 44:145 PeregrinipoZZis ------------------------------------------------- 44:16 nigepicuB ---------------------------------------------- 44:14, 16 ｰ･ｲｩ｡ｚｴｾ＠ Cedrus ----------------------------------------------- 44:17 ｰ･ｲｩｭ｡ｧｮｴｵｳｾ＠ TricoZpites ------------------------------------- 44:93

PerimonoZetes goersbachensis -------------------------------------------- 44:122 spicatus -------------------------------------------------- 44:122 ｰ･ｲｩｯｴｳｾ＠ JugZans -------------------------------------------- 44:20 PerotriZetes --------------------------------------------------- 44:140 PerotriZites ---------------------------------------------------- 44:30 Phaeoceros buZbicuZosus ---------------------------------------- 44:105 PMeocerosporites ---------------------------------------------- 44:106 baranyaensis ------------------------------------ ＴＺＱＰｾ＠ 107--108 transversus ----------------------------------------------- 44:107 Pirie-Torrens Basin

-----------------------------

44:151, 160--161, 163

PZagiochasma --------------------------------------------------- 44:140 PZanchonia crenata ---------------------------------------------------- 44:10 spectabiZis ------------------------------------------------- 44:9 Pleistocene -------------------------------- 44:46, 53, 57--58, 70, 169 Pleistocene, Lower ---------------------------------------------- 44:39 ｰｚ･ｩｳｴｯ｣ｮｵｾ＠ TubuZiJZoridites -------------------------------- 44:70 Pliocene ------------------------------ 44:101, 103--104, 119, 144, 169

44:177

Pliocene, early --------------------------------------------- 44:-18--32 Pliocene (? Upper) ----------------- 44:33, 35--43, 45, 47, 51--52, 54, 56, 59, 61, 64, 66, 68--69, 72 ｰｚｩｯ｣･ｮｵｳｾ＠ EncaZyptaesporites -------------------------------- 44:100 Poe1pena formation -------------------- 44:149--150, 154--155, 157, 164 Po1da Basin ------------------------------------------ 44:154--155, 164 PoZZenites ｣ｯｾｰｨ｡･ｵｳ＠ ------------------------------------------- 44:23 PoZycoZpites esobaZteus ----------------------------------------- 44:83 PoZyorificites -------------------------------------------------- 44:50 obZatus ---------------------------------------------------- 44:49 PoZypodiaceoisporites variabiZis -------------------------- ＴＺＱＳＲｾ＠ 134 PoZypodiidites arcus -------------------------------------------- 44:59 PoZypopodiaceoisporites trianguZus ----------------------------- 44:135

PoZyporina bipatterna --------------------------------------------- 44:51--53 chenopodiaceoides -------------------------------------- 44:52--53 fragiZis ----------------------------------------------- 44:51--53 granuZata ---------------------------------------------- ＴＺＵＳｾ＠ 52 reticuZatus ------------------------------------------------ 44:54 PoZyporopoZZenites ---------------------------------------------- 44:49 Porisporites --------------------------------------------------- 44:137 miocenicus ------------------------------------------------ 44:136 ｰｯｲｳ｡ｾ＠ Cyathea ---------------------------------------------- 44:36 Poyntz Bore ------------------------------------------ 44:147--148, 158 Prince town member ----------------------------------------- 44:154, 156

Proteacidites asperatus -------------------------------------------------- 44:84 cZintonensis ------------------------------------ ＴＺＱＷｾ＠ 159--160 concretus -------------------------------------------- ＴＺＱＹｾ＠ 156 co nfrago sus ------------------------------------------ ＴＺＱＵＰｾ＠ 163 fromensis Ｍｾ 44:152 granoratus ------------------------------------------------ 44:152 incurvatus --------------------------------- 44:147--148, 159--160 ivanhoensis ------------------------------------------------ 44:55 kopiensis ------------------------------------------------- 44:154 Zatrobensis ------------------------------------------ 44:149, 156 obscupus Ｍｾ 44:157--158 rectomarginis -------------------------------- 44:84--85, 147--148 reticuZatus Ｍｾ 44:157--158 simiZis ---------------------------------------------------- 44:55 spiniferus ------------------------------------------------- 44:86

ｰｳ･ｵ､ｯｴｲｩ｡ｾ＠

spp.

ｰｲｯｴｵ､ｩ･ｮｳｾ＠

------------------------------------------------------

44:154

symphyonemoides Ｍｾ 44:157--158 tortuosus ------------------------------------------------- 44:155 tripartitus ----------------------------------------------- 44:156 tubercuZatus ---------------------------------------------- 44:155 varius ------------------------------------------ ＴＺＱＵＷｾ＠ 159--160 wiZkatanaensis -------------------------------------------- 44:159 MYrtaceidites ----------------------------------- 44:46 TricoZporopoZZenites ---------------------------- 44:95

44:178

Pseudowintera axillaris

--------------------------------------------------

44:40 44:40 ｰｳｩｬ｡ｴｯｲｾ＠ Quintinia ----------------------------------------- 44:56 ｰｳｩｬ｡ｴｵｾ＠ Tricolpites ------------------------------------------- 44:62 ｰｳｩｬ｡ｴｵｾ＠ "Triorites" ------------------------------------------ 44:164 pterocarya ------------------------------------------------------ 44:49 Pterocaryapollenites -------------------------------------------- 44:49 ｰｵｮ｣ｴ｡ｩｬｳｾ＠ Tricolpites -------------------------------------- 44:94 Punctatosporites ellipsoideus ---------------------------------- 44:121 Quaternary ------------------------------------------------------ 44:35 Queen Charlotte Islands ------------------------------------- 44:18--32 Queensland ----------------------------------------------------- 44:145 sp.

--------------------------------------------------------

Quintinia psilatispora

-------------------------------------------

ＴＺＵＶｾ＠

62 44:56 ｲ･ｧ｡ｬｩｴｳｾ＠ Osmunda ---------------------------------------------- 44:27 Renmark beds ----------------------------------------- 44:147--148, 158 Renmark formation ---------------------------------------------- 44:150 sp.

--------------------------------------------------------

Reticulatisporites cowrensis ---------------------------------------------- 44:57--58 echinatus -------------------------------------------------- 44:58 pudens ----------------------------------------------------- 44:57 ｲ･ｴｩ｣ｵｬ｡ｳｾ＠ Polyporina ----------------------------------------- 44:54 ｲ･ｴｩ｣ｵｬｯ､ｳｾ＠ Intrapunctosporis -------------------------------- 44:127 Reticuloidosporites arcus ------------------------------------------------------ 44:59 minisporis ------------------------------------------------- 44:59 ｲ･ｴｩｦｯｭｳｾ＠ Tricolpites ----------------------------------------- 44:63 Rhodamnia ------------------------------------------------------- 44:47 argenta ---------------------------------------------------- 44:47 ｲｨｯ､｡ｭｮｩ･ｳｾ＠ ｉｾｹｲｴ｡｣･ｩ､ｳ＠ ------------------------------------- 44:47 ｲｨｯｭ｢ｩ､｡ｬｦｾｳ＠ Tricolporopollenites ------------------------- 44:96 Rhytiglossa lactea ---------------------------------------------- 44:12 Riccia ---------------------------------------------------- 44:140, 144 breidleri ------------------------------------------------- 44:111 duplex ----------------------------------------------- 44:111, 114 nigrella -------------------------------------------------- 44:111 sorocarpa ------------------------------------------------- 44:111 Ricciaesporites ------------------------------------------------ 44:112 Ill, 114--115, 144 hungaricus ---------------------------- ＴＺＱＰＹｾ＠ transdanubicuB ---------------------------------- 44:113, lIS, 144 Riella --------------------------------------------------------- 44:140 ｲｯ｢ｵｳｴｩｰｾ＠ Sparganiaceaepollenites -------------------------- 44:60 ｲｯｴｵｮ､ｳｾ＠ Sapotaceoidaepollenites ------------------------------ 44:161 Rudolphisporites mecsekensis ----------------------------------------------- 44:116 rudolphi -------------------------------------------------- 44:116 Salix albaeformis ------------------------------------------------ 44:28 apiculata -------------------------------------------------- 44:28 minuta ----------------------------------------------------- 44:29

44:179

SapotaceoidaepoZZenites potundus ------------------------------- 44:161 Sarmatian -----------------------------------------------------SaxospoPites hidasensis _______________________________________ _ 44:122 Scabpatus, SimpZicepoZZis SeZagineZZa opegona sinuites waZZacei SeZagospoPis fsp. A.'

---------------------------------------

----------------------------------------------------

44:118 44:87 44:30

44:30 44:30 44:21

seZagoides ------------------------------------------------- 44:22 Shepherds Creek ------------------------------------------------- 44:90 Siber.ia -------------------------------------------------------- 44:144 simiZis, TpicoZpopopoZZenites ----------------------------------- 44:66 simpZex, GZeicheniidites (TopidistaZispopis) ------------------- 44:124 SimpZicepoZZis ｭ･ｰｩ､ｾ｡ｮｵｳ＠ ------------------------------------------------- 44:87 scabPatus -------------------------------------------------- 44:87 simpZis, TubuZifZopidites --------------------------------------- 44:71 sinuites, SeZagineZZa ------------------------------------------- 44:30 Skonun -------------------------------------- 44:18, 21--22, 24, 26, 30 Snow River -------------------------------------------------- 44:75, 86 sooiana, VacZavipoZZis ----------------------------------------- 44:123 South America -------------------------------------------------- 44:144 southern Bohemia ----------------- 44:126, 128, 131, 135, 137, 140, 144

spapganiaceaepoZZenites bapungensis ------------------------------------------- 44:162, 60 magnoides ------------------------------------------------- 44:162 pebustispopis ---------------------------------------------- 44:60 spapganium ----------------------------------------------------- 44:162 sphaePica, Dodonaea --------------------------------------------- 44:39 spicatus, PepinomonoZetes -------------------------------------- 44:122 spicuZites, Nuphap ---------------------------------------------- 44:24 spinifepus, PPOteacidites --------------------------------------- 44:86 spinosus, MonosuZcites ------------------------------------------ 44:45 spinyspopa, Micpantheum ----------------------------------------- 44:44 spinyspoPis, HypoZepis ------------------------------------------ 44:41 StephanocoZpites costatus --------------------------------------------------- 44:61 obZatus ---------------------------------------------------- 44:61 StephanopopopoZZenites ------------------------------------------ 44:49 St. Vincent Basin ---------------------------------------------- 44:162 StyxispoPites -------------------------------------------------. 44:140 substPaitus, TpicoZpopopoZZenites ------------------------------- 44:67 SvatopZukispopites --------------------------------------------- 44:140 hymenophopus ---------------------------------------------- 44:138 Tertiary ------------------- 44:33, 35--43, 45, 47--48, 50--52, 54-.56, 59--69, 72, 75, 145--146, 148--149, 151, 153--156, 158, 160--164, 166 Tertiary, Late -------------------------- 44:2, 4, 6, 8, 10, 13, 18--32 Tertiary, Lower -----------------------------------------------.- 44:97 Tertiary, Upper ---------------------------------------- 44:15--16, 162

44:180

44:40 92, 96 44:41 44:124 44: 105 44:155 ｴｲ｡ｮｳ､ｵ｢ｩ｣ｾ＠ Ricciaesporites -------------------------------- 44: 113 ｴｲ｡ｮｳｶ･ｬｩｾ＠ Tricolporopollenites ----------------------------- 44:68 transversus, Phaeocerosporites --------------------------------- 44: 107 Tricolpites asperamarginis --------------------------------------------- 44:88 delicatulus ------------------------------------------------ 44:63 densifoveatus ---------------------------------------------- 44:89 densipunctatus --------------------------------------------- 44:90 fissilis --------------------------------------------------- 44:62 inargutus -------------------------------------------------- 44:91 inspissatus ------------------------------------------------ 44:92 matauraensis ----------------------------------------------- 44:68 pachyexinus ------------------------------------------------ 44:62 perimarginatus --------------------------------------------- 44:93 psiZatus ----------------------------------------------- 44:62, 56 punctaticulus ---------------------------------------------- 44:94 retiformis --------------------------------------------- 44:63--64 striatus' ----------------------------------------------- 44:67,95 Tricolporites ｭｩ｣ｲｯ･ｾｵｬ｡ｴｳ＠ ----------------------------------- 44:64--65, 68 valatus --------------------------------------------------- 44:163 Tricolporopollenites cooksonii ---------------------------------------------- 44:64--65 ivanhoensis ---------------------------------------- 44:65, 64, 66 latizonatus ----------------------------------------------- 44:161 pseudbstriatus ----------------------------------------- 44:95, 67 rhomboidaliformis ------------------------------------------ 44:96

ｴ･ｲ｡､ｩｳｾ＠ Drimys ---------------------------------------------Teurian Age --------------------------------- 44:80, 84--85, 88, Thelypteris leucolepis -----------------------------------------(Toridistalisporis) ｳｩｭｰｬ･ｸｾ＠ Gleicheniidites ------------------Tortonian Proteacitites -----------------------------------------------------______________________________________ _ ｴｯｲｵｳｾ＠

simiZis ------------------------------------------------ 44:66, 65 substraitus ----------------------------------------------44:67 transversaZis

------------------------------------------

44:68, 66

Trilites lachlanae ---------------------------------------- 44:104, 106 trilobites, Lycopodium -------------- .--------------------------- 44:22 Triorites canacomyricoides ------------------------------------------- 44:97 classis ---------------------------------------------------- 44:69 clavatus --------------------------------------------- 44:167--168 edwardsii -------------------------------------------- 44:167--168 harrisii ----------------------------------------------- 44:97--98 introlimbatus ---------------------------------------------- 44:99 magnificus ---------------------------- 44:147--148, 154, 167--168 scabratus ------------------------------------------------- 44:164 "Triorites" psilatus ------------------------------------------- 44:164 tripartitus, Proteacidites ------------------------------------- 44:156 Triplanosporites ------------------------------------------------ 44:38

44:181

TriporopoZZenites buZZis ---------------------------------------------------- 44:165 44:165 gemmatus Tsuga heterophyZZa ----------------------------------------------- 44:31 heterophyZZites ---------------------------------------- 44:31--32 minisaaaa -------------------------------------------------- 44:32 sieboZdii -------------------------------------------------- 44:32 TubuZij70ridites antipodiaa --------------------------------------------- 44:70--72 pZeistoaeniaus ----------------------------------------- 44:70--72 simpZis ------------------------------------------------ ＴＺＷＱｾ＠ 70 Typha sp. ------------------------------------------------------ 44:162 UZmipoZZenites -------------------------------------------------- 44:49 ｵｮ､ｚｯｳｾ＠

Ulmus

-----------------------------------------------------------

44:49 44:120 United States --------------------------------------------------- 44:18 U.S.S.R. -------------------------------------------------------- 44:18

Lusatisporis

----------------------------------------

VaaZavipoZZis paaZtovae ------------------------------------------------- 44:123 sooiana --------------------------------------------------- 44:123 ｶ｡ｚｴｵｳｾ＠ TriaoZporites ----------------------------------------- 44:163 ｶ｡ｲｩ｢ｚＸｾ＠ PoZypodiaaeoisporites ------------------------------ 44:132 ｶ｡ｲｩｵｳｾ＠ Proteaaidites ------------------------------------------ 44:157 Verruaosisporites grumosus -------------------------------- 44:100--101 ｶ･ｳｰｩｦｯｲｭｾ＠ AreoZipoZZis ---------------------------------------- 44:7 44:145 44:81 Waitaki River --------------------------------------------------- 44:90 Wani11a formation ------------------------------- 44:146, 150, 155--156 Weinmannia ------------------------------------------------------ 44:92 raaemosa --------------------------------------------------- 44:94 syZviaoZa -------------------------------------------------- 44:94 ｷｩｚｫ｡ｴｮ･ｳｾ＠ Proteaaidites ---------------------------------- 44:159 "Wi1katana formation" ---------------------- 44:150--151, 159--161, 163 ｸ｡ｮｴｨｯｭｹｲｩ､･ｳｾ＠ MYrtaaeidites ---------------------------------- 44:48 Xanthomyrtus ---------------------------------------------------- 44:48 ｺ･ｮｧｯｶ｡ｲｫｹｳｩｾ＠ Mea8eki8porites f. minor ------------------- 44:129 Zengovarkony ---------------- 44:105--106, 108, 111--112, 114--115, 117 ｷ｡ｩｴｫ･ｮｳｾ＠

Victoria

-------------------------------------------------------

MonosuZaites

--------------------------------------

ZZivisporis hungariaus -----------------------------------------------major ------------------------------------------------ ＴＺＱｾ＠ ｴｾｮｳ､｡ｵ｢ｩ＠ --------------------------------------------

44:144 143 44:144

44:182