Das Tierreich / The Animal Kingdom. Tlbd/Part 110 World Catalogue of Odonata I: Zygoptera 9783110824438, 9783110149333

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Henrik Steinmann World Catalogue of Odonata Volume I Zygoptera

DAS TIERREICH The Animal Kingdom Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen A Compilation and Characterization of the Recent Animal Groups Πάντα φεΐ

Sine systemate chaos Herausgeber/Editors

HEINZ WERMUTH

MAXIMILIAN FISCHER

(Wirbeltiere)

(Wirbellose)

Teilband

1 1 0

Part

W DE G Walter de Gruyter · Berlin · New York 1997

Henrik Steinmann

World Catalogue of Odonata Volume I Zygoptera

w DE

G Walter de Gruyter · Berlin · New York 1997

Author Dr. Henrik Steinmann Ä r a m u. 25 Η-1193 Budapest Hungary Scientific and Managing

Editor

Hofrat Mag. Dr. Maximilian Fischer Naturhistorisches Museum Wien

Citation Das Tierreich, Teilband 110 Henrik Steinmann Verlag Walter de Gruyter, Berlin · New York 1997

A-1014 Wien Austria

© printed on acid free paper which falls within the guidelines of the A N S I to ensure permanence and durability.

Library of Congress Cataloging-in-Publication

Data

Steinmann, Henrik. World catalogue of O d o n a t a / Henrik Steinmann, (Das Tierreich ; Teilbd. 100-111 = The Animal kingdom ; pt. 110-111) Includes bibliographical references (v. 2) and indexes. Contents: v. 1. Zygoptera v. 2. Anisoptera. ISBN 3-11-014933-8 (alk. paper). - ISBN 3-11-014934-6 (alk. paper) 1.Damselflies, 2.Dragonflies. I.Titel. II.Series. Tierreich ; Teilbd. 110-111. QL520.S75 1997 595.7'33-dc21 97-10528 CIP Die Deutsche Bibliothek - Cataloging-in-Publication

Data

Das Tierreich : eine Zusammenstellung und Kennzeichnung der rezenten Tierformen = The animal kingdom / Hrsg. Heinz Wermuth ; Maximilian Fischer. - Berlin ; New York : de Gruyter Teilw. hrsg. von Robert Mertens ; Willi Hennig. - Teilw. hrsg. von F. E. Schulze ... - Teilw. fortges. von R. Hesse Teilbd. 110. World catalogue of Odonata. - Vol. I. Zygoptera / Henrik Steinmann. - 1997 ISBN 3-11-014933-8 Gb.

© Copyright 1997 by Walter de Gruyter & Co., D-10785 Berlin. All rights reserved, including those of translation into foreign languages. N o part of this book may be reproduced or transmitted in any f o r m or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system, without permission in writing from the publisher. - Printed in Germany. Typesetting and Printing: Tutte Druckerei G m b H , Salzweg-Passau. Binding: Dieter Mikolai. Berlin.

Contents Introduction

VII

Systematic Index

XV

Catalogue Odonata Zygoptera Calopterygoidea Amphipterygidae Calopterygidae Chlorocyphidae Dicteriastidae Diphlebiidae Euphaeidae Polythoridae Rimanellidae Lestinoidea Chlorolestidae Lestidae Lestoideidae Megapodagrionidae Perilestidae Pseudolestidae Hemiphlebioidea Hemiphlebiidae Coenagrionoidea Coenagrionidae Isostictidae Platycnemididae Platystictidae Protoneuridae Pseudostigmatidae

1 1 2 3 3 6 44 74 75 77 89 100 101 101 107 137 138 171 174 176 176 177 177 363 368 398 417 455

Alphabetical Register

463

Introduction The world literature of dragonflies including a large number of taxonomic descriptions, by today, has become a source of many consternations. Most taxonomists regard the group as almost unmanagable. Systematic, classificatory and taxonomic research are also greatly hindered by the large number of technical papers appearing in various journals all over the world which, with the rising tendency of prices today, become unattainable for many. The number of known and described taxa has risen sharply which again forces one to restrict oneself to a particular zoogeographical or systematic unit. Consequently, the intensively studied groups are comparatively well known, while the others are almost entirely neglected. This tendency is clearly reflected by the generic revisions or by special studies dealing with a particular geographic region that were published in the past decades. So time has arrived when, after several years of compilatory work, the publication of a literature revision is unavoidable. My principal aim was to make an inventory of all the valid and invalid taxa of Odonata. This work meant the collection of world literature, the checking of data, the cross-checking of all available information and where necessary their elucidation. I laid special stress on the systematic references of the classical taxa and on any changes that occurred to them in the last two centuries. In order to fulfil this task in the great majority of the cases I looked up the original descriptions of the taxa, most of which were published in the 19th century. My fundamental time lapse falls between 1758 and 1930s. I greatly value the original systematic divisions (subgenera, species, subspecies, races, forms) of early authors (Selys Longchamps, Hagen, Brauer and others), since most of these data in subsequent papers either do not appear, or if so, they are incomplete or heavily charged with errors. The higher categories of early times were rather unreliable in contrast with the well-defined systematic ranging of infraspecific taxa. The suprageneric system of the group was elaborated much later, mostly in the works of Tillyard, Fräser, Lieftinck, Pinhey and others. In my work I adopted Davies & Tobin's schematic list without claiming to strictly adhere to a pure phylogenetical system. Consequently, I avoided to make decisions especially in yet unsettled problems, thus, the following questions have been left unanswered:

VIII - The Davies & Tobin's list frequently has species names under a genus with a remark "including" certain species, which obviously belong elsewhere. When revising the literature I found that most of the species names have already been clarified and are synonyms. In cases where no decisions had been made, it was reasonable to keep the species group together until a thorough taxonomic revision. - Selys Longchamps heavily relied on subgeneric ranging when describing a new taxon, and when having infraspecific doubts he simply described new subspecies, races and forms. Later, specialists partly rejected this system, and in his catalogue, Kirby, as early as in 1890, elevated Selys's subgenera to generic level. Today, any subgeneric classification is rather heterogeneous owing to the lack of a reliable set of characteristics either above or below the generic level. Present authors do not use subgeneric categories, or if so, they are rather rare. Consequently, I left out all such categories, of course, in the literature citations I carefully made reference to any subgeneric relationship. - Following the original description of a taxon I give the place of type deposition, but only in such case, if at least two literure references support this fact. In early descriptions frequently no holotypes were designated from the type-series. Or in other cases, the specimens were all marked with identical labels. Thus, it happened that such specimens finding their ways into various other collections started to live their independent life, and were considered in many a time as the holotype of that species. This problem can, of course, be solved only after type revisions, and not whithin the framework of the present catalogue. My work lists the suprageneric taxa according to a systematic order, while the lower categories (generic and infrageneric) in simple order of the alphabet. (The short descriptions follow Davies & Tobin). The valid taxa always begin in a new line, within each entry the original names, either valid or invalid, are set in bold face type. The line of the independent valid taxon is followed by the literature citation beginning with the year, the name of the taxon and the source of bibliography. In the case of original descriptions the sex of the type, the place of deposition and the type-locality are also given. Reference to distribution has its own entry.

A supraspecific example: Genus Devadatta

KIRBY

1859 Tetraneura SELYS, Bull. Acad. r. Belg., (2) 7: 449. - Type-species: Tetraneura argyroides SELYS, 1859 (monotypy). - 1890 Devadatta KIRBY, Syn. Cat. Neur.-Odon., London: 111 (replaced name for Tetraneura Selys, 1859). - etc.

IX A specific example: argyroides argyroides (SELYS) 1859 Tetraneura argyroides SELYS, Synopsis des Calopterygines, Add. 1: 15. Type male: Selys's Collection; type-locality: Singapore, etc.

Distribution: Malaysia.

A secondary citation: subapicalis KENNEDY 1984 Miocora subapicalis KENNEDY, 1940 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25.

Distribution: Colombia.

In the entry of the valid taxon one finds in chronological order the selected literature citations which reflect the history of the taxon, i.e. all important events. Thus, e. g. any change having taken place in its ranging or status, a new combination, redescription, recent new geographical distribution, the description of the other sex, etc. Furthermore, all those references which are important for a thorough systematic study of the taxon. Like, e. g. articles discussing a large geographical area, a big taxonomic unit, the material of significant collections or expeditions, revisionary papers, check-lists, monographs with critical analysis.

Table of the personal collections in new depositions and abbreviated localities of the citations H.J. ADAMS - British Museum (Nat. Hist.), London Agassiz Mus. - Dept. Ent. Agassiz Mus., Cambridge (Mass.) Agra Univ. - College Mus. Univ. Agra Albany Mus. - New York State Mus., Albany Alma-Ata Mus. - Zool. Inst. Acad. Sei., Alma-Ata Amsterdam Mus. - Inst. Tax. Zoöl., Amsterdam T.N. ANNANDALE - Dept. Zool. Indian Mus., Calcutta Ann Arbor Mus. - Zool. Mus. Univ. Mich., Ann Arbor A. ASAHINA - private collection, Tokyo W. S. ATKINSON - British Museum (Nat. Hist.), London E. D. AUSTEN - British Museum (Nat. Hist.), London N. BANKS - Mus. Comp. Zool. Cambridge (Mass.) Barcelona Mus. - Mus. Cienc. Nat., Barcelona G.S. BATES - Acad. Nat. Sei., Philadelphia T. BAYERN - Mus. Naturk. Berlin J. BELLE - Hope Mus., Oxford and Florida State College, Gainesville B. F. BELYSHEV - private collection, Bijsk, Altai C.H. BESCHK - Mus. Naturk. Berlin

χ G.H. BICK & J . C . BICK - Biol. Dept. Southwestern Louisiana Inst., Lafayette L. BIRO - Termeszett. Μύζ., Budapest Bishop Mus. - P.B. Bishop Mus., Honolulu Bocage Mus. - Dept. Zool. Mus., Bocage Bonn Mus. - Zool. Forschinst. Mus., Bonn D.J. BORROR - Ohio State Univ., Columbus Boston Mus. - Nat. Hist. Mus. Soc., Boston E.L. BOUVIER - Mus. Nation. Hist. Nat., Paris Brüssel Mus. - R. Mus., Bruxelles Buenos Aires Mus. - Mus. Argent. Cienc. Nat. "Bernardino Rivadaria", Buenos Aires Buitenzorg Mus. - Zool. Mus., Buitenzorg after 1954 all types in Rijksmus. natuurl. Hist., Leiden L.A. BULLA - locality unknown T.H. BUTLER - Pac. Biol. Station, Nanaimo (Canada) P. P. CALVERT - Acad. Nat. Sei. Philadelphia R. CAMMAERT - locality unknown H. CAMPION - British Museum (Nat. Hist.), London Caracas Mus. - Cienc. Nat. Mus., Caracas F. L. CARLE - destroyed Carnegie Mus. - Carnegie Nat. Hist. Mus., Pittsburgh H . - F U CHAO - Fujian Agr. Univ. Zool. Inst., Foochow T. CHARPENTIER - Mus. Naturk., Berlin and Zool. Mus., Kaliningrad S.H. CHEN - Acad. Sei. Mus. Sin., Peiping Chicago Mus. - Field Mus. Nat. Hist., Chicago M. C H U J O - Lab. Ent. Fac. Liberal Arts, Kawaga Univ., Shikoku K.A. CHRISTIANSEN - Dept. Biol. Grinnel College, Grinnel Colombo Mus. - Nat. Hist. Mus., Colombo Cornell Univ. - Dept. Ent. Cornell Univ., Ithaca Coryndon Mus. - Dept. Zool. Coryndon Memorial Mus., Nairobi V.A.E. DAECKE - Bur. Plant. Inst. Univ. Mus., Harrisburgh Dahlem Mus. - Dtsch. Ent. Inst., Berlin-Dahlem (later Eberswalde) C.W. DALE - Hope Mus., Oxford Darjeeling Mus. - Nat. Hist. Soc. Mus., Darjeeling Darmstadt Mus. - Hess. Landesmus., Darmstadt P. DEAM - locality unknown PF. DEJEAN - R. Mus., Bruxelles H. C. DELSMAN - private collection, The Hague H. D O H R N - Hist. Nat. Mus. Stettin, after 1945 in PAN, Zool. Inst., Warszawa T.W. DONELLY - Florida State College, Gainesville, and U.S. natn. Mus., Washington C.F. DREGE - Insektenhändler, Hamburg

XI Dresden Mus - Mus. Tierk., Dresden Dublin Mus. - Nat. Hist. Mus. Irel., Dublin C. DUMONT - Mus. Nation. Hist. Nat., Paris H. ESCHER - Ent. Inst. Techn. Hochsch., Zürich J.C. FABRICIUS - Zool. Mus., Kiel A. FITCHE - New York State Mus., Albany E . L . FONSCOLOMBE - Mus. Nation. Hist. Nat., Paris F. FÖRSTER - Zool. Mus. Univ. Mich., Ann Arbor and Mus. Naturk., Berlin Frankfurt/Main Mus. - Senckenberg Mus. Nat. Hist., Frankfurt am Main F. C . FRÄSER - British Museum (Nat. Hist.), London H. FRÜHSTORFER - Insektenhändler, Berlin Fujian Univ. - Fujian Agr. Univ., Foochow R.M. GABMLES - unknown locality Α. Ε. GARDNER - private collection, Banstead, Surrey GARRISON'S Coll. - destroyed C . GAUTIER - R. Mus., Bruxelles D.C. GEIJSKES - Surinams Mus., Paramaribo Ε . F. GERMAR - Zool. Mus., Halle J.C. GODEFFROY - Zool. Inst. Mus. Univ., Hamburg S. O. GODMAN - British Museum (Nat. Hist.), London M.P. GRAELLS - Mus. Cienc. Nat., Madrid C. GREGORY - British Museum (Nat. Hist.), London H . HAGEN - Mus. Comp. Zool. Cambridge (Mass.) H . HÄMÄLÄINEN - Dept. Agr. & For. Zool. Univ., Helsinki Halle Mus. - Zool. Mus., Halle Hamburg Mus. - Zool. Inst. Mus. Univ., Hamburg H.C. H A R T - British Museum (Nat. Hist.), London Harward Mus. - Zool. Inst. Mus. Univ., Harward A.C. HAUXWELL - R. Mus., Bruxelles A. HEMPEL - Inst. Biol. Univ., Säo Paulo Heude Mus. - Nat. Hist. Heude Mus., Shanghai Helsigfors Mus. - Zool. Mus. Univ., Helsinki J.S. HINE - Mus. State Univ., Columbus Hobard Mus. - Zool. Inst. Hobard Mus., Tasmania Hokkaido Univ. - Zool. Inst. & Mus. Univ., Hokkaido J.C. HOFFMANNSEGG - Mus. Tierk., Dresden Hope Mus. - Dept. Zool. Hope Univ., Oxford Indian Mus. - Dept. Zool. Indian Mus., Calcutta S.ISHIDA - unknown locality E . R . JACOBSON - Rijksmus. natuurl. Hist., Leiden F. A . KARSCH - Mus. Naturk., Berlin Kasan Mus. - Zool. Inst. Univ., Kasan

XII C.H. KENNEDY - Zool. Mus. Univ. Mich., Ann Arbor C.H. KING - unknown locality W. F. KIRBY - British Museum (Nat. Hist.), London M. KOBAYASHI - Dept. Zool. Nat. Sei. Mus., Tokyo Kobenhavn Mus. - Univ. Zool. Mus., Kobenhavn E.J. K0RMONDY - Dept. Zool. Oberlin College, Oberlin, Ohio Königsberg Mus. - Zool. Mus., Kaliningrad Kyushu Univ. - Zool. Inst. Univ., Kyushu F. F. LAIDLAW - British Museum (Nat. Hist.), London and Nation Coll. Zool. surv. India, Calcutta P . A . LATREILLE - R . M u s . , Bruxelles

Leeds Mus. - Leeds City Mus., Leeds J.J. LEGRAND - Fac. Sei. Univ. Mus., Poiters Leningrad Mus., - Zool. Inst. Acad. Sei., St. Petersburgh J.W. LEONARD - Zool. Mus. Univ. Mich., Ann Arbor M . A . LIEFTINCK - Zool. Mus. Buitenzorg and Rijksmus. natuurl. Hist., Leiden Liege Mus. - Inst. Ed. Van Beneden Univ., Liege P.L. VAN DER LINDEN - Mus. Naturk., Berlin J. F. LINDLEY - locality unknown C. LINNE - Zool. Univ. Mus., Uppsala (and all) C. E. LONGFIELD - British Museum (Nat. Hist.), London Lund Mus. - Zool. Inst. Mus. Lunds Univ., Lund A . B . M . MACHADO - private collection, Belo Horizonte, Minas Gerais Madrid Mus. - Cienc. Nat., Madrid M.S. MANIS & G.S. AMSARI - Nation. Coll. Zool. surv. India, Calcutta P. MARCHAL - Hope Mus., Oxford J. DE MARMELS - Inst. Zool. Agr. Fac. Agron. Univ. Venezuela, Caracas R. MARTIN - Riksmus. Nat. Hist., Stockholm S. MATSUMURA - Univ. Zool. Mus., Sapporo M . L . MAY - Dept. Ent. & Econ. Zool., Cook College, New Jersey R. MCLACHLAN - British Museum (Nat. Hist.), London Michigan Univ. - Zool. Mus. Univ. Mich., Ann Arbor B.E. MONTGOMERY - Zool. Inst. Pardue Univ., Lafayette B. P. MOORE - British Museum (Nat. Hist.), London H . K . MORRISON - British Museum (Nat. Hist.), London E.L. MORTON - Mus. Comp. Zool., Cambridge (Mass.) L. NAVÄS - Mus. Cienc. Nat., Barcelona J. G. NEEDHAM - Acad. Sei. Hist. Nat. Mus., Peiping and Heude Mus., Shanghai and British Museum (Nat. Hist.), London C. NIELSEN - 1st. Ent. Univ., Bologna K. OGUMA - locality unknown Ohio Mus. - Dept. Zool. & Ent. Ohio State Univ., Columbus

XIII Τ.

- private collection, Tokyo Oxford Mus. - Hope Mus., Oxford Paris Mus. - Mus. Nation. Hist. Nat., Paris Peking Mus. - Acad. Sei. Hist. Nat. mus., Peiping E . PINHEY - Zool. Dept. Nat. Mus., Bulawayo Pittsburgh Mus. - Carnegie Mus. Nat. Hist., Pittsburgh F. POEY - Amer. Ent. Soc., Philadelphia Pretoria Mus. - Dept. Zool. Transvaal Mus., Pretoria Pusa Coli. - Nation. Pusa Coll., New Delhi J . RACENIS - Univ. Centr. Esc. Biol., Caracas Rhodes Mus. - Dept. Zool. & Ent., Rhodes Univ., Grahamstown Rio de Janeiro Mus. - Dept. Zool. Nac. Cienc. Mus., Rio de Janeiro F. Ris - Senckenberg Mus., Frankfurt am Main J . P RAMBUR - R . Mus., Bruxelles M.J. ROBYNS - R . Mus., Bruxelles M.R. SAHNI - Geol. Surv. India, Calcutta S. Afr. Mus. - Div. Ent. South Africain Mus., Cape Town N.D. SANTOS - Dept. Zool. Nac. Cienc. Mus., Rio de Janeiro Sarawak Mus. - Dept. Zool. Nat. Hist. Mus., Sarawak Ε. SAUNDERS - British Museum (Nat. Hist.), London H.L.F. SAUSSURE - Naturhist Mus., Genf E. SCHMIDT - privat collection, after 1969 in Asahina's Coli. W.G. SCHNEIDER - Mus. Naturk., Berlin G . SCHUMANN - private collection, Berlin S . H . SCUDDER - Mus. Comp. Zool., Cambridge (Mass.) E . SELYS - R . Mus., Bruxelles E. SELYS & R. M C L A C H L A N - British Museum (Nat. Hist.), London C . G . SEMPER - Mus. Naturk., Berlin Senckenberg Mus. - Senckenberg Mus., Frankfurt am Main J.G. SERVILLE - R. Mus., Bruxelles B.Y. SJÖSTEDT - Naturhist. Riksmus., Stockholm M.C. SOMMER - Insektenhändler, München D . ST. QUENTIN - private collection, Wien Stettin Mus.- Hist. Nat. Mus. Stettin, after 1945 in PAN, Zool. Inst., Warszawa J.F. STEPHEN - British Museum (Nat. Hist.), London C. STEVEN - Zool. Inst. Univ. Mus., Charkow Tervuren Mus. - Mus. R. Afr. centr., Tervuren R.J. TILL YARD - Dom. Mus., Wellington and British Museum (Nat. Hist.), London Torino Mus. - Mus. Inst. Zool. sist. Univ., Torino B.J. TOUGH - private collection, Chicago Transvaal Mus. - Dept. Zool. Transvaal Mus., Pretoria OKUMURA

XIV - Zool. Inst. Acad. Sei. Sin., Peiping P. R. UHLER - Mus. Comp. Zool. Cambridge (Mass.) Z. VARGA - Biol. Inst. Kossuth Univ., Debrecen H. VIRMOND - Mus. Naturk., Berlin F. WAGNER - Insektenhändler, Wien Ε. Μ. WALKER - British Museum (Nat. Hist.), London W.S. WALL - Dept. Zool. Austral Mus., Sydney A. R. WALLACE - British Museum (Nat. Hist.), London M.E. WALSH - private collection, London A. W H I T E - Dept. Zool. S. Austr. Mus., Adelaide Warszawa Mus. - PAN Zool. Inst., Warszawa B. WATSON - Anat. Dept. Tulane Univ., New Orleans B.W. WESTERMANN - Univ. Zool. Mus., Kobenhavn M.J. WESTFALL - Dept. Zool. Univ. Florida, Gainesville E.B. WILLIAMSON - Zool. Mus. Univ. Mich., Ann Arbor W. WINTHEM - Naturhist. Mus., Wien Yerevan Mus. - Zool. Inst. Arm. Acad. Mus., Yerevan W. Z H O U - Dept. Ent. Zhejiang Mus., Hang Zhou Zürich Mus. - Ent. Inst. Techn. Hochsch., Zürich

TSAI-TSCHE-KIANG

Systematic Index subordo ZYGOPTERA

2

superfamily Calopterygoidea . . .

3

family Amphipterygidae

3

subfamily Amphipteryginae Amphipteryx Selys Devadatta Kirby Tetraneura Selys Pentaphlebia Förster

...

3 4 4 4 5

subfamily Philoganginae Philoganga Kirby Anisoneura Selys

5 6 6

family Calopterygidae Agrionidae Agriidae

6 6 7

subfamily Caliphaeinae Caliphaea Selys Notholestes McLachlan Noguchiphaea Asahina

7 7 8 8

subfamily Calopteryginae Agrioninae Vestalinae Archineura Kirby Leucopteryx Fraser Calopteryx Leach Agrion Fabricius Silphis Selys

8 8 8 8 8 9 9 9

Anaciagrion Kennedy Echo Selys Climacobasis Laidlaw Iridictyon Needham& Fischer Matrona Selys Mnais Selys Neurobasis Selys Matronides Förster Phaon Selys Prophaon Fraser Psolodesmus McLachlan . . . . Sapho Selys Umma Kirby Cleis Selys Vestalis Selys Vestalaria May Vestinus Kennedy

9 18 18 19 19 20 23 23 25 25 26 27 28 28 30 30 30

subfamily Hetaerinae Hetaerina Selys Mnesarete Cowley Lais Selys

34 34 42 42

family Chlorocyphidae Libellagidae

44 44

Africocypha Pinhey Aristocypha Laidlaw Calocypha Fraser Chlorocypha Fraser Cyrano Needham & Gyger .. Neocypha Tillyard & Fraser . Disparocypha Ris

45 45 48 48 54 54 55

XVI Heliocypha Fräser Heterocypha Laidlaw Indocypha Tillyard & Fraser Libellago Selys Micromerus Rambur Sclerocypha Fraser Melanocypha Fraser Pachycypha Lieftinck Paracypha Fraser Platycypha Fraser Rhinocypha Rambur Rhinoneura Laidlaw Sundacypha Laidlaw

55 58 58 58 58 58 62 63 63 63 65 73 73

family Dicteriastidae Heliocharatidae Cyanocharis Needham Dicterias Selys Charitopteryx Cowley Neocharis Förster Heliocharis Selys

74 74 74 74 74 74 75

family Diphlebiidae Diphlebia Selys Dineura Selys

75 76 76

family Euphaeidae Epallagidae Allophaea Fraser Anisophaea Fraser Mesophaea Fraser Anisopleura Selys Bayadera Selys Cylophaea Ris Dysphaea Selys Epallage Charpentier Euphaea Selys Pseudophaea Kirby Heterophaea Cowley Paraphaea Martin Indophaea Fraser Schmidtiphaea Asahina

77 77 77 78 78 79 80 81 81 83 83 84 88 88 88 89

family Polythoridae

89

subfamily Euthorinae Euthore Selys

90 90

subfamily Miocorinae Josocora Kennedy Kalocora Kennedy Miocora Calvert Stenocora Kennedy

91 91 92 92 92

subfamily Polythorinae Chalcopteryx Selys Cora Selys Polythore Calvert Thore Hägen

93 93 94 96 96

family Rimenellidae Rimanella Needham Rima Needham

100 100 100

superfamily Lestinoidea

101

family Chlorolestidae Synlestidae

101 101

subfamily Chlorolestinae Synlestinae Chlorolestes Selys Euchlorolestes Kennedy Ecchlorolestes Barnard Episynlestes Kennedy Phylolestes Christiansen Sinolestes Needham Synlestes Selys

101 101 101 101 103 103 . . . . 104 104 105

subfamily Chorismagrioninae . . 105 Chorismagrion Morton 105 subfamily Megalestinae Megalestes Selys

106 106

family Lestidae

107

XVII subfamily Lestinae Archilestes Selys Superlestes Williamson Cyptolestes Williamson Lestes Leach Africalestes Kennedy Agrion (Anapetes) Chapentier Chalcolestes Kennedy Icterolestes Pinhey Malgassolestes Pinhey Paralestes Schmidt Pseudochalcolestes Pinhey . . . Puella, sensu Akramowski . . Xerolestes Fraser Orolestes McLachlan Platylestes Selys Sinhalestes Fraser

107 108 108 108 109 109 109 109 109 109 109 109 109 109 127 128 128

subfamily Sympecmatinae Austrolestes Tillyard Ceylonolestes Kennedy Indolestes Fraser Ceylanicolestes Fraser Esaulestes Fraser Tricholestes Fraser Sympecma Selys Sympycna Charpentier

128 129 129 131 131 131 131 135 135

family Lestoideidae

137

subfamily Lestoideinae Lestoidea Tillyard

137 137

subfamily Coryphagrioninae . . . 165 Coryphagrion Morton 165

family Megapodagrionidae Megapodagriidae

138 138

subfamily Hypolestinae Hypolestes Gundlach Ortholestes Calvert

165 165 165

subfamily Argiolestinae Agnophilogenia Kennedy . . . Agriomorpha May Allolestes Selys Amanipodagrion Pinhey . . . .

138 138 138 139 139

subfamily Megapodagriinae . . . Podagrioninae Tatocnemidinae Arrhenocnemis Lieftinck . . . .

166 166 166 166

Archeopodagrion Kennedy . . Argiolestes Selys Wahnesia Förster Archiargiolestes Kennedy . . . . Austroargiolestes Kennedy . . Risiolestes Fraser Metagrion Calvert Bornargiolestes Kimmins . . . Burmargiolestes Kennedy . . . Caledargiolestes Kennedy . . . Caledopteryx Kennedy Calilestes Fraser Celebargiolestes Kennedy . . . Dimeragrion Calvert Heteragrion Selys Heteropodagrion Selys Mesagrion Selys Nesolestes Selys Neuragrion Karsch Neurolestes Selys Oxystigma Williamson Paraphlebia Selys Philogenia Selys Podolestes Selys Podopteryx Selys Protolestes Förster Rhinagrion Calvert Amphilestes Selys Sciotropis Racenis Trineuragrion Ris

139 139 139 140 140 140 140 146 146 147 147 147 148 148 148 153 153 153 155 155 156 156 157 160 161 162 163 163 164 164

XVIII Megapodagrion Selys Allopodagrion Förster Podagrion Selys Mesopodagrion McLachlan Tatocnemis Kirby Nesocnemis Selys

167 167 167 . 169 169 169

subfamily Philosininae Philosina Ris

170 171

subfamily Thaumatoneurinae .. 171 Thaumatoneura McLachlan . 171 family Perilestidae Nubiolestes Fraser Camerunolestes Schmidt Eolestes Fraser Perilestes Hägen Perissolestes Kennedy

171 172 . . . . 172 172 172 173

family Pseudolestidae Lestomima May Pseudolestes Kirby Rhipidolestes Ris Taolestes Needham

174 174 175 175 175

superfamily Hemiphlebioidea . . . 176 family Hemiphlebiidae

176

Hemiphlebia Selys

176

superfamily Coenagrionoidea . . . 177 family Coenagrionidae Heteropteroides Rhomboides Puellidae Agrionidae subfamily Agriocnemidinae . . . . Agriocnemis Selys Argyrocnemis Kennedy

177 177 177 177 177 178 178 178

Argiocnemis Selys Austrocnemis Tillyard Coenagriocnemis Fraser Coenargiocnemis Fraser Mortonagrion Fraser Indagrion Fraser

185 187 . . . . 188 188 188 188

subfamily Argiinae Archboldargia Lieftinck . . . . Argia Rambur Chalcargia Kennedy Cyanargia Kennedy Diargia Calvert Heliargia Kennedy Hyponeura Selys Leptargia Kennedy Micrargia Kennedy Hylaeargia Lieftinck Moroagrion Needham & Gyger Onychargia Selys Palaiargia Förster Papuargia Lieftinck

191 191 191 191 191 191 191 191 191 191 210 210 210 211 214

subfamily Coenagrioninae Agrioninae Coenagriinae Austroagrion Tillyard Austrocoenagrion Kennedy .. Cercion Naväs Chromagrion Needham Coenagrion Kirby Erythromma Charpentier . . . Pseuderythromma Kennedy . . Himalagrion Fraser Nehalennia Selys Argiallagma Selys Neoerythromma Kennedy . . . Pyrrhosoma Charpentier . . . .

214 214 215 215 216 216 218 219 227 227 228 229 229 230 231

subfamily Ischnurinae Acanthagrion Selys

232 232

XIX Myagrion Förster Acanthallagma Williamson & Williamson . Aciagrion Selys Ischnuragrion Longfield Mombagrion Sjöstedt Amorphostigma Fraser Amphiagrion Selys Amphiallagma Kennedy . . . . Andinagrion Bulla Anomalagrion Selys Archaeallagma Kennedy . . . . Argentagrion Fraser Homeoura Kennedy Austroallagma Lieftinck . . . . Calvertagrion St. Quentin .. . Cyanallagma Kennedy Enacantha Donnelly & Alayo Enallagma Charpentier Africallagma Kennedy Libyagrion Fraser Ischnallagma Kennedy Proischnura Kennedy Teleallagma Kennedy Thermagrion Förster Helveciagrion Machado Hesperagrion Calvert Ischnura Charpentier Anomalura Kennedy Boninagrion Asahina Bedfordia Mumford Celaenura Kennedy Ceratura Selys Ischnosoma Wallengren Ischnuridia Donnelly Micronympha Kirby Nanosura Kennedy Mesamphiagrion Kennedy . . Millotagrion Fraser Oreagrion Ris Oxyagrion Selys

232 237 238 238 238 243 243 244 244 245 245 245 245 246 247 247 248 248 248 248 248 248 248 248 262 263 263 263 263 263 263 263 263 263 263 263 278 278 279 279

Pacificagrion Fraser Protallagma Kennedy Oxyallagma Kennedy Rhodischnura Laidlaw Skiallagma Förster Tigriagrion Calvert Xiphiagrion Selys Zoniagrion Kennedy

282 282 282 283 283 283 284 284

subfamily Leptobasinae Antiagrion Ris Chrysobasis Racenis Diceratobasis Kennedy Inpabasis Santos Leptobasis Selys Leucobasis Racenis Mesoleptobasis Sjöstedt Metaleptobasis Calvert

284 285 285 286 286 286 288 288 288

subfamily Pseudagriinae Amphicneminae Aeolagrion Williamson Hylaeagrion Förster Amphicnemis Selys Anisagrion Selys Apanisagrion Kennedy Archibasis Kirby Stenobasis Selys Argiagrion Selys Caliagrion Tillyard Ceriagrion Selys Brachybasis Selys Palaeobasis Kennedy Nepocnemis Fraser Hylaeonympha Racenis Leptagrion Selys Megalagrion McLachlan . . . . Hawaiiagrion Kennedy Kilauagrion Kennedy Oahuagrion Kennedy Melanesobasis Donnelly . . . . Minagrion Santos

290 290 290 290 292 295 296 296 296 298 298 298 298 298 299 308 308 311 311 311 311 316 317

XX Nesobasis Selys Papuagrion Ris Pericnemis Hägen Plagulibasis Lieftinck Pseudagrion Selys Paulinagrion Fraser Seychellibasis Kennedy Stenagrion Laidlaw Teinobasis Kirby Telagrion Selys Aceratobasis Kennedy Telebasis Selys Erythragrion Selys Xanthagrion Selys Xanthocnemis Tillyard

318 319 322 323 323 323 348 349 349 357 357 358 358 362 362

family Isostictidae Austrosticta Tillyard Eurysticta Watson Isosticta Selys Neosticta Tillyard Amphisticta Sjöstedt Oristicta Tillyard Phasmosticta Lieftinck Selysioneura Förster Tanymecosticta Lieftinck . . . . Stenosticta Lieftinck

363 363 363 364 365 365 365 365 365 367 367

family Platycnemididae Micronymphidae

368 368

subfamily Calicneminae Allocnemis Selys Arabicnemis Waterston Asthenocnemis Lieftinck . . . . Calicnemia Strand Calicnemis Selys Coeliccia Kirby Trichocnemis Selys Cyanocnemis Lieftinck Idiocnemis Selys Indocnemis Laidlaw

368 369 369 369 369 369 371 371 378 379 381

Leptocnemis Selys Hemicnemis Selys Lieftinckia Kimmins Lohmaeocnemis Lieftinck . . . Mesocnemis Karsch Metacnemis Selys Oreocnemis Pinhey Paracnemis Martin Ciliagrion Sjöstedt Paramecocnemis Lieftinck . . . Rhyacocnemis Lieftinck . . . . Risiocnemis Cowley Prionocnemis Selys Hypocnemis Hägen Solomocnemis Lieftinck . . . . Stenocnemis Karsch Thaumatagrion Lieftinck . . . Torrenticnemis Lieftinck . . . . subfamily Platycnemidinae Copera Kirby Psilocnemis Selys Platycnemis Charpentier Platyscelus Wallengren Proplatycnemis Kennedy Pseudocopera Fraser family Platystictidae subfamily Palaemnematinae Palaemnema Selys

381 381 382 382 383 383 384 384 384 385 385 385 385 385 387 387 388 388

. . . . 388 388 388 . . . . 392 392 . . . . 392 392 398 . . . 398 398

subfamily Platystictinae Drepanosticta Laidlaw Ceylonosticta Fraser Platysticta Selys Protosticta Selys

402 403 403 413 413

family Protoneuridae

417

subfamily Caconeurinae Caconeura Kirby

417 417

XXI Alloneura Selys, 1860 Hypostrophoneura Munz . . . . Indoneura Laidlaw Esme Fräser Melanoneura Fraser Phylloneura Fraser

417 417 417 418 419 419

subfamily Disparoneurinae . . . . Chlorocnemis Selys Disparoneura Selys Chloroneura Laidlaw Elattoneura Cowley Isomecocnemis Cowley Nososticta Selys Notoneura Tillyard Risioneura Munz Prodasineura Cowley Alloneura Selys, 1886 Brachyneura Selys Cycloneura Fraser

419 419 422 422 423 428 429 429 429 437 437 437 437

subfamily Protoneurinae 443 Epipleoneura Williamson . . . 444 Epipotoneura Williamson . . . 446

Idioneura Selys Junix Räcenis Microneura Selys Neoneura Selys Caenoneura Kirby Peristicta Selys Phasmoneura Williamson Proneura Selys Protoneura Selys Psaironeura Williamson Roppaneura Santos family Pseudostigmatidae

446 446 446 446 446 450 . . . 450 451 451 454 455 455

subfamiliy: Pseudostigmatinae . 455 Anonisma McLachlan 455 Mecistogaster Rambur 456 Goniostigma Kennedy 456 Haplostigma Kennedy 456 Platystigma Kennedy 456 Xanthostigma Kennedy 456 Megaloprepus Rambur 459 Microstigma Rambur 459 Pseudostigma Selys 461

Catalogue

Order ODONATA Predacious insects with biting mouthparts and two equal or subequal pairs of elongate membranous wings, each wing with a complex reticulation of small crossveins and usually a conspicuous pterostigma. Eyes very large and prominent; antennae very short and filiform. Abdomen elongate, often extremely slender. Male accessory genital armature developed under 2nd and 3rd abdominal segments. Larvae usually aquatic, hemimetabolous; labium developed into a prehensile organ; respiration by means of caudal lamellae, rectal or occasionally lateral abdominal gills. 1783

Odonata FABRICIUS, Ent. Suppl.: 2 8 0 (name applied as family). - 1 8 1 5 Odonata - LEACH, Edin. Encycl., 9: 136. - 1839 Odonata - BURMEISTER, H a n d b . Ent., 2: 805. - 1842 Odonata - RAMBUR, Hist. nat. Ins. Nevropt., Paris: 24 (with families: Libellulides, Aeschnides, G o m phides, Agrionides). - 1858 Odonata - SELYS, Mem. Soc. r. Sei. Liege, 11: 261 (as suborder of order Orthoptera). - 1861 Odonata - HAGEN, Syn. Neur. N. Amer., Washington: 55 (as family of order Neuroptera). - 1868 Odonata - BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 29 (tables of genera). - 1875 Odonata - HAGEN, Proc. Boston Soc. nat. Hist., 18: 20. - 1876 Odonata - BUCHECKER, Syst. Ent. Odonata, 1:1890 - KIRBY, Syn. Cat. Neur.-Odon., London: 7. - 1893 Odonata - CALVERT, Trans. Am. ent. Soc., 2 0 : 152. - 1894 Odonata - K I R B Y , JL. Linn. Soc., London, 24: 545. - 1895 Odonata - CALVERT, Proc. Cal. Acad., (2) 4: 463. 1896. Odonata - M C L A C H L A N , Ann. Mag. nat. Hist., (6) 1 7 : 364. - 1 8 9 7 Odonata- CARPENTER, Proc. Dublin Soc., 8: 439. - 1898 Odonata - CALVERT, Trans. Amer. ent. Soc., 25: 27. - 1899 Odonata - KELLICOTT, Odon. Ohio: 114. - 1900 Odonata - WILLIAMSON, Drag. Ind. (Ind. Rep. Geol. for 1899): 229. - 1903 Odonata - NEEDHAM, Proc. U.S. nat. Mus., 26: 703 (genealogic study of wing venation). - 1904 Paraneuroptera SHIPLEY, Zool. Anz., 27: 261 (new name for Odonata). - 1905 Odonata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 714 (revision, key to suborders) - 1908 Odonata - WALKER, Ottawa Nat., 2 2 : 1 , 4 9 . - 1910 Odonata - MUTTKOWSI, Bull. Publ. Mus. Milwaukee, 1 (1): 25. - 1901-08 Odonata - CALVERT, Biol. Centr. Amer. London, 1901: 17, 1902: 73, 1903: 129, 1905: 145, 1906: 213, 1907: 309, 1908: 405. - 1909-19 Odonata - Ris, Coll. zool. Edm. de Selys Longchamps, Bruxelles, 9: 1 , 1 0 - 1 1 : 121, 1 2 - 1 3 : 385, 1 4 : 701, 1 5 : 837, 1 6 : 965, 17:1043. - 1912 Odonata - BARTENEF, Ann. Mus. zool. Acad. imp. Sei. St.-Petersbourg, 16: 409. - 1915 Odonata - BARTENEF, Faune de la Russ., Petrograd: 576. - 1919 Odonata- MUNZ, Mem. Amer. ent. soc., 3: 39 (key to suborders Anisozygoptera, Zygoptera, Anisoptera). - 1921 Odonata - Ris, Annls S. Afr. Mus., 18: 245. - 1 9 2 8 Odonata - A N D E R S , Mem. Soc. r. ent. Egypte, 3 ( 1 ) : 7 . - 1 9 2 8 Odonata - FRASER, Trans, ent. Soc. London, 76: 123. - 1928 Odonata - TILLYARD, Ree. Indian Mus., 30: 151. - 1 9 2 9 Odonata - TILLYARD, IV. Int. Congr. Ent., Ithaca: 5 4 3 . - 1 9 2 9 Odonata - LONGFIELD, Trans, ent. Soc. London, 1929: 125. - 1932 Odonata - NEEDHAM, Ree. Indian Mus., 34: 195 (key to Indian taxa). - 1934 Odonata - COWLEY, Entomologist's month. Mag., 70: 240. 1934 Odonata - Entomologist, 67: 200. - 1934 Odonata - SCHOUDETEN, Ann. Mus. Congo Beige, Zool., 3 II 3: 9. - 1935 Odonata - COWLEY, Entomologist, 68: 154. - 1935 Odonata LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 1. - 1 9 3 8 Odonata - T I L L YARD & FRASER, Austr. Zool., 9: 142 (key to suborders: Protozygoptera, Archizygoptera, Meganisoptera, Protanisoptera,

2 Anisoptera, Zygoptera, Anisozygoptera). - 1 9 3 9 Odonata - TILL YARD & FRASER, Austr. Zool., 9:195. - 1 9 4 0 Odonata-TILLYARD & FRASER, Austr. Zool., 9:359. - 1 9 4 2 Odonata—LIEFTINCK, Treubia, 18 (3): 441. - 1948 Odonata - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 117. - 1948 CWonafa-WHITEHOUSE, Trans. R. Can. Inst., 27(57): 3 . - 1 9 4 8 Odonata - ZIMMERMAN, Insects of Hawaii, 2 : 321. - 1952 Odonata - ASAHINA, Mushi, 2 3 (6): 45. - 1953 Odonata - WALKER, Toronto Univ. Press: 292. - 1954 Odonata - LIEFTINCK, Treubia, 22 Suppl.: 1. - 1954 Odonata - BRUES, MELANDER & CARPENTER, Classification of Insects, Cambridge (Mass.): 6 2 (key to suborders Anisoptera, Zygoptera, Anisozygoptera). - 1955 Odonata - LONGFIELD, Publ. cult. Comp. Diam. Angola, 2 7 : 1 1 . - 1 9 5 6 Odonata - ASAHINA, Ent. Medd., ( 4 - 5 ) : 2 0 4 . - 1 9 5 7 Odonata - FRASER, Ree. zool. Soc. N . S . Wales, 1957: 24 (with suborders: Protozygoptera, Archizygoptera, Meganisoptera, Protanisoptera, Anisoptera, Zygoptera, Anisozygoptera). 1959 Odonata - LIEFTINCK, Ent. Mitt, zool., Staatsinst., Zool. Mus. Hamburg, 1959: 21: 1. - 1959 Odonata - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 15. - 1962 Odonata LIEFTINCK, Insects of Micronesia, 5 (1): 8 (key to Micronesian suborders and families). 1962 Odonata - PINHEY, Publ. cult Comp. Diam. Angola, 59: 87 (key to African suborders and superfamilies). - 1962 Odonata - COMPTE, Publ. Inst. Biol. apl. Barcelona, 35: 33. - 1965 Odonata - ASAHINA, Kontyü, 33 (4): 493.

Suborder Zygoptera More or less delicate build, with eyes separated by a space greater than their dorsal diameter. Wings usually held closed over abdomen in repose; fore and hind wings closely alike with narrow basal area. Dc a simple quadrilateral with the proximal side formed by the lower part of the arculus. Nodus in proximal half of costa. Male with two pairs of anal appendages, the inferior pair being below the anus. Male genital armature belongs to the second abdominal segment. Females with complete ovipositor. Larva with three caudal gills. 1815 Agrionida LEACH, Brewster's Edinb., Encycl., 9: 15. - 1840 Agrionina SELYS, M o n o g r . Libell. eur., Paris: 1. - 1842 Agrionides RAMBUR, Hist. nat. Ins. Nevropt., Paris: 217 (with genera Calopteryx, Euphaea, Rhinocypha, Micromerus, Platycnemis, Lestes, Argia, Agrion, Mecistogaster, Microstima, Megaloprepus). - 1853 Zygoptera SELYS, Synopsis des Calopterygines: 6 (cited as tribe). - 1854 Zygoptera - SELYS, Mem. Soc. Liege, 9 : 1 1 . - 1 8 7 5 Zygoptera - HÄGEN, Proc. Boston Soc. nat. Hist., 18: 20. - 1890 Zygoptera - KIRBY, Syn. Cat. Neur.-Odon., London: 202. - 1895 Zygoptera - KELLICOTT, Jn. Cine. Soc. nat. Hist., 1895: 195. - 1900 Zygopterides - LUCAS, Brit. Dragonfl., London: 10. - 1903 Candibranchiata GRÜNBERG, Zool. Jb., 1 : 6 9 5 . - 1 9 0 3 Zygoptera-NEEDHAM, Bull. 6 8 Ν . Y . State Mus., 1 9 0 3 : 2 1 8 . - 1 9 1 0 Zygoptera - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 25. - 1919 Zygoptera - MUNZ, Mem. Amer. ent. Soc., 3: 8 (revision, key to families and subfamilies). - 1921 Zygopteras - Ris, Annls S. Afr. Mus., 18: 260 (with families: Calopterygidae, Agrionidae). - 1920 Zygoptera - KENNEDY, Ohio J. Sei. Columbus, 21: 83. - 1928 Zygoptera - FRASER, Trans, ent. Soc. London, 78: 123. - 1929 Zygoptera - LONGFIELD, Trans, ent. Soc. London, 77: 125. - 1934 Zygoptera - COWLEY, Entomologist, 67: 200. - 1934 Zygoptera - LIEFTINCK, Treubia, 14 (4). 377. - 1934 Zygoptera - SCHOUDETEN, Ann. Mus. Congo Beige, Zool., 3 II 3: 9. - 1937 Zygoptera - C O W L E Y , J. Soc. Bibl. nat. Hist., 1: 7 3 . - 1 9 3 8 Zygoptera - TILLYARD & FRASER, Austr. Zool., 9: 146 (key to superfamilies: Agrioidea, Hemiphlebioidea, Coenagrioidea). 1949 Zygoptera - FRASER, Trans, ent. Soc. London, 100: 135. - 1954 Zygoptera - BRUES, MELANDER & CARPENTER, Classification of Insects, Cambridge (Mass.): 6 4 (with superfamilies: Agrionoidea, Hemiphlebioidea, Coenagrionoidea). - 1954 Zygoptera - FRASER, Entomologist's mon. Mag., 9 0 : 2 1 1 . - 1 9 5 6 Zygoptera - FRASER, R . ent. Soc. Lond., 1 ( 1 0 ) : 1. - 1 9 5 7 Zygoptera - FRASER, R . Zool. Soc. N . S . Wales, 1 9 5 7 : 3 6 (with superfamilies: Agrioidea, Hemiphlebioidea, Coenagrioidea, Lestinoidea) (key to superfamilies). - 1 9 6 2 Zygoptera - PINHEY,

3 Publ. cult. Comp. Diam. Angola, 59: 87 (with superfamilies: Lestinoidea, Coenagrioidea, Agrioidea).

Superfamily Calopterygoidea Medium to large Zygoptera with broad wings, frequently coloured with metallic sheen and having closely reticulated vains. Usually 5 or more antenodals, most often numerous; postnodals usually not aligned with cross veins beneath. Dc long, rectangular and usually crossed. IRiii and Riv arising closer to arc than to nodus. Riii arising at nodus, anal vein separate from wing border from base. First thoracic suture complete (except in Amphipterygidae and Chlorocyphidae). Larva with triquetral or saccoidal (Amphipterygidae, Chlorocyphidae) anal gills; labium usually deeply indented. 1840 Normopteroides SELYS, Monogr. Libell. eur., Paris: 1 (emandation). - 1860 Calopterygina SELYS, Mem. Soc. Sei. Liege, 6:18. -1926 Agrioidea TILLYARD & FRASER, Insects of Austr.-New Zeal.: 78. - 1939 Agrioidea - TILLYARD & FRASER, Austr. Zool., 9: 202 (key to families Polythoridae, Amphipterygidae, Chlorocyphidae, Heliocharitidae, Epallagidae, Agriidae). 1954 Agrionoidea - BRUES, MELANDER & CARPENTER, Classification of Insects, Cambridge (Mass.): 65 (key to families Polythoridae, Amphipterygidae, Chlorocyphidae, Heliocharitidae, Epallagidae, Agrionidae). - 1957 Agrioidea - FRASER, R. Zool. Soc. N.S. Wales, 1957: 61 (key to families Pseudolestidae, Polythoridae, Amphipterygidae, Chlorocyphidae, Heliocharitidae, Epallagidae, Agriidae). - 1962 Agrioidea - PINHEY, Publ. cult. Comp. Diam. Angola, 59:141 (key to families Agriidae, Chlorocyphidae, Amphipterygidae). - 1965 Calopterygoidea BENEDEK, Folia ent. hung., 18: 415 (new name for Agrioidea TILLYARD, 1926). - 1984 Calopterygoidea - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 (with families Amphipterygidae, Calopterygidae, Chlorocyphidae, Dicteriastidae, Diphlebiidae, Euphaeidae, Polythoridae, Rimanellidae).

Family Amphipterygidae Medium sized forms, sturdy with long narrow petiolate wings. Some accessory antenodals (6-12); IRiii and Riv recessed close to arc\ primary antenodals evident. Base of Dc connected to radius by arc. Rii-iii not arched forward near origin. Larva with saccoid caudal gills. 1926 Amphipterygidae TILLYARD, Insects of Austr.-New Zeal.: 79. -1938 Amphipterygidae -FRASER, Proc. R. ent. Soc. London, (B) 7: 137 (with genera Amphipteryx, Devadatta, Pentaphlebia, Diphlebia, Philoganga). - 1957 Amphipterygidae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 69 (with subfamilies Philoganginae, Amphipteryginae). - 1962 Amphipterygidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 152.

Subfamily Amphipteryginae Secondary antenodals in costal space only; prox. ends of IRiii and Riv close and near arc. Dc long and narrow (not in Devadatta). Larva with gill tufts. 1939 Amphipteryginae TILLYARD & FRASER, Austr. Zool., 9: 204 (with genera Diphlebia, Amphipteryx, Devadatta, Pentaphlebia, Pseudolestes). - 1957 Amphipteryginae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 70 (with genera Diphlebia, Amphipteryx, Devadatta, Pentaphlebia,

4 Hypolestes, Pseudolestes, Dysagrion). - 1962 Amphipteryginae Diam, Angola, 59: 152.

- PINHEY, Pub. cult. Comp.

Genus Amphipteryx SELYS 1853 Amphipteryx SELYS, Synopsis des Calopterygines: 66. Type-species: Amphipteryx SELYS, 1 8 5 3 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon. London: 1 1 1 ) . phipteryx

- SELYS & HAGEN, M o n o g r . Cal.: 258 (revision). - 1890 Amphipteryx

agrioides Am-

1854

- KIRBY, Syn.

Cat. Neur.-Odon. London: 111. - 1926 Amphipteryx - TILLYARD, Insects of Austr.-New. Zeal.: 79. - 1938 Amphipteryx - FRÄSER, Proc. R. ent. Soc. London, (B) 7: 137. - 1957 Amphipteryx - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 70. - 1962 Amphipteryx - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 152.

agrioides SELYS 1853 Amphipteryx agrioides SELYS, Synopsis des Calopterygines: 66. Type female: Selys's Collection; type-locality: Colombia. 1 8 5 4 Amphipteryx agrioides - SELYS & H Ä G E N , Monogr. Cal.: 2 4 1 (redescription).

Distribution: Colombia. Genus Devadatta KIRBY 1859 Tetraneura SELYS, Bull. Acad. r. Belg., (2) 7: 449. Type-species: Tetraneura argyroides SELYS, 1859 (monotyp). - 1890 Devadatta KIRBY, Syn. Cat. Neur.-Odon. London: 111 (new name for Tetraneura SELYS, 1859 a junior primary homonymy of Tetraneura HARTIG, 1841 in Hemiptera). - 1938 Devadatta - FRASER, Proc. R. ent. Soc. London, (B) 7: 138. - 1938 Devadatta - FRASER, Proc. R. ent. soc. London, (B) 7: 138 (proposed synonymy with Devadatta K I R B Y , 1890).

argyroides argyroides (SELYS) 1859 Tetraneura argyroides SELYS, Synopsis des Calopterygines, Add. 1:15. Type male: Selys's Collection; type-locality: Singapore. 1890 Devadatta argyroides - KIRBY, Syn. Cat. Neur.-Odon. London: 111 (as generotype of Devadatta KIRBY, 1890). 1903 Devadatta argyroides - NEEDHAM, Proc. U.S. natn. Mus., 26: 53. 1927 Devadatta argyroides - Ris, Zool. Meded., Leiden, 10 (1): 3 (Sumatra). 1935 Devadatta argyroides - LIEFTINCK, Misc. Zool. Sum., 92-93: 4 (Sumatra). 1 9 3 9 Devadatta argyroides - N E E D H A M & G Y G E R , Philipp. J . Sei., 7 0 ( 3 ) : 2 5 6 (Mindanao, Luzon). 1954 Devadatta argyroides argyroides - LIEFTINCK, Treubia, 22 Suppl.: 1 (Siam, Malaya, Sumatra).

Distribution: from Thailand to Sumatra.

argyroides tiomanensis LAIDLAW 1934 Devadatta argyroides tiomanensis LAIDLAW, Stylops, 3: 102. Type not given; type-locality: Tioman Island. 1954 Devadatta argyroides tiomanensis - LIEFTINCK, Treubia, 22 Suppl.: 1 (Malaya).

Distribution: Malaysia.

ducatrix LIEFTINCK 1984 Devadatta ducatrix LIEFTINCK, 1969 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the W o r l d ,

Utrecht, 1: 2. Distribution: C h i n a .

5

filipina NEEDHAM

& GYGER

1939 Devadatta filipina NEEDHAM type-locality: Mindanao.

& GYGER,

Philipp. J. Sei.,

70

(3): 257. Type male: not given;

Distribution: Philippine Islands, multinervosa

FRASER

1933 Devadatta multinervosa FRÄSER, J. Siam Soc. nat. Hist. Suppl., 9: 129. Type male: British Museum (Nat.Hist.), London; type-locality: Laos.

Distribution: Laos. podolestoides podolestoides

LAIDLAW

1934 Devadatta podolestoides podolestoides LAIDLAW, Stylops, 3: 101. Type male: Laidlaw's Collection; type-locality: West Borneo. 1 9 3 6 Devadattapodolestoides podolestoides - K I M M I N S , J . Fed. Malay. States Mus., 18: 7 9 (Sarawak). 1954 Devadatta podolestoides podolestoides - LIEFTINCK, Treubia, 22 Suppl.: 2 (Borneo)

Distribution: Borneo. podolestoides basilanensis

LAIDLAW

1934 Devadatta podolestoides basilanensis Laidlaw, Stylops, 3:101. Type male: Laidlaw's Collection; Type-locality: Borneo.

Distribution: Borneo. Genus Pentaphlebia

FÖRSTER

1909 Pentaphlebia FÖRSTER, Jb Wiesbaden Verh. Natk., 62: 211. Type-species: Pentaphlebia stahli FÖRSTER, 1909 (original designation). - 1962 Pentaphlebia - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 153.

gamblesi 1984

PARR

Pentaphlebia gamblesi Utrecht, 1: 2.

PARR, 1 9 7 7 .

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Nigeria. stahli

FÖRSTER

1909 Pentaphlebia stahli FÖRSTER, Jb Wiesbaden Ver. Natk., 62: 213. Type male: Ann Arbor Museum; type-locality: Cameroon (as generotype of Pentaphlebia FÖRSTER, 1909 1962 Pentaphlebia stahli - PINHEY, Publ. cult. Comp. Diam. Angola: 59: 153.

Distribution: Cameroon.

Subfamily Philoganginae Secondary antenodals in costal and subcostal spaces; prox. ends of IRiii and Riv widely spaced and midway between nodus and arc. Dc broad and short. 1920

Philoganginae K E N N E D Y , Ohio J . Sei. Columbus, 21: 8 2 . Type-genus: Philoganga K I R B Y , 1 8 9 0 . 1 9 3 9 Philoganginae - T I L L Y A R D & FRASER, Austr. Zool., 9: 2 0 4 (with single genus: Philoganga). - 1 9 5 7 Philoganginae - FRASER, R . Zool. Soc. N . S . Wales, 1957: 7 0 . - 1 9 8 4 Philoganginae - DA VIES & TOBIN, The Dragonflies of thw World, Utrecht, 1: 2. -

6

Genus Philoganga

KIRBY

1859 Anisoneura SELYS, Synopsis des Calopterygines, Add. 1: 10. Type-species: Anisoneura montana HAGEN, 1859 (monotypy). - 1890 Philoganga KIRBY, Syn. Cat. Neur. - Odon. London: 111 (new name for Anisoneura SELYS, 1859, a junior primary homonymy of Anisoneura GENE, 1852 in Lepidoptera). - 1905 Philoganga - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 795. - 1912 Philoganga - Ris, Suppl. Ent., Berlin, 1: 44. - 1912 Anisoneura - Ris, Suppl. Ent., Berlin, 1: 44 (proposed synonymy with Philoganga KIRBY, 1890). - 1920 Philoganga KENNEDY, Ohio J. Sei. Columbus, 21: 82. - 1927 Philoganga - FRÄSER, Ree. Indian Mus., 29: 79. - 1938 Philoganga - FRÄSER, Proc. R. ent. Soc. London 7: 139.

loringae

FRASER

1927 Philoganga loringae FRÄSER, Ree. Indian Mus., 29: 79. Type male: British Musum (Nat. Hist.), London; type-locality: Burma.

Distribution: Burma montana

(HÄGEN)

1859 Anisoneura montana HAGEN, in SELYS, Synopsis des Calopterygines, Add. 1: 11. Type female: Hagen's Collection; type-locality: Himalaya (as generotype of Anisoneura SELYS, 1859). 1890 Philoganga montana -KIRBY, Syn.Cat.Neur.-Odon. London: 111 (as generotype of Philoganga KIRBY, 1 8 9 0 ) .

Distribution: Himalayas. robusta

NAVÄS

1936 Philoganga robusta NAVÄS, MUS. Heude Notes ent. chinoise, 3 (4): 43. Type male: not given (perhaps in Heude Museum); type-locality: China.

Distribution: China. vetusta Ris 1912 Philoganga vetusta Ris, Suppl. ent., Berlin, 1:47. Type male: not given (perhaps in Eberswalde); type-locality: China. 1935 Philoganga vetusta - MAY, Senckenbergiana, 17: 95 (description of female).

Distribution: China.

Family Calopterygidae Medium sized forms, wings usually coloured in males, less often in females. Antenodals numerous, primaries not apparent; base of Dc connected to radius by arc. Wings densely reticulated, not petiolate. Arc near wing base, its sectors arising near CuP. Stigma often reduced, especially in males. Rii + iii fused with Ri near origin. Dc long and transversed by many veins. First thoracic suture complete. 1815 Agrionidae LEACH, Edinb. Encycl., 9: 137 (for all Zygoptera). - 1842 Agrionides RAMBUR, Hist. nat. Inst. Neuropt., Paris: 216 (family defined). - 1850 Agrionidae - HAGEN & SELYS, Revue des Odonates, Paris: 133 (with tribes Calopterygines, Agrionides). - 1853 Calopterygidae SELYS, Synopsis des Calopterygines: 6 . - 1 8 6 1 Agrionina - HÄGEN, Syn. Neur. N. Amer., Washington: 56 (as tribe). - 1903 Agrionidae - NEEDHAM, Bull. 68 N.Y. State Mus.: 118 (given family rank). - 1905 Calopterygidae - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 721 (with key to subfamilies Palaeophlebiinae, Euphaeinae, Calopteryginae, Libellaginae). - 1921

7 Calopterygidae - Ris, Annls S. Afr. Mus., 18: 261 (key to South African genera Libellago, Phaon). - 1926 Agriidae TILLYARD, Insects of Austr.-New Zeal.: 79. - 1929 Calopterygidae - FRÄSER, J. Bombay nat. Hist. Soc., 33: 595. - 1939 Agriidae - T I L L Y A R D & FRASER, Austr. Zool., 9:211 (with subfamilies Hetaerinae, Agriinae, Caliphaeinae). - 1 9 5 4 Agrionidae - BRUES, M A L A N D E R & CARPENTER, Classification of Insects, Cambridge (Mass.): 67. - 1957 Agriidae - FRÄSER, R. Zool. N. S. Wales, 1957: 79 (with subfamilies Caliphaeinae, Hetaerinae, Agriinae). - 1962 Agriidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 142. - 1984 Calopterygidae - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 (with subfamilies Caliphaeinae, Calopteryginae, Hetaerinae).

Subfamily Caliphaeinae Wings hyaline and petiolate to level of anal tracheal crossing; Dc with straight upper side and crossed; CuP and IA straight. Median space uncrossed. One row of cells between anal vein and posterior border of wing. 1929 Caliphaeinae FRÄSER, J. Bombay nat. Hist. Soc., 33: 595. Type-genus: Caliphaea Selys, 1859. - 1 9 3 9 Caliphaeinae - T I L L Y A R D & FRASER, Austr. Zool., 9 : 2 1 3 (with single genus Caliphaea). - 1957 Caliphaeinae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 80.

Genus Caliphaea SELYS 1859 Caliphaea SELYS, Bull. Acad. r. Belg., (2) 7: 439. Type species: Caliphaea confusa H Ä G E N , 1859 (montypy). - 1897 Notholestes MCLACHLAN, Entomologist's month. Mag., 24: 31. Typespecies: Notholestes elwesi MCLACHLAN, 1887 (monotypy). - 1 8 9 4 Notholestes - MCLACHLAN, Ann. Mag. nat. Hist., (6) 13: 435 (proposed synonymy with Caliphaea SELYS, 1859). - 1905 Caliphaea - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 793 (key to species). - 1929 Caliphaea - FRASER, J . Bombay nat. Hist. Soc., 33: 595. - 1939 Caliphaea - T I L L Y A R D & FRASER, Austr. Zool., 9: 213. - 1957 Caliphaea - FRASER, R. Zool. Soc. N. S. Wales, 1957: 80.

confusa HÄGEN 1859 Caliphaea confusa HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 7: 440. Type male: British Museum (Nat. Hist.), London; type-locality: Nepal (as generotype of Caliphaea SELYS, 1859). 1887 Notholestes elwesi MCLACHLAN, Entomologist's month. Mag., 24: 32. Type male: British Museum (Nat.Hist.), London; type-locality: Darjeeling (as generotype of Notholestes M C L A C H LAN, 1887). 1894 Notholestes elwesi - MCLACHLAN, Ann. Mag. nat. Hist., (6) 13: 435 (proposed synonymy with confusa H Ä G E N , 1 8 5 9 ) . 1894 Caliphaea consimilis MCLACHLAN, Ann. Mag. nat. Hist., (6) 13: 435 Type male: British Museum (Nat. Hist.), London; type-locality: West China. 1 9 0 5 Caliphaea confusa - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 7 9 3 (Sikkim). 1 9 0 5 Caliphaea consimilis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 3 (China, Tibet). 1928 Caliphaea confusa - MORTON, Trans, ent. Soc. London, 76: 111 (Sikkim, Assam, Yunnan). 1930 Caliphaea consimilis - NEEDHAM, Zool. Sinica, 11 (1): 211. 1934 Caliphaea consimilis - FRASER, Fauna brit. India, Odon., 2: 149 (proposed synonymy with confusa HÄGEN, 1859). 1956 Caliphaea consimilis - ASAHINA, Ent. Medd., 27 (4-5): 205 (as valid species). 1984 Caliphaea consimilis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 (as synonym of confusa HÄGEN, 1859).

Distribution: China, Nepal, North India.

8 t h a i l a n d i c a ASAHINA 1976 Caliphaea thailandica ASAHINA, Kontyü, 44 (4): 387. Type male: Asahina's Collection; typelocality: Thailand.

Distribution: Thailand. Genus Noguchiphaea ASAHINA 1976 Noguchiphaea ASAHINA, Kontyü, 44 (4): 388. Type-species: Noguchiphaeayoshikoae 1976 (original designation). yoshikoae ASAHINA

ASAHINA,

1976 Noguchiphaea yoshikoae ASAHINA, Kontyü, 44 (4): 390 (as generotype of Noguchiphaea ASAHINA, 1976). Type male: Asahina's Collection; type-locality: Thailand. 1984 Noguchiphaea yoshikoae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3.

Distribution: Thailand. Subfamily Calopteryginae Wings broad. Sectors of arc arising from its midpoint. Basal space usually not crossed. CuP straight; wings usually coloured in males. 1854 Calopteryginae SELYS, Monogr. Cal.: 16 (as legion). Type-genus: Calopteryx LEACH, 1815. 1875 Calopterygina - HAGEN, Proc. Boston Soc. nat. Hist., 18: 20 (with genera Calopteryx, Hetaerina, Cora, Lais, Heliocharis, Dicterias, Amphipteryx, Chalcopteryx, Thore). - 1890 Agrioninae KIRBY, Syn. Cat. Neur.-Odon. London: 96. Type-genus: Agrion Fabricius, 1775. - 1903 Vestalinae NEEDHAM, Proc. U.S. natn. Mus., 26: 744. Type-genus: Vestalis SELYS, 1853. - 1905 Calopteryginae - JACOBSON & BIANKI, Pryam. lozhnos, Ross, imp.: 796 (with genera Neurobasis, Matrona, Calopteryx, Echo, Archineura, Mnais, Psolodesmus, Vestalis). - 1910 Agrioninae-MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 26. - 1 9 1 9 Agrioninae - MUNZ, Mem. Amer. ent. Soc., 3: 42 (key to genera Archineura, Echo, Climacobasis, Umma, Sapho, Agrion, Metronides, Matrona, Neurobasis, Mnais, Psolodesmus, Phaon, Vestalis, Hetaerina, Lais). - 1939 Agriinae TILLYARD & FRASER, Austr. Zool., 9: 212 (with genera Agrion, Mnais, Neurobasis, Vestalis, Matrona, Metronides, Echo, Climacobasis, Psolodesmus, Archineura, Phaon, Sapho, Umma). - 1957 Agriinae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 81 (with genera Agrion, Neurobasis, Prophaon, Vestalis, Matrona, Matronides, Echo, Climacobasis, Psolodesmus, Archineura, Phaon, Sapho, Umma, Iridictyon). - 1962 Agriinae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 142 (key to genera Phaon, Umma, Sapho). - 1984 Calopteryginae - DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d , U t r e c h t , 1: 3.

Genus Archineura KIRBY 1894 Archineura KIRBY, Ann. Mag. nat. Hist., (6) 13: 84. Type species: Archineura basilactea KIRBY, 1894 (original designation). - 1905 Archineura - JACOBSON & BIANKI, Pryam. lozhnos. Ross, i m p . : 801 (type-species: Archineura

incarnata

KARSCH, 1891). - 1933 Leucopteryx FRÄSER, J.

Siam Soc. nat. Hist., Suppl. 9: 125. Type species: Leucopteryx hataerinoides FRASER, 1933 (original designation). - 1935 Archineura - FRASER, Ree. Indian Mus., 37: 329. - 1984 Archineura - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3. - 1984 Leucopteryx - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 (as synym of Archineura KIRBY, 1894).

h e t a e r i n o i d e s (FRASER) 1933 Leucopteryx hetaerinoides FRASER, J.: Siam Soc. nat. Hist., Suppl. 9: 125. Type male: British Museum (Nat.Hist.), London; type-locality: Laos (as generotype of Leucopteryx FRASER, 1933).

9 1966 Archineura hetaerinoides - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 195.

Distribution: Laos, incarnata

(KARSCH)

1891 Echo incarnata KARSCH, Berl. ent. Z., 34: 455. Type male: Museum für Naturkunde, Berlin; type-locality: Kiating (China). 1894 Archineura basilactea KIRBY, Ann. Mag. nat. Hist., (6) 13: 86. Type male: British Museum (Nat. Hist.), London; type-locality: F o o Chow (China) (as generotype of Archineura K I R B Y , 1894). 1894 Archineura basilactea - KARSCH, Ent. Nachr., 20: 84 (proposed synonymy with incarnata KARSCH, 1891).

1896 Archineura incarnata - MCLACHLAN, Ann. Mag. nat. Hist. (6) 17: 370 (description of female). 1 9 0 5 Archineura incarnata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 1 (China).

Distribution: China. Genus Calopteryx

BURMEISTER

1775 Agrion FABRICIUS, Syst. Ent.: 425 (emendation). - 1815 Calepteryx LEACH, Edinb. Encycl.: 9: 137. Type-species: Libellula virgo LINNAEUS, 1758. - 1839 Calopteryx BURMEISTER, H a n d b . Ent., 2: 825. - 1840 Calepteryx H A G E N , Syn. Libell. europ.: 61. - 1850 Calopteryx - SELYS & HÄGEN, Revue des Odonates, Paris: 133. - 1852 Sylphis SELYS, Synopsis des Calopterygines: 8. Type-species: Calopteryx (Sylphis) angustipennis SELYS, 1853 (monotype). - 1876 Agrion - SELYS, Bull. Acad. r. Belg., (2) 41: 1 (revision, key to species). - 1890 Agrion - K I R B Y , Syn.Cat. Neur.-Odon, London: - 1905 Calopteryx - JACOBSON & BIANKI, Pryam. lozhnos, Ross, imp.: 796 (key to species). - 1910 Agrion - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 26. - 1916 Agrion - Ris, Suppl. ent. Berlin, 5: 29 (revision, key to species). - 1919 Agrion - CALVERT, Trans. Am. ent. Soc., 45: 350 (with subgenerea Nehalennia, Ischnura, Enallagma, Pyrrhosoma, Erythagrion, Leptobasis, Anomalagrion, Ceratura). - 1920 Anaciagrion KENNEDY, Ohio J. Sei. Columbus, 21: 83. Type species: Calopteryx (Calopteryx) Cornelia SELYS, 1853 (original designation). - 1950 Calopteryx - MORERA, Inst. Esp. Ent. Madrid: 72 (key to Spanish species). - 1954 Agrion - MONTGOMERY. Annls Ent. Soc. Amer., 47 (3): 471 (The Agrion-Calopteryx Problems). - 1984 Calopteryx - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 3 . - 1984 Calopteryx (Aciagrion) - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 3 .

aequabilis aequabilis SAY 1839 Calopteryx aequabilis SAY, J1 Acad. Philad., 8: 33. Type male: Boston Museum; type-locality: North America. 1873 Agrion aequabile - HAGEN, Proc., Boston Soc. nat. Hist., 15: 274. 1899 Agrion aequabile - KELLICOTT, Odon. Ohio: 9 (redescription). 1900 Agrion aequabile - WILLIAMSON, Drag. Ind.: 252. 1908 Agrion aequabile - WALKER, Ottawa Nat., 22: 3. 1910 Agrion aequabile - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 27. 1913 Agrion aequabile var. coloradicum COCKERELL, Psyche, 20: 173 (male from Colorado). 1917 Agrion aequabile aequabile - KENNEDY, Proc. U.S. natn. Mus., 52: 484.

Distribution: North America. aequabilis californica

(KENNEDY)

1917 Agrion aequabile californicum KENNEDY, Proc. U.S. natn. Mus., 52: 484. Type male: US National Museum, Washington; type-locality: California. 1984 Calopteryx aequabilis californica - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:3.

Distribution: California.

10 aequabilis hudsonica

HAGEN

1853 Calopteryx virginica SELYS, Synopsis des Calopterygines: 11. Type female: Selys's Collection; type-locality: Virginia. 1875 Calopteryx hudsonica HAGEN, Proc. Boston Soc. nat. Hist., 18: 22. Type male: Agassiz Museum; type-locality: Hudson's Bay. 1890 Agrion hudsonica - KIRBY, Syn. Cat. Neur.-Odon. London: 97. 1910 Agrion aequabile hudsonicum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 27 (Lake Superior to Hudson Bay). 1910 Calopteryx virginica - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 27 (proposed synonymy with hudsonica HÄGEN, 1875).

Distribution: U.S.A. aequabilis yakima

HÄGEN

1889 Calopteryx yakima HÄGEN, Psyche, 5: 248. Type male: Agassiz Museum; type-locality: Washington Territory. 1890 Agrion yakima - KIRBY, Syn. Cat. Neur.-Odon. London: 97. 1910 Agrion aequabile yakima - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 27 (Yakima River).

Distribution: North America, amata

HÄGEN 1889 Calopterys amata HÄGEN, Psyche, 5: 244. Type male: Agassiz Museum; type-locality: New Hampshire. 1890 Agrion amata - KIRBY, Syn. Cat. Neur.-Odon. London: 97. 1903 Agrion amatum - NEEDHAM, Bull. 68 N.Y. State Mus., 1903: 222. 1908 Agrion amatum - MUTTKOWSKI, Bull. Wise. Ν . Η . Soc., (2) 6: 66. 1910 Agrion amatum - MUTTKOWSKI, Bull. Publ. Mus., Milwaukee, 1 (1): 26 (U.S.A.: Atlantic States).

Distribution: North America. ancilla

SELYS

1853 Calopteryx ancilla SELYS, Synopsis des Calopterygines: 53. Type male: Selys's Collection; typelocality: E. Europe.

Distribution: East Europe. angustipennis

SELYS

1853 Calopteryx (Sylphis) angustipennis SELYS, Synopsis des Calopterygines: 9. Type male: British Museum (Nat.Hist.), London; type-locality: Georgia. 1853 Calopteryx (Sylphis) elegans HAGEN, in SELYS, Synopsis des Calopterygines: 9. Type female: Agassiz Museum; type-locality: Amer. septentr. (as generotype of Sylphis SELYS, 1853). 1854 Calopteryx (Sylphis) elegans - SELYS & HAGEN, Monogr. Cal.: 20 (redescription). 1890 Agrion elegans - KIRBY, Syn. Cat. Neur.-Odon., London: 96. 1910 Agrion angustipenne - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 28. 1910 Sylphis elegans - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 28 (proposed synonymy w i t h angustipennis

SELYS, 1 8 5 3 ) .

Distribution: U.S.A.: Carolina, Georgia, atrata

SELYS 1853 Calopteryx (Calopteryx) atrata SELYS, Synopsis des Calopterygines: 16. Type male: Rijksmus. Nat. Hist., Leiden; type-locality: China. 1853 Calopteryx (Calopyteryx) smaragdina SELYS, Synopsis des Calopterygines: 16. Type male: British Museum (Nat.Hist.), London; type-locality: India.

11 1854 Calopteryx 1854 Calopteryx 1890 Calopteryx

(Cal.) atrata- SELYS & HAGEN, Monogr. Cal.: 48 (redescription). (Cal.) smaragdina - SELYS & HÄGEN, Monogr. Cal.: 51 (redescription). smaragdina - KIRBY, Syn. Cat. Neur.-Odon., London: 99 (proposed synonymy

w i t h atrata

SELYS, 1 8 5 3 ) .

1931 Calopteryx atrata - SCHMIDT, Konowia, 10: 177 (Kiangs). 1932 Vestaiis tristis NAVÄS, Mus. Heude Notes ent. chinoise, 8: 8. Type not given; type-locality: China. 1935 Vestalaria smaragdina - MAY, Senckenbergiana, 17: 207 (as generotype of Vestalaria MAY, 1935). 1955 Calopteryx atrata - ASAHINA, Ent. Medd., 27 (3): 130 (China). 1962 Calopteryx atratum - CHAO, Acta ent. sinica, Suppl., 11: 27. 1962 Vestalis tristis - CHAO, Acta ertt. sinica, Suppl., 11: 27 (proposed synonymy with atrata SELYS, 1853).

Distribution: China, India. atrocyana

(FRASER)

1935 Agrion atrocyana FRÄSER, Ree. Indian Mus., 37: 330. Type male: British Museum (Nat.Hist.), London; type-locality: Tonkin. 1984 Calopteryx atrocyana - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3

Distribution: Vietnam, China. coomani

(FRASER)

1935 Agrion coomani FRASER, Ree. Indian Mus., 37: 331. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Tonkin. 1984 Calopteryx coomani - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 4.

Distribution: Vietnam. Cornelia

SELYS

1853 Calopteryx (C.) Cornelia SELYS, Synopsis des Calopterygines: 15. Type male: unknown; typelocality: Japan. 1854 Calopteryx (C.) Cornelia - SELYS & HÄGEN, M o n o g r . Cal.: 47 (redescription). 1890 Agrion Cornelia - KIRBY, Syn. Cat. Neur.-Odon. London: 99. 1 9 0 5 Calopteryx

Cornelia

- JACOBSON & BIANKI, P r y a m l o z h n o s . R o s s , i m p . : 799.

1920 Anaciagrion Cornelia - KENNEDY, Ohio J. Sei. Columbus, 21: 83 (as generotype of KENNEDY, 1920). 1 9 8 4 Calopteryx (Anaciagrion) 1: 4.

Anaciagrion

Cornelia - D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d , U t r e c h t ,

Distribution: Japan, dimidiata dimidiata

BURMEISTER

1839 Calopteryx dimidiata BURMEISTER, H a n d b . Ent., 2: 829. Type female: Agassiz Museum; typelocality: Kentucki. 1842 Calopteryx cognata RAMBUR, Hist. nat. Ins. Neuropt. Paris: 222. Type female: Selys's Collection; type-locality: Am. septentr. 1854 Calopteryx (Cal.) dimidiata - SELYS & HÄGEN, Monogr. Cal.: 24 (redescription) (Kentucki, Georgia). 1861 Calopteryx dimidiata - HAGEN, Syn. Neur. N.Amer., Washington: 57 (Kentucky, Georgia, Florida). 1890 Agrion dimidiata - KIRBY, Syn. Cat. Neur.-Odon., London: 97 (U.S.A.). 1898 Calopteryx dimidiata - CALVERT, Trans. Am. ent. Soc., 25: 50. 1910 Agrion dimidiatum dimidiatum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 28.

12 1910 Agrion cognata - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 28 (proposed synonymy with dimidiata dimidiata BURMEISTER, 1839).

Distribution: U.S.A. dimidiata apicale

BURMEISTER

1839 Calopteryx apicalis BURMEISTER, H a n d b . Ent., 2: 827. Type male: Winthem's Collection Agassiz Mus., Cambridge (Mass.); type-locality: Pennsylvania. 1853 Calopteryx (Calopteryx) apicalis - SELYS, Synopsis des Calopterygines: 10. 1854 Calopteryx (Calopteryx) apicalis - SELYS & H A G E N , Monogr. Cal.: 23 (redescription). 1890 Agrion apicalis - KIRBY, Syn. Cat. Neur.-Odon., London: 97. 1898 Agrion apicalis - CALVERT, Trans. Am. ent. Soc., 25: 48. 1910 Agrion dimidiatum apicale - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 29.

Distribution: U.S.A. exul

SELYS

1853 Calopteryx (Calopteryx) exul SELYS, Synopsis des Calopterygines: 12. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Algeria. 1 8 5 4 Calopteryx (Calopteryx) exul - SELYS & H Ä G E N , Monogr. Cal.: 3 4 (redescription). 1887 Calopteryx splendens race exul - SELYS, Ann. Soc. ent. Belg., 31: 39 (Algeria). 1890 Agrion exul - KIRBY, Syn. Cat. Neur.-Odon., London: 97.

Distribution: Algeria, Morocco, Tunisia. grandaeva

HÄGEN

1853 Calopteryx (Calopteryx) grandaeva HAGEN, in SELYS, Synopsis des Calopterygines: 16. Type female: Mus. Naturk., Berlin; type-locality: China. 1 8 5 4 Calopteryx (Calopteryx) grandaeva - SELYS & H Ä G E N , Monogr. Cal.: 5 0 (redescription). 1890 Agrion grandaeva - KIRBY, Syn. Cat. Neur.-Odon. London: 99. 1 9 0 5 Calopteryx grandaeva - JACOBSON, & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 0 (China). 1912 Calopteryx grandaeva - Ris, Suppl. ent. Berlin, 1: 54 (redescription) (Tonkin).

Distribution: China, Vietnam. haemorrhoidalis haemorrhoidalis

(VAN DER LINDEN)

1825 Agrion haemorrhoidalis VAN DER LINDEN, Monogr. Lib. Eur.: 34. Type: unknown; type-locality: Italy: Lazio. 1840 Calopteryx haemorrhoidalis - SELYS, Monogr. Libell. europ., Paris: 133. 1842 Calopteryx haemorrhoidalis - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 221. 1853 Calopteryx (Calopteryx) haemorrhoidalis -SELYS, Synopsis des Calopterygines: 14 (S. Europe, Algeria). 1 8 5 4 Calopteryx (Calopteryx) haemorrhoidalis - SELYS & H Ä G E N , Monogr. Cal.: 4 4 (redescription). 1890 Agrion haemorrhoidalis - KIRBY, Syn. Cat. Neur.-Odon., London: 99. 1940 Calopteryx haemorrhoidalis - NIELSEN, Mem. Soc. ent. ital., 19: 236 (Sardinia). 1950 Calopteryx haemorrhoidalis - MORERA, Inst. Esp., Madrid: 72 (Spain).

Distribution: South Europe, Algeria, Morocco, Tunisia. haemorrhoidalis occasi

CAPRA

1945 Calopteryx haemorrhoidalis occasi CAPRA, Ann. Mus. civ. Stor. nat., Genova, 62: 257. Type male: Mus. civ. Stor. Nat., Genova; type-locality: Italy. Distribution: Italy.

13 hyalina

MARTIN

1 9 8 4 C a l o p t e r y x h y a l i n a MARTIN, 1 9 0 9 - l o c . cit. D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 4.

Distribution: Syria. intermedia intermedia

SELYS

1887 Calopteryx intermedia SELYS, Ann. Soc. ent. Belg., 31: 41. Type female: Selys's Collection; type-locality: Syria. 1905 Calopteryx splendens intermedia - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 798. 1911 Calopteryx intermedia intermedia - BARTENEF, Warszawa Rab. zool. Kab. Univ., 1911: 131.

Distribution: Syria. intermedia cecilia

BARTENEF

1911 Calopteryx intermedia cecilia BARTENEF, Warszawa R a b . zool. Kab. Univ., 1911: 131. Type: unknown; type-locality: Transkaukasica.

Distribution: Transcaucasia. intermedia persica

BARTENEF

1912 Calopteryx intermedia persica BARTENEF, Ann. Mus. Zool. Acad. Imp., Sei. St. Petersbourg, 16: 440. Type male: Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Persia.

Distribution: Persia. japonica

SELYS

1869 Calopteryx japonica SELYS, Synopsis des Calopterygines, Add. 2: 3. Type male: Selys's Collection; type-locality: Japan. 1890 Agrion japonica - KIRBY, Syn. Cat. Neur.-Odon., London: 99. 1905 Calopteryx virgo japonica - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 797.

Distribution: Japan. laosica

FRASER

1933 Calopteryx laosica FRÄSER, J. Siam Soc. nat. Hist. Suppl., 9: 128. Type male: British Museum (Nat. Hist.), London; type-locality: Laos. 1935 Agrion laosica - FRÄSER, Ree. Indian Mus., 37: 331 (Laos, Siam). 1966 Agrion laosica - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 200.

Distribution: Thailand, Laos. maculata maculata

(BEAUVOIS)

1805 Agrion maculata BEAUVOIS, Inst. Afr. Amer.: 85. Type male: British Museum (Nat.Hist.), London; type-locality: not given. 1839 Calopteryx holosericea BURMEISTER, H a n d b . Ent., 2: 828. Type male: Agassiz Mus., Cambridge (Mass.); type-locality: Java (error). 1839 Calopteryx materna SAY, Jn Acad. Philad., 8: 32. Type male: Boston Museum; type-locality: America. 1839 Calopteryx opaca SAY, Jn Acad. Philad., 8: 32. Type male: Boston Museum; type-locality: unknown. 1842 Calopteryx papilionacea RAMBUR, Hist. nat. Ins. Neuropt., Paris: 222. Type: sex not given: Serville's Collection; type-locality: Am. septentr. 1853 Calopteryx (Calopteryx) maculata - SELYS, Synopsis des Calopterygines: 10. 1854 Calopteryx (Calopteryx) maculata - SELYS & HAGEN, Monogr. Cal.: 27 (redescription).

14 1854 Calopteryx opaca - SELYS & H A G E N , Monogr. Cal.: 27 [proposed synonymy with maculata maculata (BEAUVOIS, 1805)]. 1890 Agrion maculata - KIRBY, Syn. Cat. Neur.-Odon., London: 99. 1890 Calopteryx holosericea - KIRBY, Syn. Cat. Neur.-Odon. London: 99 [proposed synonymy with maculata maculata (BEAUVOIS, 1 8 0 5 ) ] . 1890 Calopteryx materna - KIRBY, Syn. Cat. Neur.-Odon., London: 99 [proposed synonymy with maculata

maculata

(BEAUVOIS, 1805)].

1890 Calopteryx papilionacea - KIRBY, Syn. Cat., Neur.-Odon., London: 99 [proposed synonymy with maculata maculata (BEAUVOIS, 1 8 0 5 ) ] .

Distribution: North America. maculata floridana

HUGGINS

1927 Calopteryx maculata floridana HUGGINS, Trans. Am. ent. Soc., 52: 364. Type male: Acad, nat. Sei. Philadelphia; type-locality: Florida.

Distribution: U.S.A.: Florida. melli Ris 1912 Calopteryx melli Ris, Suppl. ent. Berlin, 1: 55. Type male: Königsberg Museum; type-locality: China.

Distribution: China. okinawana

MATSUMURA

1931 Calopteryx okinawana MATSUMURA, 6000 111. Ins. Japan: 1454. Type: not given; type-locality: Japan.

Distribution: Japan. orientalis orientalis

SELYS

1887 Calopteryx splendens race orientalis SELYS, Ann. Soc. ent. Belg., 31: 40. Male from Astrabad. 1905 Calopteryx splendens orientalis -JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 798 (Persia).

Distribution: Syria. orentalis risi

SCHMIDT

1954 Calopteryx orientalis risi SCHMIDT, S.B. öst. Akad. Wiss. (Abt. I.) 163: 242. Type male: Schmidt's Collection; type-locality: Persia.

Distribution: Syria. samarcandica

(BARTENEF)

1911 Agrion samarcandica BARTENEF, Ann. Mus. Zool. Acad. Imp. Sei. St.-Petersbourg, 16: 439. Type male: Zool. Inst.Mus. Acad.Sei., St. Petersburg; type-locality: Turkestan. 1913 Calopteryx maracandica (misprint or new name for samarcandica?) BARTENEF, Rev. russ. ent. 13: 1 8 5 .

Distribution: Turkestan. splendens splendens

(HARRIS)

1782 Libellula splendens HARRIS, Exp. Engl. Ins.: 30. Type male: unknown; type-locality: unknown. 1840 Calopteryx ludoviciana SELYS, Monogr. Libell. europ. Paris: 131. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Belgium. 1840 Calopteryx parthenias CHARPENTIER, Monogr. Lib. eur., Paris: 161. Type male: Halle Museum; type-locality: Germany. 1 8 5 0 Calopteryx splendens - SELYS & H Ä G E N , Revue Odon. Libell. europ., Paris: 1 3 8 .

15 1853 Calopteryx 1853 Calopteryx splendens

(Calopteryx) splendens - SELYS, Synopsis des Calopterygines: 12. ludoviciana - SELYS, Synopsis des Calopterygines: 12 (proposed synonymy with splendens

HARRIS, 1 7 8 2 ) .

1854 Calopteryx (Calopteryx) splendens - SELYS & HAGEN, Monogr. Cal.: 36 (redescription). 1887 Calopteryx splendens - SELYS, Ann. Soc. ent. Belg., 31: 38 (with some races). 1887 Calopteryx splendens race ancilla SELYS (HÄGEN MS), Ann. Soc. ent. Belg., 31: 41. Male from Prusse. 1905 Calopteryx splendens splendens - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 798 (key to subspecies). 1911 Calopteryx splendens var. cretensis PONGRÄCZ, Annls hist.-nat. Mus. natn. hung., 9: 324. Male f r o m Crete. 1915 Calopteryx splendens var. faivrei LACROIX, Insecta, 5: 109. Male f r o m France. 1940 Calopteryx splendens - NIELSEN, Mem. ent. ital., 19: 237 (Sardinia). 1950 Calopteryx splendens - MORERA, Inst. Esp. ent., Madrid: 74 (Spain). 1956 Calopteryx splendens - FRÄSER, R. ent. Soc. Lond., 1 (10): 33 (England). 1960 Calopteryx splendens - HOFFMANN, Archives, 27: 220 (Europe, N o r t h Africa, Asia Minor).

Distribution: Europe, Asia Minor, North Africa. splendens amasina

BARTENEF

1911 Calopteryx amasina BARTENEF, Warszawa Rab. zool. Kab. Univ., 1911:138. Type: unknown; type-locality: Asia Minor.

Distribution: Asia Minor. splendens balcanica

FUDAKOWSKI

1930 Calopteryx splendens balcanica FUDAKOWSKI (APFELBECK in litt.), Ann. Mus. Zool. Pol., 9 (6): 57. Type male: Inst. Biol. Med. Ent., Sarajevo; type-locality: Mostar.

Distribution: Bosnia. splendens caprai

(CONCI)

1984 Agrion splendens caprai CONCI, 1956 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3.

Distribution: Italy. splendens cartvelica

BARTENEF

1930 Calopteryx splendens race cartvelica BARTENEF, Bull. Mus. Georgie, 5:72. Male from Caucasus. 1948 Agrion splendens cartvelica - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 143 (Armenia).

Distribution: Caucasus. splendens erevanensis

(AKRAMOVSKI)

1948 Agrion splendens erevanense AKRAMOVSKI, Zool. Zborn., Moscow, 5: 139. Type male: Zool. Inst. Mus., Yerevan; type-locality: Armenia.

Distribution: Caucasus, splendens johanseni

BELYSHEV

1961 Calopteryx splendens johanseni BELYSHEV, Frag. Faun. Warszawa, 9 (4): 36. Type male: unknown, perhaps in Inst. Zool., Warszawa; type-locality: Kazakhstan.

Distribution: Kazakhstan.

16 splendens mingrelica

SELYS

1868 Calopteryx splendens race mingrelica SELYS, Ann. Soc. ent. Belg., 12:106. Male from Mingrelia (Caucasus). 1890 Calopteryx splendens race mingrelica - KIRBY, Syn. Cat. Neur.-Odon., London: 98. 1 9 0 5 Calopteryx splendens mingrelica - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 8 .

Distribution: Caucasus. splendens pfeifferae

(GÖTZ)

1925 Agrion splendens pfeifFeri GÖTZ, Mitt, münch. ent. Ges., 13: 37. Type male: unknown, perhaps in München Museum; type-locality: Portugal.

Distribution: Portugal. syriaca Rambur 1842 Calopteryx syriaca RAMBUR, Hist. nat. Ins. Neuropt. Paris: 223. Type male: not given, perhaps in Serville's Collection; type-locality: Lebanon. 1887 Calopteryx splendens race syriaca - SELYS, Ann. Soc. ent. Belg., 31: 40 (Syria, Egypt, Mesopotamia). 1890 Calopteryx splendens syriaca - KIRBY, Syn. Cat. Neur.-Odon., London: 97 (incorrect synonymy with dimidiata BURMEISTER, 1 8 3 9 ) . 1 9 0 5 Calopteryx syriaca - JACOBSON & BIANKI, Pryam. lozhnos, Ross.imp.: 7 9 9 (Mesopotamia, Syria, Egypt). 1924 Calopteryx splendens syriaca - MORTON, Trans, ent. Soc. London, 1924: 27. 1938 Calopteryx splendens syriaca - SCHMIDT, Sitz. Akad. Wiss. Wien., (I) 147: 136 (Lebanon).

Distribution: Lebanon, Syria, Asia Minor, Egypt. taurica taurica

SELYS

1853 Calopteryx splendens var. taurica SELYS, Synopsis des Calopterygines: 13. Male from Crimea. 1887 Calopteryx splendens race traurica - SELYS, Ann. Soc. ent. Belg., 31: 40. 1890 Calopteryx splendens var. taurica - KIRBY, Syn. Cat. Neur.-Odon. London: 98 [as synonym of splendens splendens ( H A R R I S , 1 7 8 2 ) ] . 1 9 0 5 Calopteryx splendens taurica - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 8 .

Distribution: Crimea. taurica tshaldirica 1948

(BARTENEF)

Agrion splendens tshaldirica

BARTENEF, 1 9 0 9

- loc. cit.

AKRAMOVSKI,

Zool. Zborn., Moscow,

5: 142.

Distribution: Transcaucasus. terue

(ASAHINA)

1949 Agrion terue ASAHINA, Insecta matsum., 17: 33. Type male: Asahina's Collection; type-locality: Japan.

Distribution: Japan. transcaspica

BARTENEF

1912 Calopteryx transcaspica BARTENEF, Ann. Mus. Zool. Acad. Imp. Sei., St.-Petersbourg, 16:440. Type male: Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Askhabad (Transcaspian country).

Distribution: Lake Caspi.

17 unicolor

BARTENEF

1911 Calopteryx unicolor BARTENEF, Warszawa Rab. zool. Kab. Univ., 1911: 91. Type: unknown; type-locality: Novyj Marghelan.

Distribution: ? Poland, virgo virgo

(LINNAEUS) 1758 Libellula virgo LINNAEUS, Syst. Nat., 1:545. Type male: Halle Museum; type-locality: Germany. 1840 Agrion vesta CHARPENTIER, Libell. Europ. Lipsiae: 136. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Silesia. 1853 Calopteryx (Calopteryx) virgo - SELYS, Synopsis des Calopterygines: 13. 1854 Calopteryx (Calopteryx) virgo - SELYS & H A G E N , Monogr. Cal.: 40 (redescription). 1 8 9 0 Agrion virgo - K I R B Y , Syn. Cat. Neur.-Odon., London: 9 8 (as generotype of Agrion FABRICIUS, 1775). 1898 Calopteryx virgo - CALVERT, Trans. Am. ent. Soc., 25: 50. 1927 Calopteryx virgo ab. fusca VALLE, Acta Soc. Fauna Flora fenn., 56 (11): 7. Male from Finland 1950 Calopteryx virgo - MORERA, Inst. Esp. Ent., Madrid: 73 (Spain). 1956 Calopteryx virgo - FRÄSER, R. Ent. Soc. Lond., 1 (10): 33 (England). 1960 Calopteryx virgo - HOFFMANN, Archives, 27: 220 (Germany, France, Belgium, Luxembourg, Holland).

Distribution: Europe. virgo britannica

CONCI

1952 Calopteryx virgo britannica CONCI, Boll. Mus. 1st. Biol., Genova, 24: 67. Type male: Mus. civ. Stor. nat., Genova; type-locality: England. 1953 Agrion virgo brittanicum - FRASER, Entomologist's month. Mag., 89: 33.

Distribution: England. virgo festiva

(BRULLE)

1832 Agrion festiva BRÜLLE, Exped. Moree, II (1): 103. Type: unknown; type-locality: Greece. 1853 Agrion festiva - SELYS, Synopsis des Calopterygines: 13 (proposed synonymy with virgo virgo (LINNAEUS, 1 7 5 8 ) .

Calopteryx virgo festiva - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 7 (Asia Minor, Greece, Turkestan). 1924 Calopteryx virgo festiva - MORTON, Trans, ent. Soc. London, 1924: 28 (Montenegro).

1905

Distribution: Serbia, Greece, Asia Minor, Turkestan. virgo meridionalis

SELYS

1887 Calopteryx meridionalis SELYS, Ann. Soc. ent. Belg., 34: 171. Type male: Selys's Collection; type-locality: Europe. 1 9 0 5 Calopteryx virgo meridionalis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 7 .

Distribution: South Europe, waterstoni

SCHNEIDER

1984 Calopteryx waterstoni SCHNEIDER, 1984 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 6.

Distribution: Turkey. xanthostoma

(CHARPENTIER)

1825 Agrion xanthostoma CHARPENTIER, Hor. Ent., Paris: 5. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Gallia.

18 1887 Calopteryx splendens race xanthostoma - SELYS, Ann. Soc. ent. Belg., 31: 39 (Spain, France, Corse, Sicilia, Sardinia). 1905 Calopteryx splendens xanthostoma - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 798. 1984 Calopteryx xanthostoma - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 (S. W. Europe).

Distribution: Spain, France, Italy. Genus Echo 1853

SELYS

Echo SELYS, Synopsis des Calopterygines: 1 0 . Type species: Echo margarita SELYS, 1 8 5 3 (monotype). - 1 8 5 4 Echo - SELYS & H A G E N , Monogr. Cal.: 5 5 (with subgenera Echo s.str., Cleis, Sapho, Mnais). -1902 Climacobasis LAIDLAW, Proc. Zool. Soc. London, 1902: 85. Type species: Climacobasis lugens L A I D L A W , 1 9 0 2 (original designation). - 1 9 0 3 Climacobasis - L A I D L A W , Fascic. Malay. Zool., 1: 1 9 1 (proposed synonymy with Echo SELYS, 1 8 5 3 ) . - 1 9 0 5 Echo JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 1 . - 1 9 5 4 Echo - LIEFTINCK, Treubia, 22 Suppl.: 20.

margarita margarita

SELYS

1853 Echo (Echo) margarita SELYS, Synopsis des Calopterygines: 19 (as generotype of Echo SELYS, 1853). Type female: Saunder's Collection; type-locality: China. 1 8 5 4 Echo (Echo) margarita - SELYS & H Ä G E N , Monogr. Cal., 6 7 (redescription) (China). 1890 Echo margarita - KIRBY, Syn. Cat. Neur.-Odon., London: 101 (India). 1 9 0 5 Echo margarita margarita - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 1 (Bengal). 1917 Echo margarita - LAIDLAW, Ree. Indian Mus., 13: 26 (Assam).

Distribution: China, India. margarita tripartita

SELYS

1879 Echo tripartita SELYS, Ann. Soc. ent. Belg., 41: 356. Type female: Selys's Collection; typelocality: Burma. 1 9 0 5 Echo margarita tripartita - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 1 .

Distribution: Burma. modesta

LAIDLAW

1902 Echo modesta LAIDLAW, Proc. Zool. Soc. London, 1: 84. Type female: British Museum (Nat. Hist.), London; type-locality: Kwala Aring (Malaysia). 1902 Climacobasis lugens LAIDLAW, Proc. Zool. Soc. London, 1: 85 (as generotype of Climacobasis LAIDLAW, 1902). Type male: British Museum (Nat. Hist.), London; type-locality: Malay Peninsula. 1903 Climacobasis lugens - LAIDLAW, Fascic. Malay. Zool., 1: 191 (proposed synonymy with modesta LAIDLAW, 1902). 1935 Climacobasis modesta - FRÄSER, Ree. Indian Mus., 37: 332. 1954 Echo modesta - LIEFTINCK, Treubia, 22 Suppl.: 20 (Siam, Malaya). 1984 Echo modesta - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 (S.E. Asia).

Distribution: from Thailand to Malaysia. maxima

MARTIN

1904 Echo maxima MARTIN, Mission Pavie, 3: 219. Type male: Martin's Collection; type-locality: Tonkin.

Distribution: Vietnam.

19

uniformis

SELYS

1879 Echo (Echo) uniformis SELYS, Bull. Acad. r. Belg., (2) 47: 357. Type male: Selys's Collection; type-locality: Sumatra. 1890 Echo ? uniformis - KIRBY, Syn. Cat. Neur.-Odon., London: 101. 1898 Echo uniformis - SELYS, Ann. soc. ent. Belg., 42: 335 («description). 1927 Echo uniformis - Ris, Zool. Meded., Leiden, 10 (1): 10. 1948 Echo uniformis - LIEFTINCK, Treubia, 19: 284. 1954 Echo uniformis - LIEFTINCK, Treubia, 22 Suppl.: 21. 1 9 8 4 Echo iricolor K R Ü G E R , 1 8 9 8 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 ( a s s y n o n y m o f uniformis

SELYS, 1 8 7 9 ) .

Genus Iridictyon 1940 Iridictyon

Amer. Mus. Novit, no. 1081:1. Type species: Iridictyon 1940 (original designation).

N E E D H A M & FISCHER,

N E E D H A M & FISCHER,

myersi

N E E D H A M & FISCHER

myersi

N E E D H A M & FISCHER

1940 Iridictyon myersi N E E D H A M & FISCHER, Amer. Mus. Novit., no. 1081: 1 (as generotype of Iridictyon NEEDHAM & FISCHER, 1940). Type male: U.S. Nation. Mus., Washington; typelocality: British Guyana.

Distribution: Guyana. trebbaui 1984

RACENIS

Iridictyon trebbaui Utrecht, 1: 6.

RACENIS, 1 9 6 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Venezuela. Genus Matrona

SELYS

1853 Calopteryx (Matrona) SELYS, Synopsis des Calopterygines: 17. Type species -.Matrona basilaris SELYS, 1 8 5 3 (original designation). - 1 8 5 4 Calopteryx (Matrona) - SELYS & H A G E N , Monogr. Cal.: 52 (revision). - 1890 Matrona - KIRBY, Syn. Cat. Neur.-Odon., London: 100. - 1891 Matrona - KARSCH, Berl. Ent. Ztg., 34: 456. - 1897 Neurobasis (Matrona) - FÖRSTER, Ann. Soc. ent. Belg., 41: 2 0 6 . - 1 9 0 5 Matrona - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 0 (key to species). - 1 9 2 4 MatronaFRÄSER, J . Asiat. Soc. Bengal, 19: 4 5 7 . - 1 9 5 5 Matrona - ASAHINA, E n t . M e d d . , 27 (3): 130.

basilaris

SELYS

1853 Calopteryx (Matrona) basilaris SELYS, Synopsis des Calopterygines: 17 (as generotype of Matrona SELYS, 1853). Type male: Förster's Collection; type-locality: China. 1 8 5 4 Calopteryx (Matrona) basilaris - SELYS & H Ä G E N , Monogr. Cal.: 5 3 (redescription). 1897 Neurobasis (Matrona) basilaris - FÖRSTER, Ann. Soc. ent. Belg., 41: 208. 1 9 0 5 Matrona basilaris - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 0 (Tibet, Shanghai, Burma). 1916 Matrona basilaris - Ris, suppl. ent. Berl., 5: 5 (redescription) (Tonkin, China, Formosa). 1917 Matrona basilaris - LAIDLAW, Ree Indian Mus., 13: 28 (Assam). 1955 Matrona basilaris - ASAHINA, Ent. Medd., 27 (3): 130.

Distribution: China, Vietnam, Burma, India. grahami

(NEEDHAM)

1930 Agrion grahami NEEDHAM, Zool. Sinica, (A) 11: 195. Type male: unknown, perhaps in Peking Museum; type-locality: China.

20 1984 Matrona grahami - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6.

Distribution: China. japonica

FÖRSTER

1897 Matrona japonica FÖRSTER, Ann. Soc. ent. Belg., 41: 208. Type: unknown; type-locality: Japan. 1905 Matrona japonica - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 800.

Distribution: Japan. kricheldorffi

KARSCH

1891 Matrona kricheldorffi KARSCH, Berl. ent. Z., 34: 456. Type male: Mus. Naturk., Berlin; typelocality: China. 1905 Matrona kricheldorffi - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 800.

Distribution: China. nigripectus

SELYS

1879 Matrona nigripectus SELYS, Bull. Acad.r. Belg., (2) 47: 355. Type: sex not given: Selys's Collection; type-locality: Khasia Hills (India). 1 9 0 5 Matrona nigripectus - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 0 0 . 1984 Matrona basilaris f. nigripectus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6.

Distribution: India. oberthuri

(MCLACHLAN)

1894 Calopteryx oberthuri MCLACHLAN, Ann. Mag. nat. Hist., (6) 13: 433. Type male: British Museum (Nat. Hist.), London; type-locality: West China. 1 9 0 5 Calopteryx oberthuri - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 7 9 9 . 1924 Matrona oberthuri - FRÄSER, J. Asiat. Soc. Bengal, 19: 457. 1928 Matrona oberthuri - MORTON, Trans, ent. Soc. London, 76: 110 (Yunnan).

Distribution: China. Genus Mnais 1853

SELYS

Echo (Mnais) SELYS, Synopsis des Calopterygines: 2 0 . Type species: Mnais pruinosa SELYS, 1 8 5 3 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon. London: 1 0 1 ) . - 1 8 5 4 Echo (Mnais) - SELYS & HAGEN, M o n o g r . Cal.: 65 (revision). - 1890 Mnais - KIRBY, Syn. C a t . N e u r . - O d o n .

London: 1 0 1 . - 1 9 0 5 Mnais - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 0 2 (key to species). - 1 9 1 6 Mnais - Ris, Suppl. ent. Berl., 5 : 8 (key to species). - 1 9 1 7 Mnais - L A I D L A W , Ree. Indian Mus., 13: 29. - 1921 Mnais - MARTIN, Ann. Soc. ent. France, 90: 96. - 1963 Mnais - ASAHINA, Akitu, 11: 21.

andersoni

MCLACHLAN

1873 Mnais andersoni MCLACHLAN, in SELYS, Synopsis des Calopterygines, Add. 3: 8. Type male: McLachlan's Collection; type-locality: Yunnan. 1892 Mnais andersoni - SELYS, Ann. Mus. civ. Stor. nat. Genova, (2) 10: 485 (redescription) (Japan, China). 1 9 0 5 Mnais andersoni - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 2 . 1916 Mnais andersoni - Ris, Suppl, ent. Berlin, 5: 11 (Formosa). 1928 Mnais andersoni - MORTON, Trans, ent. Soc. London, 76: 110 (Yunnan). 1931 Mnais andersoni - SCHMIDT, Konowia, 10: 177 (Tschekiang). 1963 Mnais andersoni - ASAHINA, Akitu, 11: 21 (redescription).

Distribution: China, Taiwan.

21 auripennis

NEEDHAM

1930 Mnais auripennis NEEDHAM, Zool. Sinica, (A) 11: 208. Type male: perhaps in Peking Museum; type-locality: China.

Distribution: China, decolorata

BARTENEF 1912 Mnais decolorata BRATENEF, Ann. Mus. Zool., 17: 306, Type: unknown; type-locality: Kansu.

Distribution: China, earnshawi earnshawi

WILLIAMSON

1904 Mnais earnshawi WILLIAMSON, Proc. U.S. natn. Mus., 28: 185. Type male: perhaps in US natn. Mus., Washington; type-locality: Burma. 1916 Mnais earnshawi - Ris, Suppl. ent. Berlin, 5: 10 (China, Formosa). 1917 Mnais earnshawi - LAIDLAW, Ree. Indian Mus., 13: 29 (Burma). 1935 Mnais earnshawi earnshawi - MAY, Senckenbergiana, 17: 102.

Distribution: China, Taiwan, Burma.

earnshawi thoracicus MAY 1935 Mnais earnshawi thoracicus MAY, Senckenbergiana, 17:102. Type: sex not given; Senckenberg Museum; type-locality: China.

Distribution: China, edai

ASAHINA

1984

Mnais edai

ASAHINA, 1 9 7 6

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 7 .

Distribution: Japan, esakii 1984

ASAHINA

Mnais esakii A S A H I N A , 1 9 7 6 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 7 (as form of pruinosa pruinosa SELYS, 1 8 5 3 ) .

Distribution: Japan, gregoryi

FRASER

1924 Mnais gregoryi FRÄSER, J. Asiat. Soc. Bengal, 19:457. Type male: British Museum (Nat. Hist.), London; type-locality: Yunnan. 1 9 2 8 Mnais gregoryi - M O R T O N , Trans, ent. Soc. London, 7 6 : 1 1 0 (Yunnan) (description of female).

Distribution: China, icteroptera

FRASER

1929 Mnais icteroptera FRASER, J. Bombay nat. Hist. Soc., 33: 592. Type male: British Museum (Nat. Hist.), London; type-locality: Burma.

Distribution: Burma, incolor

MARTIN

1921 Mnais incolor MARTIN, Ann. Soc. ent. France, 90: 96. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Laos.

Distribution: Laos.

22 maclachlani

FRÄSER

1924 Mnais maclachlani nan.

FRÄSER, J .

Asiat. Soc. Bengal, 19: 458. Type unknown; type-locality: Yun-

Distribution: China. mneme Ris 1916 Mnais mneme Ris, Suppl. ent. Berlin, 5: 11. Type female: Naturhist. Riksmus., Stockholm; type-locality: South China.

Distribution: China. pieli

NAVÄS

1936 Mnais pieli NAVÄS, Mus. Heude Notes ent. Chinoise, 3 (4): 42. Type male: not given, perhaps in Heude Museum; type-locality: China.

Distribution: China. pruinosa pruinosa

SELYS

1853 Echo (Mnais) pruinosa SELYS, Synopsis des Calopterygines: 20. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Japan. 1854 Echo (Mnais) pruinosa - SELYS & H A G E N , Monogr. Cal.: 65 (redescription). 1 8 9 0 Mnais pruinosa - K I R B Y , Syn. Cat. Neur.-Odon. London: 1 0 1 (as generotype of Mnais SELYS, 1853).

1984 Mnais pruinosa pruinosa - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 7.

Distribution: Japan. pruinosa costalis

SELYS

1869 Mnais costalis SELYS, Synopsis des Calopterygines, A d d . 2: 7. Type male: Selys's Collection; type-locality: Japan. 1916 Mnais costalis - Ris, Suppl. ent. Berlin, 5: 9 (redescription). 1 9 8 4 Mnais pruinosa costalis f. ogumi A S A H I N A , 1 9 7 6 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 7.

Distribution: Japan. pruinosa nawai

ASAHINA

1984 Mnais pruinosa nawai A S A H I N A , 1976 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:7 with forms taketoi A S A H I N A , 1976, sahoi ASAHINA 1976, kadowakii A S A H I N A , 1976.

Distribution: Japan. semiopaca MAY 1935 Mnais semiopaca MAY, Senckenbergiana, 17: 103. Type male: Senckenberg Museum; typelocality: China.

Distribution: China. strigata

HÄGEN

1853 Echo (Mnais) strigata HAGEN, in SELYS, Synopsis des Calopterygines: 20. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Japan. 1854 Echo (Mnais) strigata - SELYS & H Ä G E N , Monogr. Cal.: 64 (redescription). 1890 Mnais strigata - KIRBY, Syn. Cat. Neur.-Odon. London: 101. 1916 Mnais strigata - Ris, Suppl. ent. Berlin, 5: 10.

Distribution: Japan.

23 tenuis

OKUMA

1913 Mnais tenuis OKUMA, Dobuts. Z. Tokyo, 25: 307. Type male: Hokkaido Univ.; type-locality: Japan. 1955 Mnais tenuis - Asahina, Ent. Medd., 27 (3): 129 (North China). 1961 Mnais tenuis - ASAHINA, Insecta matsum., 24: 59 (Taiwan).

Distribution: Japan, China, Taiwan. Genus Neurobasis

SELYS

1853 Neurobasis SELYS, Synopsis des Calopterygines: 17. Type-species: Libellula chinensis LINNAEUS, 1758 (monotypy). - 1854 Neurobasis - SELYS & HAGEN, Monogr. Cal.: 72. - 1897 Neurobasis (Matronides) FÖRSTER, Wiener ent. Ztg., 16: 101. Type species: Matronides cyaneipennis FÖRSTER, 1897 (original designation). - 1897 Neurobasis - Ann. Soc. ent. Belg., 41: 206 (with subgenera Matrona, Matronides). - 1905 Neurobasis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 796. - 1955 Neurobasis - LIEFTINCK, Nova Guinea (N.S.) 6: 155 (revision of Austroasian species).

anderssoni

SJÖSTEDT

1926 Neurobasis anderssoni SJÖSTEDT, Ent. Tidskr., 47: 247. Type male: Naturhist. Riksmus., Stockholm; type-locality: China. 1931 Neurobasis anderssoni - SCHMIDT, Konowia, 10: 177 (Tsche-kiang).

Distribution: China. apicalis

KIRBY

1891 Neurobasis apicalis KIRBY, Proc. Zool. Soc. London, 1891: 294. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon.

Distribution: Sri Lanka. australis australis

SELYS

1897 Neurobasis australis SELYS, Ann. Soc. ent. Belg., 41: 428. Type male: unknown; type-locality: New Guinea. 1898 Neurobasis paradisearum FÖRSTER, Termeszetr. Füz., Budapest, 21: 277. Type male: Termeszett. Μύζ., Budapest; type-locality: New Guinea. 1932 Neurobasis leopoldi FRASER, Mem. Mus. Hist. nat. Belg., 4 (3): 12. Type male: Fraser's Collection; type-locality: New Guinea. 1949 Neurobasis australis - LIEFTINCK, Nova Guinea (N.S.) 5: 15. 1949 Neurobasis leopoldi- LIEFTINCK, Nova Guinea (N. S.) 5:15 (proposed synonymy with australis australis

SELYS, 1 8 9 7 ) .

1955 Neurobasis australis australis - LIEFTINCK, Nova Guinea (N.S.) 6: 161. 1984 Neurobasis paradisearum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 (proposed synonymy with australis australis SELYS, 1897).

Distribution: New Guinea. australis misoolensis

LIEFTINCK

1955 Neurobasis misoolensis LIEFTINCK, Nova Guinea, (N.S.) 6: 161. Type male: Lieftinck's Collection; type-locality: Misool Island.

Distribution: Indonesia: Misool Island. chinensis chinensis

(LINNAEUS)

1758 Libellula chinensis LINNAEUS, Syst. Nat., 1: 545. Type male: unknown; type-locality: Tibet.

24 1776 Agrion notabilitata FABRICIUS, Gen. Ins.: 248. Type male: Univ. zool. Mus., Kobenhavn; typelocality: India. 1842 Calopteryx disparilis RAMBUR, Hist. nat. Ins. Neuropt., Paris: 224. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: not given. 1842 Calopteryx chinensis - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 226. 1853 Neurobasis chinensis - SELYS, Synopsis des Calopterygines: 18 (as generotype of Neurobasis SELYS, 1853).

1853 Calopteryx sinensis var. florida WALKER, List. Neur. Ins. B.M., 4: 603. Male from Ceylon. 1 8 5 4 Neurobasis chinensis - SELYS & H A G E N , Monogr. Cal.: 7 2 (redescription). 1869 Neurobasis florida - SELYS, Synopsis des Calopterygines, Add. 2: 4 (Ceylon). 1897 Neurobasis chinensis - SELYS, Ann. Soc. ent. Belg., 41: 428 (with some races). 1 9 0 5 Neurobasis chinensis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 6 . 1924 Neurobasis chinensis - FRÄSER, Ree. Indian Mus., 26: 479 (West India). 1927 Neurobasis chinensis - Ris, Zool. Meded., Leiden, 10 (1): 10 (Sumatra). 1934 Neurobasis chinensis chinensis - SCHMIDT, Arch. Hydrob. Suppl., 13: 331 (Sumatra). 1934 Neurobasis chinensis florida - SCHMIDT, Arch. Hydrob. Suppl., 13: 331 (Java). 1934 Neurobasis chinensis florida - LIEFTINCK, Treubia, 14 (4): 383 (Java). 1935 Neurobasis chinensis chinensis - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 3 (Sumatra). 1941 Neurobasis chinensis - LIEFTINCK, Treubia, 17: 340 (key to Malaysian subspecies). 1954 Neurobasis chinensis chinensis - LIEFTINCK, Treubia, 22 Suppl.: 23 (Siam, Malaysia, Sumatra). 1954 Neurobasis chinensis florida - LIEFTINCK, Treubia, 22 Suppl.: 24 (Java). 1955 Neurobasis chinensis chinensis - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69 (Ceylon). 1964 Neurobasis chinensis chinensis - SCHMIDT, Ent. Tidskr., 85: 153 (Burma) 1965 Neurobasis chinensis chinensis - ASAHINA, Kontyü, 33 (4): 498 (Hong-Kong). 1984 Neurobasis chinensis f. florida - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8.

Distribution: Hong-Kong, South China, India, Sri Lanka, Malaysia, Sumatra, Java. chinensis longipes

HÄGEN

1887 Neurobasis longipes HAGEN, Abh. Zool.-Bot. Ges. Wien, 37:648. Type male: unknown, perhaps in Naturhist. Mus., Wien; type-locality: Borneo. 1936 Neurobasis longipes - COOMANS DE R U I T E R , Trop. Natuur, 25: 72. 1941 Neurobasis chinensis longipes - LIEFTINCK, Treubia, 17: 338. 1954 Neurobasis chinensis longipes - LIEFTINCK, Treubia, 22 Suppl.: 24 (Borneo).

Distribution: Borneo. cyaneipennis

FÖRSTER

1897 Neurobasis Matronides 1897 Neurobasis of female). 1912 Matronides 1954 Neurobasis

(Matronides) cyaneipennis FÖRSTER, Wiener ent. Z., 16: 101 (as generotype of FÖRSTER, 1897). Type male: Naturhist. Mus., Wien; type-locality: Borneo. (Matronides) cyaneipennis - FÖRSTER, Ann. Soc. ent. Belg., 41: 207 (description cyaneipennis - LAIDLAW, J. Str. Br. R. Asiat. Soc., 63: 93 (Borneo). cyaneipennis - LIEFTINCK, Treubia, 22 Suppl.: 24 (Borneo).

Distribution: Borneo. ianthinipennis

LIEFTINCK

1949 Neurobasis ianthinipennis LIEFTINCK, Nova Guinea (N.S.) 5: 22. Type male: Lieftinck's Collection; type-locality: New Guinea.

Distribution: New Guinea.

25 kaupi kaupi

BRAUER

1867 Neurobasis kaupi BRAUER, Verh. Zool.-bot. Ges. Wien, 17: 293. Type male: Darmstadt Museum; type-locality: Celebes. 1897 Neurobasis chinensis race kaupi - SELYS, Ann. Soc. ent. Belg., 41: 429. 1898 Neurobasis chinensis race kaupi - SELYS, Ann. Soc. ent. Belg., 42: 338. 1955 Neurobasis kaupi kaupi - LIEFTINCK, Nova Guinea (N.S.) 6: 158.

Distribution: Celebes. kaupi pavo

LIEFTINCK

1955 Neurobasis kaupi pavo LIEFTINCK, Nova Guinea (N.S.) 6: 158. Type male: Lieftinck's Collection; type-locality: Celebes.

Distribution: Celebes. kimminsi

LIEFTINCK

1955 Neurobasis kimminsi LIEFTINCK, Nova Guinea (N.S.) 6:163. Type male: Lieftinck's Collection; type-locality: New Guinea.

Distribution: New Guinea. luzoniensis

SELYS

1879 Neurobasis luzoniensis SELYS, Bull. Acad. r. Belg., (2) 47: 360. Type male: Selys's Collection; type-locality: Luzon. 1882 Neurobasis kaupi race luzoniensis - SELYS, Ann. Soc. Esp. Hist, nat., 11: 19. 1939 Neurobasis luzoniensis - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 248 (Luzon).

Distribution: Philippine Islands. Genus Phaon

SELYS

1853 Phaon SELYS, Synopsis des Calopterygines: 23. Type species: Calopteryx iridipennis BURMEISTER, 1839 ( m o n o t y p y ) . - 1854 Phaon - SELYS & HAGEN, M o n o g r . Cal.: 68 (with s u b g e n e r a

Phaon s.str., Neurobasis). - 1890 Phaon - KIRBY, Syn. Cat. Neur.-Odon. London: 101. 1941 Prophaon FRÄSER, Proc. R. ent. Soc. London, (B) 10:40. Type species: Phaon camerunensis SJÖSTEDT, 1900 (original designation). - 1962 Phaon - PINHEY, Publ. cult. Comp. Diam. Angola, 59:143. - 1962 Prophaon - PINHEY, Publ. cult. Comp. Diam. Angola, 59:143 (proposed s y n o n y m y w i t h Phaon

camerunensis

SELYS, 1853).

(SJÖSTEDT)

1900 Libellago camerunensis SJÖSTEDT, Bih. Svenska Akad. Handl., 25 (2): 56. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1941 Prophaon camerunensis - FRASER, Proc. R. ent. Soc. London, (B) 10: 40 (as generotype of Prophaon

FRASER, 1941).

1949 Libellago camerunensis -FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 47 (among incertae sedis). 1962 Phaon camerunensis - PINHEY, Publ. cult. comp. Diam. Angola, 59: 144 (Cameron, Nigera, Liberia). 1962 Phaon fraseri PINHEY, Publ. cult. Comp. Diam. Angola, 59: 143. Type male: unknown; typelocality: Cameroon. 1984 Phaon fraseri - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 [proposed synonymy with camerunensis (SJÖSTEDT, 1900)].

Distribution: Cameroon, Nigeria, Liberia.

26 iridipennis iridipennis

(BURMEISTER)

1839 Calopteryx iridipennis BURMEISTER, H a n d b . Ent., 2: 827. Type male: Drege's Collection in Halle Museum; type-locality: Port Natal. 1842 Euphaea iridipennis - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 232. 1853 Phaon iridipennis - SELYS, Synopsis des Calopterygines: 24 (as generotype of Phaon SELYS, 1853). 1854 Phaon (Phaon) iridipennis - SELYS & H A G E N , Monogr. Cal.: 78 (redescription). 1898 Phaon iridipennis - CALVERT, Trans. Am. ent. Soc., 25: 49. 1906 Sapho iridipennis occidentalis FÖRSTER, Wiesbaden Jb Ver. Natk., 59: 330. Type: unknown. 1921 Phaon iridipennis - Ris, Annls S. Afr. Mus., 18: 264 (Zululand, Transvaal) (redescription). 1958 Phaon iridipennis - PINHEY, Stuttg. Beitr. Naturk., 1958 (10): 2 (trop. and subtropical Africa). 1959 Phaon iridipennis - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 30. 1962 Phaon iridipennis - PINHEY, Publ. cult. Comp. Diam. Angola, 59:143 (tropical and subtropical Africa).

Distribution: tropical and subtropical Africa, iridipennis fuliginosus

SELYS

1879 Phaon iridipennis race fuliginosus SELYS, Bull. Acad. r. Belg., (2) 47: 359. Male from Madagascar. 1962 Phaon iridipennis fuliginosus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144. 1984 Phaon iridipennis f.fuliginosus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8.

Distribution: Madagascar. rasoherinae

FRASER

1949 Phaon rasoherinae FRASER, Mem. Inst. sei. Madagascar, 3A: 32. Type male: Inst. Rech. Sei., Madagascar; type-locality: Madagascar. 1962 Phaon rasoherinae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144.

Distribution: Madagascar. Genus Psolodesmus

MCLACHLAN

1870 Psolodesmus MCLACHLAN, Trans, ent. Soc. London, 1870: 165. Type species: Psolodesmus mandarinus M C L A C H L A N , 1 8 7 0 (monotypy). - 1 8 7 3 Echo (Psolodesmus) - SELYS, Synopsis des Calopterygines, Add.: 3: 9. - 1890 Psolodesmus - KIRBY, Syn. Cat. Neur.-Odon., London: 1 0 1 . - 1 9 0 5 Psolodesmus - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 0 2 . - 1 9 1 6 Psolodesmus - Ris, Suppl. ent. Berlin, 5: 1 1 . - 1961 Psolodesmus - ASAHINA, Insecta matsum., 24: 5 9 .

mandarinus mandarinus

MCLACHLAN

1870 Psolodesmus mandarinus MCLACHLAN, Trans, ent. Soc. London, 1870: 165 (as generotype of Psolodesmus MCLACHLAN, 1870). Type male: McLachlan's Collection; type-locality: China. 1873 Echo (Psolodesmus) mandarinus - SELYS, Synopsis des Calopterygines, Add. 3: 10. 1890 Psolodesmus mandarinus - KIRBY, Syn. Cat. Neur. - Odon., London: 101. 1 9 0 5 Psolodesmus mandarinus - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 3 (China). 1984 Psolodesmus mandarinus mandarinus—DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8.

Distribution: China, mandarinus dorothea

WILLIAMSON

1904 Psolodesmus dorothea WILLIAMSON, Ent. News, Philad., 15: 249. Type female: Williamson's Collection; type-locality: Formosa. 1916 Psolodesmus dorothea - Ris, Suppl. ent. Berlin, 5: 11 (Formosa).

27 1984 Psolodesmus mandarinus dorothea - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 .

Distribution: Taiwan, mandarinus kuroiwae 1913 Psolodesmus 5: 153. Type 1950 Psolodesmus 1961 Psolodesmus

MATSUMURA

dorothea kuroiwae MATSUMURA, in OGUMA, J. Coll. agric. Tohuku, Imp. Univ., male: Hokkaido Univ.; type-locality: Japan. mandarinus kuroiwae - CHEN, Quart. J. Taiwan Mus., 3: 28. mandarinus kuroiwae - ASAHINA, Insecta matsum., 24: 59.

Distribution: Japan. Genus Sapho

SELYS

1853 Echo (Sapho) SELYS, Synopsis des Calopterygines: 21. Type species:Agrion ciliata FABRICIUS, 1781 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 100). - 1854 Echo (Sapho) - SELYS & HAGEN, Monogr. Cal.: 57 (revision). - 1869 Sapho - MCLACHLAN, Entomologist's month. Mag., 6: 27. - 1890 Sapho - KIRBY, Syn. Cat. Neur.-Odon., London: 100. - 1919 Sapho - MUNZ, Mem. Amer. ent. Soc., 3 (1): 43. - 1962 Sapho - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145.

bicolor

SELYS

1853 Echo (Sapho) bicolor SELYS, Synopsis des Calopterygines: 21. Type male: R. Mus., Brussels; type-locality: Guinea. 1854 Echo (Sapho) bicolor - SELYS & HÄGEN, Monogr. Cal.: 61 (redescription). 1890 Sapho bicolor - KIRBY, Syn.Cat.Neur.-Odon., London: 100. 1917 Sapho superba SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 10. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1962 Sapho bicolor - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (Guinea, Ghana). 1962 Sapho superba - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (Cameroon, Liberia). 1984 Sapho superba - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 (proposed synonymy with bicolor SELYS, 1853).

Distribution: West Africa. ciliata

(FABRICIUS)

1781 Agrion ciliata FABRICIUS, Spec. Ins., 1: 528. Type female: British Museum (Nat. Hist.), London; type-locality: Sierra Leone. 1853 Echo (Sapho) ciliata - SELYS, Synopsis des Calopterygines: 22 (Guinea, Sierra Leone). 1853 Echo (Sapho) ciliata ab. fumipenne SELYS, Synopsis des Calopterygines: 22. Male from Sierra Leone. 1854 Echo (Sapho) ciliata - SELYS & HÄGEN, Monogr. Cal.: 58 (redescription). 1890 Sapho ciliata - KIRBY, Syn. Cat. Neur.-Odon., London: 100 (as generotype of Sapho SELYS, 1853).

1947 Sapho ciliata - Fraser, Trans. R. ent. Soc. London, 98: 21 (Ivory Coast). 1962 Sapho ciliata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (Sierra Leone, Ivory Coast, Gold Coast, Togo, Nigeria).

Distribution: West Africa. fumosa

LONGFIELD

1932 Sapho fumosa LONGFIELD, Stylops, 1: 206. Type male: British Museum (Nat. Hist.), London; type-locality: Sierra Leone.

28 1962 Sapho fumosa - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (Sierra Leone, Portuguese Guinea, French Guinea).

Distribution: West Africa. gloriosa SELYS 1873 Sapho gloriosa SELYS (MCLACHLAN in litt.), Synopsis des Calopterygines, Add. 2: 58. Type male: McLachlan' Collection; type-locality: Gabon. 1917 Sapho gloriosa - SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 11 (Cameroon). 1934 Sapho gloriosa - SCHOUDETEN, Ann. Mus. Congo Beige, Zool., (3) II 3: 72 (Belgian Congo). 1962 Sapho gloriosa — PINHEY, Publ. cult. Comp. Diam. Angola, 59:46 (Gabon, French Cameroon, Belgian Congo).

Distribution: West Africa. orichalcea M C L A C H L A N 1869 Sapho orichalcea MCLACHLAN, Entomologist's month. Mag., 6: 27. Type male: McLachlan's Collection [in British Museum (Nat. Hist.), London]; type-locality: West Africa. 1869 Sapho orichalcea - SELYS, Synopsis des Calopterygines, Add. 2: 6 (redescription). 1941 Sapho orichalcea - FRÄSER, Proc. R. ent. Soc. London, (B) 10: 40 (Cameroon). 1962 Sapho orichalcea - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (Belgian Congo, Cameroon, South Nigeria, Spanish Guinea).

Distribution: West Africa. venusta KARSCH 1889 Sapho venusta KARSCH, Ent. Nachr., 15 (15): 233. Type female: Mus. Naturk., Berlin; typelocality: Sierra Leone. 1962 Sapho venusta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (Cameroon).

Distribution: Sierra Leone, Cameroon. Genus Umma KIRBY 1853 Cleis SELYS, Synopsis des Calopterygines: 22. Type species: Cleis cincta SELYS, 1853 (monotype). - 1890 Umma KIRBY, Syn. Cat. Neur.-Odon., London: 100 (new name for Cleis SELYS, 1853, a junior primary homonymy of Cleis GUERIN, 1831 in Lepidoptera et Cleis MULSANT, 1850 in Coleoptera). - 1922 Umma - NAVAS, Treb. Mus. Ci. nat. Barcelona, 4: 113. - 1933 Umma - LONGFIELD, Stylops, 2: 139. - 1934 Umma - SCHOUDETEN, Ann. Mus. Congo Beige, Zool., (3) II 3: 72. - 1962 Umma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144.

cincta (HÄGEN) 1853 Phaon (Cleis) cincta HAGEN, in SELYS, Synopsis des Calopterygines: 23. Type series of males, not designated: R. Mus. Brussels; type-locality: Guinea. 1853 Calopteryx cincta - WALKER, List. Neur. Ins. B.M., 4: 607. 1890 Umma cincta - KIRBY, Syn. Cat. Neur.-Odon., London: 100 (as generotype of Umma KIRBY, 1890).

1917 Umma cincta - SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 7 (Congo). 1962 Umma cincta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144 (Guinea, Ivory Coast, Gold Coast, Togo, Nigeria, Belgian Congo, Uganda).

Distribution: tropical Africa. declivia FÖRSTER 1906 Umma declivia FÖRSTER, Jb. Mannheim Ver. Natk., 71-72: 51. Type male: Ann Arbor Museum; type-locality: Usambara.

29 1962 Umma declivia - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144 (Tanganyika, Togo).

Distribution: Central Africa. distincta

LONGFIELD

1933 Umma distincta LONGFIELD, Stylops, 2:139. Type male: British Museum (Nat. Hist.), London; type-locality: North Rhodesia. 1962 Umma distincta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144 (Belgian Congo, Angola, North Rhodesia).

Distribution: Central Africa. electa

LONGFIELD

1933 Umma electa type-locality: 1934 Umma electa 1962 Umma electa

LONGFIELD, Stylops, 2: 139. Type male: British Museum (Nat.Hist.), London; Belgian Congo. - SCHOUDETEN, Ann. Mus. Congo Beige, Zool., (3) II 3: 73 (Belgian Congo). - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 144 (Congo, Angola).

Distribution: West Africa femina

LONGFIELD

1947 Umma femina LONGFIELD, Arq. Mus. Bocage, 16:20. Type female: British Museum (Nat. Hist.), London; type-locality: Angola. 1962 Umma femina - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145.

Distribution: Angola. infumosa

FRÄSER

1951 Umma infumosa FRÄSER, Rev. franc. Ent., 18; 98. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: French Guinea. 1962 Sapho infumosa - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 [a race of form o f f u m o s a LONGFIELD, 1 9 3 2

(Sapho)!].

Distribution: French Guinea. longistigma

(SELYS)

1869 Sapho longistigma SELYS, Bull. Acad. r. Belg., (2) 27: 650. Type series: in R. Mus. Brussels, holotype not designated; type-locality: Old Calabar. 1899 Umma longistigma - KARSCH, Ent. Nachr., 25: 161 (Cameroon). 1941 Umma longistigma - FRÄSER, Proc. R. ent. Soc. London, (B) 10: 40 (Cameroon). 1962 Umma longistigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145 (Cameroon, Belgian Congo, South Nigeria, Spanish Guinea).

Distribution: West Africa. mesostigma

(SELYS)

1879 Phaon (Cleis) mesostigma SELYS, Bull. Acad. r. Belg., (2) 47: 358. Type male: R. Mus., Brussels; type-locality: Cameroon. 1879 Phaon (Cleis) monostigma SELYS, Bull. Acad. r. Belg., (2) 47: 358. Type female: R. Mus., Brussels; type-locality: Cameroon. 1890 Umma monostigma - KIRBY, Syn. Cat. Neur.-Odon., London: 100 (lapsus calami). 1899 Umma fuscomarginis SJÖSTEDT, Bih. Svenska Vet. Akad., 25 (2): 45. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1922 Umma splendida NAVÄS, Treb. Mus. Ci. nat., Barcelona, 4:113. Type male: Mus. Cienc. Nat., Madrid; type-locality: Fernando Po. 1934 Umma fuscipleuris SCHOUDETEN, Ann. Mus. Congo Beige, Zool., 3 (1): 73 (error pro fuscomarginis SJÖSTEDT, 1899).

30 1950 Umma fuscomarginea comarginis

- FRÄSER, Bull. Inst, franc. Afr. noire, 12: 618 (lapsus calami pro fus-

SJÖSTEDT, 1 8 9 9 ) .

1962 Umma mesostigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145 (Congo, Cameroon, Spanish Guinea, Dahomey). 1962 Umma splendida - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145 (Spanish Guinea). 1984 Umma splendida - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 [proposed s y n o n y m y w i t h mesostigma

(SELYS, 1879)].

Distribution: tropical Africa, puella

SJÖSTEDT

1917 Umma puella SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 7. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1950 Sapho puella - FRÄSER, Bull. Inst, franc. Afr. noire, 12: 618. 1962 Umma puella - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145 (Cameroon, Congo).

Distribution: Central Africa. purpurea P I N H E Y I960 Umma purpurea Pinhey, Entomologist's month. Mag., 96: 264. Type male: N a t n . Mus., Bulawayo; type-locality: British Cameroon.

Distribution: Cameroon. saphirina

FÖRSTER

1916 Umma saphirina FÖRSTER, Neue Beitr. syst. Insektenst., 1: 23. Type male: Förster's Collection; type-locality: Cameroon. 1950 Umma saphirina - KIMMINS, Ann. Mag. nat. Hist., (12) 3: 172 (redescription). 1962 Umma saphirina - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 145 (Uganda, Congo).

Distribution: Central Africa. Genus Vestalis

SELYS

1853 Vestalis SELYS, Synopsis des Calopterygines: 24. Type species: Calopteryx luctuosa BURMEISTER, 1839 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 102). - 1 8 5 4 Vestalis SELYS & HAGEN, Monogr. Cal.: 79 (revision). - 1890 Vestalis - KIRBY, Syn. Cat. Neur.-Odon., London: 102. - 1905 Vestalis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 803 (key to species). - 1920 Vestinus KENNEDY, Ohio J. Sei. Columbus, 21: 83. Type species: Calopteryx gracilis RAMBUR, 1842 (original designation). - 1929 Vestalis - FRÄSER, J. Bombay nat. Hist. Soc., 33: 581 (key to Indian species). - 1935 Vestalaria MAY, Senckenbergiana, 17: 207. Type species: Vestalis smaragdina SELYS, 1897 (original designation). - 1954 Vestalis - LIEFTINCK, Treubia, 22 Suppl.: 2 1 . - 1 9 5 4 Vestinus-LIEFTINCK, Treubia, 22 Suppl.: 21 (proposedsynonymy w i t h Vestalis

SELYS, 1 8 5 3 ) . - 1 9 8 4 Vestalis

(Vestalis)

- DAVIES & TOBIN, T h e D r a g o n f l i e s o f

the World, Utrecht, 1: 9. - 1984 Vestalis (Vestinus) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9. - 1984 Vestalis (Vestalaria) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9.

amabilis

LIEFTINCK

1965 Vestalis amabilis LIEFTINCK, Tijdschr. Ent., 108: 349. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1984 Vestalis (Vestinus) amabilis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9.

Distribution: Borneo.

31 amaryllis

LIEFTINCK

1965 Vestalis amaryllis LIEFTINCK, Tijdschr. Ent., 108: 345. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Indonesia: Bangka Island. 1984 Vestalis (Vestinus) amaryllis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9.

Distribution: Indonesia: Bangka Island. amethystina

LIEFTINCK

1965 Vestalis amethystina LIEFTINCK, Tijdschr. Ent., 108: 343. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Thailand. 1984 Vestalis (Vestinus) amethystina - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9. (Borneo).

Distribution: Thailand, ? Borneo. amnicola

LIEFTINCK

1965 Vestalis amnicola LIEFTINCK, Tijdschr. Ent., 108: 349. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1 9 8 4 Vestalis ( Vestinus) amnicola - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 .

Distribution: Borneo. amoena

HÄGEN

1853 Vestalis amoena HAGEN, in SELYS, Synopsis des Calopterygines: 25. Type male: Schneider & Westerman's Collection; type-locality: Sumatra. 1 8 5 4 Vestalis amoena - SELYS & H A G E N , Monogr. Cal.: 82 (redescription). 1920 Vestinus amoena - KENNEDY, Ohio J. Sei. Columbus, 21: 83. 1927 Vestalis amoena - Ris, Zool. Meded., Leiden, 10 (1): 10. 1934 Vestinus amoenus - SCHMIDT, Arch. Hydrob., Suppl., 13: 333. 1935 Vestalis amoena „ S E L Y S " (misprint) - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 3. 1954 Vestalis amoena - LIEFTINCK, Treubia, 22 Suppl.: 21 (Siam, Sumatra, Borneo, Malaya). 1965 Vestalis amoena - LIEFTINCK, Tijdschr. Ent., 108: 339 (redescription, revision, and key to amoenagroup). 1984 Vestalis (Vestinus) amoena - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 (S.E. Asia).

Distribution: Southeast Asia. anacolosa

LIEFTINCK

1965 Vestalis anacolosa LIEFTINCK, Tijdschr. ent., 108: 353. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: North Borneo. 1 9 8 4 Vestalis ( Vestinus) anacolosa - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 .

Distribution: Borneo. apicalis apicalis

SELYS

1873 Vestalis apicalis SELYS, Bull. Acad. r. Belg., (2) 47: 362. Type male: Oxford Museum; typelocality: East India. 1 9 0 5 Vestalis apicalis - JACOBSON & BIANKI, Pryam. lozhnos, Ross, imp.: 8 0 3 (India). 1924 Vestalis apicalis - FRÄSER, Ree. Indian Mus., 26: 479. 1 9 8 4 Vestalis ( Vestinus) apicalis apicalis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 .

Distribution: India.

32 apicalis nigrescens

FRÄSER

1929 Vestalis nigrescens FRÄSER, J. Bombay nat. Hist. Soc., 33: 584. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1955 Vestalis (Vestinus) apicalis nigrescens - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69.

Distribution: Sri Lanka. apicalis submontana

FRASER

1929 Vestalis apicalis amoena FRÄSER. J. Bombay nat. Hist. Soc., 33: 584. Type male: British Museum (Nat. Hist.), London; type-locality: Nilgiris (India). 1934 Vestalis apicalis submontana FRASER, F a u n a brit. India, Odon., 2: 130 (new name for Vestalis apicalis amoena FRASER, 1929, a junior secondary homonymy of Vestalis gracilis amoena FRASER, 1929).

Distribution: India. atropha

LIEFTINCK

1965 Vestalis atropha LIEFTINCK, Tijdschr. Ent., 108: 351. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1984 Vestalis (Vestinus) atropha - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9.

Distribution: Borneo. beryllae

LAIDLAW

1915 Vestalis beryllae LAIDLAW, J. Sarawak Mus., 2: 273. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1 9 2 3 Vestinus beryllae - L A I D L A W , J . Mal. Br. R . Asiatic Soc., 1: 3 3 2 . 1954 Vestalis beryllae - LIEFTINCK, Treubia, 22 Suppl.: 22 (Borneo). 1965 Vestalis beryllae - LIEFTINCK, Tijdschr. Ent., 108: 355 (Borneo). 1 9 8 4 Vestalis (Vestinus) beryllae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 .

Distribution: Borneo. gracilis gracilis

(RAMBUR)

1842 Calopteryx gracilis RAMBUR, Hist. nat. Ins. Neuropt., Paris: 224. Type: sex not given: Serville's Collection; type-locality: Bombay (India). 1853 Vestalis gracilis - SELYS, Synopsis des Calopterygines: 26. 1854 Vestalis gracilis - SELYS & HAGEN, Monogr. Cal.: 84 (redescription) (Tibet, Conchinchina, Bombay). 1 9 0 5 Vestalis gracilis - JACOBSON & BIANKI, Pryam. lozhnos Ross, imp.: 8 0 3 (India). 1920 Vestalis ( Vestinus) gracilis- KENNEDY, Ohio J. Sei. Columbus, 21: 83 (as generotype of Vestinus KENNEDY, 1 9 2 0 ) .

1924 1934 1954 1984

Vestalis Vestalis Vestalis Vestalis 1: 10.

gracilis - FRASER, Ree. Indian Mus., 26: 479 (W. India). gracilis gracilis - FRASER, F a u n a brit. India, Odon., 2: 126. gracilis gracilis - LIEFTINCK, Treubia, 22, Suppl.: 22 (Siam, Malaya). (Vestinus) gracilis gracilis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht,

Distribution: East Asia, gracilis montana

FRASER

1929 Vestalis gracilis amoena FRASER, J. Bombay nat. Hist. Soc., 33: 583. Type: not given; typelocality: India. 1934 Vestalis gracilis montana FRASER, F a u n a brit. India Odon., 2: 128 (new name for Vestalis gracilis amoena FRASER, 1929, a junior secondary homonymy of Vestalis apicalis amoena FRASER, 1929).

33 1984 Vestaiis ( Vestinus) gracilis amoena - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 10.

Distribution: India, luctuosa (BURMEISTER) 1839 Calopteryx luctuosa BURMEISTER, Handb. Ent., 2: 828. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Java. 1842 Calopteryx formosa RAMBUR, Hist. nat. Ins. Neuropt., Paris: 225. Type female: Serville's Collection; type-locality: Java. 1853 Vestaiis luctuosa - SELYS, Synopsis des Calopterygines: 25. 1854 Vestaiis luctuosa - SELYS & H A G E N , Monogr. Cal.: 80 (redescription). 1890 Vestaiis luctuosa - KIRBY, Syn. Cat. Neur.-Odon., London: 102 (as generotype of Vestaiis SELYS, 1 8 5 3 ) .

1934 1934 1935 1954

Vestaiis Vestaiis Vestaiis Vestaiis

luctuosa luctuosa luctuosa luctuosa

- SCHMIDT, Arch. Hydrob., Suppl.: 13: 331. - LIEFTINCK, Treubia, 14 (4): 383 (Java, Sumatra). - LIEFTINCK, Misc. Zool. Sum.: 9 2 - 9 3 : 3 (Sumatra). - LIEFTINCK, Treubia, 22 Suppl.: 22 (Sumatra, Java, Bali).

Distribution: Sumatra, Java, Bali. lugens SELYS 1879 Vestaiis lugens SELYS, Bull. Acad. r. Belg. (2) 47: 361. Type male: Selys's Collection; typelocality: Sumatra. 1927 Vestaiis lugens - Ris, Zool. Meded., Leiden, 10 (1): 11. 1934 Vestaiis lugens - SCHMIDT, Arch. Hydrob. Suppl., 13: 332. 1935 Vestaiis lugens - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 3. 1954 Vestaiis lugens - LIEFTINCK, Treubia, 22 Suppl.: 23 (Malaya, Sumatra, Mentawei Island).

Distribution: Mentawai Islands, Malaysia, Sumatra. melania SELYS 1873 Vestaiis melania SELYS, Bull. Acad. r. Belg. (2) 35: 474. Type male: McLachlan's Collection; type-locality: Luzon. 1891 Vestaiis melania - SELYS, Ann. Soc. Esp. Hist, nat., 20: 213. 1 9 3 9 Vestaiis melania - N E E D H A M & G Y G E R , Philipp. J . Sei. 7 0 ( 3 ) : 2 4 9 (Luzon, Mindanao).

Distribution: Philippine Islands. smaragdina smaragdina SELYS 1879 Vestalis smaragdina SELYS, Bull. Acad. r. Belg., (2) 47: 362. Type male: R. Mus., Brussels; type-locality: Khasia Hills (Assam). 1935 Vestalaria smaradgina - MAY, Senckenbergiana, 17: 207 (as generotype of Vestalaria MAY, 1935). 1948 Vestalis smaragdina smaragdina-LiErrmcK., Ark. Zool., Uppsala, 41A (10): 1 (Burma, Tibet). 1984 Vestalis ( Vestalaria) smaragdina smaragdina - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 10.

Distribution: India, Burma, Tibet. smaragdina velata Ris 1912 Vestalis smaragdina velata Ris, Suppl. ent. Berlin, 1: 56. Type male: Königsberg Museum; type-locality: China. 1984 Vestalis (Vestalaria) smaragdina velata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 10.

Distribution: China.

34 virens NEEDHAM 1930 Vestalis virens NEEDHAM, Zool. Sinica, (A) 11: 199. Type: unknown; type-locality: China. 1984 Vestalis (Vestalaria) virens - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:10.

Distribution: China.

Subfamily Hetaerinae Medium sized with narrow wings. Usually red at wing base in males (not in Mnesarate). Sectors of arc arising from lowest point of arc, basal space transversed; Dc with strongly curved upper side; CuP and anal vein sigmoid. 1939 Hetaerinae TILL YARD & FRASER, Austr. Zool., 9:212 (in family Agriidae) (for genera Hetaerina and Mnesarete). Type-genus: Hetaerina SELYS, 1853. - 1957 Hetaerinae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 81 (with genera Hetaerina, Mnesarate).

Genus Hetaerina SELYS 1853 Hataerina (Hetaerina) SELYS, Synopsis des Calopterygines: 30. Type species: Libellula caja D R U R Y , 1 7 7 3 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 0 4 . - 1 8 5 4 Hetaerina (Hetaerinaj - SELYS & H A G E N , Monogr. Cal.: 9 6 (revision). - 1 8 9 0 Hetaerina - K I R B Y , Syn Cat. Neur.-Odon. London: 104. - 1916 Hetaerina - Ris, Arch. Naturg., (A) 82: 47 (key to species). - 1939 Hetaerina - TILL YARD & FRASER, Austr. Zool., 9: 212. - 1946 Hetaerina - FRASER, Trans. R. ent. Soc. London, 96: 32. - 1957 Hetaerina - FRASER, R. Zool. Soc. N.S. Wales, 1957: 81.

albistigma HÄGEN 1853 Hetaerina albistigma HAGEN, in SELYS, Synopsis des Calopterygines: 45. Type: unknown; typelocality: Puerto Cabello.

Distribution: Venezuela. amazonica SJÖSTEDT 1917 Hetaerina amazonica SJÖSTEDT, Ark. Zool., Uppsala, 11(15): 36. Type male: Riksmus. Naturh., Stockholm; type-locality: Manaos.

Distribution: Brazil. americana (FABRICIUS) 1798 Agrion americana FABRICIUS, Ent. Syst. Suppl.: 287. Type male: Mus. Comp. Zool., Cambridge (Mass.); type-locality: Am. septentr. 1839 Calopteryx americana - BURMEISTER, Handb. Ent., 2: 826. 1839 Lestes basalis SAY, J1 Acad. Philad., 8: 35. Type male: Boston Museum; type-locality: Philadelphia. 1853 Hetaerina (Hetaerina) americana - SELYS, Synopsis des Calopterygines: 41 (Mexico and Sou,th U.S.A.). 1854 Hetaerina (Hetaerina) americana - SELYS & HÄGEN, Monogr. Cal.: 151 (redescription) (Massachussets, Missouri, Indiana, Mexico). 1859 Hetaerina californica HAGEN, in SELYS, Synopsis des Calopterygines, Add. 1: 6. Type male: Mus. Comp. Zool., Cambridge (Mass.); type-locality: California. 1861 Hetaerina basalis HÄGEN, Syn. Neur. N. Amer., Washington: 60. Type: not given; type-locality: Texas. 1 8 6 2 Agrion basalis - F L I N T & H A R R I S , Ins. Injur. Veget., 3 : 1 2 . 1 8 6 3 H e t a e r i n a p s e u d a m e r i c a n a WALSH, P r o c . E n t . S o c . P h i l a d . , 2: 2 2 3 . T y p e :

unknown.

35 1863 Hetaerina texana WALSH, Proc. Ent. Soc. Philad., 2: 227. Type: unknown; type-locality: Texas. 1890 Hetaerina americana - KIRBY, Syn. Cat. Neur.-Odon., London: 106. 1890 Lestes basalis - KIRBY, Syn. Cat. Neur.-Odon., London: 106. [as synonym of americana (FABRICIUS, 1 7 9 8 ) ] .

1890 Hetaerina californica - KIRBY, Syn. Cat. Neur.-Odon., London: 106 [proposed synonymy with americana (FABRICIUS, 1798)]. 1890 Hetaerinapseudamericana-KIRBY, Syn. Cat. Neur.-Odon., London: 106 [proposed synonymy with americana (FABRICIUS, 1798)].

Distribution: North America. auripennis

(BURMEISTER)

1839 Calopteryx auripennis BURMEISTER, H a n d b . Ent., 2: 827. Type female: Winthem's Collection; type-locality: not given. 1853 Hetaerina (Hetaerina) auripennis - SELYS, Synopsis des Calopterygines: 33. 1 8 5 4 Hetaerina (Hetaerina) auripennis - SELYS & H A G E N , Monogr. Cal.: 1 0 9 (redescription). 1890 Hetaerina auripennis - KIRBY, Syn. Cat. Neur.-Odon. London: 104 (Brazil).

Distribution: Brazil. aurora Ris 1916 Hetaerina aurora Ris, Arch. Naturg., (A) 82: 58. Type male: unknown; type-locality: Colombia.

Distribution: Colombia. borchgravii

SELYS

1869 Hetaerina borchgravii SELYS, Bull. Acad. r. Belg., (2) 27: 658. Type male: unknown; typelocality: Brazil.

Distribution: Brazil. brightwelli

(KIRBY)

1825 Agrion brightwelli KIRBY, Trans. Linn. Soc. London, 14: 107. Type male: unknown; typelocality: Brazil. 1842 Calopteryx brightwelli - BURMEISTER, H a n d b . Ent., 2: 827. 1953 Hetaerina (Hetaerina) brightwelli - SELYS, Synopsis des Calopterygines: 46. 1854 Hetaerina (Hetaerina) brightwelli - SELYS & H Ä G E N , Monogr. Cal.: 148 (redescription). 1890 Hetaerina brightwelli - KIRBY, Syn. Cat. Neur.-Odon., London: 107. 1898 Hetaerina brightwelli - CALVERT, Trans. Am. ent. Soc., 25: 48.

Distribution: Brazil. caja

(DRURY)

1773 Libellula caja DRURY, 111. Ex. Ent., 2: 45. Type male: Hagen's Collection; type-locality: Venezuela. 1842 Calopteryx caja - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 226. 1853 Hetaerina (Hetaerina) caja - SELYS, Synopsis des Calopterygines: 32 (as generotype of Hetaerina SELYS, 1853). 1 8 5 4 Hetaerina (Hetaerina) caja - SELYS & H Ä G E N , Monogr. Cal.: 1 0 4 (redescription). 1890 Hetaerina caja - KIRBY, Syn. Cat. Neur.-Odon., London: 104. 1916 Hetaerina caja - Ris, Arch. Naturg., (A) 82: 54 (Panama). 1932 Hetaerina caja - GEIJSKES, Zool. Meded., Leiden, 14: 235 (Trinidad).

Distribution: Central America.

36

capitalis

SELYS

1873 Hetaerina capitalis SELYS, Bull. Acad. r. Belg., (2) 35: 482. Type male: not given; type-locality: unknown. 1916 Hetaerina capitalis - Ris, Arch. Naturg., (A) 82: 57 (Costa Rica, Colombia). 1923 Hetaerina capitalis var. colombiana NAVÄS, Mem. Pont. Acc. n. Line., (2) 6: 10. Male from Colombia.

Distribution: Colombia, Costa Rica, carnifex

HÄGEN

1853 Hetaerina (Hetaerina) carnifex HAGEN, in SELYS, Synopsis des Calopterygines: 37. Type male: Hagen's Collection; type-locality: Brazil. 1853 Hetaerina (Hetaerina) carnifex race fulgens SELYS, Synopsis des Calopterygines: 39. Male from Brazil. 1854 Hetaerina (Hetaerina) carnifex - SELYS & HAGEN, Monogr. Cal.: 125 (redescription). 1890 Hetaerina carnifex - KIRBY, Syn. Cat. Neur.-Odon., London: 105. 1948 Hetaerina carnifex - FRASER, Acta Zool. lilloana, 5: 64 (Argentina).

Distribution: Brazil, Argentina. charca

CALVERT

1909 Hetaerina charca CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 86. Type male: Calvert's Collection; type-locality: Bolivia. 1916 Hetaerina charca - Ris, Arch. Naturg., (A) 82: 60 (Peru). 1923 Hetaerina charca - WILLIAMSON, Occas. Pap. Mus. Zool. Univ. Michigan, 130: 14. 1943 Hetaerina charca - SCHMIDT, Beitr. Fauna Perus, 2: 251 (Peru, Bolivia).

Distribution: Bolivia, Peru. crassidens

DE

MARMELS

1984 Hetaerina crassidens DE MARMELS, 1982 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 11.

Distribution: Venezuela. dominula

HÄGEN

1953 Hetaerina (Hetaerina) dominula HAGEN, in SELYS, Synopsis des Calopterygines: 33. Type male: Mus. Naturk., Berlin; type-locality: Br. Guiana. 1854 Hetaerina (Hetaerina) dominula - SELYS & HÄGEN, Monogr. Cal.: 107 (redescription). 1890 Hetaerina dominula - KIRBY, Syn. Cat. Neur.-Odon., London: 104. 1948 Hetaerina dominula - FRASER, Acta Zool. lilloana, 5: 63 (Argentina). 1984 Hetaerina caja dominula - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 11.

Distribution: British Guyana, Argentina. donna

SELYS

1873 Hetaerina (Hetaerina) donna SELYS, Bull. Acad. r. Belg., (2) 35: 478. Type male: Selys's Collection; type-locality: Argentina.

Distribution: Argentina, dupex

SELYS

1869 Hetaerina (Hetaerina) dupex SELYS, Bull., Acad. r. Belg., (2) 27: 656. Type male: Selys's Collection; type-locality: Bogota. 1916 Hetaerina dupex - Ris, Arch. Naturg., (a) 82: 58 (Colombia).

37 1984 Hetaerina duplex (misprint) - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:11.

Distribution: Colombia. fuscibasis

CALVERT

1909 Hetaerina fuscibasis CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 81. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil. fuscoguttata

SELYS

1878 Hetaerina fuscoguttata SELYS, C.R. SOC. Ent. Belg., 21: 21. Type male: R. Mus., Brussels; type-locality: Panama. 1916 Hetaerina fuscoguttata - Ris, Arch. Naturg., (A) 82: 53 (Costa Rica).

Distribution: Costa Rica, Panama. hebe

SELYS

1853 Hetaerina (Hetaerina) hebe SELYS, Synopsis des Calopterygines: 34. Type male: Selys & Hagen's Collection; type-locality: Brazil. 1854 Hetaerina (Hetaerina) hebe - SELYS & HAGEN, Monogr. Cal.: 112 (redescription). 1890 Hetaerina hebe - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: Brazil. infecta

CALVERT

1901 Hetaerina infecta CALVERT, Biol. Centr. Amer. Neuropt.: 38. Type male: Smith & Schumann's Collection; type-locality: Mexico.

Distribution: Mexico, Costa Rica, klugi

SCHMIDT

1943 Hetaerina klugi SCHMIDT, Beitr. Fauna Perus, 2: 252. Type male: Mus. Naturk, Berlin; typelocality: Peru.

Distribution: Peru. laesa

HÄGEN

1853 Hetaerina (Hetaerina) laesa HAGEN, in SELYS, Synopsis des Calopterygines: 36. Type male: Mus. Naturk., Berlin; type-locality: Suriname. 1854 Hetaerina (Hetaerina) laesa - SELYS & HÄGEN, Monogr. Cal.: 119 (redescription). 1890 Hetaerina laesa - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: Suriname. longipes

HÄGEN

1853 Hetaerina (Hetaerina) longipes HAGEN, in SELYS, Synopsis des Calopterygines: 37. Type male: Hagen's Collection; type-locality: Brazil. 1854 Hetaerina (Hetaerina) longipes - SELYS & HÄGEN, Monogr., Cal. 121 (redescription). 1890 Hetaerina longipes - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: Brazil. luteola

(RAMBUR)

1842 Calopteryx luteola RAMBUR, Hist. Nat. Ins. Neuropt., Paris: 223. Type female: Serville's Collection; type-locality: Martinique.

38 1842 Calopteryx cruentata RAMBUR, Hist. Nat. Ins. Neuropt., Paris: 228. Type male: unknown; type-locality: unknown. 1853 Hetaerina (Hetaerina) cruentata - SELYS, Synopsis des Calopterygines: 39 (Brazil, Colombia, Mexico, Martinique). 1853 Hetaerina (Hetaerina) brasiliensis SELYS, Synopsis des Calopterygines: 40. Type male: Selys's Collection; type-locality: Brazil. 1854 Hetaerina (Hetaerina) cruentata - SELYS & HAGEN, Monogr. Cal. 127 (redescription) (Venezuela, Suriname, Colombia, Mexico). 1854 Hetaerina (Hetaerina) luteola - SELYS & HÄGEN, Monogr. Cal.: 127 [proposed synonymy with cruentata (RAMBUR, 1842)]. 1854 Hetaerina (Hetaerina) lineata HAGEN, in SELYS & HÄGEN, Monogr. Cal.: 129. Type: unknown. 1890 Hetaerina luteola - KIRBY, Syn. Cat. Neur.-Odon., London: 105. 1916 Hetaerina cruentata - Ris, Arch. Naturg., (A) 82: 53 (Mexico).

Distribution: Central and South America. macropus

SELYS

1853 Hetaerina (Hetaerina) macropus SELYS, Synopsis des Calopterygines: 44. Type male: Selys's Collection; type-locality: Central America. 1853 Hetaerina (Hetaerina) occisa HAGEN, in SELYS, Synopsis des Calopterygines: 44. Type male: Selys's Collection; type-locality: Colombia. 1854 Hetaerina (Hetaerina) macropus - SELYS & HÄGEN, Monogr. Cal.: 141 (redescription) (Mexico, Honduras). 1873 Hetaerina macropus var. asticta SELYS, Bull. Acad. r. Belg., (2) 35: 481 (as variety of macropus SELYS, 1 8 5 3 ) .

1873 Hetaerina macropus var. heterosticta SELYS, Bull. Acad. r. Belg., (2) 35: 481 (as variety of macropus

SELYS, 1 8 5 3 ) .

1873 Hetaerina occisa var. sublimata SELYS, Bull. Acad. r. Belg., (2) 36: 613 (as variety of occisa HÄGEN, 1 8 5 3 ) .

1890 Hetaerina macropus - KIRBY, Syn. Cat. Neur.-Odon., London: 107. 1890 Hetaerina occisa - KIRBY, Syn. Cat. Neur.-Odon., London: 107. 1916 Hetaerina macropus - Ris, Arch. Naturg., (A) 82: 56 (Mexico, Guatemala, Panama, Colombia, Peru). 1923 Hetaerina macropus - WILLIAMSON, Occas. Pap. Mus. Zool. Univ. Michigan, 130: 16. 1932 Hetaerina macropus - GEIJSKES, Zool. Meded., Leiden, 14: 236 (Trinidad). 1943 Hetaerina macropus - SCHMIDT, Beitr. Fauna Perus, 2: 251 (Peru, Venezuela, Trinidad, Colombia, Mexico).

Distribution: from Mexico to Peru. majuscula

SELYS

1853 Hetaerina (Hetaerina) majuscula SELYS, Synopsis des Calopterygines: 47. Type male: Selys's Collection; type-locality: Colombia. 1854 Hetaerina (Hetaerina) majuscula - SELYS & HÄGEN, Monogr. Cal.: 151 (redescription) (Colombia, Suriname). 1890 Hetaerina majuscula - KIRBY, Syn. Cat. Neur.-Odon., London: 107. 1916 Hetaerina majuscula - Ris, Arch. Naturg., (A) 82: 57 (Costa Rica, Colombia).

Distribution: Costa Rica, Colombia. maxima

MCLACHLAN

1879 Hetaerina maxima MCLACHLAN, Entomologist's month. Mag., 15: 244. Type male: McLachlan's Collection; type-locality: Costa Rica.

Distribution: C o s t a Rica.

39

medinai 1984

RACENIS

Hetaerina medinai Utrecht, 1: 11.

RACENIS, 1 9 6 8

- loc. cit.

DAVIS & TOBIN,

The Dragonflies of the World,

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Venezuela. mendezi 1984

JURZITA

Hetaerina mendezi Utrecht 1: 11.

JURZITA, 1 9 8 2

Distribution: Argentina. miniata

SELYS

1879 Hetaerina miniata SELYS, Bull. Acad. r. Belg., (2) 47: 370. Type male: Selys's Collection; typelocality: Chiriqui.

Distribution: South America, moribunda

HÄGEN

1853 Hetaerina (Hetaerina) moribunda HAGEN, in SELYS, Synopsis des Calopterygines: 42. Type male: Hagen's Collection; type-locality: Guyana. 1 8 5 4 Hetaerina (Hetaerina) moribunda - SELYS & H A G E N , Monogr. Cal.: 1 5 4 (redescription). 1890 Hetaerina moribunda - KIRBY, Syn. Cat. Neur.-Odon., London: 106.

Distribution: Guyana, mortua

HÄGEN

1853 Hetaerina (Hetaerina) mortua HAGEN, in SELYS, Synopsis des Calopterygines: 35. Type male: Hagen's Collection; type-locality: Guyana. 1 8 5 4 Hetaerina (Hetaerina) mortuna - SELYS & H Ä G E N , Monogr. Cal.: 1 1 7 (redescription). 1890 Hetaerina mortua - KIRBY, Syn. Cat. Neur.-Odon., London: 105. Distribution: Guyana.

papaverina

FRASER

1946 Hetaerina papaverina FRASER, Trans. R. ent. Soc. London, 96: 33. Type male: Leeds City Museum; type-locality: Peru. 1966 Hetaerina papavarina - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 209 (lectotype male: British Museum (Nat. Hist.), London).

Distribution: Peru, pilula

CALVERT

1901 Hetaerina pilula CALVERT, Biol. Centr. Amer. Neuropt.: 33. Type: unknown; type-locality: Mexico.

Distribution: Mexico. proxima

SELYS

1853 Hetaerina (Hetaerina) proxima SELYS, Synopsis des Calopterygines: 38. Type male: Selys's Collection; type-locality: Brazil. 1 8 5 4 Hetaerina (Hetaerina) proxima - SELYS & H Ä G E N , Monogr. Cal.: 1 2 5 (redescription). 1890 Hetaerina proxima - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: Brazil.

40 rosea

SELYS

1853 Hetaerina (Hetaerina) rosea SELYS, Synopsis des Calopterygines: 31. Type male: Selys & Hagen's Collection; type-locality: Brazil. 1 8 5 4 Hetaerina (Hetaerina) rosea - SELYS & H A G E N , Monogr. Cal.: 1 0 4 (redescription). 1890 Hetaerina rosea - KIRBY, Syn. Cat. Neur.-Odon., London: 104. 1916 Hetaerina rosea - Ris, Arch. Naturg., (A) 82: 55 (Argentina). 1948 Hetaerina rosea - FRASER, Acta Zool. lilloana, 5: 64 (Argentina).

Distribution: Brazil, Argentina. rudis

CALVERT

1901 Hetaerina rudis CALVERT, Biol. Centr. Amer. Neuropt.: 40. Type male: Calvert's Collection; type-locality: Central America.

Distribution: Central America, sempronia

HÄGEN

1853 Hetaerina (Hetaerina) sempronia HAGEN, in SELYS, Synopsis des Calopterygines: 45. Type male: Mus. Naturk., Berlin; type-locality: Mexico. 1 8 5 4 Hetaerina (Hetaerina) sempronia - SELYS & H Ä G E N , Monogr. Cal.: 1 4 7 (redescription). 1890 Hetaerina sempronia - KIRBY, Syn. Cat. Neur.-Odon., London: 107. 1916 Hetaerina sempronia - Ris, Arch. Naturg., (A) 82: 54.

Distribution: Mexico. sanguinea

SELYS

1853 Hetaerina (Hetaerina) sanguinea SELYS, Synopsis des Calopterygines: 31. Type male: Selys's Collection; type-locality: Para. 1854 Hetaerina (Hetaerina) sanguinea - SELYS & H Ä G E N , Monogr. Cal.: 100 (redescription). 1879 Lais brevistyla SELYS, Bull. Acad. r. Belg., (2) 47: 367. Type male: Selys's Collection; typelocality: Rio N a p o . 1890 Hetaerina sanguinea - KIRBY, Syn. Cat. Neur.-Odon., London: 104. 1916 Hetaerina sanguinea - Ris, Arch. Naturg., (A) 82: 56 (Colombia, Peru). 1917 Hetaerina sanguinea - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 39 (Amazonas) 1943 Hetaerina sanguinea - SCHMIDT, Beitr. F a u n a Perus, 2: 251 (Peru, Colombia, Amazonas). 1948 Hetaerina sanguinea - FRASER, Acta Zool. lilloana, 5: 65 (Argentina).

Distribution: Peru, Colombia, Brazil, Argentina. sanguinolenta

HÄGEN

1853 Hetaerina (Hetaerina) sanguinolenta HAGEN, in SELYS, Synopsis des Calopterygines: 35. Type male: Hägen & Selys's Collection; type-locality: Brazil. 1 8 5 4 Hetaerina (Hetaerina) sanguinolenta - SELYS & H Ä G E N , Monogr. Cal.: 1 1 5 (redescription). 1890 Hetaerina sanguinolenta - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: Brazil. septentrionalis

SELYS

1853 Hetaerina (Hetaerina) septentrionalis SELYS, Synopsis des Calopterygines: 36. Type male: British Museum (Nat. Hist.), London; type-locality: Georgia. 1 8 5 4 Hetaerina (Hetaerina) septentrionalis - SELYS & H Ä G E N , Monogr. Cal.: 1 1 9 (redescription). 1890 Hetaerina septentrionalis - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: U.S.A.: Georgia.

41 simplex

SELYS

1853 Hetaerina (Hetaerina) simplex SELYS, Synopsis des Calopterygines: 30. Type male: Selys's Collection; type-locality; Brazil. 1 8 5 4 Hetaerina (Hetaerina) simplex - SELYS & H A G E N , Monogr. Cal.: 9 8 (redescription). 1869 Hetaerina perplex SELYS, Bull. Acad. r. Belg., (2) 27: 655. Type male: Selys's Collection; typelocality: Para. 1890 Hetaerina simplex - KIRBY, Syn. Cat. Neur.-Odon., London: 104.

Distribution: Brazil. tolteca Calvert 1901 Hetaerina tolteca CALVERT, Biol. Centr. Amer., Neuropt.: 40. Type male: Mus. Comp. Zool., Cambridge (Mass.); type-locality: Mexico.

Distribution: Mexico. titia

(DRURY)

1773 Libellula titia DRURY, 111. Ex. Ent., 2: 45. Type: lost; type-locality: not given (? Honduras). 1839 Calopteryx tricolor BURMEISTER, Handb. Ent., 2: 827. Type male: Mus. Comp. Zool., Cambridge (Mass.), type-locality: Pennsylvania. 1842 Calopteryx titia - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 227. 1853 Hetaerina (Hetaerina) tricolor - SELYS, Synopsis des Calopterygines: 42 (Georgia) 1853 Hetaerina (Hetaerina) titia - SELYS, Synopsis des Calopterygines: 43. 1853 Hetaerina (Hetaerina) limbata SELYS, Synopsis des Calopterygines: 43. Type male: Walsh's Collection; type-locality: Illionois. 1854 Hetaerina (Hetaerina) tricolor - SELYS & H Ä G E N , Monogr. Cal.: 156 (redescription). 1854 Hetaerina (Hetaerina) titia - SELYS & H Ä G E N , Monogr. Cal.: 158 (redescription). 1862 Hetaerina rupinsulensis WALSH, Proc. Acad. nat. Sei. Philad., 1862: 383. Type: destroyed; typelocality: Rock Island. 1862 Hetaerina rupamnensis WALSH, Proc. Ent. Soc. Philad., 2: 230. Type: destroyed; type-locality: unknown. 1873 Hetaerina bipartite SELYS, Bull. Acad. r. Belg., (2) 35: 481 (as race of titia). 1890 Hetaerina titia - KIRBY, Syn. Cat. Neur.-Odon., London: 106 (Honduras, Texas, Mexico). 1890 Hetaerina tricolor - KIRBY, Syn. Cat. Neur.-Odon., London: 106. 1898 Hetaerina titia - CALVERT, Trans. Am. ent. Soc., 25: 47. 1910 Hetaerina tricolor limbata - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 34 (Texas, Illinois). 1912 Hetaerina titia - WILLIAMSON, Ent. News, 23: 101. 1 9 1 2 Calopteryx tricolor - WILLIAMSON, Ent. News, 2 3 : 1 0 1 (proposed synonymy with titia ( D R U R Y , 1773).

Distribution: South U.S.A., Central America, vulnerata

HÄGEN

1853 Hetaerina (Hetaerina) vulnerata HAGEN, in SELYS, Synopsis des Calopterygines: 40. Type male: Mus. Naturk., Berlin; type-locality: Brazil. 1854 Hetaerina (Hetaerina) vulnerata-SELYS & HÄGEN, Monogr. Cal.: 150 (redescription) (Mexico, Brazil, Colombia). 1890 Hetaerina vulnerata - KIRBY, Syn. Cat. Neur.-Odon., London: 105.

Distribution: from Mexico to Brazil. westfalli 1984

RACENIS

Hetaerina westfalli Utrecht, 1: 12.

RACENIS, 1 9 6 8

Distribution: Venezuela.

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

42 Genus Mnesarete

COWLEY

1853 Hetaerina (Lais) SELYS, Synopsis des Calopterygines: 27. Type species: Lais hyalina SELYS, 1853 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 103). - 1854 Hetaerina (Lais)

- SELYS & HAGEN, M o n o g r . Cal.: 87. - 1890 Lais - KIRBY, Syn. C a t . N e u r . - O d o n . ,

London: 103. - 1916 Lais - Ris, Arch. Naturg., (A) 82: 40 (revision, key to species). - 1934 Mnesarete COWLEY, Entomologist, 67: 201 (new name for Lais SELYS, 1853, a junior primary homonymy of Lais GISTL, 1848 in Tunicata).

aenea

(SELYS)

1853 Hetaerina (Lais) aenea SELYS, Synopsis des Calopterygines: 28. Type male: Dale's Collection; type-locality: Para. 1854 Hetaerina (Lais) aenea - SELYS & HÄGEN, Monogr. Cal.: 91 (redescription). 1890 Lais aenea - KIRBY, Syn. Cat. Neur.-Odon., London: 103 (Amazons). 1984 Mnesarete aenea - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 12.

Distribution: Brazil. cupraea

(SELYS)

1853 Hetaerina (Lais) cupraea SELYS, Synopsis des Calopterygines: 28. Type male: British Museum (Nat. Hist.), London; type-locality: Brazil. 1890 Lais cupraea - KIRBY, Syn. Cat. Neur.-Odon., London: 103. 1946 Mnesarete cupraea - FRASER, Trans. R. ent. Soc. London, 96: 31 (Brazil, Peru).

Distribution: Brazil, Peru. devilli 1880 1881 1890 1946

(SELYS)

Lais devilli SELYS, C.R. Soc. ent. Belg., 23: 49. Type: unknown; type-locality: Ecuador. Lais devillei (misprint) - MCLACHLAN, Trans, ent. Soc. London, 1881: 26. Lais devilli - KIRBY, Syn. Cat. Neur.-Odon., London: 103. Mnesarete devillei (misprint) - FRASER, Trans. R. ent. Soc. London, 96: 30 (Peru, Ecuador).

Distribution: Ecuador, Peru. globifer

(SELYS)

1853 Hetaerina (Lais) globifer SELYS, Synopsis des Calopterygines: 27. Type male: Hagen's Collection; type-locality: Brazil. 1854 Hetaerina (Lais) globifer - SELYS & HÄGEN, Monogr. Cal.: 88 (redescription). 1890 Lais globifer - KIRBY, Syn. Cat. Neur.-Odon., London: 103. 1984 Mnesarete globifer - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:12.

Distribution: Brazil. grisea (Ris) 1916 Lais grisea Ris, Arch. Naturg., (A) 82: 41. Type male: not given; type-locality: Argentina. 1948 Mnesarete grisea - FRASER, Acta Zool. lilloana, 5: 65.

Distribution: Argentina. guttifera (SELYS) comb. nov. 1873 Lais guttifera SELYS, Bull. Acad. r. Belg., (2) 35: 475. Type male: Selys's Collection; typelocality: Brazil. Distribution: Brazil.

43 h a u x w e l l i (SELYS) 1869 Lais hauxwelli SELYS, Bull. Acad. r. Belg., (2) 27: 654. Type male: Hauxwell's Collection; type-locality: Pebas. 1879 Lais fulgida SELYS, Bull. Acad. r. Belg., (2) 47: 365. Type male: Selys's Collection; type-locality: Rio Napo. 1943 Lais hauxwelli - SCHMIDT, Beitr. Fauna Perus, 2: 250 (Ecuador). 1943 Lais fulgida - SCHMIDT, Beitr. Fauna Perus, 2: 250 (Peru). 1946 Mnesarete hauxwelli - FRASER, Trans. R. ent. Soc. London, 96: 28 (redescription) (Peru, Colombia).

Distribution: Brazil, Peru, Colombia. hincksi FRASER 1946 Mnesarete hincksi FRASER, Trans. R. ent. Soc. London, 96: 26. Type male: Leeds Museum; type-locality: Ecuador. 1966 Mnesarete hincksi-KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18:196 [type in British Museum (Nat.Hist.), London],

Distribution: Ecuador.

hyalina (HÄGEN) 1853 Hetaerina (Lais) hyalina HAGEN, in SELYS, Synopsis des Calopterygines: 28. Type male: Naturh. Mus., Wien; type-locality: Brazil. 1890 Lais hyalina - KIRBY, Syn. Cat. Neur.-Odon., London: 104 (as generotype of Lais, SELYS, 1853).

1934 Mnesarete hyalina - COWLEY, Entomologist, 67: 201 (as generotype of Mnesarete COWLEY, 1934).

Distribution: Brazil, imperatrix (MCLACHLAN) comb. nov. 1878 Mnais imperatrix MCLACHLAN, Trans. Ent. Soc. London, 1878: 85. Type female: British Museum (Nat. Hist.), London; type-locality: Ecuador. 1916 Lais imperatrix - Ris, Arch. Naturg., (A) 82: 43 (Colombia).

Distribution: Ecuador, Colombia. marginata (SELYS) comb. nov. 1879 Lais marginata SELYS, Bull. Acad. r. Belg., (2) 47: 366. Type male: Selys's Collection; typelocality: Peru. 1890 Lais marginata - KIRBY. Syn. Cat. Neur.-Odon., London: 104. 1943 Lais marginata - SCHMIDT, Beitr. Fauna Perus, 2: 251 (Peru).

Distribution: Peru. metallica (SELYS) comb. nov. 1869 Lais metallica SELYS, Bull. Acad. r. Belg., (2) 27:654. Type male: unknown; type-locality: Bahia.

Distribution: South America. pruinosa (HÄGEN) 1853 Hetaerina (Lais) pruinosa HAGEN, in SELYS, Synopsis des Calopterygines: 28. Type male: Mus. Naturk., Berlin; type-locality: Brazil. 1854 Hetaerina (Lais) pruinosa - SELYS & HAGEN, Monogr. Cal.: 95 (redescription). 1890 Lais pruinosa - KIRBY, Syn. Cat. Neur.-Odon., London: 104.

44 1948 Mnesarete pruirtosa - FRÄSER, Acta Zool. lilloana, 5: 65 (Argentina).

Distribution: Brazil, Argentina, pudica (HAGEN) comb. nov. 1853 Hetaerina (Lais) pudica HAGEN, in SELYS, Synopsis des Calopterygines: 29. Type male: N a turhist. Mus., Wien; type-locality: Brazil. 1854 Hetaerina (Lais) pudica - SELYS & HAGEN, Monogr. Cal.: 95 (redescription). 1890 Lais pudica - KIRBY, Syn. Cat. Neur. Odon., London: 104.

Distribution: Brazil. regina (Ris) 1916 Lais regina Ris, Arch. Naturg., (A) 82: 46. Type male: not given; type-locality: Bolivia. 1984 Mnesarete regina - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 3 .

Distribution: Bolivia. rollinati (MARTIN) 1897 Lais rollinati MARTIN, Ann. Soc. ent. France, 66: 592. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Bolivia. 1916 Lais rollinati - Ris, Arch. Naturg., (A) 82: 45. 1984 Mnesarete rollinati - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 3 .

Distribution: Bolivia. saltuum (Ris) 1916 Lais saltuum Ris, Arch. Naturg., (A) 82: 44. Type male: not given; type-locality: Peru. 1943 Lais saltuum - SCHMIDT, Beitr. F a u n a Perus, 2: 251. 1 9 8 4 M n e s a r e t e s c i n t i l l a RACENIS, 1 9 6 8 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 13 (Venezuela).

Distribution: Peru, Venezuela. smaragdina (SELYS) 1869 Lais smaragdina SELYS, Bull. Acad. r. Belg., (2) 27: 654. Type male: Bates's Collection; typelocality: Santarem. 1984 Mnesarete smaragdina - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 3 .

Distribution: Brazil. Family Chlorocyphidae Small stout species with petiolate wings usually longer than abdomen. Clypeus prominent. Primary antenodals evident, 5 to 10 secondaries but none proximal to primaries, IRiii and Riv recessed. Rii-iii usually strongly arched forward near origin. Upper segment of arc touching radius. Intercalary sectors present. IA absent in Disparocypha. 1905 Libellaginae JACOBSON & BIANKI, Pryam. lozhnos. ross, imp.: 803 (with genera Rhinocypha, Micromerus - in family Calopterygidae). - 1934 Libellagidae LIEFTINCK, Treubia, 14.(4): 385. - 1937 Chlorocyphidae COWLEY, Trans. R. ent. Soc. London, 86: 1 (defined as family). Typegenus: Chlorocypha FRASER, 1928. - 1939 Chlorocyphidae TILLYARD & FRASER, Austr. Zool., 9: 205 (with genera Libellago, Rhinoneura, Disparocypha, Indocypha, Neocypha, Rhinocypha, Chlorocypha, Calocypha). - 1950 Chlorocyphidae - LAIDLAW, Trans. R. ent. Soc. London, 101:258 (revision, with genera Libellago, Melanocypha, Rhinoneura, Rhinocypha, Chlorocypha,

45 Indocypha, Sundacypha, Heterocypha, Aristocypha, Disparocypha, Melanocypha).-1957 Chlorocyphidae- FRÄSER, R . Zool. Soc. N . S . Wales, 1957: 7 0 (key to genera Libellago, Pachycypha, Melanocypha, Sclerocypha, Disparocypha, Calocypha, Rhinocypha, Cyrano, Indocypha, Chlorocypha, Platycypha, Paracypha, Heliocypha). - 1962 Chlorocyphidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 (key to genera Chlorocypha, Platycypha).

Genus Africocypha

PINHEY

1961 Chlorocypha (Africocypha) PINHEY, Entomologist's month. Mag., 96: 261. Type species: Chlorocypha (Africocypha) greyi PINHEY, 1 9 6 1 (original designation). - 1 9 8 4 Africocypha - DAVIES & TOBIN, the Dragonflies of the World, Utrecht, 1: 13.

lacus-elephantum

(KARSCH)

1899 Libellago lacus-elephantum'KARSCH, Ent. Nachr., 25 (11): 165. Type female: Mus. Naturk., Berlin; type-locality: Cameroon. 1949 Libellago lacus-elephantum - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 48 (in incertae sedis). 1961 Chlorocypha (Africocypha) greyi PINHEY, Entomologist's month. Mag., 96: 261. Type male: Nat. Mus. Bulawayo; type-locality: South Africa (as generotype of Africocypha PINHEY, 1 9 6 1 ) . 1961 Chlorocypha (Africocypha) natali PINHEY, Entomologist's month. Mag., 96: 263. Type female: Nat. Mus., Bulawayo; type-locality: South Africa. 1962 Chlorocypha lacus-elephantum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149 (Cameroon). 1984 Africocypha lacuselephantum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 15. 1 9 8 4 Chlorocypha (Africocypha) greyi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 3 (proposed synonymy with lacuselephantum (KARSCH, 1899). 1 9 8 4 Chlorocypha (Africocypha) natali- DAVIES & TOBIN. The Dragonflies of the World, Utrecht, 1: 1 3 [proposed synonymy with lacuselephantum (KARSCH, 1899)].

Distribution: Cameroon, South Africa. Genus Aristocypha

LAIDLAW

1950 Aristocypha LAIDLAW, Trans. R. ent. Soc. London, 101: 273. Type species-.Libellago (Rhinocypha) quadrimaculata Selys, 1853 (original designation). - 1954 Aristocypha - LIEFTINCK, Treubia, 22 Suppl.: 7 (proposed synonymy with Rhinocypha RAMBUR, 1842). - 1984 Aristocypha - D A VIED & TOBIN, The Dragonflies of the World, Utrecht, 1: 13 (as good genus).

cuneata

(SELYS)

1853 Libellago (Rhinocypha) cuneata SELYS, Synopsis des Calopterygines: 60. Type male: Selys's Collection; type-locality: Tibet. 1853 Libellago cuneata - WALKER, List. Neur. Ins. B . M . 4: 650. 1854 Libellago (Rhinocypha) cuneata - SELYS & H A G E N , Monogr. Cal.: 206 (redescription). 1890 Libellago cuneata - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1903 Rhinocypha adamantina FÖRSTER, Annls hist.-nat. Mus. natn. hung., 1: 547. Type male: destroyed; type-locality: Sikkim. 1904 Rhinocypha adamantina - WILLIAMSON, Proc. U.S. natn. Mus., 28: 174. 1917 Rhinocypha cuneata - LAIDLAW, Ree. Indian Mus., 13: 36 (India). 1950 Aristocypha cuneata - LAIDLAW, Trans. R. ent. Soc. London, 101: Til. 1950 Rhinocypha adamantina -LAIDLAW, Trans. R. ent. Soc. London, 101: 277 [proposed synonymy with cuneata (SELYS, 1853)].

Distribution: India, Tibet. fenestrella

(RAMBUR)

1842 Rhinocypha fenestrella RAMBUR, Hist. nat. Ins. Neuropt., Paris: 236. Type: sex not given; typelocality: not given.

46 1853 Libellago (Rhinocypha) fenestrella - SELYS, Synopsis des Calopterygines: 60 (Malaysia). 1853 Libellago fenestrella - WALKER, List. Neur. Ins. Β. M., 4: 650. 1854 Libellago (Rhinocypha) fenestrella -SELYS & HAGEN, Monogr. Cal.: 204 (redescription) (type male: Mus. Nation. Hist. Nat., Paris; type-locality: Malacca). 1890 Rhinocypha fenestrella - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1917 Rhinocypha fenestrella - LAIDLAW, Ree. Indian Mus., 13: 35 (as species-group). 1931 Rhinocypha baibarana MATSUMURA, 6000 111. Ins. Japan: 1458. Type male: Hokkaido Univ.; type-locality: Japan. 1950 Aristocypha fenestrella - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Rhinocypha baibarana - LAIDLAW, Trans. R. ent. Soc. London, 101: 277 [proposed synonymy w i t h fenestrella

(RAMBUR, 1 8 4 2 ) ] .

1954 Rhinocypha fenestrella 1961 Rhinocypha fenestrella

- LIEFTINCK, Treubia, 22 Suppl.: 10 (Siam, Malaya). - ASAHINA, Insecta matsum., 24: 58.

Distribution: from Japan to Malaysia.

fulgidipennis

(GUERIN)

1831 Agrion fulgidipennis GUERIN, Mag. Zool. Ins., 1: 15. Type male: Selys's Collection; type-locality: Cochin-China. 1842 Rhinocypha fulgidipennis - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 233. 1853 Libellago (Rhinocypha) fulgidipennis - SELYS, Synopsis des Calopterygines: 59. 1853 Libellago fulgidipennis - WALKER, List Neur. Ins. B.M., 4: 645. 1854 Libellago (Rhinocypha) fulgidipennis - SELYS & HÄGEN, Monogr. Cal.: 200 (redescription). 1890 Rhinocypha fulgidipennis - KIRBY, Syn. Cat. Neur.-Odon., London: 112. 1900 Rhinocypha fulgidipennis - MCLACHLAN, Ent. Mag., 36: 114. 1950 Aristocypha fulgidipennis - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1984 Aristocypha fulgipennis (misprint) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 14.

Distribution: Vietnam, hilaryae

(FRASER)

1927 Rhinocypha (Nat. Hist.), 1950 Aristocypha 1966 Aristocypha

hilaryae FRÄSER, Ree. Indian. Mus., 29: 83. Type male: British Museum London; type-locality: Burma. hilaryae - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. hilaryae - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 195.

Distribution: Burma. immaculata

(SELYS)

1879 Rhinocypha immaculata SELYS, Bull. Acad. r. Belg., (2) 47: 385. Type male: Selys's Collection; type-locality: Khasia Hill (Assam). 1890 Rhinocypha unimaculata (misprint) - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1917 Rhinocypha immaculata - LAIDLAW, Ree Indian Mus., 13: 34 (Assam). 1950 Aristocypha immaculata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: India. iridea

(SELYS)

1892 Rhinocypha iridea SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 492. Type male: Mus. civ. Stor. nat., Genova; type-locality: Burma. 1917 Rhinocypha iridea - LAIDLAW, Ree. Indian Mus., 13: 37. 1933 Rhinocypha iridea - FRÄSER, J. Siam Soc. nat. Hist. Suppl., 9: 121. 1950 Rhinocypha iridea - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

47 1984

Aristocypha

iridea -

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 1 4 .

Distribution: Burma. kerri

(FRASER)

1933 Rhinocypha (Nat. Hist.), 1950 Aristocypha 1966 Aristocypha 1984 Aristocypha

kerri FRÄSER, J. Siam. Soc. nat. Hist. Suppl. 9: 121. Type male: British Museum London; type-locality: China. kerri - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. iridea kerri - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 199. iridea f. kerri - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 4 .

Distribution: China. quadrimaculata

(SELYS)

1853 Libellago (Rhinocypha) quadrimaculata SELYS, Synopsis des Calopterygines: 60. Type male: Selys's Collection; type-locality: India. 1853 Libellago quadrimaculata - WALKER, List Neur. Ins. B.M., 4: 651. 1 8 5 4 Libellago (Rhinocypha) quadrimaculata - SELYS & H A G E N , Monogr. Cal.: 2 0 2 (redescription). 1890 Rhinocypha quadrimaculata - KIRBY, Syn. Cat. Neur.-Odon., London: 112. 1917 Rhinocypha fenestrella quadrimaculata - LAIDLAW, Ree. Indian Mus., 13: 36. 1922 Rhinocypha bifenestrata FRASER, Mem. Dept. Agr. India, 7: 63. Type male: British Museum (Nat. Hist.), London; type-locality: Darjeeling. 1922 Rhinocypha quadrimaculata hemihyalina FRASER, Mem. Dept. Agr. India, 7: 64. Type male: British Museum (Nat. Hist.), London; type-locality: Assam. 1950 Aristocypha quadrimaculata - LAIDLAW, Trans. R. ent. Soc. London, 101: 273 (as generotype o f Aristocypha

(LAIDLAW, 1950).

Rhinocypha bifenestrata - L A I D L A W , Trans. R . ent. Soc. London, 1 0 1 : 2 7 3 [proposed synonymy with quadrimaculata (SELYS, 1853)]. 1950 Rhinocypha quadrimaculata hemihyalina - LAIDLAW, Trans. R. ent. Soc. London, 101: 273 [proposed synonymy with quadrimaculata (SELYS, 1853)]. 1964 Rhinocypha quadrimaculata - SCHMIDT, Ent. Tidschr., 85: 153 (Burma). 1950

Distribution: India, Burma. spuria

(SELYS)

1879 Rhinocypha spuria SELYS, Bull. Acad. r. Belg., (2) 47: 388. Type male: Selys's Collection; type-locality: Khasia Hills. 1917 Rhinocypha fenestrella spuria - LAIDLAW, Ree. Indian Mus., 13: 37 (India). 1950 Aristocypha spuria - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

Distribution: India, Burma. trifasciata

(SELYS)

1853 Libellago (Rhinocypha) trifasciata SELYS, Synopsis des Calopterygines: 61. Type male: Selys's Collection; type-locality: India. 1853 Libellago trifasciata - WALKER, List Neur. Ins. Β. M., 4: 530. 1 8 5 4 Libellago (Rhinocypha) trifasciata - SELYS & H Ä G E N , Monogr. Cal.: 2 0 7 (redescription). 1879 Rhinocypha bifasciata SELYS, Bull. Acad. r. Belg., 48: 386. Type male: Selys's Collection; typelocality: Darjeeling. 1890 Rhinocypha trifasciata - KIRBY, Syn. Cat. Neur.-Odon. London: 113. 1917 Rhinocypha trifasciata - LAIDLAW, Ree. Indian Mus., 13: 34 (India, Burma, Assam). 1917 Rhinocypha bifasciata - LAIDLAW, Ree. Indian Mus., 13: 34 (India). 1950 Rhinocypha trifasciata - LAIDLAW, Trans. R. ent. Soc. London, 101: 34 (India, Burma). 1950 Aristocypha bifasciata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

48 1984 Aristocypha

bifasciata - DAVIES & TOBIN, the Dragonflies of the World, 1 : 1 3 (proposed syn-

o n y m y w i t h trifasciata

SELYS, 1 8 5 3 ) ] .

Distribution: India, Burma. Genus Calocypha

FRASER

1928 Calocypha FRÄSER, J. Bombay nat. Hist. Soc., 32: 457. Type species:Rhinocypha laidlawi FRÄSER, 1924 (monotypy). - 1984 Calocypha - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 14.

laidlawi

(FRASER)

1924 Rhinocypha laidlawi FRASER, Ree. Indian Mus., 26: 482. Type male: British Museum (Nat. Hist.), London; type-locality: South India. 1924 Rhinocypha laidlawi - FRASER, Ree. Indian Mus., 26: 519 (description of female). 1928 Calocypha laidlawi - FRASER, J. Bombay nat. Hist. Soc., 32: 457 (as generotype of Calocypha FRASER, 1 9 2 4 ) .

1931 Calocypha (sic? Calocypha) laidlawi - FRASER, Ree. Indian Mus., 33: 463 (Malabar). 1950 Calocypha laidlawi - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1966 Calocypha laidlawi - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 200.

Distribution: South India. Genus Chlorocypha

FRASER

1928 Chlorocypha (Libellago) FRASER, J. Bombay nat. Hist. Soc., 32: 864 (new name for Libellago auct. nec SELYS, 1854). Type species: Agrion dispar BEAUVOIS, 1805 (designated by FRASER, 1934, F a u n a brit. India Odon., 2: 56). - 1934 Chlorocypha-FRASER, F a u n a brit. India Odon., 2: 56. - 1949 Chlorocypha - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 17 (revision). - 1936 Chlorocypha - LONGFIELD, Trans. R. ent. Soc. London, 85: 467 (proposed synonymy with Libellago SELYS, 1854). - 1950 Chlorocypha - LAIDLAW, Trans. R. ent. soc. London, 101: 270. - 1962 Chlorocypha - PINHEY, Publ.cult. Comp. Diam. Angola, 59: 147.

aphrodite (LE ROI) 1915 Libellago aphrodite LE ROI, Ergeb. d. Zentr. Afr. Exp. Zool., 1: 331. Type male: Senckenberg Museum; type-locality: Belgian Congo. 1949 Chlorocypha aphrodite - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 46. 1962 Chlorocypha aphrodite - PINHEY, Publ. cult. Comp. Diam. Angola, 49: 147.

Distribution: Congo Republic. basilewskyi basilewskyi 1955 Chlorocypha Mus. r. Afr. 1962 Chlorocypha 1984 Chlorocypha 1: 14.

FRASER

basilewskyi FRASER, Ann. Mus. Congo Beige, 8, Sei. zool., 36: 23. Type male: centr. Tervuren; type-locality: R u a n d a - U r u n d i . basilewskyi - Pinhey, Publ. cult. Comp. Diam. Angola, 59: 147 (Belgian Congo). basilewskyi basilewskyi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht,

Distribution: Congo Republic, Ruanda-Urundi. basilewskyi jacksoni

PINHEY

1952 Chlorocypha jacksoni PINHEY, OCC. Pap. Coryndon Mus., 3: 13. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1 9 6 2 Chlorocypha jacksoni - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 4 9 .

49 1984 Chlorocypha 1: 15.

basilewskyi jacksoni - DA VIES & TOBIN, The Dragonflies of the World, Utrecht,

Distribution: Uganda, crocea crocea

LONGFIELD

1947 Chlorocypha croceus LONGFIELD, Arq. Mus. Bocage, 16: 17. Type male: Bocage Museum; type-locality: Angola. 1949 Chlorocypha croceus - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 50 (in incertae sedis).

Distribution: Angola. crocea bamptoni 1984

PINHEY

Chlorocypha crocea bamptoni the World, Utrecht, 1: 14.

PINHEY, 1 9 7 5

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of

Distribution: Central Africa. cancellata

(SELYS)

1879 Libellago cancellata SELYS, Bull. Acad. r. Belg., (2) 47: 383. Type male: Selys's Collection; type-locality: Old Calabar. 1914 Libellago hintzi GRÜNBERG, Ent. Rundsch., 31: 53. Type male: Mus. Naturk., Berlin; typelocality: Cameroon. 1941 Chlorocypha cancellata - FRÄSER, Proc. R. ent. soc. London, (B) 10: 39 (Cameroon). 1949 Chlorocypha cancellata - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 26 (redescription). 1949 Libellago hintzi - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 26 [proposed synonymy with cancellata (SELYS, 1879)]. 1950 Chlorocypha cancellata - LAIDLAW, Trans. R. ent. Soc. London, 101: 270 (Cameroon). 1952 Chlorocypha hintzi - PINHEY, Entomologist's month. Mag., 88: 172 (as valid species, redescription). 1962 Chlorocypha hintzi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149. 1969 Chlorocypha cancellata insulana PINHEY, Occ. Pap. natn. Mus. Rhod., 4 (29 B): 223. Type male: British Museum (Nat. Hist.), London; type-locality: Fernando Po

Distribution: West Africa centripunctata

GAMBLES

1974 Chlorocypha centripunctata GAMBLES, Entomologist's month. Mag., 110: 106. Type male: Gambles's Collection; type-locality: Nigeria.

Distribution: Nigeria, curta

(HÄGEN)

1853 Libellago curta HAGEN, in SELYS, Synopsis des Calopterygines: 58. Type male: Naturhist. Mus., Wien; type-locality: Guinea. 1854 Libellago curta - SELYS & H A G E N , Monogr. Cal.: 229 (amplified description of both sexes; female from Sierra Leone). 1890 Libellago curta - KIRBY, Syn. Cat. Neur.-Odon., London: 112. 1893 Libellago decorata KARSCH, Berl.ent. Z.,38:34. Type male: Mus. Naturk., Berlin; type-locality: West Africa. 1936 Chlorocypha curta - LONGFIELD, Trans. R. ent. Soc. London, 85: 467. 1936 Libellago decorata - LONGFIELD, Trans. R. ent. Soc. London, 85: 467 [proposed synonymy with curta ( H Ä G E N , 1 8 5 3 ) ] . 1941 Chlorocypha curta race bicolor FRASER, Proc. R. ent. Soc. London, (B) 10: 39. Male from Uganda.

50 1949 Chlorocypha 1950 Chlorocypha

curia - FRÄSER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 25 (redescription). curia - LAIDLAW, Trans. R. ent. Soc. London, 101: 270 (Sierra Leone).

Distribution: from Sierra Leone to Uganda. eyanifrons

(SELYS)

1873 Libellago eyanifrons SELYS, Bull. Acad. r. Belg., (2) 35: 493. Type female: unknown; typelocality: G a b o o n . 1900 Libellago glaucifrons SJÖSTEDT, Bih. Svenska Akad. Handl., 25 (4,2): 51. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1947 Chlorocypha dispar race cordosa FRASER, Trans. R. ent. Soc. London, 98:23. Male f r o m G a b o n . 1949 Chlorocypha eyanifrons - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 33 (redescription). 1966 Chlorocypha dispar cordosa - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 187.

Distribution: West Africa. dahli

FRASER

1956 Chlorocypha dahli FRASER, Bull. Inst, franc. Afr. noire, (A) 18: 940. Type male: unknown; type-locality: Cameroon.

Distribution: Cameroon. dispar dispar

(BEAUVOIS)

1805 Agrion dispar BEAUVOIS, Ins. Afr.-Amer. (Neur.), 7: 2: 85. Type male: British Museum (Nat. Hist.), London; type-locality: Sierra Leone. 1853 Libellago (Libellago) dispar - SELYS, Synopsis des Calopyterygines: 58 (Sierra Leone). 1 8 5 4 Libellago (Libellago) dispar - SELYS & HAGEN, Monogr. Cal.: 2 2 6 (redescription). 1890 Libellago dispar - KIRBY, Syn. Cat. Neur.-Odon., London: 112 (as generotype of Libellago auct.) 1934 Chlorocypha dispar - FRASER, Trans. R. ent. Soc. London, 98: 21. 1934 Chlorocypha (Agrion) dispar - FRASER, F a u n a brit. India, Odon., 2: 56 (as generotype of Chlorocypha FRASER, 1934). 1947 Chlorocypha dispar - FRASER, Trans. R. ent. Soc. London, 98: 21 (Ivory Coast). 1949 Chlorocypha dispar - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 36 (redescription). 1950 Chlorocypha dispar - LAIDLAW, Trans. R. ent. Soc. London, 101: 270 (Gaboon). 1951 Chlorocypha fraseri SCHMIDT, Arq. Mus. Bocage, 20: 161. Type male: Schmidt's Collection; type-locality: Portuguese Guinea. 1962 Chlorocypha dispar dispar - PINHEY, Publ. cult. Comp. Diam. Angola, 59:149 (Belgian Congo, Ivory Coast, Cameroon, Sierra Leone, Portuguese Guinea, Liberia, G h a n a , Nigeria, Togo). 1962 Chlorocypha dispar fraseri - PINHEY, Publ. cult. Comp. Diam, Angola, 59: 149.

Distribution: West Africa. dispar ovulosa

FRASER

1947 Chlorocypha dispar ovulosa FRASER, Trans. R. ent. Soc. London, 98: 23. Type male: originally in McLachlan's Collection, now unknown; type-locality: Old Calabar. 1962 Chlorocypha dispar ovulosa - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149. 1984 Chlorocypha dispar f. ovulosa - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:15.

Distribution: Cameroon. dispar pyriformosa

FRASER

1947 Chlorocypha dispar pyriformosa FRASER, Trans. R. ent. Soc. London, 98: 23. Type male: British Museum (Nat. Hist.), London; type-locality: Ivory Coast. 1 9 6 2 Chlorocypha

dispar

pyriformosa

— PINHEY, P u b l . c u l t . C o m p . D i a m . A n g o l a , 5 9 : 1 4 9 .

51 1984

Chlorocypha dispar f. pyriformosa

- DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 1 5 .

Distribution: Ivory Coast. frigida 1984

PINHEY

Chlorocypha frigida Utrecht, 1: 15.

PINHEY, 1 9 6 1

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: North Rhodesia. ghesquierei

FRASER

1959 Chlorocypha ghesquierei FRASER, Rev. Zool. Bot. afr., 60: 308. Type male: Mus. r. Afr. centra., Tervuren; type-locality: Belgian Congo. 1962 Chlorocypha ghesquierei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149.

Distribution: Congo Republic. glauca glauca

(SELYS)

1879 Libellago glauca SELYS, Bull. Acad. r. Belg., (2) 47: 380. Type male: ? Mus. Naturk., Berlin; type-locality: Cameroon (Mountains). 1949 Chlorocypha glauca - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 23. 1959 Chlorocypha glauca glauca - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 28.

Distribution: Cameroon. glauca radix

LONGFIELD

1959 Chlorocypha glauca radix LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 28. Type male: British Museum (Nat. Hist.), London; type-locality: Nigeria. 1973 Chlorocypha pavonis LIEFTINCK, Odonatologica, 5 (2): 34. Type: unknown; type-locality: Togo.

Distribution: Nigeria, Togo. grandis

(SJÖSTEDT)

1899 Libellago grandis SJÖSTEDT, Bih. Svenska Akad. Handl., 25 (4): 58. Type female: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1899 Libellago gracilis KARSCH, Ent. Nachr., 25 (11): 3. Type male: Mus. Naturk., Berlin; typelocality: Cameroon. 1949 Libellago grandis - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 48 (in incertae sedis). 1949 Chlorocypha gracilis - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 36 [incorrect synonymy with dispar dispar (BEAUVOIS, 1805)]. 1962 Chlorocypha grandis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149. 1962 Chlorocypha gracilis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149 (CAMEROON, Belgian Congo, French Guinea). 1 9 8 4 Chlorocypha grandis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 5 [proposed synonymy with gracilis ( K A R S C H , 1 8 9 9 ) ] .

Distribution: Cameroon, Congo Republic, French Guinea. helenae 1984

LEGRAND

Chlorocypha helenae Utrecht, 1: 15.

LEGRAND, 1 9 8 4

Distribution: Gabon.

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

52 luminosa

(KARSCH)

1893 Libellago luminosa KARSCH, Berl. ent. Z., 38: 33. Type male: Mus. Naturk., Berlin; typelocality: Togo. 1898 Libellago jejuna KARSCH, Ent. Nachr., 24 (22): 345. Type male: Mus. Naturk., Berlin; typelocality: Togo. 1899 Libellago consueta KARSCH, Ent. Nachr., 25 (24): 376. Type male: Mus. Naturk., Berlin; typelocality: Nyassa. 1941 Chlorocypha luminosa - FRÄSER, Proc. R. ent. Soc. London, (B) 10: 39. 1941 Libellago jejuna - FRASER, Proc. R. ent. Soc. London, (B) 10: 39 [proposed synonymy with luminosa

(KARSCH, 1893)].

1950 Chlorocypha luminosa - LAIDLAW, Trans. R. ent. Soc. London, 101: 270 (Togoland). 1962 Chlorocypha luminosa - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 149 (Togo, Belgian, Congo, South Tanganyika, Nyasaland, Rhodesia, Mozambique). 1962 Libellago consueta - PINHEY, Publ. cult. Comp.Diam. Angola, 59: 149 [proposed synonymy with luminosa ( K A R S C H , 1 8 9 3 ) ] .

Distribution: tropical Africa. molindica

FRASER

1934 Libellago trifaria SCHOUDETEN (nec K A R S C H , 1899), Ann. Mus. Congo Beige, Zool., 3 ( 3 ) 1: 76. 1948 Chlorocypha molindica FRASER, Proc. R. ent. Soc. London, (B) 17: 9. Type male: Mus.r. A f t . centr., Tervuren; type-locality: Belgian Congo. 1950 Chlorocypha molindica - LAIDLAW, Trans. R. ent. Soc. London, 101: 270 (Congo). 1960 Chlorocypha hasta PINHEY, Occ. Pap. nat. Mus. S. Rhodesia, 3: 510. Type: unknown; typelocality: Tanganyika. 1962 Chlorocypha molindica - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150 (Belgian Congo, Uganda, Ruanda-Urundi). 1 9 6 2 Libellago graueri SCHMIDT, 1 9 6 2 - loc. cit. as Chlorocypha graueri - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 151 (as type male in Naturhist. Mus., Wien, and proposed synonymy with molindica FRASER, 1 9 4 8 ) .

Distribution: tropical Africa. neptuna

(SJÖSTEDT)

1899 Libellago neptunus SJÖSTEDT, Bih. Svenska, Akad. Handl., 25 (2): 54. Type male: Naturhist. Riksmus., Stockholm; type-locality: Bonge (Cameroon). 1949 Chlorocypha neptunus - FRASER, Bull. Inst. Sei. nat. Belg., 25 (6): 38. 1950 Chlorocypha neptuna - LAIDLAW, Trans. R. ent. Soc. London, 101: 270 (Cameroon). 1962 Chlorocypha neptuna - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150.

Distribution: Cameroon. nigeriensis

GAMBLES

stat. nov.

1974 Chlorocypha selysi nigeriensis GAMBLES, Entomologist's month. Mag., 110: 115. Type male: Gambles's Collection; type-locality: Cameroon.

Distribution: Cameroon. rubida

(HÄGEN)

1853 Libellago rubida HAGEN, in SELYS, Synopsis des Calopterygines; 58. Type male: Univ. zool. Mus., Kobenhavn; type-locality: Guinea. 1854 Libellago (Libellago) rubida - SELYS & H A G E N , Monogr. Cal.: 228 (redescription). 1890 Libellago rubida - KIRBY, Syn. Cat. Neur.-Odon., London: 112. 1949 Chlorocypha rubida - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 20 (redescription). 1950 Chlorocypha

rubida - LAIDLAW, Trans. R . ent. Soc. L o n d o n , 101: 270 ( C a m e r o o n ) .

53 1962 Chlorocypha rubida - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150 (Guinea, Cameroon, Belgian Congo, Uganda).

Distribution: Equatorial Africa. rubriventris

PINHEY

1 9 8 4 C h l o r o c y p h a r u b r i v e n t r i s PINHEY, 1 9 7 5 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e

World, Utrecht, 1: 15.

Distribution: Rhodesia. schmidti

PINHEY

1984 Chlorocypha schmidti PINHEY, 1967 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 15.

Distribution: Congo. selysi

(KARSCH)

1899 Libellago selysi KARSCH, Ent. Nachr., 25: 165. Type male: Mus. Naturk., Berlin; type-locality: Belgian Congo. 1899 Libellago camerunensis SJÖSTEDT, Bih. Svenska Akad. Handl., 25(2): 56. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1936 Chlorocypha selysi - LONGFIELD, Trans. R. ent. Soc. London, 85: 468. 1941 Chlorocypha dispar selysi - FRÄSER, Proc. R. ent. Soc. London, (B) 10: 39 (Cameroon). 1949 Chlorocypha selysi - FRASER, Bull. Inst. Sei. nat. Belg., 25 (6): 26. 1962 Chlorocypha selysi - PINHEY, Publ. cult. Comp. Diam. Angola, 59; 150 (Cameroon, Congo, U g a n d a , Nigeria).

Distribution: West Africa. seydeli

FRASER

1958 Chlorocypha seydeli FRASER, Rev. Zool. Bot. afr., 57: 109. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1962 Chlorocypha seydeli - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150.

Distribution: Congo Republic. sharpae

PINHEY

1971 Chlorocypha sharpae PINHEY, Arnoldia, Rhodesia, 5 (18): 1. Type male: Pinhey's Collection; type-locality: Liberia.

Distribution: Liberia. tenuis

LONGFIELD

1912 Libellago lanceolata MARTIN, - MS name only. 1912 Libellago alata MARTIN, MS name only from Ruwenzori. 1936 Chlorocypha tenuis LONGFIELD, Trans. R. ent. Soc. London, 85: 468. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1962 Chlorocypha tenuis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150.

Distribution: Uganda. trifaria

(KARSCH)

1899 Libellago trifaria KARSCH (nec SCHOUDETEN, 1934), Ent. Nachr., 25 (24): 378. Type female: Mus. Naturk., Berlin; type-locality: Central Africa. 1914 Libellago trifascia (misprint) - GRÜNBERG, Ent. Rundsch., 31: 53. 1949 Libellago trifaria - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 47 (in incertae sedis).

54 1949 Chlorocypha straeleni FRÄSER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 28. Type male: R. Mus., Brussels; type-locality: Uganda. 1962 Chlorocypha straelini- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150 (Uganda, Congo). 1 9 6 2 Chlorocypha straeleni bwamba LONGFIELD, 1 9 5 9 - loc. cit. PINHEY, Publ. cult. Comp. Diam. Angola, 59: 150 [male in Sommer's Collection - British Museum (Nat. Hist.), London]. 1963 Chlorocypha trifaria - PINHEY, J. ent. Soc. S. Afr., 26: 152. 1963 Chlorocypha straeleni- PINHEY, J . ent. Soc. S. Afr., 2 6 : 152 [proposed synonymy with trifaria (KARSCH, 1899)].

Distribution: Central Africa. victoriae (FÖRSTER) 1914 Libellago rubida victoriae FÖRSTER, Arch. Natur., 80 (A): 61: Type male: Förster's Collection; type-locality: Uganda. 1929 Libellago victoriae - SJÖSTEDT, Revue Zool.-bot. Afr., 17: 55. 1949 Chlorocypha victoriae - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 30. 1959 Chlorocypha victoriae - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 29. 1962 Chlorocypha victoriae - PINHEY, Publ. cult. Comp. Diam. Angola 59: 150 (Uganda, Belgian Congo, Angola).

Distribution: Uganda, Congo Republic, Angola, wittei FRASER 1955 Chlorocypha wittei FRASER, Explor. Pare nat. Upemba, Miss, de Witte, 38 (1): 10. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1962 Chlorocypha wittei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 151. 1 9 8 4 Chlorocypha fabamacula PINHEY, 1 9 6 1 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 4 (proposed synonymy with wittei FRASER, 1 9 5 5 ) .

Distribution: Congo Republic. Genus Cyrano NEEDHAM & GYGER 1939 Cyrano

Philipp. J. Sei., 7 0 (3): 250. Type species-.Rhinocypha unicolor 1869 (original designation). - 1939 Neocypha TILLYARD & FRASER, Austr. Zool., 9 : 207 (nom. nud.). - 1950 Cyrano - FRASER, Bull. Inst. Sci.nat. Belg., 26 (18): 17 (type species: Libellago asiatica SELYS, 1879). - 1950 Cyrano - LAIDLAW, Trans. R. ent. Soc. London, 101: 270. - 1957 Cyrano - FRASER, R. Zool. Soc. N.S. Wales, 1 9 5 7 : 74. - 1957 Neocypha - FRASER, R. Zool. Soc. N.S. Wales, 1 9 5 7 : 74 (proposed synonymy with Cyrano NEEDHAM & G Y G E R , 1939). NEEDHAM & G Y G E R ,

HÄGEN,

unicolor (HÄGEN) 1869 Rhinocypha unicolor HAGEN, in SELYS, Synopsis des Calopterygines, Add.: 21. Type female: not given; type-locality: Manila. 1879 Libellago asiatica SELYS, Bull. Acad. r. Belg., (2) 47: 384. Type male: Selys's Collection; typelocality: Luzon. 1891 Libellago asiatica - SELYS, Ann. Soc. Esp. Hist, nat., 20: 216. 1939 Cyrano unicolor - NEEDHAM & G Y G E R , Philipp. J. Sei., 7 0 (3): 250 (as generotype of Cyrano NEEDHAM & G Y G E R , 1939). 1939 Libellago asiatica - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 255. 1950 Cyrano unicolor - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1950 Libellago asiatica - FRASER, Bull. Inst. Sei. nat. Belg., 26 (18): 17 (as generotype of Cyrano NEEDHAM & G Y G E R , 1 9 3 9 ) . 1984

Libellago asiatica - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, synonymy with unicolor ( H Ä G E N , 1 8 6 9 ) ] .

Distribution: Philippine Islands.

1:16

[proposed

55 Genus Disparocypha Ris 1916 Disparocypha Ris, Ent. Mitt. Berlin, 5: 313. Type species: Disparocypha biedermanni Ris, 1916 (original designation). - 1950 Disparocypha - LAIDLAW, Trans. R. ent. Soc. London, 101: 274.

biedermanni Ris 1916 Disparocypha biedermanni Ris, Ent. Mitt. Berlin, 5: 313. Type male: Mus. Naturk., Berlin; type-locality: Celebes.

Distribution: Celebes. Genus Heliocypha

FRASER

1949 Heliocypha FRASER, Bull. Inst. r. Sei. nat. Belg., 25: 75. Type species-.Rhinocypha bisignata HÄGEN, 1853. - 1950 Heliocypha - LAIDLAW, Trans. R. ent. Soc. London, 101: 272. - 1954 Heliocypha - LIEFTINCK, Treubia, 22 Suppl.: 7 (incorrect synonymy with Rhinocypha RAMBUR, 1842). - 1984 Heliocypha - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 17.

angusta

(HÄGEN)

1853 Libellago (Rhinocypha) angusta HAGEN, in SELYS, Synopsis des Calopterygines: 62. Type male: Univ. Zool. Mus., Kebenhavn.; type-locality: Sumatra. 1854 Libellago (Rhinocypha) angusta - SELYS & HAGEN, Monogr. Cal.: 212 (redescription). 1890 Rhinocypha angusta - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1898 Rhinocypha apicalis KRÜGER, Stettiner ent. Ztg., 59: 79. Type male: unknown, perhaps in Warszawa Museum; type-locality: Sumatra. 1903 Rhinocypha apicalis - LAIDLAW, Fasc. Malayenses Zool., 1: 196 (Perak). 1926 Rhinocypha angusta - FRASER, Treubia, 8 (3-4): 485. 1927 Rhinocypha angusta - Ris, Zool. Meded., Leiden, 10 (1): 6 (Sumatra). 1934 Rhinocypha angusta - SCHMIDT, Arch. Hydrob. Suppl., 13: 322. 1935 Rhinocypha angusta - LIEFTINCK, Misc. Zool. Sum., 92-93: 4. 1935 Rhinocypha apicalis - LIEFTINCK, Misc. Zool. Sum., 92-93: 4 [proposed synonymy with angusta (HÄGEN, 1853)]. 1947 Rhinocypha angusta oceanis LIEFTINCK, Tijdschr. Ent., 88: 220. Type male: unknown; typelocality: Engano Island. 1950 Heliocypha angusta angusta - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1950 Heliocypha angusta oceanis - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1984 Heliocypha angusta f. oceanis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 17.

Distribution: Sumatra, Engano Island. biforata

(SELYS)

1859 Libellago (Rhinocypha) biforata SELYS, Synopsis des Calopterygines, Add. 1:12. Type male: Selys's Collection; type-locality: Malacca. 1890 Rhinocypha biforata - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1917 Rhinocypha biforata - LAIDLAW, Ree. Indian Mus., 13: 37 (Malay Peninsula to Burma). 1922 Rhinocypha beesoni FRASER, Mem. Dept. Agr. India, 7: 61. Type male: British Museum (Nat. Hist.), London; type-locality: North India. 1928 Rhinocypha biforata abbreviate FRASER, J. Bombay nat. Hist. Soc., 32: 454. Type male: British Museum (Nat. Hist.), London; type-locality: Assam. 1950 Rhinocypha biforata abbreviata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Heliocypha biforata delimbata LAIDLAW, Trans. R. ent. Soc. London, 101: 277 (after SELYS, 1879).

1950 Rhinocypha beesoni - LAIDLAW, Trans. R. ent. Soc. London, 101: 277 [proposed synonymy w i t h biforata

(SELYS, 1859)].

1954 Rhinocypha biseriata biforata - LIEFTINCK, Treubia, 22 Suppl.: 9 (Siam, Malaya). 1966 Heliocypha biforata abbreviata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 176.

56 1966 Heliocypha beesoni - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 181 (as good species). 1984 Heliocypha biforata f. abbreviata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 16. 1984 Heliocypha biforata f. beesoni - Davies & Tobin, The Dragonflies of the World, Utrecht, 1:16.

Distribution: East Asia, biseriata

(SELYS) 1859 Libellago (Rhinocypha) biseriata SELYS, Synopsis des Calopterygines, Add. 1: 11. Type male: Selys's Collection; type-locality: Borneo. 1890 Rhinocypha biseriata - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1932 Rhinocypha biseriata anambae LAIDLAW, Bull. Raffles Mus., 7: 99. Type: unknown; typelocality: Anamba Island. 1950 Heliocypha biseriata biseriata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Heliocypha biseriata anambae - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1984 Heliocypha biseriata f. anambae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 16.

Distribution: Borneo, Anamba Island, bisignata

(HÄGEN) 1853 Libellago (Rhinocypha) bisignata HAGEN, in SELYS, Synopsis des Calopterygines: 62. Type male: Westermann's Collection; type-locality: Bengal. 1853 Libellago bisignata - WALKER, List Neur. Ins. Β. M., 4: 468. 1854 Libellago (Rhinocypha) bisignata - SELYS & HAGEN, Monogr. Cal.: 214 (redescription). 1890 Rhinocypha bisignata - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1917 Rhinocypha bisignata - LAIDLAW, Ree. Indian Mus., 13: 38 (Conchin State). 1924 Rhinocypha bisignata - FRÄSER, Ree. Indian Mus., 26: 483 (West India). 1949 Heliocypha bisignata- FRASER, Bull. Inst. r. Sei. nat. Belg., 25: 75 (as generotype of Heliocypha FRASER, 1 9 4 9 ) .

1950 Heliocypha bisignata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: India. fenestrata fenestrata (BURMEISTER) 1839 Calopteryx fenestrata BURMEISTER, Handb. Ent., 2: 826. Type male: Mus. Naturk., Berlin; type-locality: Java. 1842 Rhinocypha vitrella RAMBUR, Hist. nat. Ins. Neuropt., Paris: 234. Type male: Serville's Collection; type-locality: Java. 1842 Rhinocypha infumata RAMBUR, Hist. nat. Ins. Neuropt., Paris: 237. Type: sex not given; Serville's Collection; type-locality: Java. 1853 Libellago (Rhinocypha) fenestrata - SELYS, Synopsis des Calopterygines: 62. 1854 Libellago (Rhinocypha) fenestrata - SELYS & HÄGEN, Monogr. Cal.: 216 (redescription). 1890 Rhinocypha fenestrata - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1898 Rhinocypha fenestrata - CALVERT, Trans. Am. ent. Soc., 25: 47. 1934 Rhinocypha fenestrata - SCHMIDT, Arch. Hydrobiol., Suppl.: 13: 322. 1947 Rhinocypha fenestrata fenestrata - LIEFTINCK, Tijdsch. Ent., 88: 217. 1950 Heliocypha fenestrata fenestrata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1954 Rhinocypha fenestrata fenestrata - LIEFTINCK, Treubia, 22: Suppl.: 10.

Distribution: Java. fenestrata cornelii

(LIEFTINCK)

1947 Rhinocypha fenestrata cornelii LIEFTINCK, Tijdschr. Ent., 88: 218. Type male: Buitenzorg Museum; type-locality: Bali. 1 9 5 4 Rhinocypha fenestrata cornelii — L I E F T I N C K , Treubia, 2 2 Suppl.: 1 0 .

57 1984 Heliocypha fenestrata

cornelii-

DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:16.

Distribution: Bali, mariae

(LIEFTINCK)

1930 Rhinocypha mariae Sumatra. 1934 Rhinocypha mariae 1950 Heliocypha mariae 1954 Rhinocypha mariae

LIEFTINCK, Treubia, 12:136. Type male: Buitenzorg Museum; type-locality: - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 323. - LAIDLAW, Trans. R. ent. soc. London, 101: 278. - LIEFTINCK, Treubia, 22 Suppl.: 11 (Sumatra).

Distribution: Sumatra. nubecula

LIEFTINCK

1948 Rhinocypha nubecula LIEFTINCK, Treubia, 19 (2): 266. Type male: not given; type-locality: Sumatra. 1950 Heliocypha nubecula - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1954 Rhinocypha nubecula - LIEFTINCK, Treubia, 22: Suppl.: 12 (Sumatra).

Distribution: Sumatra, perforata

(PERCHERON)

1835 Agrion perforatus PERCHERON, Gen. Ins. Neuropt.: 2. Type male: unknown; type-locality: Cochin-China. 1842 Rhinocypha perforata - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 235. 1853 Libellago (Rhinocypha) perforata - SELYS, Synopsis des Calopterygines: 63. 1853 Libellago perforata - WALKER, List Neur. Ins. Β. M., 4: 647. 1 8 5 4 Libellago (Rhinocypha) perforata - SELYS & H A G E N , Monogr. Cal.: 2 1 9 (redescription). 1879 Rhinocypha perforata var. limbata SELYS, Bull. Acad. r. Belg., (2) 47: 392. Male from Assam. 1890 Rhinocypha perforata - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1900 Rhinocypha whiteheadi KIRBY, Ann. Mag. nat. Hist., (7) 5: 536. Type male: British Museum (Nat. Hist.), London; type-locality: Hainan. 1902 Rhinocypha inas LAIDLAW, Proc. Zool. Soc. London, 1: 88. Type male: Laidlaw's Collection; type-locality: Malay Peninsula. 1914 Rhinocypha quattuordecimmaculata O G U M A , J . coll. Agr. Tohuku imp. Univ., 5: 159. Type male: H o k k a i d o Univ.; type-locality: Japan. 1917 Rhinocypha perforata whiteheadi - LAIDLAW, Ree. Indian Mus., 13: 38. 1927 Rhinocypha perforata beatifica FRÄSER, Ree. Indian Mus., 29: 86. Type male: British Museum (Nat. Hist.), London; type-locality: Assam. 1931 Rhinocypha perforata limbata - LAIDLAW, J. Fed. Mai. States Mus., 16: 181. 1 9 3 1 Rhinocypha albistigma MATSUMURA (nec SELYS, 1 8 7 3 ) , 6 0 0 0 111. Ins. Japan (as species nova after Zool. Ree.). 1950 Heliocypha perforata perforata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Heliocypha perforata limbata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Rhinocypha inas - LAIDLAW, Trans. R. ent. Soc. London, 101: 277 [proposed synonymy with perforata perforata (PERCHERON, 1 8 3 5 ) ] . 1950 Rhinocypha quattuordecimmaculata - LAIDLAW, Trans. R. ent. Soc. London, 101: 278 [proposed synonymy with perforata perforata (PERCHERON, 1 8 3 5 ) ] . 1950 Heliocypha perforata beatifica - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1954 Rhinocypha perforata perforata - LIEFTINCK, Treubia, 22, Suppl.: 13 (Siam, Malaya). 1961 Rhinocypha perforata - ASAHINA, Insecta matsum., 24: 58. 1966 Heliocypha perforata beatifica - KIMMINS, Bull. Brit. Mus. Nat. Hist. (Ent.), 18: 181. 1984 Heliocypha perforata f. beatifica - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 16.

Distribution: China, Vietnam, Burma, Malaysia, India.

58 Genus Heterocypha LAIDLAW 1950 Heterocypha LAIDLAW, Trans. R. ent. Soc. London, 101: 273. Type species: Rhinocypha vitrinella Fräser, 1935 (original designation).

vitrinella (FRÄSER) 1935 Rhinocypha (Nat. Hist.), 1950 Heterocypha Heterocypha

vitrinella FRÄSER, Ree. Indian Mus., 37: 332. Type male: British Museum London; type-locality: Assam. vitrinella - LAIDLAW, Trans. R. ent. soc. London, 101: 273 (as generotype of LAIDLAW, 1950).

Distribution: Assam. Genus Indocypha TILLYARD & FRÄSER 1939 Indocypha T I L L Y A R D & FRÄSER, Austr. Zool., 9: 206 (nom. nud.). - 1949 Indocypha FRÄSER, Bull. Inst. R. Sei. nat. Belg., 25: 75. Type species: Libellago vittata SELYS, 1891 (original designation). - 1950 Indocypha - LAIDLAW, Trans. R. ent. Soc. London, 101: 271.

svenhedini (SJÖSTEDT) 1933 Rhinocypha sven-hedini SJÖSTEDT, Ark. Zool., Uppsala, 25A (5): 15. Type male: Naturhist. Riksmus., Stockholm; type-locality: China. 1950 Indocypha svenhedini - LAIDLAW, Trans. R. ent. Soc. London, 101: 271 (Annam). 1984 Indocypha vittata f. svenhedini - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:17.

Distribution: China, Vietnam. vittata (SELYS) 1891 Libellago vittata SELYS, Ann. Soc. ent. Belg., 30: 58. Type male: Selys's Collection; type-locality: Burma. 1949 Indocypha vittata - FRÄSER, Bull. Inst. R. Sei. nat. Belg., 25: 75 (as generotype of Indocypha T I L L Y A R D & FRÄSER, 1939). 1950 Indocypha vittata - LAIDLAW, Trans. R. ent. Soc. London, 101: 271 (Burma).

Distribution: Burma.

Genus Libellago SELYS 1840 Libellago SELYS, Monogr. Libell. europ., Paris: 200. Type species: Calopteryx lineata BURMEISTER, 1 8 3 9 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 1 2 ) . - 1 8 4 2 Micromerus RAMBUR, Hist. nat. Ins. Neuropt., Paris: 238. Type species: Calopteryx lineata BURMEISTER, 1 8 3 9 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 1 5 ) . - 1 8 5 3 Libellago - SELYS, Synopsis des Calopterygines: 57 (with subgenera Libellago s.str., Rhinocypha). - 1854 Libellago - SELYS, Mem. Soc. r. Sei. Liege, 9:198 (revision, key to subgenera). - 1 8 5 4 Libellago - SELYS & H Ä G E N , Monogr. Cal.: 1 9 2 (revision, key to subgenera). - 1 8 9 0 Libellago - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 1 2 . - 1 9 0 5 Micromerus - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 6 . - 1 9 2 8 Micromerus - FRÄSER, J . Bombay nat. Hist. Soc., 32: 6 8 4 (proposed synonymy with Libellago SELYS, 1 8 4 0 ) . - 1 9 3 4 Micromerus - SCHMIDT, Arch. Hydrob. Suppl., 13: 324 (key to species). - 1934 Libellago - COWLEY, Entomologist's month. Mag., 70: 243. - 1934 Micromerus - COWLEY, Entomologist's month. Mag., 70: 243 (proposed synonymy with Libellago SELYS, 1 8 4 0 ) . - 1 9 4 9 Sclerocypha FRASER, Bull. Inst. R. Sei. nat. Belg., 25 ( 6 ) : 7 . Type species: Micromerus bisignatus M C L A C H L A N , 1 8 7 0 (original designation). - 1954 Libellago - LIEFTINCK, Treubia, 22 Suppl.: 2. - 1984 Libellago (Sclerocypha) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 7 .

59 adami

FRÄSER

1939 Libellago adami FRÄSER, Proc. R. ent. Soc. London, (B) 8: 24. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1940 Libellago miae LIEFTINCK, Spolia zeylan., 22: 84. Type male: unknown; type-locality: Ceylon. 1955 Libellago adami - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 68 (Ceylon). 1966 Drepanosticta adami - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 177. 1984 Libellago adami - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 17.

Distribution: Sri Lanka. andamanensis

(FRÄSER)

1924 Micromerus andamanensis FRÄSER, Ree. Indian Mus., 26: 410. Type male: British Museum (Nat. Hist.), London; type-locality: Andaman Island. 1984 Libelago andamensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:17.

Distribution: Andaman Island. asclepiades (Ris) 1916 Micromerus asclepiades Ris, Ent. Mitt. Berlin, 5: 303. Type male: Mus. Naturk., Berlin; typelocality: Celebes. 1984 Libellago asclepiades - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:17.

Distribution: Celebes. aurantiaca

(SELYS)

1859 Micromerus aurantiacus SELYS, Synopsis des Calopterygines, Add. 1: 14. Type male: Selys's Collection; type-locality: Malacca. 1903 Micromerus annandalei LAIDLAW, Fase. Malayenses Zool., 1: 197. Type male: Laidlaw's Collection; type-locality: Malay States. 1932 Libellago aurantiaca - LIEFTINCK, Konowia, 11:2. 1937 Libellago aurantiaca - LIEFTINCK, Treubia, 16 (1): 57 (Sumatra, Burma, Tonkin, Siam, Borneo). 1954 Libellago aurantiaca - LIEFTINCK, Treubia, 22 Suppl.: 2 (Siam, Borneo, Sumatra). 1954 Micromerus annandalei- LIEFTINCK, Treubia, 22 Suppl.: 2 [proposed synonymy with aurantiaca (SELYS, 1859)].

Distribution: Sumatra, Burma, Vietnam, Thailand, Borneo. bisignata

(MCLACHLAN)

1870 Micromerus bisignatus MCLACHLAN, Trans. Ent. Soc. London, 1870: 168. Type male: British Museum (Nat. Hist.), London; type-locality: Celebes. 1873 Micromerus bisignatus - SELYS, Synopsis des Calopterygines, Add. 3: 31 (redescription). 1949 Sclerocypha bisignatus - FRÄSER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 7 (as generotype of Sclerocypha FRÄSER, 1949). 1984 Libellago (Sclerocypha) bisignata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 17.

Distribution: Celebes. collarti

NAVÄS

1929 Libellago collarti NAVÄS, Rev. Zool. Bot. afr., 18 (1) 20. Type female: unknown; type-locality: Belgian Congo. 1949 Libellago collarti - FRASER, Bull. Inst. r. Sei. nat. belg., 25 (6): 47 (among incertae sedis).

Distribution: Congo Republic.

60 dorsocyana

LIEFTINCK

1937 Libellago dorsodyana LIEFTINCK, Treubia, 16 (1): 56. Type male: Buitenzorg Museum; typelocality: South Borneo. 1950 Libellago dorsocyana - LAIDLAW, Trans. R. ent. Soc. London, 101: 269. 1954 Libellago dorsocyana - LIEFTINCK, Treubia, 22 Suppl.: 3 (Borneo).

Distribution: Borneo. finalis

(HÄGEN)

1869 Micromerus finalis HAGEN, in SELYS, Synopsis des Calopterygines, Add. 1: 21. Type male: not given; type-locality: Ceylon. 1890 Micromerus finalis - KIRBY, Syn. Cat. Neur.-Odon., London: 115. 1917 Micromerus finalis - LAIDLAW, Ree. Indian Mus., 13: 39 (Ceylon). 1955 Libellago finalis - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 68.

Distribution: Sri Lanka. greeni

(LAIDLAW)

1924 Micromerus greeni LAIDLAW, Spol. zeylan., 12: 352. Type male: unknown; type-locality: Ceylon. 1955 Libellago greeni - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 68 (Ceylon).

Distribution: Sri Lanka. hyalina

(SELYS)

1859 Micromerus hyalinus SELYS, Synopsis des Calopterygines, Add. 1: 13. Type male: Selys's Collection; type-locality: Malacca. 1927 Micromerus hyalinus - Ris, Zool. Meded., Leiden, 10 (1): 10 (Sumatra). 1932 Libellago hyalina - LIEFTINCK, Konowia, 11: 2. 1935 Libellago hyalina - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 , 5. 1954 Libellago hyalina - LIEFTINCK, Treubia, 22: Suppl.: 3 (Malaya, Sumatra, Borneo). 1 9 8 4 Libellago malayana S T . Q U E N T I N , 1 9 6 6 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 17.

Distribution: Malaysia, Sumatra, Borneo, lineata lineata

(BURMEISTER) 1839 Calopteryx lineatus Burmeister, H a n d b . Ent., 2: 826. Type male: Winthem's Collection (perhaps in Mus. comp. Zool. Cambridge); type-locality: Java. 1842 Micromerus lineatus - RAMBUR, Hist. nat. Ins. Neuropt., Paris; 238 (as generotype of Micromerus RAMBUR, 1842). 1842 Micromerus uxor RAMBUR, Hist. nat. Ins. Neuropt., Paris: 239. Type female: unknown; typelocality: Java. 1853 Micromerus blandus HAGEN, in SELYS, Synopsis des Calopterygines: 64. Type male: Univ. zool. Mus., Kobcnhavn; type-locality: Nicobar. 1854 Micromerus blandus - SELYS & H A G E N , Monogr. Cal.: 254 (redescription). 1854 Micromerus lineatus - SELYS & H Ä G E N , Monogr. Cal.: 256 (redescription). 1886 Micromerus obscurus KIRBY, Proc. Zool. Soc. London, 1886: 328. Type male: British Museum (Nat. Hist.), London; type-locality: Hassan Abdal. 1898 Micromerus signatus KRÜGER, Stettiner ent. Ztg., 59: 86. Type male: ? Warszawa Museum; type-locality: Pehang. 1924 Micromerus lineatus - FRÄSER, Ree. Indian Mus., 26: 483 (West India). 1932 Libellago lineata - LIEFTINCK, Konowia, 11 (1): 9. 1932 Micromerus signatus - LIEFTINCK, Konowia, 11 (1): 9 [proposed synonymy with lineata lineata (BURMEISTER, 1 8 3 9 ) ] .

1934 Micromerus

lineatus lineatus -

SCHMIDT,

Arch. Hydrob. Suppl.,

13.

325 (Java, Sumatra).

61 1934 Libellago lineata lineata - LIEFTINCK, Treubia, 14 (4): 387 (Java). 1 9 3 5 Libellago lineata lineata - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 5 (Sumatra). 1 9 3 9 Libellago lineata - N E E D H A M & G Y G E R , Philipp J . Sei., 7 0 ( 3 ) : 2 5 6 . 1942 Libellago lineata - FRÄSER, Proc. R. ent. Soc. London, (B) 11: 98 (Indo-China, Java). 1954 Libellago lineata lineata - LIEFTINCK, Treubia, 22 Suppl.: 3 (Siam, Malaya, Sumatra, Java, Borneo). 1964 Micromerus lineatus - SCHMIDT, Ent. Tidschr., 85: 153 (Burma).

Distribution: East Asia, lineata indica

(FRASER)

1928 Micromerus lineatus indicus FRÄSER, J. Bombay nat. Hist. Soc., 32: 686. Type male: British Museum (Nat. Hist.), London; type-locality: South India. 1931 Micromerus lineatus indicus - FRÄSER, Ree. Indian Mus., 33: 463. 1955 Libellago indica - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 68 (Ceylon). 1966 Libellago lineata indica - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 197.

Distribution: Sri Lanka, South India, naias

LIEFTINCK

1932 Libellago naias LIEFTINCK, Konowia, 11 (1): 4. Type male: Buitenzorg Museum; type-locality: Flores. 1936 Libellago naias - LIEFTINCK, Revue suisse Zool., 43 (5): 116 (Flores, Sumba). 1953 Libellago naias - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 137.

Distribution: Flores, Sumba. phaethon (LAIDLAW) 1931 Micromerus phaethon LAIDLAW, J. Fed. Mai. States Mus., 16: 245. Type male: Laidlaw's Collection; type-locality: North Borneo. 1932 Libellago phaethon - LIEFTINCK, Konowia, 11:2. 1937 Libellago phaethon - COWLEY, Trans. R. ent. Soc. London, 86: 4. 1954 Libellago phaethon - LIEFTINCK, Treubia, 22 Suppl.: 4.

Distribution: Borneo. rufescens

(SELYS)

1873 Micromerus rufescens SELYS, Synopsis des Calopterygines, Add.: 64. Type male: British Museum (Nat. Hist.), London; type-locality: Celebes. 1984 Libellago rufescens - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 18.

Distribution: Celebes. semiopaca

(SELYS)

1873 Micromerus semiopacus SELYS, Synopsis des Calopterygines, Add.: 64. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1890 Micromerus semiopacus - KIRBY, Syn. Cat. Neur.-Odon., London: 115. 1891 Micromerus martinae KARSCH, Ent. Nachr., 17 (16). 243. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1902 Micromerus affinis LAIDLAW, Proc. Zool. Soc. London, 1902, 1: 90. Type male: unknown; type-locality: Kwala Aring. 1932 Libellago mima LIEFTINCK, Konowia, 11: 2. Type male: R. Mus., Brussels; type-locality: Borneo. 1934 Micromerus semiopacus martinae - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 325. 1935 Libellago semiopaca - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 5 (Sumatra).

62 1935

Micromerus martinae - LIEFTINCK, Misc. Zool. Sum., semiopaca (SELYS, 1 8 7 3 ) ] .

92-93: 5

[proposed synonymy with

Distribution: Borneo, Sumatra. stictica

(SELYS)

1859 Micromerus stigmatizans race sticticus SELYS, Synopsis des Calopterygines, Add. 1: 14. Male from Borneo. 1869 Micromerus sticticus - SELYS, Synopsis des Calopterygines, Add. 2: 21. 1932 Libellago stictica - LIEFTINCK, Konowia, 11 (1): 10. 1954 Libellago stictica - LIEFTINCK, Treubia, 22 Suppl.: 5 (Borneo).

Distribution: Borneo. stigmatizans

(SELYS)

1859 Micromerus stigmatizans SELYS, Synopsis des Calopterygines, Add. 1:13. Type male: R. Mus., Brussels; type-locality: Malacca. 1932 Libellago stigmatizans - LIEFTINCK, Konowia, 11 (1): 9. 1936 Libellago stigmatizans - LIEFTINCK, Treubia, 16 (1): 58 (Sumatra). 1954 Libellago stigmatizans - LIEFTINCK, Treubia, 22 Suppl.: 6 (Malaya, Sumatra).

Distribution: Malaysia, Sumatra, sumatrana

(SELYS)

1879 Micromerus sumatranus SELYS, Bull. Acad. r. Belg., (2) 47: 397. Type male: Selys's Collection; type-locality: Sumatra. 1927 Micromerus sumatranus - RlS, Zool. Meded., Leiden, 10 (1): 10 (Sumatra). 1931 Libellago sumatrana - LIEFTINCK, Misc. Zool. Sum., 59: 3. 1934 Micromerus aurantiacus sumatranus - SCHMIDT, Arch. Hydrobiol Suppl., 13: 326. 1935 Libellago sumatrana - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 5. 1954 Libellago sumatrana - LIEFTINCK, Treubia, 22: Suppl.: 6 (Mentawai Island, Sumatra, Java). 1964 Micromerus aurantiacus sumatranus - SCHMIDT, Ent. Tidschr., 85: 153.

Distribution: Sumatra, Java, Mentawei Island, xanthocyana

(SELYS) 1869 Micromerus xanthocyanus SELYS, Synopsis des Calopterygines, Add. 2: 22. Type male: Rijksmus. Natuurl. Mus., Leiden; type-locality: Celebes. 1984 Libellago xanthocyana - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 18.

Distribution: Celebes. Genus Melanocypha

FRASER

1949 Melanocypha FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 7. Type species: Micromerus snellemanni Selys, 1879 (original designation). - 1950 Melanocypha - LAIDLAW, Trans. R. ent. Soc. London, 101: 269. - 1954 Melanocypha - LIEFTINCK, Treubia, 22 Suppl.: 2 (incorrect synonymy with Libellago SELYS, 1 8 4 0 ) .

snellemanni snellemanni

(SELYS)

1879 Micromerus snellemanni SELYS, Bull. Acad. r. Belg., (2) 47: 398. Type male: Selys's Collection; type-locality: Sumatra. 1932 Libellago snellemanni - LIEFTINCK, Konowia, 11: 2. 1934 Micromerus snellemanni - SCHMIDT, Arch.Hydrob. Suppl., 13: 325. 1935 Libellago snellemanni - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 5. 1949 Melanocypha snellemanni - FRASER, Bull. Inst. R. nat. Belg., 3325 (6): 11 (as generotype of Melanocypha

FRASER, 1 9 4 9 ) .

63 1950 Melanocypha snellemanni snellemanni - LAIDLAW, Trans. R. ent. Soc. London, 101: 269. 1954 Libellago snellemanni snellemanni - LIEFTINCK, Treubia, 22 Suppl.: 5.

Distribution: Sumatra. snellemanni javana

LAIDLAW

1950 Melanocypha snellemanni javana LAIDLAW, Trans. R. ent. Soc. London, 101: 274. Type male: British Museum (Nat. Hist.), London; type-locality: Java. 1954 Libellago snellemanni javana - LIEFTINCK, Treubia, 22: Suppl.: 5 (West Java).

Distribution: Java. Genus Pachycypha

LIEFTINCK

1950 Pachycypha LIEFTINCK, Treubia, 20 (3): 631. Type species: Pachycypha aurea LIEFTINCK, 1950 (original designation). - 1954 Pachycypha - LIEFTINCK, Treubia, 22 Suppl.: 2.

aurea

LIEFTINCK

Pachycypha aurea LIEFTINCK, Treubia, 20 (3): 634. (as generotype of Pachycypha LIEFTINCK, 1 9 5 0 ) . Type male: not given, perhaps in Buitenzorg Museum; type-locality: Borneo. 1954 Pachycypha aurea - LIEFTINCK, Treubia, 22 Suppl.: 6. 1950

Distribution: Borneo. Genus Paracypha

FRASER

1949 Paracypha FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 11. Type species: Rhinocypha culata SELYS, 1853 (original designation).

unimaculata

unima-

(SELYS)

1853 Libellago (Rhinocypha) unimaculata SELYS, Synopsis des Calopterygines: 61. Type male: Selys's Collection; type-locality: India. 1853 Libellago unimaculata - WALKER, List Neur. Ins. Β. M., 4: 649. 1854 Libellago (Rhinocypha) unimaculata - SELYS & H A G E N , Monogr. Cal.: 209 (redescription). 1890 Rhinocypha unimaculata - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1917 Rhinocypha unimaculata - LAIDLAW, Ree. Indian Mus., 13: 35 (Assam). 1949 Paracypha unimaculata - FRASER, Bull. Inst. Sei. nat. Belg., 25 (6): 11 (as generotype of Paracypha SELYS, 1 8 5 3 ) . 1950 Paracypha unimaculata - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

Distribution: India. Genus Platycypha

FRASER

1949 Platycypha FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 10. Type species: Libellago caligata SELYS, 1 8 5 3 (original designation). - 1 9 5 0 Platycypha - L A I D L A W , Trans. R. ent. soc. London, 101: 270. - 1950 Platycypha - FRASER, Bull. Inst. R. Sei. nat. Belg., 26 (18): 2 (key to species). - 1 9 5 8 Platycypha - PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, 23B: 1 0 4 (key to species, revision). - 1962 Platycypha - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 151.

amboniensis

(MARTIN)

1915 Libellago amboniensis MARTIN, Voy. Alluaud et Jeannel Afr., Odonata: 42. Type male: A n n A r b o r Museum; type-locality: Kenya. 1949 Libellago amboniensis- FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 49 (among incertae sedis). 1 9 5 8 Platycypha amboniensis - PINHEY, OCC. Pap. nat. Mus. S . Rhodesia, 2 2 B : 1 0 5 .

64 1962 Platycypha

amboniensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 151.

Distribution: Kenya. auripes

(FÖRSTER)

1906 Libellago auripes FÖRSTER, Mannheim Jb. Verk. Natk., 71/72: 54. Type male: A n n A r b o r Museum; type-locality: U s a m b a r a . 1949 Libellago auripes - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 49 (in incertae sedis). 1950 Platycypha greenwayi PINHEY, in FRASER, Bull. Inst. R. Sei. nat. Belg., 26 (18): 2. Type male: British Museum (Nat. Hist.), London; type-locality: Tanganyika. 1 9 5 8 Platycypha auripes - PINHEY, OCC. Pap. nat. Mus. S . Rhodesia, 2 2 B : 1 0 5 . 1 9 5 8 Platycypha greenwayi - PINHEY, OCC. Pap. nat. Mus. S . Rhodesia, 2 2 B : 1 0 5 [proposed synonymy with auripes (FÖRSTER, 1 9 0 6 ) ] . 1962 Platycypha

auripes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 151.

Distribution: Tanzania. caligata caligata

(SELYS)

1853 Libellago (Libellago) caligata SELYS, Synopsis des Calopterygines: 57. Type female: Riksmus. Naturhist., Stockholm; type-locality: Natal. 1854 Libellago (Libellago) caligata - SELYS & HAGEN, Monogr. Cal.: 231 (redescription) (South Africa, and Abyssinia). 1890 Libellago caligata - KIRBY, Syn. Cat. Neur.-Odon., London: 112. 1891 Libellago ambigua GERSTAECKER, Jb. H a m b . wiss. Aust., 9: 7. Type female: lost; type-locality: Zanzibar. 1898 Libellago hartmanni FÖRSTER, Ent. Nachr., 24: 166. Type male: Mus. Naturk., Berlin; typelocality: Transvaal. 1906 Libellago hartmanni - FÖRSTER, Wiesbaden Jb. Ver. Natk., 59: 331 [proposed synonymy with caligata

caligata

(SELYS, 1853)].

1921 Libellago caligata - Ris, Annls S. Afr. Mus., 18: 262 (Pretoria, Zululand, Transvaal, Natal). 1949 Platycypha caligata - FRASER, Bull. Inst. R. Sei. nat. Belg., 25 (6): 10 (as generotype of Platacypha FRASER, 1949). 1958 Platycypha caligata - PINHEY, Stuttg. Beitr. Naturk., 1958 (10): 2 (Ethiopian Region). 1 9 5 8 Libellago sulphuripes M A R T I N , ined (Transvaal) - loc. cit. PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, 2 2 B : 1 0 5 [as syn. with caligata caligata (SELYS, 1 8 5 3 ) ] . 1959 Platycypha caligata caligata - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 27. 1984 Platycypha caligata f. lacus PINHEY, 1982 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 19.

Distribution: South Africa and Ethiopia. caligata angolense

LONGFIELD

1959 Platycypha caligata angolense LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 27. Type male: not given [perhaps in British Museum (Nat. Hist.), London]; type-locality: Angola. 1962 Platycypha caligata angolense - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 152.

Distribution: Angola. fitzsimonsi fitzsimonsi

(PINHEY)

1950 Chlorocypha fitzsimonsi PINHEY, Ann. Transvaal. Mus., 21: 270. Type male: Transvaal M u seum; type-locality: Natal. 1958 Chlorocypha fitzsimonsi - PINHEY, Occ. Pap. nat. Mus. S. Rhodesia, 22B: 105 (Rhodesia). 1962 Platycypha fitzsimonsi fitzsimonsi- PINHEY, Publ. cult. Diam. Angola, 59:152 (Natal, S. Rhodesia).

Distribution: South Africa.

65 fitzsimonsi inyangae

PINHEY

1958 Platycypha fitzsimonsi inyangae PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, 22B: 106. Type male: British Museum (Nat. Hist.), London; type-locality: Rhodesia. 1962 Platycypha fitzsimonsi inyangae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 159 (form or subsp.?).

Distribution: Rhodesia. lacustris lacustris (FÖRSTER) 1914 Libellago caligata lacustris FÖRSTER, Arch. Naturg., 80 (a): 61. Type male: Förster's Collection or lost; type-locality: Uganda. 1940 Chlorocypha armageddoni FRASER, Annls ent. soc. Am., 33: 551. Type male: British Museum (Nat. Hist.), London; type-locality: East Africa. 1950 Platycypha lacustris - LAIDLAW, Trans. R. ent. Soc. London, 101: 270. 1958 Platycypha lacustris - PINHEY, Occ. Pap. nat. Mus. S. Rhodesia, 22B: 105. 1958 Chlorocypha armageddoni - PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, 22B: 105 [proposed synonymy with lacustris lacustris (FÖRSTER, 1914)]. 1962 Platycypha lacustris lacustris - PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, (B) 3: 904.

Distribution: Uganda, West Kenya. lacustris chingolae PINHEY 1962 Platycypha lacustris chingolae PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, (B) 3: 904. Type male: unknown; type-locality: North Rhodesia.

Distribution: North Rhodesia. rufitibia rufitibia (PINHEY) 1961 Chlorocypha (Africocypha) rufitibia PINHEY, Entomologist's month. Mag., 96: 263. Type male: National Museum; Bulawayo; type-locality: South Africa. 1984 Platycypha rufitibia rufitibia - DA VIES & TOBIN, The Dragonlies of the World, Utrecht, 1:19.

Distribution: South Africa. rufitibia lucalaensis PINHEY 1984 Platycypha rufitibia lucalaensis PINHEY, 1967 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 19.

Distribution: Luanda. pinheyi FRASER 1950 Platycypha pinheyi FRASER, Bull. Inst. R. Sei. nat. Belg., 26: 6. Type male: British Museum (Nat. Hist.), London; type-locality: Tanganyika. 1962 Platycypha pinheyi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 152.

Distribution: Tanzania. picta (PINHEY) comb. nov. 1962 Chlorocypha picta PINHEY, J. ent. Soc. S. Afr., 25: 30. Type male: Pinhey's Collection; typelocality: Congo.

Distribution: Congo Republic. Genus Rhinocypha RAMBUR 1842 Rhinocypha RAMBUR, Hist. nat. Ins. Neuropt.: 232. Type species·.Rhinocypha tincta RAMBUR, 1842 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 114.). - 1853 Libellago

66 (Rhinocypha)

- SELYS, Synopsis des Calopterygines: 59. - 1854 Libellago (Rhinocypha)

SELYS & HÄGEN, M e m . S o c . r. Sei. Liege, 9: 198 ( r e v i s i o n ) . - 1890 Rhinocypha

-

- KIRBY, S y n .

Cat. Neur.-Odon, London: 114). - 1904 Rhinocypha - WILLIAMSON, Proc. U.S. natn. Mus., 28: 173 (revision). - 1905 Rhinocypha - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 803 (key to species). - 1917 Rhinocypha - LAIDLAW, Ree. Indian Mus., 13: 33 (key to species groups). - 1934 Rhinocypha - SCHMIDT, Arch. Hydrobiol. Suppl.: 13: 317 (key to species). 1939 Rhinocypha - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 252 (key to Philippian species). - 1950 Rhinocypha - LAIDLAW, Trans. R. ent. Soc. London, 101: 270. - 1954 Rhinocypha LIEFTINCK, Treubia, 22: Suppl.: 7. - 1954 Heliocypha - LIEFTINCK, Treubia, 22 Suppl.: 7 (incorrect synonymy with Rhinocypha RAMBUR, 1842). - 1954 Aristocypha - LIEFTINCK, Treubia, 22 Suppl.: 7 (incorrect synonymy with Rhinocypha RAMBUR, 1842). a l b i s t i g m a SELYS 1873 Libellago (Rhinocypha) albistigma SELYS (nec MATSUMURA, 1931), Synopsis des Calopterygines, Add. 62. Type male: British Museum (Nat. Hist.), London; type-locality: Malaysia. 1950 Rhinocypha albistigma - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: Malaysia. a n i s o p t e r a SELYS 1879 Rhinocypha anisoptera SELYS, Bull. Acad. r. Belg., (2) 47: 394. Type male: Selys's Collection; type-locality: Sumatra. 1934 Rhinocypha anisoptera - LIEFTINCK, Treubia, 14 (4): 385 (Java). 1935 Rhinocypha anisoptera - LIEFTINCK, Misc. Zool. Sum., 92-93: 4. 1950 Rhinocypha anisoptera - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1954 Rhinocypha anisoptera - LIEFTINCK, Treubia, 22 Suppl.: 8. Sumatra, Java.

Distribution: Sumatra, Java, a u r o f u l g e n s LAIDLAW 1931 Rhinocypha aurofulgens LAIDLAW, J. Fed. Malay. Stat. Mus., 16: 242. Type male: Laidlaw's Collection; type-locality: Borneo. 1950 Rhinocypha aurofulgens - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1954 Rhinocypha aurofulgens - LIEFTINCK, Treubia, 22 Suppl.: 8 (Borneo).

Distribution: Borneo. aurulenta FÖRSTER 1903 Rhinocypha aurulenta FÖRSTER, Annls hist.-nat. Mus. natn. hung., 1903: 547. Type male: Termeszett. Muz., Budapest; type-locality: Buru Island. 1950 Rhinocypha aurulenta - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: Buru. c o g n a t a KIMMINS 1936 Rhinocypha cognata KIMMINS, Ann. Mag. nat. Hist., (10) 18: 84. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1950 Rhinocypha cognata - LAIDLAW, Trans. R. ent. Soc. London, 101: 270. 1954 Rhinocypha cognata - LIEFTINCK, Treubia, 22 Suppl.: 9 (Borneo).

Distribution: Borneo. colorata HÄGEN 1869 Libellago (Rhinocypha) colorata HAGEN, in SELYS, Synopsis des Calopterygines, Add. 2: 20. Type male: Semper's Collection; type-locality: Manila.

67 1890 Rhinocypha 1939 Rhinocypha 1950 Rhinocypha

colorata - KIRBY, Syn. Cat. Neur.-Odon., London: 114. colorata - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 254 (Luzon, Cebu). colorata - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: Philippine Islands. cucullata

SELYS

1873 Libellago (Rhinocypha) cucullata SELYS, Synopsis des Calopterygines, Add. 3: 28. Type male: Selys & Hagen's Collection; type-locality: Labuan (Borneo). 1890 Rhinocypha cucullata - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1937 Rhinocypha cucullata - COWLEY, Trans. R. ent. Soc. London, 86: 11. 1954 Rhinocypha cucullata - LIEFTINCK, Treubia, 22 Suppl.: 9 (Borneo).

Distribution: Borneo. dorsosanguinea

LIEFTINCK

1961 Rhinocypha dorsosanguinea LIEFTINCK, Fieldiana (Zool.) 42: 124. Type male: unknown; typelocality: Basilian I.

Distribution: Basilian Island. drusilla

NEEDHAM

1930 Rhinocypha drusilla NEEDHAM, Zool. Sinica, (A) 11: 221. Type male: unknown, perhaps in Peking Museum; type-locality: China. 1950 Rhinocypha drusilla - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: China. frontalis

SELYS

1873 Libellago (Rhinocypha) frontalis SELYS, Synopsis des Calopterygines, Add. 3: 26. Type male: Selys's Collection; type-locality: Moluccas. 1890 Rhinocypha frontalis - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1916 Rhinocypha frontalis - Ris, Ent. Mitt., 5: 311 (Celebes). 1935 Rhinocypha frontalis - LIEFTINCK, Treubia, 15 (2): 176 (Celebes). 1950 Rhinocypha frontalis - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: Celebes. hageni

KRÜGER

1898 Rhinocypha hageni KRÜGER, Stettiner ent. Ztg., 59: 135. Type male: unknown, perhaps in Warszawa Museum; type-locality: Jolo. 1950 Rhinocypha hageni - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: Philippine Islands. heterostigma

RAMBUR

1842 Rhinocypha heterostigma RAMBUR, Hist. nat. Ins. Neuropt., Paris: 236. Type male: Serville's Collection; type-locality: Java. 1853 Libellago (Rhinocypha) heterostigma - SELYS, Synopsis des Calopterygines: 63. 1853 Libellago heterostigma - WALKER, List Neur. Ins. Β. M., 4: 646. 1854 Libellago (Rhinocypha) heterostigma - SELYS & HAGEN, Monogr. Cal.: 221 (redescription). 1890 Rhinocypha heterostigma - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1926 Rhinocypha heterostigma - FRASER, Treubia, 8 ( 3 - 4 ) : 484 (Java). 1926 Rhinocypha io FRASER, Treubia, 8 ( 3 - 4 ) : 485. Type male: not given; type-locality: Sumatra. 1934 Rhinocypha heterostigma - SCHMIDT, Arch. Hydrob. Suppl., 13: 320. 1950 Rhinocypha heterostigma heterostigma - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Rhinocypha heterostigma io - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

68 1954 Rhinocypha 1954 Rhinocypha

heterostigma - LIEFTINCK, Treubia, 22 Suppl.: 11 (Java). io - LIEFTINCK, Treubia, 22 Suppl.: 11 (proposed synonymy with

heterostigma

RAMBUR, 1 8 4 2 ) .

1984 Rhinocypha heterostigma

f. io - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:19.

Distribution: Java. humeralis

SELYS

1873 Libellago (Rhinocypha) humeralis SELYS, Synopsis des Calopterygines, Add. 3: 24. Type male: Selys's Collection; type-locality: L a b u a n (Borneo). 1873 Libellago (Rhinocypha) eximia MCLACHLAN, in SELYS, Synopsis des Calopterygines, Add. 3: 24. Type male: McLachlan's Collection; type-locality: Borneo. 1890 Rhinocypha humeralis - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1890 Rhinocypha eximia - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1950 Rhinocypha humeralis - LAIDLAW, Trans. R. ent. Soc. London, 101: 277. 1950 Rhinocypha eximia - LAIDLAW, Trans. R. ent. soc. London, 101: 277. 1954 Rhinocypha humeralis - LIEFTINCK, Treubia, 22 Suppl.: 11 (Borneo). 1954 Rhinocypha eximia - LIEFTINCK, Treubia, 22 Suppl.: 11 (proposed synonymy with humeralis SELYS, 1 8 7 3 ) .

Distribution: Borneo. ignipennis

SELYS

1879 Rhinocypha ignipennis SELYS, Bull. Acad. r. Belg., (2) 47: 389. Type male: Selys's Collection; type-locality: Khasia Hill (Assam). 1890 Rhinocypha ignipennis - KIRBY, Syn. Cat. Neur.-Odon., London: 113. 1905 Rhinocypha ignipennis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 805 (India, Burma). 1917 Rhinocypha ignipennis - LAIDLAW, Ree. Indian Mus., 13: 35 (Assam). 1950 Rhinocypha ignipennis - LAIDLAW, Trans. R. ent. Soc. London, 101: 277.

Distribution: India, Burma. katharina

NEEDHAM

1930 Rhinocypha katharina NEEDHAM, Zool. Sinica, (A) 11: 222. Type male: unknown, perhaps in Peking Museum; type-locality: China. 1984 Rhinocypha katherina (misprint) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 19.

Distribution: China, latimacula

LIEFTINCK

1974 Rhinocypha latimacula LIEFTINCK, Steenstrupia, 3 (12): 115. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Tawi-Tawi.

Distribution: Philippine Islands, liberata

LIEFTINCK

1949 Rhinocypha liberata LIEFTINCK, N o v a Guinea (N.S.), 5: 27. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Solomon Island. 1949 Rhinocypha liberata - LIEFTINCK, Treubia, 20 (2): 330 (Guadalcanal, Ugi). 1950 Rhinocypha liberata - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1957 Rhinocypha liberata - KIMMINS, Bull. Brit. Mus. (Nat. Hist.), Ent., 5: 311 (redescription).

Distribution: Solomon Islands.

69 maculata 1931

MATSUMURA

Rhinocypha maculata Japan.

M A T S U M U R A , 6 0 0 0 111.

Ins., Japan: ?. Type: unknown; type-locality:

Distribution: Japan. monochroa

SELYS

1873 Libellago (Rhinocypha) monochroa SELYS, Synopsis des Calopterygines, Add. 3: 61. Type male: British Museum (Nat. Hist.), London; type-locality: Celebes. 1890 Rhinocypha monochroa - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1950 Rhinocypha monochroa - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

Distribution: Celebes. moultoni

LAIDLAW

1915 Rhinocypha moultoni LAIDLAW, Proc. Zool. Soc. London, 1915: 35. Type male: Laidlaw's Collection; Type-locality: Borneo. 1950 Rhinocypha moultoni - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1954 Rhinocypha moultoni - LIEFTINCK, Treubia, 22: Suppl.: 12.

Distribution: Borneo. ogasawarensis

MATSUMURA & OGUMA

1913 Rhinocypha (sic) ogasawarensis M A T S U M U R A & O G U M A , Zool. Mag., Tokyo, 25: 319. Type male: Hokkaido Univ.; type-locality: Ogasawara. 1950 Rhinocypha ogasawarensis - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1952 Rhinocypha ogasawarensis - ASAHINA, Mushi, 23 (6): 48 (redescription) (Bonin Island). 1962 Rhinocypha ogasawarensis - LIEFTINCK, Insecta of Micronesia, 5 (1): 12 (endemic - Bonin Island).

Distribution: Bonin Island. pagenstecheri pagenstecheri

FÖRSTER

1897 Rhinocypha pagenstecheri FÖRSTER, Ent. Nachr., 23: 333. Type male: Mus. Naturk., Berlin; type-locality: Lombok. 1898 Libellago lombockensis MCLACHLAN, Ent. Mag., 1898: 273. Type male: British Museum (Nat. Hist.), London; type-locality: Lombok. 1916 Rhinocypha pagenstecheri - Ris, Ent. Mitt., 5: 316 (Lombok, Flores). 1936 Rhinocypha pagenstecheri pagenstecheri- LIEFTINCK, Revue suisse Zool., 43 (5): 111 (Sumba). 1950 Rhinocypha pagenstecheri - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1984 Libellago lombockensis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 20 (as syn. with pagenstecheri pagenstecheri FÖRSTER, 1897).

Distribution: Lombok, Flores, Sumba. pagenstecheri pusilla

LIEFTINCK

1953 Rhinocypha pagenstecheri pusilla LIEFTINCK, Verh. naturf. Ges. Basel, 64: 136. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Damar. 1984 Rhinocypha pagenstecheri f. pusilla - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 0 .

Distribution: Indonesia: Damar Island. pagenstecheri timorana

LIEFTINCK

1936 Rhinocypha pagenstecheri timorana LIEFTINCK, Revue suisse Zool., 43 (5). 111. Type male: unknown; type-locality: Timor.

70 1953 Rhinocypha pagenstecheri timorana - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 135 (Timor). 1984 Rhinocypha pagenstecherii. timorana- DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 0 .

Distribution: Timor, pallidifrons Ris 1927 Rhinocypha pallidifrons Ris, Zool. Meded., Leiden, 10 (1): 9. Type female: Rijksmus. Natuurl. Hist., Leiden; type-locality: Sumatra. 1950 Rhinocypha pallidifrons - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1954 Rhinocypha pallidifrons - LIEFTINCK, Treubia, 22 Suppl.: 12 (Sumatra).

Distribution: Sumatra. pelops

LAIDLAW

1936 Rhinocypha pelops LAIDLAW, J. Fed. Mai. States Mus., 13:60. Type male: Laidlaw's Collection; type-locality: Perak. 1948 Rhinocypha pelops - LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 2 (Burma). 1950 Rhinocypha pelops - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1954 Rhinocypha pelops - LIEFTINCK, Treubia, 22 Suppl.: 12 (Malaya).

Distribution: Malaysia. Phantasma

LIEFTINCK

1935 Rhinocypha phantasma LIEFTINCK, Treubia, 15 (2): 175. Type male: not given, perhaps in Buitzenzorg Museum; type-locality: Celebes. 1950 Rhinocypha phantasma - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

Distribution: Celebes. sanguinolenta

LIEFTINCK

1961 Rhinocypha sanguinolenta LIEFTINCK, Fieldiana (Zool.), 42: 125. Type male: unknown; typelocality: Minadanao.

Distribution: Philippine Islands. selysi

KRÜGER

1898 Rhinocypha selysi KRÜGER, Stettines ent. Ztg., 59: 81. Type male: Mus. Naturk., Berlin; typelocality: Sumatra. 1927 Rhinocypha selysi - Ris, Zool. Meded., Leiden, 10 (1): 6. 1934 Rhinocypha selysi - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 319. 1934 Rhinocypha selysi - LIEFTINCK, Treubia, 14 (4): 386 (Java). 1950 Rhinocypha selysi - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1954 Rhinocypha selysi - LIEFTINCK, Treubia, 22 Suppl.: 13.

Distribution: Sumatra, Java. spinifer

LAIDLAW

1931 Rhinocypha spinifer LAIDLAW, J. Fed. Mai. States Mus., 16: 241. Type male: Laidlaw's Collection; type-locality: Borneo. 1937 Rhinocypha spinifer - COWLEY, Trans. R. ent. Soc. London, 86: 8. 1954 Rhinocypha spinifer - LIEFTINCK, Treubia, 22 Suppl.: 13 (Borneo).

Distribution: Borneo.

71 stygia

FÖRSTER

1897 Rhinocypa stygia FÖRSTER, Ann. Soc. ent. Belg., 51: 210. Type male: Mus. Naturk., Berlin; type-locality: Borneo. 1950 Rhinocypha stygia - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1954 Rhinocypha stygia - LIEFTINCK, Treubia, 22 Suppl.: 14 (Borneo).

Distribution: Borneo. sumbana

FÖRSTER

1897 Rhinocypha pagenstecheri race sumbana FÖRSTER, Ent. Nachr., 23: 335. Female from Sumba. 1898 Rhinocypha braueri KRÜGER, Stettiner ent. Ztg., 1898: 133. Type male: Mus. Naturk., Berlin; type-locality: Sumba. 1936 Rhinocypha sumbana - LIEFTINCK, Revue suisse Zool., 43 (5): 114. 1 9 5 0 Rhinocypha sumbana - L A I D L A W Trans. R. ent. Soc. London, 101: 2 7 7 . 1950 Rhinocypha braueri - LAIDLAW, Trans. R. ent. Soc. London, 101: 277 (proposed synonymy with sumbana FÖRSTER, 1 8 9 7 ) .

Distribution: Sumba. terminata

SELYS

1869 Libellula (Rhinocypha) terminata SELYS, Synopsis des Calopterygines, Add. 2: 18. Type male: Selys's Collection; type-locality: Moluccas. 1878 Libellula (Rhinocypha) ustulata SELYS, Mitt. Zool. Mus., Dresden, 3: 269. Type male: Mus. Tierk., Dresden; type-locality: Amboina. 1890 Rhinocypha terminata - K I R B Y Syn. Cat. Neur.-Odon., London: 114. 1890 Rhinocypha ustulata - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1950 Rhinocypha ustulata - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. 1950 Rhinocypha terminata - LAIDLAW, Trans. R. ent. Soc. London: 101: 278 (incorrect synonymy with ustulata SELYS, 1878).

Distribution: Indonesia: Moluccas. tincta tincta

RAMBUR

1842 Rhinocypha tincta RAMBUR, Hist. nat. Ins. Neuropt., Paris: 237. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: not given („Java"). 1853 Libellago (Rhinocypha) tincta - SELYS, Synopsis des Calopterygines: 64 (Manilla, type in Mus. Naturk., Berlin). 1 8 5 4 Libellago (Rhinocypha) tincta - SELYS & H A G E N , Monogr. Cal.: 2 2 3 (redescription). 1890 Rhinocypha tincta - K I R B Y , Syn. Cat. Neur.-Odon., London: 114 (as generotype of Rhinocypha RAMBUR, 1842).

1907 1934 1938 1950

Rhinocypha Rhinocypha Rhinocypha Rhinocypha

tincta tincta tincta tincta

- TILLYARD, Proc. Linn. Soc. N.S. Wales, 32: 399 (Australia). - SCHMIDT, Arch. Hydrob., Suppl.: 13: 323. tincta - LIEFTINCK, Nova Guinea (N.S.), 2: 63. tincta - LAIDLAW, Trans. R. ent. Soc. London, 101: 278.

Distribution: Australia, Java. tincta adusta

LIEFTINCK

1949 Rhinocypha tincta adusta LIEFTINCK, Treubia, 20 (2): 332. Type male: British Museum (Nat. Hist.), London; type-locality: Solomon Islands. 1984 Rhinocypha tincta f. adusta - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 19.

Distribution: Solomon Islands.

72

tincta amanda 1938

1984

LIEFTINCK

Rhinocypha tincta amanda LIEFTINCK, Nova Guinea, (N.S.), 2 : 6 8 . Type male: Buitenzorg Museum; type-locality: New Guinea. Rhinocypha tincta f. amanda - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 1 9 .

Distribution: New Guinea. tincta dentiplaga

LIEFTINCK

1938 Rhinocypha tincta dentiplaga LIEFTINCK, Nova Guinea, (N.S.), 2: 69. Type male: Ann Arbor Museum; type-locality: New Guinea. 1950 Rhinocypha tincta dentiplaga - L A I D L A W , Trans. R . ent. Soc. London, 1 0 1 : 277. 1984 Rhinocypha tincta f. dentiplaga- DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:19.

Distribution: New Guinea. tincta retrograda

LIEFTINCK

1938 Rhinocypha tincta retrograda LIEFTINCK, Nova Guinea (N.S.), 2: 65. Type male: Rijksmus. Natuurl. His., Leiden; type-locality: New Guinea. 1 9 5 0 Rhinocypha tincta retrograda - L A I D L A W , Trans. R . ent. Soc. London, 1 0 1 : 2 7 8 . 1 9 8 4 Rhinocypha tincta f. retrograda- DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 2 0 .

Distribution: New Guinea, tincta sagitta

LIEFTINCK

1938 Rhinocypha tincta sagitta LIEFTINCK, Nova Guinea (N.S.), 2: 66. Type male: Lieftinck's Collection; type-locality: New Guinea. 1950 Rhinocypha tincta sagitta - L A I D L A W , Trans. R . ent. Soc. London, 1 0 1 : 278. 1984 Rhinocypha tincta f. sagitta - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 20.

Distribution: New Guinea. tincta semitincta

SELYS

1869 Libellago (Rhinocypha) semitincta SELYS, Synopsis des Calopterygines, Add. 1: 20. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Gilolo. 1 8 9 0 Rhinocypha semitincta - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 1 4 (Philippines, New Guinea). 1891 Rhinocypha semitincta - SELYS, Ann. Soc. Esp. Hist, nat., 2 0 : 213 (Mindanao). 1 9 3 8 Rhinocypha tincta semitincta - LIEFTINCK, Nova Guinea (N.S.), 2: 5 9 (Moluccas, Key, Bismarck Archipelago, New Guinea). 1939 Rhinocypha semitincta - NEEDHAM & G Y G E R , , Philipp. J. Sei., 70 (3): 254 (Mindanao). 1 9 4 9 Rhinocypha tincta semitincta - LIEFTINCK, Treubia, 2 0 ( 2 ) : 3 3 1 (New Britain). 1984 Rhinocypha tincta f. semitincta - DAVIES & TROBIN, The Dragonflies of the World, Utrecht, 1: 2 0 .

Distribution: from Philippine Islands to New Guinea, trimaculata

SELYS

1853 Libellago (Rhinocypha) trimaculata SELYS, Synopsis des Calopterygines: 62. Type male: Selys's Collection: type-locality: Tibet. 1853 Libellago trimaculata - WALKER, List Neur. Ins. Β. M., 4: 649. 1 8 5 4 Libellago (Rhinocypha) trimaculata - SELYS & H A G E N , Monogr. Cal.: 2 1 1 (redescription). 1 8 9 0 Rhinocypha trimaculata - KIRBY, Syn. Cat. Neur.-Odon., London: 1 1 3 . 1917 Rhinocypha trimaculata - L A I D L A W , Ree. Indian Mus., 1 3 : 3 5 (Tibet, Assam). 1950 Rhinocypha trimaculata - L A I D L A W , Trans. R . ent. Soc. Lodon, 1 0 1 : 278. Distribution: C h i n a , I n d i a .

73

turconii SELYS 1891 Rhinocypha ippines. 1939 Rhinocypha 1950 Rhinocypha 1961 Rhinocypha

turconii SELYS, Ann. Soc. Esp., 20: 215. Type male: unknown; type-locality: Philturconii - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 253 (Mindanao, Luzon). turconii - LAIDLAW, Trans. R. ent. Soc. London, 101: 278. turconii - LIEFTINCK, Fieldiana (Zool.), 42: 122 (Mindanao).

Distribution: Philippine Islands. uenoi ASAHINA 1964 Rhinocypha uenoi ASAHINA, Kontyü, 32 (2): 299. Type male: Asahina's Collection; type-locality: Iriomote.

Distribution: Iriomote Island. xanthe Ris 1927 Rhinocypha xanthe Ris, Zool. Meded., Leiden, 10 (1): 7. Type male: Rijksmus. Naturl. Hist., Leiden; type-locality: Sumatra. 1950 Rhinocypha xanthe - LAIDLAW, Trans. R. ent. Soc. London, 101: 270. 1954 Rhinocypha xanthe - LIEFTINCK, Treubia, 22 Suppl.: 14 (Sumatra).

Distribution: Sumatra. Genus Rhinoneura LAIDLAW 1915 Rhinoneura LAIDLAW, Proc. zool. Soc. London, 1915: 33. Type species:Rhinoneura villosipes L A I D L A W , 1 9 1 5 (original designation). - 1 9 5 4 Rhinoneura - LIEFTINCK, Treubia, 2 2 : Suppl.: 1 5 .

caerulea KIMMINS 1936 Rhinoneura caerulea KIMMINS, J. Fed. Mai. States Mus., 18: 79. Type male: unknown; typelocality: Borneo. 1950 Rhinoneura caerulea - LAIDLAW, Trans. R. ent. soc. London, 101: 270. 1954 Rhinoneura caerulea - LIEFTINCK, Treubia, 22 Suppl.: 15 (Borneo).

Distribution: Borneo. villosipes LAIDLAW 1915 Rhinoneura villosipes LAIDLAW, Proc. zool. Soc. London, 1915: 33. Type male: unknown; type-locality: Borneo (as generotype of Rhinoneura LAIDLAW, 1915). 1950 Rhinoneura villosipes - LAIDLAW, Trans. R. ent. Soc. London, 101: 270. 1954 Rhinoneura villosipes - LIEFTINCK, Treubia, 22 Suppl.: 15 (North Borneo).

Distribution: Borneo. Genus Sundacypha LAIDLAW 1950 Sundacypha LAIDLAW, Trans. R. ent. Soc. London, 101: 272. Type species: Libellago (Rhinocypha) petiolata SELYS, 1 8 5 9 (original designation). - 1 9 5 4 Sundacypha - LIEFTINCK, Treubia, 22 Suppl.: 14.

petiolata (SELYS) 1859 Libellago (Rhinocypha) petiolata SELYS, Synopsis des Calopterygines, Add. 1: 12. Type: sex not given: Selys's Collection; type-locality: Malacca. 1890 Rhinocypha petiolata - KIRBY, Syn. Cat. Neur.-Odon., London: 114. 1898 Rhinocypha karschi KRÜGER, Stettiner ent. Ztg., 59: 83. Type male: Mus. Naturk., Berlin; type-locality: Sumatra - Borneo. 1902 Micromerus rubropictus MARTIN, Bull. Mus. Paris, 7: 508. Type male: 508. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Borneo.

74 1934 Calocypha karschi- LAIDLAW, Stylops, 3: 99. 1950 Sundacypha petiolata - LAIDLAW, Trans. R. ent. Soc. London, 101: 272 (as generotype of Sundacypha LAIDLAW, 1950). 1950 Rhinocypha karschi - LAIDLAW, Trans. R. ent. Soc. London, 101: 272 [proposed synonymy with petiolata (SELYS, 1859)]. 1950 Micromerus rubropictus - LAIDLAW, Trans. R. ent. Soc. London, 101: 272 [proposed synonymy with petiolata (SELYS, 1859)].

Distribution: Malaysia, Sumatra, Borneo. Family Dicteriastidae Medium sized forms. Primary antenodals evident; IRiii and Riv recessed: Rii-iii strongly arched forward near origin; basal antenodals present. Legs long with no spines. 1939 Heliocharitidae TILLYARD & FRASER, Austr. Zool., 9: 207 (with genera Heliocharis, Charitopteryx, Cyanocharis). - 1957 Heliocharitidae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 74 (with genera Heliocharis Dicterias, Charitopteryx, Cyanocharis). - 1984 Dicteriastidae DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 21 (with genera Cyanocharis, Dicterias, Heliocharis).

Genus Cyanocharis

NEEDHAM

1903 Cyanocharis NEEDHAM, Proc. Soc. Washington, 16: 55. Type species: Cyanocharis valga NEEDHAM, 1903 (monotypy)

valga

NEEDHAM

1903 Cyanocharis valga NEEDHAM, Proc. Soc. Washington, 16: 56 (as generotype of NEEDHAM, 1903). Type male: unknown; type-locality: Brazil.

Cyanocharis

Distribution: Brazil. Genus Dicterias

SELYS

1853 Dicterias SELYS, Synopsis des Calopterygines: 55. Type species: Dicterias atrosanguinea Selys, 1853 (monotype). - 1854 Dicterias - SELYS & HAGEN, Monogr. Cal.: 189 (revision). - 1906 Neocharis FÖRSTER, Mannheim Jb. Ver. Natk., 71-72: 68. Type species: Neocharis cothurnata FÖRSTER, 1906 (original designation). - 1918 Dicterias - Ris. Arch. Naturg. (A) 82: 10 (revision). - 1934 Charitopteryx COWLEY, Entomologist, 67: 201 (new name for Neocharis FÖRSTER, 1906, a junior primary homonymy of Neocharis SHARP, 1877 in Coleoptera).

atrosanguinea

SELYS

1853 Dicterias atrosanguinea (DALE in MS) SELYS, Synopsis des Calopterygines: 56 (as generotype of Dicterias Selys, 1853). Type male: Dale's Collection; type-locality: Para.

Distribution: Brazil. cothurnata

(FÖRSTER)

1906 Neocharis cothurnata FÖRSTER, Mannheim Jb. Ver. Natk., 71-72: 70 (as generotype of Neocharis FÖRSTER, 1906). Type male: unknown; type-locality: Suriname. 1934 Charitopteryx cothurnata - COWLEY, Entomologist, 67: 201 (as generotype of Charitopteryx COWLEY, 1 9 3 4 ) .

1948 Dicterias cothurnata - CALVERT, Zoologica, 33: 84.

Distribution: Suriname.

75

Genus Heliocharis 1853

SELYS

Heliocharis SELYS, Synopsis des Calopterygines: 5 5 . Type species: Heliocharis amazona SELYS, 1853 (monotypy). - 1854 Heliocharis - SELYS & HAGEN, Monogr. Cal.: 187 (revision). - 1890 Heliocharis - KIRBY, Syn. Cat. Neur.-Odon., London: 110.

amazona amazona

SELYS

1853 Heliocharis amazona SELYS, Synopsis des Calopterygines: 55 (as generotype of Heliocharis SELYS, 1853). Type male: Bates's Collection; type-locality: Amazonas. 1853 Dicterias amazona - WALKER, List Neur. Ins. B.M., 4: 642. 1854 Heliocharis amazona - SELYS & H A G E N , Monogr. Cal.: 188 (redescription). 1916 Dicterias umbra Ris, Arch,. Naturg., (A) 82:12. Type male: unknown; type-locality: Colombia. 1920 Dicterias peruviana NAVÄS, An. Soc. Cient. Argentina, 90: 34. Type male: unknown; typelocality: Peru. 1943 Heliocharis amazona amazona - SCHMIDT, Beitr. Fauna Perus, 2: 240. 1943 Dicterias umbra - SCHMIDT, Beitr. Fauna Perus, 2: 240 (proposed synonymy with amazona amazona Selys, 1853). 1984 Dicterias peruviana - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 21 (as syn. with amazona amazona SELYS, 1 8 5 3 ) .

Distribution: South America. amazona libera

SELYS

1869 Heliocharis libera SELYS, Synopsis des Calopterygines, Add. 2: 17. Type male: Bates's Collection; type-locality: Santarem. 1943 Heliocharis amazona libera - SCHMIDT, Beitr. Fauna Perus, 2: 240. 1948 Heliocharis libera - FRÄSER, Acta Zool. lilloana, 5: 63 (Argentina).

Distribution: Brazil, Argentina. brasiliensis

HAGEN

1859 Heliocharis brasiliensis HAGEN, in SELYS, Synopsis des Calopterygines, Add. 1: 9. Type male: British Museum (Nat. Hist.), London; type-locality: Bahia.

Distribution: Brazil. procera

HAGEN

1859 Dicterias procera HAGEN, in SELYS, Synopsis des Calopterygines, Add. 1: 10. Type sex not given: British Museum (Nat. Hist.), London; type-locality: Santarem. 1 9 8 4 Heliocharis procera - DAVIES & TOBIN, the Dragonflies of the World, Utrecht, 1: 2 1 (as syn. ? with amazona libera SELYS, 1 8 6 9 ) .

Distribution: Santarem.

Family Diphlebiidae Rather large sturdy Zygoptera. Males nonmetallic blue with black markings. First thoracic interpleural suture incomplete. Wings narrow, petiolated to halfway between primary antenodals. Primaries evident, one above and one proximal to arc. Secondary antenodals (3-6) all distal to primaries and in costal space only; postnodals not coinciding with cross veins beneath. Median space uncrossed; space below R1 free from arc to nodus; proximal ends of Riii and

76

close and near to arc; Riii arising at nodus; Dc long and narrow, uncrossed; many intercalaries. Superior anal appendages forcipate. Riv

1984 Diphlebiidae DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 21. Type-genus: Diphlebia

SELYS, 1 8 6 9 .

Genus Diphlebia

SELYS

1859 Dineura SELYS, Synopsis des Calopterygines, Add. 1: 16. Type species: Amphipteryx lestoides SELYS, 1853 (original designation). - 1869 Diphlebia SELYS, Bull. Acad. r. Belg., (2) 27: 662 (new name for Dineura SELYS, 1859, a junior primary homonymy of Dineura DAHLBOM, 1835 in Hymenoptera). Type species: Amphipteryx lestoides SELYS, 1853 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 111). - 1890 Dineura - KIRBY, Syn. Cat. Neur.-Odon., Lond o n : 111 ( p r o p o s e d s y n o n y m y w i t h Diphlebia

SELYS, 1 8 6 9 ) . - 1 9 1 1 Diphlebia

-

TILLYARD,

Proc. Linn Soc. N.S. Wales, 36: 584 (revision).

coerulescens

TILLYARD

1914 Diphlebia euphoeoides coerulesces TILLYARD, Proc. Linn. Soc. N.S. Wales, 38: 235. Type male: Tillyard's Collection; type-locality: Queensland. 1984 Diphlebia coerulescens - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22.

Distribution: Australia. euphoeoides

TILLYARD

1907 Diphlebia euphoeoides TILLYARD, Proc. Linn. Soc. N.S. Wales, 32: 394. Type male: not given; type-locality: Queensland. 1910 Diphlebia reinholdi FÖRSTER, Wien. ent. Ztg, 29: 55. Type male: Naturhist. Mus., Wien; typelocality: New Guinea. 1917 Diphlebia euphoeoides - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 23.

Distribution: Australia, New Guinea. hybridoides

TILLYARD

1911 Diphlebia hybridoides TILLYARD, Proc. Linn. Soc. N.S. Wales, 36: 584. Type male: Tillyard's Collection; type-locality: Queensland. 1914 Diphlebia hybridoides - TILLYARD, Proc. Linn. Soc. N.S. Wales, 38: 236. 1917 Diphlebia hybridoides - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 24.

Distribution: Australia. lestoides lestoides

(SELYS)

1853 Amphipteryx lestoides SELYS, Synopsis des Calopterygines: 67. Type male: British Museum (Nat. Hist.) London; type-locality: Australasia. 1854 Amphiteryx lestoides - SELYS & HAGEN, Monogr. Cal.: 245 (redescription). 1859 Dineura lestoides - SELYS, Synopsis des Calopterygines, Add. 1: 16 (as generotype of Dineura SELYS, 1 8 5 9 ) .

1869 Diphlebia lestoides-

SELYS, Bull. Acad. r. Belg., (2) 27: 662 (as generotype of Diphlebia SELYS,

1869).

1890 1901 1917 1956

Diphlebia Diphlebia Diphlebia Diphlebia

lestoides lestoides lestoides lestoides

- KIRBY, Syn. Cat. Neur.-Odon., London: 111 (tropical America ?). - Martin, Mem. Soc. zool. France, 14: 243 (Victoria, New South Wales). - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 22 (Queensland). lestoides - FRASER, Entomologist's month. Mag., 92: 305.

Distribution: Australia.

77 lestoides tillyardi FRÄSER 1956 Diphlebia lestoides tillyardi FRASER, Entomologist's month. Mag., 92: 305. Type male: British Museum (Nat. Hist.), London; type-locality: New South Wales.

Distribution: Australia. nexans CALVERT 1903 Diphlebia nexans CALVERT, Biol. Centr.-Amer. Neur.: 84. Type: unknown; type-locality: Brazil.

Distribution: Brazil. nymphoides TILLYARD 1911 Diphlebia nymphoides TILLYARD, Proc. Linn. Soc. N.S. Wales, 36: 588. Type male: Tillyard's Collection; type-locality: New South Wales. 1917 Diphlebia nymphoides - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 24 (Queensland).

Distribution: Australia. Family Euphaeidae Medium sized forms, wings not petiolate; antenodals numerous (12-20); Rii and Riii not fused with Ri near origin. Dc short, rarely transversed, base of cell connected to radius by arc. First thoracic suture incomplete. Stigma well developed. Larva with lateral abdominal gills, in addition to caudal sacs. 1905

Euphaeinae JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 3 (as subfamily). - 1 9 3 9 Epallagidae T I L L Y A R D & FRASER, Austr. Zool., 9: 2 0 9 (with genera Epallage, Dysphaea, Euphaea, Allophaea, Indophaea, Anisophaea, Heterophaea, Anisolpleura, Bayadera, Cyclophaea).- 1957 Epallagidne - FRASER, R. Zool. Soc. N. S. Wales, 1 9 5 7 : 7 8 (with subfamily Epallaginae FRASER, 1928. - 1984 Epallaginidae - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22.

Genus Allophaea FRASER 1929 Allophaea FRASER, J. Bombay nat. Hist. Soc., 33:288 (for Euphaea pars). Type species: Euphaea (Euphaea) ochracea SELYS, 1859 (original designation). - 1 9 5 4 Allophaea - LIEFTINCK, Treubia, 22 Suppl.: 15 (incorrect synonymy with Euphaea SELYS, 1840).

lara lara KRÜGER) 1898 Euphaea lara KRÜGER, Stettiner ent. Ztg, 59: 131. Type male: Mus. Naturk., Berlin; typelocality: Soemba. 1936 Euphaea lara lara - LIEFTINCK, Revue suisse Zool., 43 (5): 106 (Sunda). 1953 Euphaea lara lara - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 139 (Sumba). 1 9 8 4 Allophaea lara lara - DAVIS & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 2 .

Distribution: Sunda. lara balica (MCLACHLAN) 1898 Euphaea lara var. balica MCLACHLAN, Entomologist's month. Mag., (2) 9:272. Male from Bali. 1936 Euphaea lara balica - LIEFTINCK, Revue suisse Zool., 43 (5): 108 (Sunda). 1984 Allophaea lara balica - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22.

Distribution: Sunda, Bali.

78

lara lombokensis

(MCLACHLAN)

1898 Euphaea lara var. lombokensis MCLACHLAN, Entomologist's month. Mag., (2) 9: 272. Male from Lombok. 1936 Euphaea lara lombokensis - LIEFTINCK, Revue suisse Zool., 43 (5): 108 (Flores, Lombok). 1953 Euphaea lara lombokensis - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 139 (Lombok, Flores, Sumbawa). 1984 Allophaea lara lombokensis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22.

Distribution: Lombok, Bali, Flores. ochracea ochracea

(SELYS)

1959 Euphaea (Euphaea) ochracea SELYS, Synopsis des Calopterygines, Add. 1: 8. Type male: Selys's Collection; type-locality: Malacca. 1890 Pseudophaea ochracea - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1929 Allophaea ochracea - FRÄSER, J. Bombay nat. Hist. Soc., 33: 288 (as generotype of Allophaea FRASER, 1929).

1935 Euphaea ochracea ochracea - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 4 (Sumatra). 1942 Allophaea ochracea - FRÄSER, Proc. R. ent. Soc. London, (B) 11: 98. 1954 Euphaea ochracea ochracea - LIEFTINCK, Treubia, 22 Suppl.: 17 (Siam, Malaya).

Distribution: Malaysia, Thailand, Sumatra. ochracea brunnea

(SELYS)

1984 Allophaea ochracea brunnea World, Utrecht, 1: 22.

SELYS,

1897 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: India. Genus Anisophaea

FRASER

1928 Mesophaea FRASER, J. Bombay nat. Hist. Soc., 33: 49. No type-species. 1934 Anisophaea FRASER, Fauna brit. India, Odon., 2: 75 (new name for Mesophaea FRASER, 1928, a junior primary homonymy of Mesophaea PASCOE, 1869 in Coleoptera). -1939 Anisophaea - T I L L YARD & FRASER, Austr. Zool., 9: 210 (as n. nom. for Mesophaea FRASER, 1928). - 1984 Anisophaea - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22 (type species: Euphaea (Euphaea) decorata HÄGEN, 1853).

decorata

(HÄGEN)

1853 Euphaea (Euphaea) decorata HAGEN, in SELYS, Synopsis des Calopterygines: 51. Type male: Univ. zool. Mus., Kabenhavn; type-locality: China. 1 8 5 4 Euphaea (Euphaea) decorata - SELYS & H A G E N , Monogr. Cal.: 172 (redescription) (HongKong). 1890 Pseudophaea decorata - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1935 Euphaea decorata - MAY, Senckenbergiana, 17: 98 (description of female). 1965 Euphaea decorata - ASAHINA, Kontyü, 33 (4): 498 (Hongkong). 1984 Anisophaea decorata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22.

Distribution: Hong Kong. ornata

(CAMPION)

1924 Pseudophaea ornata CAMPION, Ann. Mag. nat. Hist., (9) 13: 174. Type male: British Museum (Nat. Hist.), London; type-locality: Hainan. 1 9 8 4 Anisophaea ornata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 2 .

Distribution: China.

79 Genus Anisopleura

SELYS

1853 Euphaea (Anisopleura) SELYS, Synopsis des Calopterygines: 48. Type species: Anisopleura lestoides SELYS, 1 8 5 3 (monotypy). - 1 8 5 4 Euphaea (Anisopleura) - SELYS & H A G E N , Monogr. Cal.: 158. - 1 8 9 0 Anisopleura-KIRBY, Syn. Cat. Neur.-Odon., London: 108. - 1 9 0 5 Anisopleura - JACOBSON & B I A N K I , Pryam lozhnos. Ross, imp.: 7 9 3 .

comes

HAGEN

1880 Anisopleura comes HAGEN, C . R . Soc. Ent. Belg., 23: 113. Type male: R. Mus., Brussels; typelocality: Himalaya. 1 9 0 5 Anisopleura comes „ H A G E N , 1 8 7 3 " - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 7 9 4 . 1928 Anisopleura comes - MORTON, Trans, ent. Soc. London, 76: i l l (Yunnan). 1984 Anisopleura comes - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 22 (India).

Distribution: India, China. furcata

SELYS

1892 Anisopleura furcata SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 488. Type male: Mus. civ. Stor. nat. Genova; type-locality: Burma.

Distribution: Burma. kusumi

SAHNI

1965 Anisopleura kusumi SAHNI, Indian J. Ent., 27: 212. Type male: Sahni's Collection; type-locality: Nepal.

Distribution: Nepal. lestoides

SELYS

1853 Euphaea (Anisopleura) sopleura SELYS, 1853). 1854 Euphaea (Anisopleura) 1890 Anisopleura lestoides 1 9 0 5 Anisopleura lestoides -

lestoides SELYS, Synopsis des Calopterygines: 48 (as generotype of AniType male: Selys's Collection; type-locality: India. lestoides - SELYS & H A G E N , Monogr. Cal.: 159 (redescription). KIRBY, Syn. Cat. Neur.-Odon., London: 108. JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 7 9 4 .

Distribution: India.

quigyuanensis 1984

ZHON

Anisopleura quigyuanensis World, Utrecht, 1: 22.

ZHON, 1 9 8 2

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: China. subplatystyla

FRÄSER

1927 Anisopleura subplatystyla FRÄSER, Ree. Indian Mus., 29: 81. Type male: British Museum (Nat. Hist.), London; type-locality: Khasia Hills.

Distribution: India, vallei

ST. QUENTIN

1937 Anisopleura vallei ST. QUENTIN, Konowia, 16 (1): 86. Type male: Naturhist. Mus., Wien; typelocality: Assam.

Distribution: Assam.

80 Genus Bayadera 1853

SELYS

Bayadera SELYS, Synopsis des Calopterygines: 4 9 . Type species: Epallage indica SELYS, 1 8 5 3 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 0 8 ) . - 1 8 9 0 Bayadera - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 0 8 . - 1 9 0 5 Bayadera - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 794 (key to species). - 1912 Bayadera - Ris, Suppl. Ent. Berlin, 1: 48 (revision, key to speces).

bidentata

NEEDHAM

1930 Bayadera bidentata NEEDHAM, Zool. Sinica, (A) 11: 218. Type male: Cornell Univ.; typelocality: China. 1955 Bayadera bidentata - ASAHINA, Ent. Medd., 27 ( 4 - 5 ) : 207 (China).

Distribution: China. brevicauda brevicauda

FRASER

1934 Bayadera brevicauda FRASER, F a u n a brit. India, Odon. 2: 83. Type male: British Museum; type-locality: Formosa.

Distribution: Taiwan. brevicauda ishigakiana

ASAHINA

1964 Bayadera brevicauda ishigakiana ASAHINA, Kontyu, 32 (1): 1. Type male: Asahina's Collection; type-locality: Ishigaki Island.

Distribution: Ishigaki Island. continentalis 1984

ASAHINA

Bayadera continentalis World, Utrecht, 1: 22.

ASAHINA, 1 9 7 3

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: China. fasciata

SJÖSTEDT

1933 Bayadera fasciata SJÖSTEDT, Ark. Zool., Uppsala, 25Α (5): 14. Type female: Riksmus. N a turhist., Stockholm; type-locality: Szechwan.

Distribution: China. forcipata

NEEDHAM

1930 Bayadera forcipata NEEDHAM, Zool. Sinica, (A) 11: 217. Type male: Needham's Collection; type-locality: China.

Distribution: China. hyalina

SELYS

1879 Bayadera hyalina SELYS, Bull. Acad. r. Belg., (2) 47: 373. Type male: Selys's Collection; typelocality: Khasia Hills. 1912 Bayadera hyalina - Ris, Suppl. ent. Berlin, 1: 50 (redescription) (Formosa).

Distribution: India, Taiwan. indica

(SELYS)

1853 Euphaea (Epallage) indica SELYS, Synopsis des Calopterygines: 49. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1854 Euphaea (Epallage) indica - SELYS & H A G E N , Monogr. Cal.: 153 (redescription).

81 1890 Bayadera indica - KIRBY, Syn. Cat. Neur.-Odon., London: 108 (as generotype of Bayadera SELYS, 1 8 5 3 ) .

Distribution: India. kali

COWLEY

1936 Bayadera kali COWLEY, Ann. Mag. nat. Hist., (10) 18: 477. Type male: British Museum (Nat. Hist.), London; type-locality: Assam.

Distribution: Assam. longicauda

FRASER

1928 Bayadera longicauda FRÄSER, J. Bombay nat. Hist. Soc., 33: 53. Type male: originally in Darjeeling Museum, perhaps destroyed; type-locality: Sikkim.

Distribution: Sikkim. melania

NAVAS

1934 Bayadera melania NAVÄS, M U S . Heude Notes ent. chinoise, perhaps in Mus. Heude; type-locality: China.

2

(1): 3. Type male: not given,

Distribution: China. melanopteryx Ris 1912 Bayadera melanopteryx Ris, Suppl. ent. Berlin, 1: 49. Type male: Königsberg Museum; typelocality: China. 1931 Bayadera melanopteryx - SCHMIDT, Konowia, 10: 177. 1934 Caliphaea nitens NAVÄS, Mus. Heude Notes ent. chinoise, 2 (1): 2. Type male: not given, perhaps in Mus. Heude; type-locality: China. 1956 Bayadera melanopteryx - ASAHINA, Ent. Medd., 27 (4-5): 206. 1962 Bayadera melanopteryx - CHAO, Acta ent. sinica, Suppl., 11: 27. 1962 Caliphaea nitens - CHAO, Acta ent. sinica, Suppl., 11: 27 (proposed synonymy with melanopteryx

RIS, 1912).

Distribution: China. Genus Cyclophaea Ris 1930 Cyclophaea Ris, Mitt, münch. ent. Ges., 20: 80. Type species: Cyclophaea cyanifrons Ris, 1930 (monotyp).

cyanifrons Ris 1930 Cyclophaea cyanifrons Ris, Mitt, münch. ent. Ges., 20: 81 (as generotype of Cyclophaea Ris, 1 9 3 0 ) . Type male: unknown; type-locality: Philippines. 1939 Cyclophaea cyanifrons - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 248.

Distribution: Philippine Islands. Genus Dysphaea

SELYS

1853 Euphaea (Dysphaea) SELYS, Synopsis des Calopterygines: 53. Type species: Dysphaea dimidiata SELYS, 1 8 5 3 (monotypy). - 1 8 5 4 Euphaea (Dysphaea) - SELYS & H A G E N . Monogr. Cal.: 1 8 5 . - 1 8 9 0 Dysphaea- KIRBY, Syn. Cat. Neur.-Odon., London: 110. - 1954 Dysphaea - LIEFTINCK, Treubia, 22: Suppl.: 19.

82

basitincta

MARTIN

1904 Dysphaea basitincta MARTIN, Mission Pavie, 3: 218. Type: unknown; type-locality: Tonkin.

Distribution: China, Vietnam. dimidiata

SELYS

1853 Euphaea (Dysphaea) dimidiata SELYS, Synopsis des Calopterygines: 54 (as generotype of Dysphaea SELYS, 1853). Type male: Selys's Collection; type-locality: Java. 1854 Euphaea (Dysphaea) dimidiata - SELYS & HAGEN, Monogr. Cal.: 185 (redescription). 1854 Euphaea (Dysphaea) limbata SELYS, Synopsis des Calopterygines, Add. 1: 9. Type male: Selys's Collection; type-locality: Malacca. 1873 Euphaea (Dysphaea) semilimbata SELYS, Synopsis des Calopterygines, Add. 3: 22. Type male: Selys's Collection; type-locality: Borneo. 1890 Dysphaea dimidiata - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1890 Dysphaea limbata - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1890 Dysphaea semilimbata - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1934 Dysphaea dimidiata - SCHMIDT, Arch. Hydrobiol. Suppl.: 13: 330 (Sumatra). 1935 Dysphaea dimidiata - LIEFTINCK, Misc. Zool. Sum., 92-93: 4 (Sumatra). 1935 Euphaea limbata - LIEFTINCK, Misc. Zool. Sum., 92-93: 4 (proposed synonymy with dimidiata SELYS, 1853).

1954 Dysphaea dimidiata - LIEFTINCK, Treubia, 22 Suppl.: 19 (Siam, Borneo, Malaya, Sumatra). 1954 Euphaea semilimbata - LIEFTINCK, Treubia, 22 Suppl.: 19 (proposed synonymy with dimidiata SELYS, 1853).

Distribution: from Malaysia to Sumatra. ethela

FRASER

1924 Dysphaea ethela FRÄSER, Ree. Indian Mus., 26: 480. Type male: British Museum (Nat. Hist.), London; type-locality: South India.

Distribution: South India. gloriosa

FRASER

1938 Dysphaea gloriosa FRASER, Proc. R. ent. Soc. London, (B) 7: 197. Type male: British Museum (Nat. Hist.), London; type-locality: Siam.

Distribution: Thailand. lugens

SELYS

1873 Euphaea (Dysphaea) lugens) SELYS, Synopsis des Calopterygines, Add. 3:21. Type male: Selys's Collection; type-locality: Borneo. 1890 Dysphaea lugens - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1911 Dysphaea lugens - Ris, Ann. Soc. ent. Belg., 55: 232. 1954 Dysphaea lugens - LIEFTINCK, Treubia, 22 Suppl.: 20 (Borneo).

Distribution: Borneo. walli

FRASER

1927 Dysphaea walli FRASER, Ree. Indian Mus., 29: 82. Type male: British Museum (Nat. Hist.), London; type-locality: Burma.

Distribution: Burma.

83 Genus Epallage 1840

CHARPENTIER

Agrion (Epallage) CHARPENTIER, Libell. europ. Lipsiae: 1 6 . Type species: Agrion fatime C H A R PENTIER, 1 8 4 0 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 0 8 ) . - 1 8 5 0 Euphaea (Epallage) - SELYS & H A G E N , Revue des Odonates, Paris: 1 4 3 . - 1 8 5 3 Euphaea (Epallage) - SELYS, Synopsis des Calopterygines: 4 9 . - 1 8 5 4 Euphaea (Epallage) - SELYS & HAGEN, Monogr. Cal.: 162 (revision). - 1890 Epallage - KIRBY, Syn. Cat. Neur. Odon., London: 1 0 8 . - 1 9 0 5 Epallage - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 4 (key to species).

alma

SELYS

1879 Euphaea (Epallage) alma SELYS, Bull. Acad. r. Belg., (2) 47: 372. Type male: Morton's Collection; type-locality: Astrabad. 1887 Epallage alma - SELYS, Ann. Soc. ent. Belg., 31: 42 (Astrabad). 1890 Epallage alma - KIRBY, Syn. Cat. Neur.-Odon., London: 108. 1 9 0 5 Epallage alma - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 7 9 3 (Persia). 1924 Epallage alma - MORTON, Trans, ent. Soc. London, 1924: 29 (redescription) (Caspian region).

Distribution: Caspian region, Turkmenia, Iran, Afghanistan. fatime fatime

(CHARPENTIER)

1840 Agrion (Epallage) fatime CHARPENTIER, Libell. europ. Lipsae: 132 (as generotype of Epallage CHARPENTIER, 1 8 4 0 ) . Type male: Mus. Hist, nat., Geneve; type-locality: Turkish Europe. 1 8 5 0 Euphaea fatime - SELYS ( & H Ä G E N ) , Revue des Odonates, Paris: 1 4 3 (redescription). 1853 Euphaea (Epallage) fatime - SELYS, Synopsis des Calopterygines: 50 (Asia Minor, Greece). 1 8 5 4 Euphaea (Epallage) fatime - SELYS & H Ä G E N , Monogr. Cal: 1 6 5 (redescription). 1887 Epallage fatime - SELYS, Ann. Soc. ent. Belg., 31: 41 (Asia Minor, Greece, Turkey). 1890 Epallage fatime - KIRBY, Syn. Cat. Neur.-Odon., London: 108. 1 9 0 5 Epallage fatime - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 4 (Turkey, Greece, Cyprus, Asia Minor, Syria). 1917 Epallage fatima (misprint) - LAIDLAW, Ree. Indian Mus., 13: 40 (Kashmir). 1938 Epallage fatime - SCHMIDT, Sitz.Ber. Akad. Wiss., Wien, (I) 147: 136 (Lebanon). 1948 Epallage fatime - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 136 (Armenia). 1984 Epallage fatime - STEINMANN, Fauna hung., 160: 51 (Hungary).

Distribution: from Kashmir to Southcentral Europe. fatime amasina

SELYS

1887 Epallage fatime amasina SELYS, Ann. Soc. ent. Belg., 31: 42 (doubtful date). Type male: unknown; type-locality: Asia Minor. 1 9 0 5 Epallage fatime amasina - JACOBSON & BIANKI, Pryam, lozhnos. Ross, imp.: 7 9 5 . 1912 Epallage fatime amasina - BARTENEF, Mitt. Kaukas. Mus., 7 (1): 113.

Distribution: Asia Minor. fatime anatolica

SELYS

1887 Epallage fatime race anatolica SELYS, Ann. Soc. ent. Belg., 31: 42 (doubtful date). Male from Asia Minor. 1 9 0 5 Epallage fatime anatolica - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 7 9 5 . 1984 Epallage fatime f. anatolica - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 23.

Distribution: Anatolia and Azerbaijan. Genus Euphaea 1840

Euphaea SELYS (nec lopteryx holosericea

Monogr. Libell. europ., Paris: 2 0 0 . Type species: Ca(original designation). - 1 8 5 0 Euphaea - SELYS & H Ä G E N ,

RAMBUR, 1 8 4 2 ) , SELYS, 1 8 4 0

SELYS

84 Revue des Odonates, Paris: 143 (with syn. Epallage CHARPENTIER, 1840). - 1853 Euphaea SELYS, Synopsis des Calopterygines: 48 (with subgenera Anisopleura, Epallage, Euphaea s. Str., Dysphaea). - 1854 Euphaea - SELYS & HAGEN, Monogr. Cal.: 152 (revision, key to subgenera). - 1890 Euphaea - KIRBY, Syn. Cat. Neur.-Odon., London: 96 (incorrect synonymy with Agrion FABRICIUS, 1775). - 1890 Pseudophaea KIRBY, Syn. Cat. Neur.-Odon., London: 1 0 9 ( r e p l a c e d n a m e t o Euphaea

RAMBUR, 1 8 4 2 - n e c SELYS, 1 8 4 0 ) . -

1 9 0 5 Pseudophaea

-

JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 795 (partim) (key to species). - 1934 Euphaea - SCHMIDT, Arch. Hydrobiol., Suppl., 13: 326 (key to species). - 1934 Pseudophaea - COWLEY, Entomologist, 67: 242 (proposed synonymy with Euphaea SELYS, 1840). - 1954 Euphaea LIEFTINCK, Treubia, 22 Suppl.: 15. - 1954 Allophaea - LIEFTINCK, Treubia, 22 Suppl.: 15 (incorrect synonymy with Euphaea SELYS, 1840). - 1954 Indophaea - LIEFTINCK, Treubia, 22 Suppl.: 15 (incorrect synonymy with Euphaea SELYS, 1840).

amphicyana Ris 1930 Euphaea amphicyana Ris, Mitt, münch. ent. Ges., 20: 89. Type male: unknown; type-locality: Mindanao. 1939 Euphaea amphicyana - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 247.

Distribution: Philippine Islands. aspasia

SELYS

1853 Euphaea (Euphaea) aspasia SELYS, Synopsis des Calopterygines: 52. Type male: Halle Museum; type-locality: Sumatra. 1854 Euphaea (Euphaea) aspasia - SELYS & HÄGEN, Monogr. Cal.: 173 (redescription) (type female: Rijksmus. Natuurl. Hist., Leiden, from Java). 1890 Pseudophaea aspasia - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1927 Euphaea aspasia - Ris, Zool. Meded., Leiden, 10 (1): 5. 1934 Euphaea aspasia - SCHMIDT, Arch. Hydrobiol., Suppl., 13: 330. 1935 Euphaea aspasia - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : (Sumatra). 1954 Euphaea aspasia - LIEFTINCK, Treubia, 22 Suppl.: 15 (Sumatra).

Distribution: Sumatra, Java. basalis

(LAIDLAW)

1915 Pseudophaea basalis LAIDLAW, Proc. Zool. Soc. London, 1915: 32. Type male: unknown; typelocality: Borneo. 1920 Pseudophaea basalis - LAIDLAW, Proc. Zool. Soc. London, 1920: 326. 1930 Euphaea basalis - Ris, Mitt, münch. ent. Ges., 20: 88. 1954 Euphaea basalis - LIEFTINCK, Treubia, 22 Suppl.: 16.

Distribution: Borneo. bocki

MCLACHLAN

1880 Euphaea bocki MCLACHLAN, Entomologist's month. Mag., 16: 204. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1890 Pseudophaea bocki - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1927 Euphaea bocki - Ris, Zool. Meded., Leiden, 10 (1): 5. 1934 Euphaea bocki - SCHMIDT, Arch. Hydrob. Suppl., 13: 330. 1935 Euphaea bocki - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 3. 1954 Euphaea bocki - LIEFTINCK, Treubia, 22 Suppl.: 16.

Distribution: Sumatra. brunnea

SELYS

1879 Euphaea (Euphaea) brunnea SELYS, Bull. Acad. r. Belg., (2) 47: 374. Type male: unknown; type-locality: Khasia Hills.

85 1890 Pseudophaea brunnea - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1 9 0 5 Pseudophaea brunnea - JACOBSON & BIANKI, Pyram. lozhnos. Ross, imp.:

795

(India, Burma).

Distribution: India, Bangladesh, Manipur, Burma. compar

MCLACHLAN

1870 Euphaea compar MCLACHLAN, Trans, ent. Soc. London, 1870:167. Type male: British Museum (Nat. Hist.), London; type-locality: Amoy. 1873 Euphaea (Euphaea) compar - SELYS, Synopsis des Calopterygines, Add. 3: 20. 1890 Pseudophaea compar - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1 9 0 5 Pseudophaea compar - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 7 9 5 (China). 1984 Euphaea formosa f. compar - DA VIES & TROBIN, The Dragonflies of the World, Utrecht, 1: 23.

Distribution: China: Guangdong, Macau. cora Ris 1930 Euphaea cora Ris, Mitt, münch. ent. Ges., 20: 86. Type male: Mus. Naturk., Berlin; typelocality: Philippines. 1939 Euphaea cora - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 245.

Distribution: Philippine Islands. formosa

HÄGEN

1869 Euphaea (Euphaea) formosa HAGEN, in SELYS, Synopsis des Calopterygines, Add. 2: 16. Type male: Hagen's Collection; type-locality: Formosa. 1890 Pseudophaea formosa - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1912 Euphaea formosa - Ris, Suppl. ent. Berlin, 1: 52 (redescription).

Distribution: Taiwan. guerini guerini

RAMBUR

1842 Euphaea (Euphaea) guerini RAMBUR, Hist. nat. Ins. Neuropt., Paris: 229. Type male: Guerin's Collection; type-locality: Conchin China. 1853 Euphaea (Euphaea) guerini - SELYS, Synopsis des Calopterygines: 53. 1 8 5 4 Euphaea (Euphaea) guerini - SELYS & H A G E N , Monogr. Cal.: 1 7 9 (redescription). 1890 Pseudophaea guerini - KIRBY, Syn. Cat. Neur.-Odon., London: 110.

Distribution: Vietnam, Laos, Thailand. guerini inonei 1984

ASAHINA

Euphaea guerini inonei World, Utrecht, 1: 24.

ASAHINA, 1 9 7 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Vietnam. impar

SELYS

1859 Euphaea (Euphaea) impar SELYS, Synopsis des Calopterygines, Add. 1: 7. Type male: Selys's Collection; type-locality: Malacca. 1859 Euphaea (Euphaea) inaequipar SELYS, Synopsis des Calopterygines, Add. 1: 7. Type male: Selys's Collection; type-locality: Borneo. 1890 Pseudophaea impar - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1890 Pseudophaea inaequipar - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1920 Pseudophaea impar inaequipar - LAIDLAW, Proc. Zool. Soc. London, 1920: 327. 1935 Euphaea impar impar - LIEFTINCK, Misc. Zool. Sum., 92-93: 3 (Sumatra). 1954 Euphaea impar - LIEFTINCK, Treubia, 22 Suppl.: 16 (Malaya, Sumatra, Borneo).

Distribution: Malaysia, Sumatra, Borneo.

86 laidlawi

KIMMINS

1936 Euphaea laidlawi KIMMINS, J. Fed. Mai. States Mus., 13: 77. Type male: Fed. Malay. States Mus.: type-locality: Malaysia. 1954 Euphaea laidlawi - LIEFTINCK, Treubia, 22 Suppl.: 17 (Borneo).

Distribution: Borneo, Malaysia. masoni

SELYS

1879 Euphaea (Euphaea) masoni SELYS, Bull. Acad. r. Belg., (2) 47: 377. Type' male: Selys's Collection; type-locality: Burma. 1890 Pseudophaea masoni - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1903 Euphaea masoni - LAIDLAW, Fase. Malayenses Zool., 1: 194. 1904 Euphaea masoni - WILLIAMSON, Proc. U.S. natn. Mus., 28: 182 (Tonkin). 1924 Pseudophaea masoni - LAIDLAW, J. Mal. Br. R. Asiatic Soc., 2: 299. 1954 Euphaea masoni - LIEFTINCK Treubia, 22 Suppl.: 17 (Siam, Assam, Burma, Indo-China).

Distribution: Vietnam, Burma, Laos, Thailand. modigliani

SELYS

1898 Euphaea modigliani SELYS, Ann. Soc. ent. Belg., 42: 336. Type male: Selys's Collection; typelocality: Mentawi Island. 1927 Euphaea modiglianii- Ris, Zool. Meded., Leiden, 10 (1): 5 (Sumatra). 1935 Euphaea modigliani - LIEFTINCK, Misc. Zool. Sum., 92-93, 4 (Sumatra). 1954 Euphaea modigliani - LIEFTINCK, Treubia, 22 Suppl.: 17 (Mentawei Is., Sumatra).

Distribution: Mentawai Island, Sumatra. opaca

SELYS

1853 Euphaea (Euphaea) opaca SELYS, Synopsis des Calopterygines: 53. Type male: Selys's Collection; type-locality: China. 1854 Euphaea (Euphaea) opaca- SELYS & HAGEN, Monogr. Cal.: 183 (redescription). 1890 Pseudophaea opaca - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1905 Pseudophaea opaca - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 795 (China). 1935 Euphaea opaca - MAY, Senckenbergiana, 17: 97.

Distribution: China. refulgens

HÄGEN

1853 Euphaea (Euphaea) refulgens HAGEN, in SELYS, Synopsis des Calopterygines: 53. Type male: Naturhist. Mus., Wien; type-locality: Manila. 1854 Euphaea (Euphaea) refulgens - SELYS & HÄGEN, Monogr. Cal.: 181 (redescription). 1890 Pseudophaea refulgens - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1939 Euphaea

refulgens

- NEEDHAM & GYGER, P h i l i p p . J . Sei., 70 (3): 246.

Distribution: Philippine Islands: Luzon, Mindoro. semperi

SELYS

1879 Euphaea (Euphaea) semperi SELYS, Bull. Acad. r. Belg., (2) 47: 376. Type male: R. Mus., Brussels; type-locality: Manila. 1890 Pseudophaea semperi - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1984 Euphaea semperi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 24 (Luzon). Distribution: P h i l i p p i n e I s l a n d s .

87 splendens

HAGEN

1853 Euphaea (Euphaea) splendens HAGEN, in SELYS, Synopsis des Calopterygines: 52. Type male: Naturhist. Mus., Wien; type-locality: Ceylon. 1 8 5 4 Euphaea (Euphaea) splendens - SELYS & H A G E N , Monogr. Cal.: 1 7 8 (redescriptiop). 1890 Pseudophaea splendens - KIRBY, Syn. Cat. Neur.-Odon., London: 110. 1894 Pseudophaea carissima KIRBY, J. Linn. Soc., London, 24: 559. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1 8 9 4 Pseudophaea viridissima K I R B Y , J . Linn. Soc. London, 24: 5 5 9 . Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1955 Euphaea splendens - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69.

Distribution: Sri Lanka. subcostalis

SELYS

1873 Euphaea (Euphaea) subcostalis SELYS, Synopsis des Calopterygines, Add. 3: 19. Type male: Selys & McLachlan's Collection; type-locality: Lebuan. 1890 Pseudophaea subcostalis - KIRBY, Syn. Cat. Neur. Odon., London: 109. 1915 Pseudophaea subcostalis - LAIDLAW, Proc. Zool. Soc. London, 1915: 32. 1930 Euphaea subcostalis - Ris, Mitt, münch. ent. Ges., 20: 88. 1934 Euphaea subcostalis - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 326. 1954 Euphaea subcostalis - LIEFTINCK, Treubia, 22 Suppl.: 18 (Malaya, Borneo).

Distribution: Malaysia, Borneo. subnodalis

(LAIDLAW)

1915 Pseudophaea subnodalis LAIDLAW, Proc. Zool. Soc. London, 1915: 31. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1930 Euphaea subnodalis - Ris, Mitt, münch. ent. Ges., 20: 88. 1954 Euphaea subnodalis - LIEFTINCK, Treubia, 22 Suppl., 18 (North Borneo).

Distribution: Borneo. superba 1984

KIMMINS

Euphaea superba Utrecht, 1: 24.

KIMMINS, 1 9 3 6

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: China. tricolor

SELYS

1859 Euphaea (Euphaea) tricolor SELYS, Synopsis des Calopterygines, Add. 1: 8. Type male: Selys's Collection; type-locality: Borneo. 1890 Pseudophaea tricolor - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1920 Pseudophaea tricolor - LAIDLAW, Proc. Zool. Soc. London, 1920: 326. 1930 Euphaea tricolor - Ris, Mitt, münch. ent. Ges., 20: 87. 1954 Euphaea tricolor - LIEFTINCK, Treubia, 22 Suppl.: 18 (Borneo).

Distribution: Borneo. variegata variegata

RAMBUR

Euphaea holosericea SELYS (nec BURMEISTER, 1 8 3 9 ) , Monogr. Libell. europ., Paris: 2 0 0 . Type: unknown; type-locality: unknown. 1842 Euphaea variegata RAMBUR, Hist. nat. Ins. Neuropt., Paris: 229. Type male: Serville's Collection; type-locality: Java. 1853 Euphaea (Euphaea) variegata - SELYS, Synopsis des Calopterygines: 52. 1 8 5 4 Euphaea (Euphaea) variegata - SELYS & H Ä G E N , Monogr. Cal.: 1 7 3 (redescription). 1840

88 1890 Pseudophaea variegata - KIRBY, Syn. Cat. Neur.-Odon., London: 109 (as generotype of Pseudophaea K I R B Y , 1 8 9 0 ) . 1927 Euphaea variegata - Ris, Zool. Meded., Leiden, 10 (1): 6 (Sumatra). 1934 Euphaea variegata variegata - SCHMIDT, Arch. Hydrob. Suppl., 13: 328 (Java). 1954 Euphaea variegata - LIEFTINCK, Treubia, 22 Suppl.: 19 (Sumatra, Java, Bali).

Distribution: Java, Bali, Sumatra. variegata intermedia

KRÜGER

1898 Euphaea intermedia KRÜGER, Stettiner ent. Ztg, 59: 76. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1934 Euphaea variegata intermedia - SCHMIDT, Arch. Hydrob. Suppl., 13: 329. 1954 Euphaea intermedia - LIEFTINCK, Treubia, 22 Suppl.: 19 (incorrect synonymy with variegata RAMBUR, 1842).

Distribution: Sumatra. yayeyamana

MATSUMURA & OKUMA

1913 Euphaea yayeyamana M A T S U M U R A & O K U M A , Dobuts. Z. Tokyo, 25: 307. Type male: Hokkaido Univ.; type-locality: Japan. 1961 Euphaea yayeyamana - ASAHINA, Insecta matsum., 24: 58.

Distribution: Japan. Genus Heterophaea

COWLEY

1903 Paraphaea M A R T I N , Bull. Mus. Paris, 8: 507. Type species: Paraphaea barbarata M A R T I N , 1903 (original designation). - 1934 Heterophaea COWLEY, Entomologist, 67: 201 (new name for Paraphaea MARTIN, 1903, a junior primary homonymy of Paraphaea BATES, 1873 in Coleoptera).

barbarata

(MARTIN)

1903 Paraphaea barbarata M A R T I N , Bull. Mus. Paris, 8: 507 (as generotype of Paraphaea M A R T I N , 1903). Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Manila. 1934 Heterophaea barbarata - COWLEY, Entomologist, 67: 201 (as generotype of Heterophaea COWLEY, 1934). 1939 Heterophaea barbarata - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 247.

Distribution: Manila, Luzon, Mindoro. ruficollis (Ris) 1930 Paraphaea ruficollis Ris, Mitt, miinch. ent. Ges., 20: 90. Type male: unknown; type-locality: Luzon. 1939 Heterophaea ruficollis - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 247.

Distribution: Luzon. Genus Indophaea 1929

FRASER

Indophaea FRÄSER, J . Bombay nat. Hist. Soc., 33: 2 9 3 . Type species: Euphaea dispar R A M B U R , 1842 (original designation). - 1954 Indophaea - LIEFTINCK, Treubia, 22 Suppl.: 15 (proposed synonymy with Euphaea SELYS, 1 8 4 0 ) .

cardinalis

(FRASER)

1924 Pseudophaea cardinalis FRASER, Ree. Indian Mus., 26: 512. Type male: British Museum (Nat. Hist.), London; type-locality: West India. 1929 Indophaea

cardinalis

— FRASER, J. B o m b a y n a t . Hist. Soc., 33: 295.

89 1931 Indophaea cardinalis - FRÄSER, Ree. Indian Mus., 33: 463. 1966 Indophaea cardinalis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 185.

Distribution: West India. dispar

(RAMBUR)

1842 Euphaea dispar RAMBUR, Hist. nat. Ins. Neuropt., Paris: 230. Type female: unknown; typelocality: India. 1853 Euphaea (Euphaea) dispar - SELYS, Synopsis des Calopterygines: 51. 1 8 5 4 Euphaea (Euphaea) dispar - SELYS & H A G E N , Monogr. Cal.: 1 6 9 (redescription) (Bengal). 1890 Pseudophaea dispar - KIRBY, Syn. Cat. Neur.-Odon., London: 109. 1924 Pseudophaea dispar - FRÄSER, Ree. Indian Mus., 26: 479 (West India). 1929 Indophaea dispar - FRÄSER, J. Bombay nat. Hist. Soc., 33: 293 (as generotype of Indophaea FRASER, 1929).

Distribution: India. fraseri

(LAIDLAW)

1920 Pseudophaea fraseri LAIDLAW, Ree. Indian Mus., 19: 23. Type male: Indian Museum ?; typelocality: India. 1924 Pseudophaea fraseri fraseri - FRÄSER, Ree. Indian Mus., 26: 480. 1929 Indophaea fraseri - FRÄSER, J. Bombay nat. Hist. Soc., 33: 296.

Distribution: India. wynaadensis (FRASER) stat. nov. 1922 Pseudophaea fraseri race wynaadensis FRASER, Ree. Indian Mus., 24: 8. Type male: originally in Fraser's Collection, now unknown; type-locality: India. 1924 Pseudophaea fraseri wynaadensis - FRASER, Ree. Indian Mus., 26: 480. 1966 Indophaea fraseri wynaadensis - KIMMINS, Burr. Brit. Mus. nat. Hist. (Ent.); 18: 219 (as syn. with fraseri ( L A I D L A W , 1 9 2 0 ) . 1 9 8 4 Indophaeafraser f. wynaadensis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 2 4 .

Distribution: India. Genus Schmidtiphaea

ASAHINA

1978 Schmidtiphaea ASAHINA, Proc. Jap. Soc. syst. Zool., 14: 44. Type species: schmidti ASAHINA, 1978 (original designation).

schmidti

Schmidtiphaea

ASAHINA

1978 Schmidtiphaea schmidti ASAHINA, Proc. Jap. Soc. syst. Zool., 14:44 (as generotype of Schmidtiphaea ASAHINA, 1978). Type male: Asahina's Collection; type locality: Assam.

Distribution: India. Family Polythoridae Medium to large forms with broad, often coloured wings. Accessory antenodals present, IRiii and Riv recessed, usually only distal primary antenodal evident and strengthened to support Dc instead of support by arc; sectors of arc arising from its costal angle making irregular shaped Dc. Base of Dc connected to radius without interposition of arc. Dc proximal side longer than distal. Stigma large. Larva with lateral abdominal gills.

90 1939 Polythoridae TILLYARD & FRÄSER, Austr. Zool., 9 : 208 (with genera Miocora, Cora, Euthore, Polythore, Chalcopteryx). - 1957 Polythoridae - FRÄSER, R . Zool. N . S . Wales, 1 9 5 7 : 75 (with subfamilies Euthorinae, Miocorinae, Polythorinae).

Subfamily Euthorinae Both primary antenodals present. De in fore wing with 5 - 6 cross veins. Wings clear or with white and brown bands. 1984 Euthorinae DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (with genus Euthore SELYS, 1869).

Genus Euthore

SELYS

1869 Thore (Euthore) SELYS, Bull. Acad. r. Belg., (2) 2 7 : 668. Type species: Thore fasciata SELYS, 1853 (original designation). - 1918 Euthore - Ris, Arch. Naturg., (A) 82: 25 (revision, key to species).

fasciata

(HÄGEN)

1853 Thore (Thore) fasciata HAGEN, in SELYS, Synopsis des Calopterygines: 70. Type male: Hagen's Collection; type-locality: Venezuela. 1854 Thore (Thore) fasciata - SELYS & H A G E N , Monogr. Cal.: 259 (redescription). 1 8 6 9 Thore (Euthore) fasciata - SELYS, B U L L . Acad. r. Belg., ( 2 ) 2 7 : 6 7 6 . 1873 Thore (Euthore) plagiata SELYS, Synopsis des Calopterygines, Add. 3: 37. Type female: McLachlan's Collection; type-locality: Rio Negro. 1909 Euthore fasciata fasciata - CALVERT, Ann. Carnegie Mus., Pittsburg, 6: 91. 1916 Euthore fasciata - Ris, Arch. Naturg., (A) 82: 27 (Colombia). 1984 Euthore fasciata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25. 1 9 8 4 Euthore plagiata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 5 [proposed synonymy with fasciata ( H Ä G E N , 1 8 5 3 ) ] .

Distribution: Venezuela, Colombia. fassli Ris 1914 Euthore fassli Ris, Ent. Mitt., 3:282. Type male: Mus. Naturk., Berlin; type-locality: Colombia. 1916 Euthore fassli - Ris, Arch. Naturg., (A) 82: 27.

Distribution: Colombia. fastigiata

(SELYS)

1859 Thore (Thore) fastigiata SELYS, Synopsis des Calopterygines, Add. 1: 16. Type male: Selys's Collection; type-locality: Bogota. 1869 Thore (Euthore) fastigiata - SELYS, Bull. Acad. r. Belg., (2) 27: 677. 1916 Euthore fastigiata - Ris, Arch. Naturg., (A) 82: 28 (Colombia).

Distribution: Colombia. hyalina

(SELYS)

1853 Thore (Thore) hyalina SELYS, Synopsis des Calopterygines: 71. Type male: Selys's Collection; type-locality: Bahia. 1854 Thore (Thore) hyalina - SELYS & H Ä G E N , Monogr. Cal.: 261 (redescription). 1869 Thore (Euthore) hyalina - SELYS, Bull. Acad. r. Belg., (2) 27: 677. 1916 Euthore

hyalina

- R i s , A r c h . N a t u r g . , (A) 82: 26 ( C o l o m b i a ) .

91 1943 Euthore hyalina - SCHMIDT, Beitr. F a u n a Perus, 2: 245 (Colombia).

Distribution: Colombia and Brazil. inlactea

CALVERT

1909 Euthore inlactea CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 91. Type male: Carnegie Museum; type-locality: Colombia - Peru. 1943 Euthore fasciata inlactea - SCHMIDT, Beitr. F a u n a Perus, 2: 245 (Ecuador, Colombia). 1984 Euthore inlactea - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (Peru).

Distribution: Colombia, Ecuador, Peru. leroii Ris 1916 Euthore leroii Ris, Arch. Naturg., (A) 82: 28. Type male: not given; type-locality: Colombia.

Distribution: Colombia. meridana

SELYS

1879 Euthore meridana SELYS, Bull. Acad. r. Belg., (2) 47: 404. Type male: Selys's Collection; typelocality: Venezuela.

Distribution: Venezuela. mirabilis

MCLACHLAN

1878 Euthore mirabilis MCLACHLAN, Trans, ent. Soc. London, 1878: 87. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador.

Distribution: Ecuador. montgomeryi

RACENIS

1984 Euthore montgomeryi RACENIS, 1968 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25.

Distribution: Venezuela. terminalis

(MCLACHLAN)

1878 Cora terminalis MCLACHLAN, Trans, ent. Soc. London, 1878: 92. Type male: British Museum (Nat. Hist.), London; type-locality: Bolivia. 1914 Cora terminalis terminalis - FÖRSTER, Arch. Naturg., 80 (A): 60 (with subspecies). 1943 Cora terminalis - SCHMIDT, Beitr. F a u n a Perus, 2: 241. 1984 Euthore terminalis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25.

Distribution: Bolivia, and Peru. Subfamily Miocorinae Only distal primary antenodals present; anal vein simple or no intercalary if bifurcate. Wings clear or with brown tip. Dc in fore wing with 3 - 4 cross veins. 1984 Miocorinae - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (with genera Josocora, Kalocora, Miocora and Stenocora).

Genus Josocora

KENNEDY

1940 Josocora KENNEDY, Ann. ent. Soc. Amer., 33: 408. Type species: Cora jocosa MCLACHLAN, 1881 (original designation) (in subfamily Polythorinae). - 1984 Josocora - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (transferred to subfamily Miocorinae).

92

jocosa

(MCLACHLAN)

1881 Cora jocosa McLachlan, Entomologist's month. Mag., 1881: 30. Type male: McLachlan's Collection; type-locality: Ecuador. 1940 Josocora jocosa - KENNEDY, Ann. ent. Soc. Amer., 33: 408 (as generotype of Josocora KENNEDY, 1 9 4 0 ) .

Distribution: Ecuador. klenei

(KARSCH)

1891 Cora klenei KARSCH, Soc. Ent., 6: 113. Type male: Mus. Naturk., Berlin; type-locality: Ecuador. 1984 Josocora klenei - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25.

Distribution: Ecuador. Genus Kalocora

KENNEDY

1940 Kalocora KENNEDY, Ann. ent. Soc. Amer., 33: 407. Type species: Cora aurea Ris, 1916 (original designation) (in subfamily Polythorinae). - 1984 Kalocora - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (transferred to subfamily Miocorinae).

aurea (Ris) 1916 Cora aurea Ris, Arch. Naturg., (A) 82: 24. Type male: not given; type-locality: Colombia. 1940 Kalocora aurea-KENNEDY, Ann. ent. Soc. Amer., 33:407 (as generotype of Kalocora K E N N E D Y , 1940).

Distribution: Colombia. Genus Miocora 1917

CALVERT

Miocora CALVERT, Ent. News., 28: 2 5 9 . Type species: Miocora peraltica CALVERT, 1 9 1 7 (original designation). - 1941 Miocora - KENNEDY, Ann. ent. Soc. Amer., 33: 417 (key to species). - 1 9 8 4 Miocora - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 5 .

pellucida

KENNEDY

1941 Miocora pellucida KENNEDY, Ann. ent. Soc. Amer., 33: 418. Type male: Kennedy's Collection; type-locality: Peru.

Distribution: Peru. peraltica

CALVERT

1917 Miocora peraltica CALVERT, Ent. News, 28: 260. Type male: unknown; type-locality: Costa Rica (as generotype of Miocora CALVERT, 1917).

Distribution: Costa Rica. subapicalis 1984

KENNEDY

Miocora subapicalis Utrecht, 1: 25.

KENNEDY,

1940 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Colombia. Genus Stenocora

KENNEDY

1940 Stenocora KENNEDY, Ann. ent. Soc. Amer., 33: 411. Type species: Stenocorapercornuta NEDY, 1940 (original designation).

KEN-

93 percornuta KENNEDY 1940 Stenocora percornuta Kennedy, Ann. ent. Soc. Amer., 33: 411. (as generotype of Stenocora KENNEDY, 1940). Type male: Acad. Sei. Philadelphia; type-locality: Ecuador. 1961 Stenocora percornuta - CALVERT, Proc. Acad. nat. Sei. Philad., 113: 2 (redescription) (Peru).

Distribution: Ecuador, Peru.

Subfamily Polythorinae Only distal primary antenodal present; anal vein bifurcate with intercalary vein ( = trifurcate). Dc in fore wing with 6 - 8 cross veins. Wings usually coloured. 1919 Polythorinae MUNZ, Mem. Amer. ent. Soc., 3: 42 (key to genera Polythore, Euthore, Chalcopteryx, Cora, Miocora). - 1 9 8 4 Polythorinae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (with genera Chalcopteryx, Cora, Polythore).

Genus Chalcopteryx SELYS 1853 Thore (Chalcopteryx) SELYS, Synopsis des Calopterygines: 69. Type species: Rhinocypha rutilans RAMBUR, 1 8 4 2 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 1 6 ) . 1 8 5 4 Thore (Chalcopteryx) - SELYS & H A G E N , Monogr. Cal.: 2 5 0 (revision). - 1 8 9 0 Chalcopteryx - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 1 6 . - 1 9 4 3 Chalcopteryx - SCHMIDT, Beitr. Fauna Perus, 2: 242 (key to species).

radians Ris 1916 Chalcopteryx radians Ris, Ent. Mitt., 3: 284. Type male: Senckenberg Museum; type-locality: Mato Grosso. 1943 Chalcopteryx radians - SCHMIDT, Beitr. Fauna Perus, 2: 244 (in key) (Amazons).

Distribution: Brazil and Peru. rutilans rutilans (RAMBUR) 1842 Rhinocypha rutilans RAMBUR, Hist. nat. Ins. Neuropt., Paris: 233. Type: sex not given: Mus. Nation. Hist. Nat., Paris; type-locality: not given. 1853 Thore (Chalcopteryx) rutilans - SELYS, Synopsis des Calopterygines: 68 (Para). 1 8 5 4 Thore (Chalcopteryx) rutilans - SELYS & H Ä G E N , Monogr. Cal.: 2 5 1 (redescription). 1890 Chalcopteryx rutilans - KIRBY, Syn. Cat. Neur.-Odon., London: 116 (as generotype of Chalcopteryx SELYS, 1853) (Para, Santerem). 1943 Chalcopteryx rutilans rutilans - SCHMIDT, Beitr. Fauna Perus, 2: 245 (Bolivia, Mato Grosso, Para).

Distribution: Bolivia, Brazil. rutilans confluens SCHMIDT 1943 Chalcopteryx rutilans confluens SCHMIDT, Beitr. Fauna Perus, 2: 245 (in key) from Manicore. 1984 Chalcopteryx rutilans f. confluens - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 6 .

Distribution: Brazil. scintillans M C L A C H L A N 1870 Chalcopteryx scintillans MCLACHLAN, Trans, ent. Soc. London, 1870: 169. Type male: McLachlan's Collection; type-locality: Säo Paulo. 1917 Chalcopteryx scintillans - SJOSTEDT, Ark. Zool., Uppsala, 11 (15): 35 (Amazonas).

94 1943 Chalcopteryx

scintillans - SCHMIDT, Beitr. F a u n a Perus, 2: 242 (redescription) (Peru).

Distribution: Brazil, Peru. seabrai 1984

SANTOS & MACHADO

Chalcopteryx seabrai SANTOS & of the World, Utrecht, 1: 26.

MACHADO, 1961

- loc. cit.

D A VIES & TOBIN,

The Dragonflies

Distribution: Brazil. Genus Cora Selys 1853

Thore (Cora) SELYS, Synopsis des Calopterygines: 7 1 . Type species: Cora cyanea SELYS, 1 8 5 3 (monotypy). - 1 8 5 4 Thore (Cora) - SELYS & H A G E N , Monogr. Cal.: 2 6 2 . - 1 8 9 0 Cora KIRBY, Syn. Cat. Neur.-Odon., London: 118. - 1918 Cora - Ris, Arch. Naturg., (A) 82: 14 (key to species). - 1943 Cora - SCHMIDT, Beitr. F a u n a Perus, 2: 241.

alcyone

SELYS

1873 Thore (Cora) alcyone SELYS, Synopsis des Calopterygines, Add. 3: 39. Type male: Selys's Collection; type-locality: Bogota. 1890 Cora alcyone - KIRBY, Syn. Cat. Neur.-Odon., London: 118.

Distribution: Colombia. bogotensis

FÖRSTER

1914 Cora terminalis race bogotensis FÖRSTER, Arch. Naturg. (A) 80: 60. Male from Colombia. 1946 Cora terminalis bogotensis - FRASER, Trans. R. ent. Soc. London., 96: 24 [incorrect synonymy with Euthore modesta (SELYS, 1869)]. 1984 Cora bogotensis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 6 .

Distribution: Colombia.

brasiliensis

HÄGEN

1869 Thore (Cora) brasiliensis HAGEN, in SELYS, Synopsis des Calopterygines, Add. 2: 34. Type male: Hagen's Collection; type-locality: Brazil. 1890 Cora brasiliensis - KIRBY, Syn. Cat. Neur.-Odon., London: 118. 1894 Cora brasiliensis „ M O N T G O M E R Y , 1967 ( 5 : 1 4 3 ) " (misprint) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 26.

Distribution: Brazil. chirripa

CALVERT

1907 Cora chirripa CALVERT, Biol. Centr.-Amer., London: 348. Type male: unknown; type-locality: Costa Rica. 1916 Cora chirripa - Ris, Arch. Naturg., (A) 82: 19.

Distribution: Costa Rica. confusa

KENNEDY

1940 Cora confusa KENNEDY, Ann. ent. Soc. Amer., 33: 430. Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador.

95 cyane

SELYS

1853 Thore (Cora) cyane SELYS, Synopsis des Calopterygines: 71 (as generotype of Cora SELYS, 1853). Type male: Selys's Collection; type-locality: Venezuela. 1854 Thore (Cora) cyane - SELYS & H A G E N , Monogr. Cal.: 265 (redescription). 1869 Thore (Cora) incana HAGEN, in SELYS, Synopsis des Calopterygines, Add. 2: 35. Type male: Selys & Hagen's Collection; type-locality: Venezuela. 1890 Cora cyane - KIRBY, Syn. Cat. Neur.-Odon., London: 118. 1890 Cora incana - KIRBY, Syn. Cat. Neur.-Odon., London: 118.

Distribution: Venezuela. dualis

MCLACHLAN

1878 Cora dualis MCLACHLAN, Trans, ent. Soc. London, 1878: 90. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador.

Distribution: Ecuador, inca

SELYS

1873 Thore (Cora) inca SELYS, Synopsis des Calopterygines, Add. 3: 39. Type male: R. Mus., Brussels; type-locality: Quito. 1890 Cora inca - KIRBY, Syn. Cat. Neur.-Odon., London: 118. 1916 Cora inca - Ris, Arch. Naturg., (A) 82: 17 (Colombia).

Distribution: Colombia. irene Ris 1916 Cora irene Ris, Arch. Naturg., (A) 82: 19. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Bolivia. 1943 Cora irene - SCHMIDT, Beitr. Fauna Perus, 2: 241 (Peru).

Distribution: Bolivia, Peru. lugubris

NAVÄS

1934 Cora lugubris NAVÄS, Broteria, 30 (3): 135. Type male: not given; type-locality: Colombia.

Distribution: Colombia. marina

SELYS

1869 Thore (Cora) marina SELYS, Synopsis des Calopterygines, Add. 2: 34. Type male: Selys's Collection; type-locality: Mexico. 1890 Cora marina - KIRBY, Syn. Cat. Neur.-Odon., London: 118. 1916 Cora marina - Ris, Arch. Naturg., (A) 82: 19 (Panama).

Distribution: Mexico, Panama. modesta

SELYS

1869 Thore (Cora) modesta SELYS, Synopsis des Calopterygines, Add. 2: 36. Type female: Selys's Collection; type-locality: Colombia. 1890 Cora modesta - KIRBY, Syn. Cat. Neur.Odon., London: 118. 1914 Cora terminalis race bogotensis FÖRSTER, Arch. Naturg., (A) 80: 60. Male from Colombia. 1916 Cora modesta - Ris, Arch. Naturg., (A) 82: 22 (redescription). 1946 Euthore modesta - FRASER, Trans. R. ent. Soc. London, 96: 24. 1946 Cora terminalis race bogotensis - FRASER, Trans. R. ent. Soc. London, 96: 24 (proposed synonymy with modesta SELYS, 1869).

Distribution: Colombia.

96 munda

MCLACHLAN

1878 Cora munda MCLACHLAN, Trans, ent. Soc. London, 1878: 91. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador.

Distribution: Ecuador. notoxantha Ris 1916 Cora notoxantha Ris, Arch. Naturg., (A) 82:20. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Panama.

Distribution: Panama. obscura Ris 1916 Cora obscura Ris, Arch. Naturg., (A) 82: 21. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Panama.

Distribution: Panama. semiopaca

SELYS

1878 Cora semiopaca SELYS, C.R. Soc. ent. Belg., 21: 21. Type male: R. Mus., Brussels; type-locality: Panama. 1914 Cora terminalis semiopaca - FÖRSTER, Arch. Naturg., (A) 80: 60. 1916 Cora semiopaca - Ris, Arch. Naturg., (A) 82: 22.

Distribution: Panama. subfumata

FÖRSTER

1914 Cora semiopaca race subfumata FÖRSTER, Arch. Naturg., (A) 80: 60. Male from Peru. 1943 Cora semiopaca subfumata - SCHMIDT, Beitr. Fauna Perus, 2: 241. 1 9 8 4 Cora subfumata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 2 6 .

Distribution: Peru. skinneri

CALVERT

1907 Cora skinneri CALVERT, Biol. Centr.-Amer., London: 349. Type male: unknown; type-locality: Costa Rica.

Distribution: Costa Rica. xanthostoma Ris 1916 Cora xanthostoma Ris, Arch. Naturg., (A) 82: 16. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Colombia.

Distribution: Colombia. Genus Polythore

CALVERT

1853 Thore H A G E N , in SELYS, Synopsis des Calopterygines: 69. Type species: Thore gigantea SELYS, 1853 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 116). - 1 8 5 4 Thore (Thore) - SELYS & H A G E N , Monogr. Cal.: 249 (with subgenera Chalcopteryx, Cora, Thore). - 1861 Thore - HÄGEN, Syn. Neur. N. Amer., Washington: 307. - 1890 Thore - KIRBY, Syn. Cat. Neur.-Odon., London: 1 1 6 . - 1903 Thore - NEEDHAM, Proc. U.S. natn. Mus., 26: 7 1 7 . - 1 9 1 7 Polythore CALVERT, Ent. News, 28: 262. Type species: Thore gigantea SELYS, 1853 (original designation) (new name for Thore H Ä G E N , 1853, a junior primary homonymy of Thore K O C H , 1850 in Arachnoidea). - 1918 Thore - Ris, Arch.Naturg., (A) 82: 29 (key to species). - 1943 Thore - SCHMIDT, Beitr. Fauna Perus, 2: 245 (key to species). - 1943 Polythore - SCHMIDT, Beitr. Fauna Perus, 2 : 2 4 5 (incorrect synonymy with Thore H Ä G E N , 1 8 5 3 ) . - 1 9 4 8 Polythore

97 - FRÄSER, Trans. R. ent. Soc. London, 96: 12 (revision, key to species of group concinna). 1 9 8 4 Thore - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 26 (proposed synonymy with Poly thore CALVERT, 1 9 1 7 ) .

acostai (NAVAS) comb. nov. 1924 Thore acostai NAVÄS, Mem. R. Acad. Ci. Barcelona, 3 (18): 9. Type male: Madrid Museum; type-locality: Peru.

Distribution: Peru. aurora

(SELYS)

1879 Thore aurora SELYS, Bull. Acad. r. Belg., (2) 47:401. Type male: R. Mus., Brussels; type-locality: Ecuador. 1924 Thore batesi N A V A S (nec SELYS, 1869), Mem. R. Acad. Ci. Barcelona, 3 (18): 319 (emandation). 1943 Thore aurora - SCHMIDT, Beitr. Fauna Perus, 2: 247. 1 9 4 3 Thore batesi - SCHMIDT, Beitr. Fauna Perus, 2 : 2 4 7 (proposed synonymy with aurora (SELYS, 1879).

1946 Polythore aurora - FRASER, Trans. R. ent. Soc. London, 96: 23.

Distribution: Ecuador, Peru. batesi (SELYS) comb nov. 1869 Thore (thore) batesi SELYS (nec NAVÄS, 1924), Synopsis des Calopterygines, Add. 2: 29. Type male: Bates's Collection; type-locality: Säo Paulo. 1890 Thore batesi - KIRBY, Syn. Cat. Neur.-Odon., London: 117.

Distribution: Brazil. beata

(MCLACHLAN)

1869 Thore beata MCLACHLAN, Entomologist's month. Mag., 1869: 28. Type male: British Museum (Nat. Hist.), London; type-locality: Peru. 1869 Thore (Thore) beata - SELYS, Synopsis des Calopterygines, Add. 2: 30. 1869 Thore (Thore) inaequalis SELYS, Synopsis des Calopterygines, Add. 2: 30. Type female: Selys's Collection; type-locality: Amazonas. 1890 Thore beata - KIRBY, Syn. Cat. Neur.-Odon., London: 117. 1890 Thore inaequalis - KIRBY, Syn. Cat. Neur.-Odon., London: 117. 1920 Thore beata - NAVAS, An. Soc. Cient., Argentina, 90: 34. 1943 Thore beata - SCHMIDT, Beitr. Fauna Perus, 2: 347. 1946 Polythore beata - FRASER, Trans. R. ent. Soc. London, 96: 23.

Distribution: Brazil, Peru. boliviana (MCLACHLAN) comb. nov. 1878 Thore boliviana MCLACHLAN, Trans, ent. Soc. London, 1878: 89. Type male: British Museum (Nat. Hist.), London; type-locality: Bolivia. 1916 Thore boliviana - Ris, Arch. Naturg., (A) 82: 32 (Peru). 1943 Thore boliviano boliviana - SCHMIDT, Beitr. Fauna Perus, 2: 247.

Distribution: Bolivia, Peru. concinna

(MCLACHLAN)

1881 Thore concinna MCLACHLAN, Entomologist's month. Mag., 1881: 28. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador. 1889 Sapho pulchella KIRBY, Proc. Zool. Soc. London, 1889: 300. Type male: British Museum (Nat. Hist.), London; type-locality: ? Cameroon. 1946 Polythore concinna - FRASER, Trans. R. ent. Soc. London, 96: 22 (Ecuador, Colombia).

98 1962 Sapho pulchella - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 146 [proposed synonymy with concinna ( M C L A C H L A N , 1 8 8 1 ) ] .

Distribution: Ecuador, Colombia. derivata derivata

(MCLACHLAN)

1881 Thore derivata MCLACHLAN, Trans.ent. Soc. London, 1881: 81. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador. 1916 Thore derivata - Ris, Arch. Naturg., (A) 82: 36 (redescription) (Peru). 1943 Thore picta derivata - SCHMIDT, Beitr. Fauna Perus, 2: 250 (Peru). 1946 Polythore derivata race originate FRASER, Trans. R. ent. Soc. London, 96:18. Male from Peru. 1 9 4 6 Polythore derivata race terminate FRASER, Trans. R . ent. Soc. London, 9 6 : 1 8 . Male from Peru. 1966 Polythore derivata originata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 208. [Holotype male: British Museum (Nat. Hist.), London)]. 1966 Polythore derivata terminata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 208 ([Holotype male: British Museum (Nat.Hist.), London)]. 1984 Polythore derivata f. originata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:27. 1984 Polythore derivata derivata f. terminata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 27.

Distribution: South America. derivata adjuncta

FRASER

stat. nov.

1946 Polythore derivata race adjucta FRASER, Trans. R. ent. Soc. London, 96: 18. Male from Colombia.

Distribution: Colombia. derivata ambigua

FRASER

stat. nov.

1946 Polythore derivata race ambigua FRASER, Trans. R. ent. Soc. London, 96:19. Male from Peru.

Distribution: Peru. gigantea

(SELYS)

1853 Thore (Thore) gigantea SELYS, Synopsis des Calopterygines: 69 (as generotype of Thore SELYS, 1853). Type male: Selys's Collection; type-locality: Colombia. 1854 Thore (Thore) gigantea - SELYS & H A G E N , Monogr. Cal.: 254 (redescription). 1890 Thore gigantea - KIRBY, Syn. Cat. Neur.-Odon., London: 116. 1916 Thore gigantea - Ris, Arch. Naturg., (A) 82: 38 (Colombia, Ecuador). 1917 Polythore gigantea - CALVERT, Ent. News, 28:263 (as generotype of Polythore CALVERT, 1917). 1946 Polythore gigantea - FRASER, Trans. R. ent. Soc. London, 96: 15 (Amazonas).

Distribution: Brazil, Colombia, Ecuador. montana

(FÖRSTER)

1914 Thore montana FÖRSTER, Arch. Naturg., (A) 80: 60. Type female: Förster's Collection; typelocality: Peru. 1943 Thore montana - SCHMIDT, Beitr. F a u n a Perus, 2: 248 [incorrect synonymy with pozuzina (FÖRSTER, 1914)].

1984 Polythore montana species).

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht, 1: 27 (as good

Distribution: Peru. mutata

(MCLACHLAN)

1881 Thore mutata MCLACHLAN, Entomologist's month. Mag., 1881: 29. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador.

99 1946 Polythore mutata - FRÄSER, Trans. R. ent. Soc. London, 96: 24 (Ecuador, Colombia).

Distribution: Ecuador, Colombia. ornata

(SELYS)

1879 Thore ornata Peru. 1890 Thore ornata 1916 Thore ornata 1943 Thore ornata

comb. nov. SELYS, Bull. Acad. r. Belg., (2)47:400. Type male: R. Mus., Brussels; type-locality: - KIRBY Syn. Cat. Neur.-Odon., London: 116. - Ris, Arch. Naturg., (A) 82: 34. ornata - SCHMIDT, Beitr. Fauna Perus, 2: 247.

Distribution: Peru. picta

(RAMBUR)

1842 Euphaea picta RAMBUR, Hist. nat. Ins. Neuropt., Paris: 231. Type male: Marchal's Collection; type-locality: Guyana. 1853 Thore (Thore) picta - SELYS, Synopsis des Calopterygines: 70. 1853 Thore (Thore) saundersii SELYS, Synopsis des Calopterygines: 70. Type male: Saunder's Collection; type-locality: Amazons. 1873 Thore (Thore) picturata SELYS, Synopsis des Calopterygines, Add. 3: 35. Type male: Selys's Collection; type-locality: Cayenne. 1890 Thore picta - KIRBY, Syn. Cat. Neur.-Odon., London: 116. 1890 Thore saundersii - KIBY, Syn. Cat. Neur.-Odon., London: 117. 1943 Thore picta picta - SCHMIDT, Beitr. F a u n a Perus, 2: 248 (key to subspecies). 1946 Polythore picta - FRASER, Trans. R. ent. Soc. London, 46: 14 (Colombia, Peru, Ecuador, Guyana).

Distribution: Colombia, Peru, Brazil, Ecuador, Guyana, pozuzina

(FÖRSTER)

1914 Thore pozuzina FÖRSTER, Arch. Naturg., (A) 80: 59. Type male: Mus. Naturk., Berlin; typelocality: Peru. 1943 Thore ornata pozuzina - SCHMIDT, Beitr. Fauna Perus, 2: 248. 1984 Polythore pozuzina - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 27.

Distribution: Peru. procera

(SELYS)

comb. nov.

1869 Thore (Thore) procera SELYS (as gigantea race procera), Synopsis des Calopterygines, Add. 2: 27. Type male: Selys's Collection; type-locality: Bogota. 1890 Thore procera - KIRBY, Syn. Cat. Neur.-Odon., London: 116. 1916 Thore procera - Ris, Arch. Naturg., (A) 82: 34 (Colombia). 1 9 4 8 Thore procera - FRASER, Trans, r. ent. Soc. London, 9 6 : 1 4 [as syn. with picta ( R A M B U R , 1 8 4 2 ) ] .

Distribution: Colombia. tincta

(NAVÄS)

1924 Thore tincta NAVAS, Mem. R. Acad. Ci. Barcelona, 3 (18): 7. Type male: unknown; typelocality: Peru. 1943 Thore tincta - SCHMIDT, Beitr. F a u n a Perus, 2: 250 [proposed synonymy with vittata (SELYS, 1869)]. 1984 Polythore tincta - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 25 (as good species).

Distribution: Peru.

100 vittata

(SELYS)

1869 Thore (Thore) picta race vittata SELYS, Synopsis des Calopterygines, Add. 2: 29. Male from Amazons. 1873 Thore (thore) picta race aequatorialis SELYS, Synopsis des Calopterygines, Add. 3: 36. Female from Ecuador. 1873 Thore albovittata SELYS, Bull. Acad. r. Belg., (2) 36: 618. Type male: R. Mus., Brussels; typelocality: Amazonas. 1890 Thore vittata - KIRBY, Syn. Cat. Neur.-Odon., London: 117. 1916 Thore vittata - Ris, Arch. Naturg., (A) 82: 37 (Peru). 1916 Thore aequatorialis - Ris, Arch. Naturg., (A) 82: 31 [incorrect synonymy with victoria (MCLACHLAN, 1869)].

1943 Thore vittata - SCHMIDT, Beitr. Fauna Perus, 2: 250. 1943 Thore albovittata - SCHMIDT, Beitr. Fauna Perus, 2: 250 [proposed synonymy with vittata (SELYS, 1869)].

1946 Polythore vittata - FRASER, Trans. R. ent. Soc. London, 96: 21 (Ecuador).

Distribution: Ecuador, Brazil. victoria

(MCLACHLAN)

comb. nov.

1869 Thore victoria MCLACHLAN, Entomologist's month. Mag., 6: 28. Type female: McLachlan's Collection; type-locality: Bolivia. 1869 Thore (Thore) victoria - SELYS, Synopsis des Calopterygines, Add. 2: 25 (redescription). 1890 Thore victoria - KIRBY, Syn. Cat. Neur.-Odon., London: 116. 1916 Thore victoria - Ris, Arch. Naturg., (A) 82: 31 (Peru).

Distribution: Bolivia, Peru. williamsoni

(FÖRSTER)

1903 Thore williamsoni FÖRSTER, Insekten-Börse, 20, sep.: 2. Type male: Mus. Naturk., Berlin; type-locality: Peru. 1943 Thore boliviana williamsoni - SCHMIDT, Beitr. Fauna Perus, 2: 247. 1984 Polythore williamsoni - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 27.

Distribution: Peru. Family Rimanellidae Medium sized, non-metallic Zygoptera with lestine facies. First thoracic interpleural suture incomplete. Wings narrow, petiolated to first primary antenodal; distal primary just distal to arc, no secondary antenodals. Postnodals numerous, not co-inciding with cross veins beneath. Median space uncrossed; space below Rl free from arc to nodus. IRiii and Riv substantially recessed; Riii arising almost at nodus. Dc short and bread, especially in forewing and angled sharply down distally; several intercalaries. Appendages sub-forcipate. One species only, isolated in north of S. America. 1984 Rimanellidae DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 27. Type-genus: Rimanella

NEEDHAM, 1934.

Genus Rimanella

NEEDHAM

1933 Rima NEEDHAM, Amer. Mus. Novit., 664: 3. Type species: Rima arcana NEEDHAM, 1933 (original designation). - 1934 Rimanella NEEDHAM, Ent. News, 45: 50 (new name for Rima N E E D HAM, 1 9 3 3 , a j u n i o r p r i m a r y h o m o n y m y o f Rima

PELLARY, 1 8 9 5 i n M o l l u s c a ) .

101 arcana

(NEEDHAM)

1933 Rima arcana NEEDHAM, Amer. Mus. Novit., 664: 3. Type male: Needham's Collection; typelocality: Venezuela (as generotype of Rima NEEDHAM, 1933). 1934 Rimanella arcana -NEEDHAM, Ent. News, 45: 50 (as generotype of Rimanella NEEDHAM, 1934).

Distribution: Venezuela. Superfamily Lestinoidea Small to medium sized Zygoptera with petiolate wings; usually only 2 antenodals, postnodals in line with the cross veins below them; anal vein fused with posterior border of wings; Dc closed at base (except Chorismagrion); arc at level of second antenodal or distal to it; IRiii & Riv usually begin closer to arc than to nodus; usually oblique vein joints Riii and IRiii. Supplementary sectors (intercalaries) present; pterostigma long. Male genital hamules elongate. 1957

Lestinoidea FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 5 3 (for families Perilestidae, Chlorolestidae, Lestidae, Megapodagriidae). - 1962 Lestinoidea - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 88 (key to African families).

Family Chlorolestidae Rather robust forms with very petiolate wings. Stigma wide, intercalated veins present, Riv origin at or just proximal to subnodus. CuP curves sharply upwards from origin. Dc not always closed at base. 1926

Synlestidae T I L L Y A R D , Insects Austr. N. Zeal.: 7 6 . - 1 9 3 8 Synlestidae - T I L L Y A R D & FRASER, Austr. Zool., 9: 154 (with key to subfamilies Chorismagriinae, Perilestinae, Synlestinae, Megalestinae). - 1 9 5 1 Chlorolestidae FRASER, Ent. News, 62: 6 3 . - 1 9 6 2 Synlestidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 89. - 1962 Synlestidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 89 (proposed synonymy with Chlorolestidae). - 1984 Chlorolestidae DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:29 (with subfamilies Chlorolestinae, Chorismagrioninae, Megalestinae).

Subfamily Chlorolestinae Wings sometimes coloured in males. IRiii & Riv origin nearer nodus than to arc, anal vein leaves wing border below Dc and far distal to Ac; Dc normally closed at base. 1926 Synlestinae TILLYARD, Insects of Austr. and New Zeal.: 78 (with genera Synlestes, Chlorolestes). - 1938 Synlestinae - T I L L Y A R D & FRASER, Austr. Zool., 9: 155 (with genera Synlestes, Chlorolestes). - 1957 Chlorolestinae FRASER, R. Zool. Soc. N.S. Wales, 1957: 55. - 1962 Synlestinae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 90 (with genera Chlorolestes, Ecchlorolestes). - 1984 Chlorolestinae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 29 (with genera Chlorolestes, Ecchlorolestes, Episnylestes, Euchlorolestes, Phylolestes, Sinolestes, Synlestes).

Genus Chlorolestes

SELYS

1862 Chlorolestes SELYS, Bull. Acad. r. Belg., (2) 14: 33. Type species: Chlorolestes conspicuus SELYS, 1862 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 125). - 1890 Chlorolestes - KIRBY, Syn. Cat. Neur.-Odon., London: 125. - 1920 Euchlorolestes KENNEDY, Ohio J. Sei.

102 Columbus, 21: 84. Type species: Agrion fasciata BURMEISTER, 1839 (original designation). 1921 Chlorolestes - Ris, Annls S. Afr. Mus., 18: 279 (key to South Africa species). - 1962 Chlorolestes - PINHEY, Publ. cult. C o m p . Diam. Angola, 59: 90 (with subgenera Chlorolestes, s.str., Euchlorolestes K E N N E D Y .

apricans

WILMOT

1984 Chlorolestes apricans WILMOT, 1975 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the World,

Utrecht, 1: 29.

Distribution: South Africa. conspicuus

HÄGEN

1862 Chlorolestes conspicua HAGEN, in SELYS, Synopsis des Agrionines, 3: 32. Type male: unknown; type-locality: C a p of G o o d Hope. 1890 Chlorolestes conspicua - KIRBY, Syn. Cat. Neur.-Odon., London: 125 (as generotype of Chlorolestes

SELYS, 1 8 6 2 ) .

1921 Chlorolestes conspicuus - RIS, Annls S. Afr. Mus., 18: 280. 1955 Chlorolestes conspicuus - BRINCK, South African Animal Lifte, 2: 192 (Cape Province). 1962 Chlorolestes (Chlorolestes) conspicuus - PINHEY, Pub. cult. Comp. Diam. Angola, 59: 91 (types not designated).

Distribution: South Africa. draconicus

BALINSKY

1956 Chlorolestes draconicus BALINSKY, Ann. Transvaal Mus., 22: 511. Type male: Transvaal M u seum; type-locality: Natal. 1962 Chlorolestes (Euchlorolestes) draconicus- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91.

Distribution: South Africa. elegans

PINHEY

1950 Chlorolestes elegans PINHEY, Ann. Transvaal Mus., 21: 260. Type male: Transvaal Museum; type-locality: South Rhodesia. 1958 Chlorolestes elegans - PINHEY, Occ. Pap. nat. Mus. S. Rhodesia, 22B: 99 (Zambezi, Rhodesia). 1962 Chlorolestes (Euchlorolestes) elegans - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91 (S.Africa).

Distribution: South Africa, fasciatus

(BURMEISTER)

1839 Agrion fasciata BURMEISTER, H a n d b . Ent., 2: 822. Type male: Halle Museum; type-locality: Natal. 1862 Chlorolestes fasciata - SELYS, Synopsis des Agrionines, 3: 34. 1898 Chlorolestes fasciata - CALVERT, Trans. Am. ent. Soc., 25: 44. 1920 Euchlorolestes fasciata - KENNEDY, Ohio J. Sei. Columbus, 21: 84 (as generotype of Euchlorolestes

KENNEDY, 1 9 2 0 ) .

1921 Chlorolestesfasciata - Ris, Annls S. Afr. Mus., 18:284 (redescription) (Cape, Natal, Zululand). 1955 Chlorolestes fasciata - BRINCK, South African Animal Life, 2:192 (Basutoland, Natal, Transvaal). 1962 Chlorolestes (Euchlorolestes) fasciata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91. 1 9 8 4 Chlorolestes fasciatus nigerrimus PINHEY, 1 9 8 0 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 29 (Transvaal). 1984 Chlorolestes fasciatus f. nigerrimus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 29.

Distribution: South Africa.

103 tessellatus

(BURMEISTER)

1839 Agrion tessellata BURMEISTER, H a n d b . Ent., 2: 823. Type male: Halle Museum; type-locality: Cape of Good Hope. 1839 Agrion longicauda BURMEISTER, H a n d b . Ent., 2: 823. Type male: Halle Museum; type-locality: Cape of Good Hope. 1862 Chlorolestes tessellata - SELYS, Synopsis des Agrionines, 3: 33. 1862 Chlorolestes longicauda - SELYS, Synopsis des Agrionines, 3: 33. 1921 Chlorolestes tessellata - Ris, Annls S. Afr. Mus., 18: 286 (redescription) (S. Africa). 1962 Chlorolestes (Euchlorolestes) tesselatus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91. 1962 Agrion longicauda - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91 [proposed synonymy with tessellatus (BURMEISTER, 1 8 3 9 ) ] .

Distribution: South Africa. umbratus

HÄGEN

1862 Chlorolestes umbrata Hagen, in SELYS, Synopsis des Agrionines, 3: 35. Type male: Hagen's Collection; type-locality: Cape of Good Hope. 1921 Chlorolestes umbrata - Ris, Annls S. Afr. Mus., 18: 283 (redescription). 1962 Chlorolestes (Chlorolestes) umbrata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91 (types not designated).

Distribution: South Africa. Genus Ecchlorolestes

BARNARD

1937 Ecchlorolestes BARNARD, Annls S. Afr. Mus., 32: 190. Type species: Chlorolestes perengueyi Ris, 1921 (original designation). - 1962 Ecchloroletes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 91.

nylephtha

(BARNARD)

1937 Chlorolestes nylephtha BARNARD, Annls S. Afr. Mus., 32: 201. Type male: South African Museum; type-locality: Cape. 1962 Chlorolestes (Ecchloroletes) nylephtha - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 92. 1984 Ecchlorolestes nylephtha - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 29.

Distribution: South Africa. perengueyi (Ris) 1921 Chlorolestes perengueyi Ris, Annls S. Afr. Mus., 18: 282. Type male: South African Museum; type-locality: Cape Colony. 1937 Ecchlorolestes perengueyi - BARNARD, Annls S. Afr. Mus., 32: 190 (as generotype of Ecchlorolestes BARNARD, 1937). 1962 Ecchlorolestes perengueyi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 92.

Distribution: South Africa. Genus Episynlestes

KENNEDY

1920 Episynlestes KENNEDY, Ohio J. Sei. Columbus, 21: 84 Type species: Synlestes TILLYARD, 1914 (original designation).

albicauda

albicauda

(TILLYARD)

1914 Synletes albicauda TILLYARD, Proc. Linn. Soc. N.S. Wales, 38: 238. Type male: Tillyard's Collection; type-locality: Queensland. 1917 Synlestes albicauda - SJOSTEDT, Ark. Zool., Uppsala, 11 (11). 26.

104 1920 Episynlestes albicauda - KENNEDY, Ohio J. Sei. Columbus, 2J: 84 (as generotype of Episynlestes KENNEDY, 1 9 2 0 ) .

Distribution: Australia. cristatus

WATSON & MOULDS

1977 Episynlestes cristatus WATSON & M O U L D S , J1 Austr. ent. Soc., Collection; type-locality: Queensland.

16

(3): 257. Type male: Watson's

Distribution: Queensland. Genus Phylolestes 1948 Phylolestes CHRISTIANSEN, Psyche, 1948 (original designation).

CHRISTIANSEN ethalae

CHRISTIANSEN,

1948 Phylolestes ethalae CHRISTIANSEN, Psyche, 5 4 : 2 5 8 . (as generotype of Phylolestes 1948). Type male: Christiansen's Collection; type-locality: Haiti.

CHRISTIANSEN,

ethalae

54:

256. Type species: Phylolestes

CHRISTIANSEN

Distribution: Haiti. Genus Sinolestes

NEEDHAM

1930 Sinolestes NEEDHAM, Zool. Sinica, (A) 11: 243. Type species: Sinolestes editus NEEDHAM, 1930 (original designation). - 1933 Sinolestes - MAY, Senckenbergiana, 15: 256. - 1934 Sinolestes - COWLEY, Entomologist, 67: 253 (type species: Sinolestes truncatus NEEDHAM, 1930).

editus

NEEDHAM

1930 Sinolestes editum NEEDHAM, Zool. Sinica, (A) 11: 243 (as generotype of Sinolestes 1930). Type female: unknown, perhaps in Peking Museum; type-locality: China. 1943 Sinolestes editum - SCHMIDT, Mitt. D. ent. Ges., 11: 109 (Tsche-kiang). 1947 Sinolestes edita - CHAO, Biol. Bull. Fukien Christ. Univ., 6: 21. 1956 Sinolestes editus - ASAHINA, Ent. Medd., 27 (4-5): 217 (C. China).

NEEDHAM,

Distribution: China. ornatus

NEEDHAM

1930 Sinolestes ornatus NEEDHAM, Zool. Sinica, (A) 11:244. Type male: unknown, perhaps in Peking Museum; type-locality: China. 1 9 5 6 Sinolestes ornata - ASAHINA, Ent. Medd., 27 ( 4 - 5 ) : 2 1 7 (incorrect synonymy with editus N E E D HAM, 1 9 3 0 ) . 1984

Sinolestes ornatus -

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 3 0 .

Distribution: China. truncatus

NEEDHAM

1930 Sinolestes truncatus Needham, Zool. Sinica, (A) 11: 243. Type: unknown, perhaps in Peking Museum; type-locality: China. 1934 Sinolestes truncatus - COWLEY, Entomologist, 6 7 : 253 (as generotype of Sinolestes NEEDHAM, 1930).

Distribution: China.

105 Genus Synlestes SELYS 1869 Synlestes SELYS, C.R. SOC. ent. Belg., 11: 122. Type species: Synlestes weyersi SELYS, 1869 (monotype). - 1917 Synlestes - TILL YARD, Proc. Linn. Soc. N.S. Wales, 42: 469 (revision, key to species). - 1926 Synlestes - TILLYARD, Insects of Austr.-New Zeal.: 78 (in Synlestinae). - 1 9 3 8 Synlestes - TILLYARD & FRASER, Austr. Zool., 9: 1 5 5 . - 1 9 5 7 Synlestes - FRÄSER, R . Zool. Soc. N.S. Wales, 1957: 5 5 (in subfamily Chlorolestinae). - 1 9 6 2 Synlestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 9 0 (in subfamily Synlestinae). - 1 9 8 4 Synlestes - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 29 (in subfamily Chlorolestinae).

nigrescens TILL YARD 1917 Synlestes nigrescens TILLYARD, Proc. Linn. Soc. N.S. Wales, 42: 472. Type male: Tillyard's Collection; type-locality: New South Wales.

Distribution: Australia. selysi TILLYARD 1917 Synlestes selysi TILLYARD, Proc. Linn. Soc. N.S. Wales, 42: 473. Type male: Tillyard's Collection; type-locality: New South Wales.

Distribution: Australia, tillyardi

FRASER 1948 Synlestes tillyardi FRASER, Bull. Mus. Hist. nat. Belg., 34 (16): 4. Type male: R. Mus., Brussels; type-locality: S.E. Australia.

Distribution: Australia, tropicus

TILLYARD 1917 Synlestes tropicus TILLYARD, Proc. Linn. Soc. N.S. Wales, 42: 475. Type male: Tillyard's Collection; type-locality: Kuranda.

Distribution: North Queensland, weyersi SELYS 1869 Synlestes weyersi SELYS, C.R. Soc. ent. Belg., 11: 123. Type female: unknown, perhaps in R. Mus., Brussels; type-locality: Queensland (as generotype of Synlestes SELYS, 1 8 6 9 ) . 1901 Synlestes weyersi - MARTIN, Mem. Soc. zool. France, 14: 244 (Victoria). 1917 Synlestes weyersi - TILLYARD, Proc. Linn. Soc. N.S. Wales, 42: 472 (key to subspecies) (New South Wales). 1917 Synlestes weyersi - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 26 (Queensland). 1948 Synlestes weyersi - FRASER, Bull. Mus. Hist. nat. Belg., 24 (16): 2 (redescription).

Distribution: Australia. Subfamily Chorismagrioninae Dc of fore wing open at base; Riv origin at level of subnodus. 1938

Chorismagrioninae TILLYARD & FRASER, Austr. Zool., 9: 1 5 5 (in family Synlestidae) (with genus Chorismagrion M O R T O N , 1 9 1 4 ) . - 1 9 8 4 Chorismagrioninae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 30 (in family Chlorolestidae).

Genus Chorismagrion

MORTON

1914 Chorismagrion MORTON, Trans, ent. Soc. London, 1914: 170. Type species: Chorismagrion risi MORTON, 1914 (original designation) (for subfamily Synlestinae). - 1938 Chorismagrion - TILLYARD & FRASER, Austr. Zool., 9: 155 (in subfamily Chorismagrioninae).

106 risi

MORTON

1914 Chorismagrion risi MORTON, Trans, ent. Soc. London, 1914: 170 (as generotype of Chorismagrion MORTON, 1914). Type male: Morton's Collection; type-locality: Queensland. 1917 Chorismagrion risi - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 39 (redescription).

Distribution: Australia. Subfamily Megalestinae Wings hyaline, IRiii & Riv arise nearer to arc than to nodus; anal vein leaves wing border at or prox. to Ac. Dc closed at base. 1938 Megalestinae TILLYARD & FRASER, Austr. Zool., 9: 155 (in family Synlestidae) (with genera Megalestes, Orolestes). - 1984 Megalestinae - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 30 (in family Chlorolestidae) (with genus Megalestes).

Genus Megalestes

SELYS

1862 Lestes (Megalestes) SELYS, Bull. Acad. r. Belg., (2) 13: 293. Type species: Lestes (Megalestes) major SELYS, 1862 (monotypy). - 1890 Megalestes - KIRBY, Syn. Cat. Neur.-Odon., London: 159. - 1905 Megalestes - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 806. - 1938 Megalestes

chengi

- TILLYARD & FRASER, A u s t r . Z o o l . , 9: 155.

CHAO

1947 Megalestes chengi CHAO, Biol. Bull. Fukien Univ., 6:20. Type male: Fujian Univ.; type-locality: Fukien.

Distribution: China. distans

NEEDHAM

1930 Megalestes distans NEEDHAM, Zool. Sinica, (A) 11: 231. Type male: unknown, perhaps in Peking Museum; type-locality: China.

Distribution: China. heros

NEEDHAM

1930 Megalestes heros NEEDHAM, Zool. Sinica, (A) 11: 229. Type male: unknown, perhaps in Peking Museum; type-locality: China.

Distribution: China. irma

FRASER

1926 Megalestes irma FRASER, J. Darjeeling nat. Hist. Soc., 1: 32. Type male: originally in Darjeeling Museum, now unknown; type-locality: Sikkim.

Distribution: India. lieftincki

LAHIRI

1984 Megalestes lieftincki LAHIRI, 1979 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 30. Distribution: I n d i a .

107 maai

CHEN

1947 Megalestesmaai CHEN, Biol. Bull. Fukien Univ., 6:27. Type male: „Coll. Tokyo"; type-locality; Formosa. 1956 Megalestes maai - ASAHINA, Ent. Medd., 27 (4-5): 216 (description of female) (C. China).

Distribution: China, Taiwan. major

SELYS

1862 Lestes (Megalestes) type-locality: India 1890 Megalestes major 1916 Megalestes maior 1905 Megalestes major 1944 Rhinagrion major -

major SELYS, Synopsis des Agrionines, 2: 9. Type male: Selys's Collection; (as generotype of Megalestes SELYS, 1862). KIRBY, Syn. Cat. Neur.-Odon., London: 159. Ris, Suppl. ent. Berlin, 5: 11 (Formosa). JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 806. NEEDHAM, Bull. Mus. comp. Zool. Harv., 94: 163.

Distribution: India, Taiwan. micans

NEEDHAM

1930 Megalestes micans NEEDHAM, Zool. Sinica, (A) 11: 230. Type male: unknown, perhaps in Peking Museum; type-locality: China.

Distribution: China. riccii

NAVÄS

1935 Megalestes riccii NAVÄS, Mus. Heude ent. chinoise, 2 (5): 89. Type male: not given, perhaps in Heude Museum; type-locality: China. 1962 Megalestes riccii - CHAO, Acta ent. sinica, Suppl., 11: 28.

Distribution: China. suensoni

ASAHINA

1956 Megalestes suensoni ASAHINA, Ent. Medd., 27 (4-5): 213. Type female: Univ. zool. Mus., Kobenhavn; type-locality: China.

Distribution: China. Family Lestidae Dc well removed from wing border and with very acute distal angle; oblique vein between Riii & TRiii; anal vein leaves posterior border of wing at level of arc (not in Chalcolestes). 1 9 0 3 L e s t i d a e NEEDHAM, B u l l . 6 8 N . Y . S t a t e M u s . : 2 1 6 . - 1 9 0 5 Lestidae

- JACOBSON & BIANKI,

Pryam. lozhnos. Ross, imp.: 750. - 1938 Lestidae - TILLYARD & FRASER, Austr. Zool., 9: 156 (with subfamilies Hypolestinae, Lestinae). - 1951 Lestidae - FRASER, Ent. News, 62: 61 (classification). - 1962 Lestidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 92. - 1984 Lestidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 30 (with subfamilies Lestinae, Sympecmatinae).

Subfamily Lestinae Wings usually partly open at rest. Dc of all wings same shape and size; stigma rather elongated. 1903 Lestinae NEEDHAM, Bull. 68 N.Y. State Mus.: 216 (subfamily first defined). Type-genus: Lestes LEACH, 1815). - 1910 Lestinae - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 35 (with

108 genera Ortholestes, Archilestes, Lestes) (in family C o e n a g r i o n i d a e ) . - 1919 Lestinae - MUNZ, M e m . A m e r . ent. Soc., 3: 48 (key to genera Pseudolestes, Ortholestes, Archilestes, Megalestes, Orolestes, Platylestes, Sympycna, Austrolestes, Lestes). - 1938 Lestinae-TILLYARD & FRASER, A u s t r . Zool., 9: 157 (with genera Austrolestes, Africalestes, Archilestes, Superlestes, Cyptolestes, Platylestes, Ceylonolestes, Indolestes, Sympecma, Lestes).

Genus Archilestes

SELYS

1862 Lestes (Archilestes) SELYS, Synopsis des Agrionines, 2: 10. Type species: Lestes grandis RAMBUR, 1842 (original designation). - 1921 Superlestes WILLIAMSON, OCC. P a p . M u s . Z o o l . U n i v . M i c h i g a n , 96: 2. Type species: Lestes (Lestes) exoletus HÄGEN, 1862 (original designation). - 1921 Cyptolestes WILLIAMSON, Occ. P a p . M u s . Z o o l . U n i v . M i c h i g a n , 96: 4. Type-species: Cyptolestes tuberculatus WILLIAMSON, 1921 (original designation). - 1984 Archilestes (Cyptolestes) - DAVIES & TOBIN, T h e Dragonflies of the World, U t r e c h t , 1: 30.

californicus

MCLACHLAN

1895 Archilestes califomica MCLACHLAN, A n n . M a g . nat. Hist., (6) 16: 20. Type male: British M u seum ( N a t . Hist.), L o n d o n ; type-locality: California. 1915 Archilestes

californicus

- KENNEDY, Proc. U.S. n a t n . mus., 49: 260 (life-history).

Distribution: U.S.A.: California. exoletus

(HÄGEN)

1862 Lestes (Lestes) exoleta HAGEN, in SELYS, Synopsis des Agrionines, 2: 26. Type male: M u s . N a t u r k . , Berlin; type-locality: Brazil. 1890 Lestes exoletus - KIRBY, Syn. C a t . N e u r . - O d o n . , L o n d o n : 161. 1921 Superlestes exoletus - WILLIAMSON, Occ. P a p . M u s . U n i v . M i c h i g a n , 96: 2 (as g e n e r o t y p e of Superlestes

WILLIAMSON, 1 9 2 1 ) .

1984 Archilestes

exoletus

- DAVIES & TOBIN, T h e Dragonflies of the World, U t r e c h t , 1: 30.

Distribution: Brazil. grandis

(RAMBUR)

1842 Lestes grandis RAMBUR, Hist. n a t . Inst. N e u r o p t . , Paris: 244. Type male: M a r c h a l ' s Collection; type-locality: C o l o m b i a . 1861 Lestes grandis - HAGEN, Sys. N e u r . N . A m e r . , W a s h i n g t o n : 66 (Mexico, C o l o m b i a , Venezuela). 1862 Lestes (Archilestes) grandis - SELYS, Synopsis des Agrionines, 2: 10 (as g e n e r o t y p e of Archilestes SELYS, 1862). 1890 Archilestes grandis - KIRBY, Syn. C a t . N e u r . - O d o n . , L o n d o n : 159. 1916 Archilestes grandis - Ris, A r c h . N a t u r g . , (A) 82: 61 (Mexico, C o s t a Rica, C o l o m b i a , Texas).

Distribution: North-central America. guayaraca

D E MARMELS

1 9 8 4 A r c h i l e s t e s g u a y a r a c a D E MARMELS, 1 9 8 2 - l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e

World, U t r e c h t , 1: 30.

Distribution: Venezuela. latialatus

DONNELLY

1 9 8 4 A r c h i l e s t e s l a t i a l a t u s DONNELLY, 1 9 8 1 - l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f

World, U t r e c h t , 1: 30.

Distribution: Central America.

the

109 neblina

GARRISON

1982 Archilestes neblina GARRISON, OCC. Pap. Mus. Zool. Univ. Mich., 702: 2. Type male: Agassiz Mus., Cambridge (Mass.); type-locality: Costa Rica.

Distribution: Costa Rica, regalis

GLOYD

1944 Archilestes regalis GLOYD, OCC. Pap. Mus. Zool. Univ. Michigan, 483: 1. Type: unknown; type-locality: Mexico.

Distribution: Mexico. tuberculatus

(WILLIAMSON)

1921 Cyptolestes tuberculatus WILLIAMSON, OCC. Pap. Mus., Univ. Michigan, 96: 6. Type male: Williamson's Collection; type-locality: Venezuela (as generotype of Cyptolestes WILLIAMSON, 1921).

1984 Archilestes (Cyptole' es) tuberculatus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 30.

Distribution: Venezuela. Genus Lestes

LEACH

1815 Lestes LEACH, in BREWSTER'S Edinb. Encycl., 9 (1): 137. Type species: Agrion barbara FABRICIUS, 1798. - 1832 Puella BRÜLLE, Expl. Morae, 2 (1): 104 with species: Libellula puella LINNAEUS, 1758. -1840 Agrion (Anapetes) CHARPENTIER, Libell. europ. Lipsiae: 18. Type species:not given, first emandation: Agrion forcipula CHARPENTIER, 1825. - 1862 Lestes (Lestes) - SELYS, Synopsis des Agrionines, 2: 11 (with subgenera Megalestes, Archilestes, Lestes s.str., Synpycna, Platylestes). - 1850 Lestes - SELYS & HÄGEN, Revue des Odonates, Paris: 146 (revision). - 1890 Puella - KIRBY, Syn. Cat. Neur.-Odon., London: 160 (proposed synonymy with Lestes LEACH, 1815 - without generotype). - 1905 Lestes - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 807 (key to species) [with generotype Lestes sponsa (HANSEMANN, 1823], - 1918 Lestes - Ris, Arch. Naturg., (A) 82: 61. - 1920 Africalestes KENNEDY, Ohio J. Sei. Columbus, 21: 84. Type species: Agrion virgatum BURMEISTER, 1839 (original designation). 1920 Chalcolestes KENNEDY, Ohio J. Sei. Columbus, 21: 84. Type species: Agrion viridis VAN DER LINDEN, 1825 (original designation). - 1932 Lestes - LIEFTINCK, Nova Guinea, 15 (5): 7 (revision, key to New Guinean species). - 1934 Puella - COWLEY, Entomologist, 67: 204 (type Agrion barbara FABRICIUS, 1798). - 1934 Anapetes - COWLEY, Entomologist, 67: 250 type Agrion forcipula CHARPENTIER, 1825). - 1948 Puella - AKRAMOWSKI, Zool. Zborn., Moscow, 5: 153. - 1950 Lestes - MORERA, Inst. Esp. Ent. Madrid: 74 (key to Spanish species). - 1951 Lestes - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 121 (key to Malgasy species). - 1951 Paralestes SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 124. Type species: Lestes (Lestes) praemorsus HÄGEN, 1862 (original designation). - 1951 Xerolestes FRASER, Ent. News. 62: 69. Type species: Lestes pallidus RAMBUR, 1842 (nec NAVÄS, 1924) (original designation). - 1954 Lestes - LIEFTINCK, Treubia, 22 Suppl.: 25. - 1954 Anapetes - LIEFTINCK, Treubia, 22 Suppl.: 25 (proposed synonymy with Lestes LEACH, 1815). - 1954 Paralestes - LIEFTINCK, Treubia, 22 Suppl.: 25 (proposed synonymy with Lestes LEACH, 1815). - 1956 Lestes - FRÄSER, R. Ent. Soc. London, 1 (10): 33 (key to British species). - 1962 Lestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 93 (revision). - 1962 Africalestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 93 (proposed synonymy with Lestes LEACH, 1815). - 1962 Lestes (Chalcolestes) - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94. - 1962 Lestes (Paralestes) PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94. - 1962 Lestes (Xerolestes) - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. - 1984 Lestes - STEINMANN, Fauna Hung., 160: 40 (key to Hungarian species). - 1984 Lestes - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 31. - 1984 Lestes (Icterolestes) PINHEY, 1980. Type species: Lestes intericus GERSTAECKER, 1869 (original designation) - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht,

110 1 : 3 1 . - 1 9 8 4 Lestes (Malgassolestes) PINHEY, 1 9 8 0 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 1 . - 1 9 8 4 Lestes (Pseudochalcolestes) PINHEY, 1 9 8 0 . Type species: Lestes auripennis FRASER, 1 9 5 5 (original designation) - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 31.

alacer

HÄGEN

1861 Lestes alacer HAGEN, Syn. Neur. N. Amer., Washington: 67. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Texas. 1862 Lestes (Lestes) alacer - SELYS, Synopsis des Agrionines, 2: 20. 1890 Lestes alacer - KIRBY, Syn. Cat. Neur.-Odon., London: 161. 1910 Lestes alacer - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 36.

Distribution: Central America. albofasciatus

FRASER

1943 Lestes albofasciatus FRASER, Proc. R. ent. Soc. London, (B) 12: 113. Type male: British Museum (Nat. Hist.), London; type-locality: Buru. 1984 Lestes quercifolia f. albofasciatus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 1 .

Distribution: Buru. amicus

MARTIN

1910 Lestes amicus MARTIN, Ann. Soc. ent. France, 79: 91. Type male: unknown; type-locality: Katanga. 1921 Lestes amicus - Ris, Annls S. Afr. Mus., 18: 275 (Rhodesia). 1962 Lestes (Lestes) amicus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 93 (Rhodesia, Angola, Belg. Congo, Tanganyika).

Distribution: Congo Republic, Tanzania, Angola, Rhodesia. angularis

FRASER

1929 Lestes angularis FRASER, J. Bombay nat. Hist. Soc., 33: 848. Type male: Pusa Collection; type-locality: India.

Distribution: India. apollinaris

NAVÄS

1934 Lestes apollinaris NAVÄS, Broteria, 30 (3): 136. Type male: Barcelona Museum; type-locality: Colombia.

Distribution: Colombia. auripennis

FRASER

1955 Lestes auripennis FRASER, Nat. malgache, 7: 39. Type male: Inst. Rech. Sei. Madagascar; type-locality: Madagascar. 1962 Lestes (Lestes) auripennis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 93. 1 9 8 4 Lestes (Pseudochalcolestes) auripennis - PINHEY (as generotype of Pseudochalcolestes PINHEY, 1980) - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 31.

Distribution: Madagascar. auritus

HÄGEN

1862 Lestes (Lestes) aurita HAGEN, in SELYS, Synopsis des Agrionines, 2: 28. Type male: Mus. Naturk., Berlin; type-locality: Brazil.

Ill 1890 Lestes auritus - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: Brazil, barbarus

(FABRICIUS)

1798 Agrion barbara FABRICIUS, Suppl. Ent. Syst.: 286. Type male: unknown; type-locality: North Africa (Barbary). 1815 Lestes barbarus - LEACH, in Brewster's Edinb. Encycl., 9 (1): 137 (as generotype of Lestes LEACH, 1 8 1 5 ) .

1823 Agrion nympha HANSEMANN (nec SELYS, 1840, and STEPHENS, 1836), Widern. Zool. Mag., 2 (1): 161 (emendation). 1825 Lestes barbarus - VAN DER LINDEN, Mon. Lib. Europ.: 36. 1862 Lestes (Lestes) barbara - SELYS, Synopsis des Agrionines, 2: 34 (Mediterranean territory, Asia Minor, Algeria). 1887 Lestes barbara - SELYS, Ann. Soc. ent. Belg., 31: 43 (Europe, Algeria, Egypt, Mesopotamia, Turkestan). 1890 Lestes barbarus - KIRBY, Syn. Cat. Neur.-Odon., London: 162. 1928 Lestes barbarus - ANDRES, Mem. Soc. R. ent. Egypt, 3 (1): 21 (Egypt). 1940 Lestes barbara - NIELSEN, Mem. Soc. ent. ital., 19: 237 (Sardinia). 1948 Lestes barbara - AKRAMOVSKI, Zool. Zbom., Moscow, 5: 147 (Armenia). 1950 Lestes barbarus - MORERA, Inst. Esp. Ent., Madrid: 76 (Spain). 1960 Lestes barbarus - HOFFMANN, Archives, 27: 222 (Central Europe, Mediterranean 1961 Lestes barbara - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 37 (Kazakhstan). 1984 Lestes barbarus - STEINMANN, Fauna Hung., 160: 42 (from Kashmir to Central Europe).

Distribution: Central and South Europe, West Asia to Mongolia, and India. bipupillatus

CALVERT

1909 Lestes bipupillatus CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 95. Type male: unknown; type-locality: Brazil.

Distribution: Brazil, boninensis

(ASAHINA)

1952 Austrolestes boninensis ASAHINA, Mushi, 23:49. Type male: Asahina's Collection; type-locality: Bonin Island. 1962 Lestes boninensis - LIEFTINCK, Insects of Micronesia, 5 (1): 12 (endemic).

Distribution: Bonin Island. chromatus

MARTIN

1910 Lestes chromatus MARTIN, Ann. Soc. ent. France, 79: 94. Type male: ? Mus. Nation. Hist. Nat., Paris; type-locality: Senegal. 1962 Lestes (Xerolestes) chromatus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1984 Lestes pallidus f. chromatus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33.

Distribution: Senegal. cineraceus

MARTIN

1910 Lestes cineraceus MARTIN, Ann. Soc. ent. France, 79: 93. Type male: unknown; type-locality: Senegal. 1962 Lestes (Xerolestes) cineraceus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1984 Lestes pallidus f. cineraceus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33.

Distribution: Senegal.

112 concinnus

HAGEN

1862 Lestes (Lestes) concinna HAGEN, in SELYS, Synopsis des Agrionines, 2: 37. Type male: Mus. Naturk., Berlin; type-locality: Batavia. 1862 Lestes (Lestes) amata HAGEN, in SELYS, Synopsis des Agrionines, 2: 37. Type male: Mus. Naturk., Berlin; type-locality: Java. 1890 Lestes concinnus - KIRBY, Syn. Cat. Neur.-Odon., London: 162 (China, Java). 1906 Lestes paludosus TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 181. Type male: Tillyard's Collection; type-locality: Queensland. 1913 Austrolestes paludosus - TILLYARD, Proc. Linn. Soc. N.S. Wales, 7: 427. 1917 Lestes paludosus - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 26 (redescription). 1930 Lestes concinna - NEEDHAM, Man. Dragonflies of China: 235 (China). 1934 Lestes concinna - LIEFTINCK, Treubia, 14 (14): 387 (Java). 1936 Lestes concinnus - LIEFTINCK, Revue suisse Zool., 43 (5): 116 (Sumba). 1939 Lestes concinna - NEEDHAM & GYGER, Philipp. J. Sei. 70 (3): 261 (Philippines). 1953 Lestes concinna - LIEFTINCK, Verh. Naturf. Ges. Basel, 64: 139 (Flores, Sumbawa). 1954 Lestes concinna - LIEFTINCK, Treubia, 22 Suppl.: 25. 1954 Lestes (Lestes) amata - LIEFTINCK, Treubia, 22 Suppl.: 25 (proposed synonymy with concinnus HÄGEN, 1862). 1960 Lestes concinnus - LIEFTINCK, Mem. Soc. ent. ital., 38: 232 (redescription). 1962 Lestes (Xerolestes) concinna - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1962 Lestes paludosus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95 (proposed synonymy with concinnus H Ä G E N , 1 8 6 2 ) .

Distribution: from China to Queensland, congener

HÄGEN

1861 Lestes congener HÄGEN, Syn. Neur. N. Amer., Washington: 67. Type male: Mus. comp. Zool., Cambridge (Mass.); type-locality: New York. 1862 Lestes (Lestes) congener - SELYS, Synopsis des Agrionines, 2: 32 (Delaware, New York, Texas). 1890 Lestes congener - KIRBY, Syn. Cat. Neur.-Odon., London: 162. 1946 Lestes congener - LA RIVERS, Ent. News, 57: 217 (Nevada).

Distribution: U.S.A. dajakanus

LIEFTINCK

1948 Lestes dajakanus LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 3. Type male: Naturhist. Riksmus., Stockholm; type-locality: Borneo. 1954 Lestes dajakanus - LIEFTINCK, Treubia, 22 Suppl.: 25.

Distribution: Borneo. dichrostigma

CALVERT

1909 Lestes dichrostigma CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 102. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil. disarmatus

FRASER

1961 Lestes disarmatus FRASER, in PINHEY, Publ. Brit. Mus. (Nat. Hist.) Ent., 11: 221. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1962 Lestes (Xerolestes) disarmatus - PINHEY, Publ. cult. Comp. Diam. Angola., 59: 95.

Distribution: Uganda.

113 disjunctus disjunctus

SELYS

1862 Lestes (Lestes) disjuncta SELYS, Synopsis des Agrionines, 2: 18. Type male: British Museum (Nat. Hist.), London; type-locality: Amer. merid. 1890 Lestes disjunctus - KIRBY, Syn. Cat. Neur.-Odon., London: 161 (United States). 1952 Lestes disjunctus disjunctus - WALKER, Trans. Am. ent. Soc., 78: 64.

Distribution: U.S.A. disjunctus australis

WALKER

1952 Lestes disjunctus australis WALKER, Trans. Am. ent. Soc., 78: 64. Type male: Walker's Collection; type-locality: U.S.A..

Distribution: U.S.A. dissimulans

FRASER

1955 Lestes dissimulans FRASER, Revue Zool. Bot. afr., 51: 38. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1962 Lestes (Paralestes) dissimulans - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94.

Distribution: Congo Republic. dorothea

FRASER

1924 Lestes dorothea FRASER, Ree. Indian Mus., 26: 484. Type male: British Museum (Nat. Hist.), London; type-locality: South India.

Distribution: South India. dryas 1840 1861

1862 1862 1890 1890

1935 1935

1948 1950 1956 1958 1960 1960

1961 1962 1984

KIRBY

Lestes nympha SELYS (nec HANSEMANN, 1 8 2 3 , and STEPHENS, 1 8 3 6 ) , Monogr. Libell. europ., Paris: 141 (emandation). Lestes forcipata H A G E N , (nec RAMBUR, 1 8 4 2 ) , Syn. Neur. N.Amer., Washington: 7 1 Type male: Hagen's Collection; type-locality: Chicago. Lestes (Lestes) hamata SELYS (nec H Ä G E N , 1861), Synopsis des Agrionines, 2: 16. Type male: Selys's Collection; type-locality: Chicago. Lestes (Lestes) nympha - SELYS, Synopsis des Agrionines, 2: 17 (Europe). Lestes uncatus K I R B Y , Syn. Cat. Neur.-Odon., London: 1 6 0 . (new name for forcipata H Ä G E N , 1861). Lestes dryas K I R B Y , Syn. Cat. Neur.-Odon., London: 1 6 0 (new name for Lestes nympha SELYS, 1 8 4 0 , a junior secondary homonymy of Lestes nympha STEPHENS, 1 8 3 6 ) . Lestes dryas - COWLEY, Entomologist, 68: 155. Lestes uncatus - C O W L E Y , Entomologist, 6 8 : 1 5 5 (proposed synonymy with drys K I R B Y , 1 8 9 0 ) . Lestes dryas - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 148 (Armenia). Lestes dryas - MORERA, Inst. Esp. Ent., Madrid: 78 (Spain). Lestes dryas - FRASER, R. Ent. Soc. London, 1 (10): 33 (England). Lestes dryas - ASAHINA, Insecta matsum., 22: 64 (Kurile Island). Lestes dryas - HOFFMANN, Archives, 27: 222 (Germany, Belgium, France, Holland, Luxemburgh). Lestes uncatus m o r p h a skaloni BELYSHEV & G A G I N A , Fragm. faun. Warszawa, 8 : 1 7 3 . Male from Lake Baikal region). Lestes dryas - BELYSHEV, Fragm. faun. Warszawa, 9 (4): 37 (Kazakhstan). Lestes dryas - BENEDEK, Reichenbachia, 11 (17): 185 (Mongolia). Lestes dryas - STEINMANN, F a u n a Hung., 160: 41 (Hungary).

Distribution: Holarctic species, found in North America as well as in Europe and Asia.

114 elatus

HAGEN

1862 Lestes (Lestes) elata H A G E N , in SELYS, Synopsis des Agrionines, 2: 35. Type male: Hagen's Collection; type-locality: Ceylon. 1 8 9 0 Lestes elatus - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 6 2 (India, Ceylon). 1 9 2 4 Lestes elata - FRÄSER, Ree. Indian Mus., 26: 4 8 4 (West India). 1 9 5 1 Paralestes elatus - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 1 2 4 . 1 9 5 5 Lestes elata - LIEFTINCK, Zool. Meded., Leiden, 34 ( 5 ) : 6 9 (Ceylon). 1 9 8 4 Lestes (Paralestes) elatus - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 2 .

Distribution: Sri Lanka, India. eurinus SAY 1839 Lestes eurinus SAY, J1 Acad. nat. Sei. Philad., 8: 36. Type male: Boston Museum; type-locality: U.S.A.. 1 8 6 1 Lestes eurina - H A G E N , Syn. Neur. N.Amer., Washington: 7 0 (Massachusetts). 1 8 6 2 Lestes (Lestes) eurina - SELYS, Synopsis des Agrionines, 2: 3 2 . 1 8 9 0 Lestes eurina - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 6 2 . 1 8 9 1 Lestes eurinus - SCUDDER, Psyche, 6: 6 6 (redescription).

Distribution: falcifer 1917

U.S.A.

SJÖSTEDT

Lestes falcifer SJÖSTEDT, Ark. Zool., Uppsala, 11 Stockholm; type-locality: Amazonas.

(15): 2.

Type male: Naturhist. Riksmus.,

Distribution: Brazil. forcipatus

RAMBUR

1842 Lestes forcipatus RAMBUR (nec H Ä G E N , 1861), Hist. nat. Ins. Neuropt., Paris: 246. Type male: Serville's Collection; type-locality: Amer. sept. 1861 Lestes hamata H Ä G E N (nec SELYS, 1862), Syn. Neur. N. Amer., Washington: 70. Type: sex not given; type-locality: New Jersey. 1862 Lestes (Lestes) forcipata - SELYS, Synopsis des Agrionines, 2: 19 (New Jersey, Georgia, Illinois). 1890 Lestes forcipatus - K I R B Y , Syn. Cat. Neur.-Odon., London: 161.

Distribution: forficula

U.S.A.

RAMBUR

1842 Lestes (Lestes) forficula RAMBUR, Hist. nat. Ins. Neuropt., Paris: 247. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: unknown. 1890 Lestes forficula - K I R B Y , Syn. Cat. Neur.-Odon., London: 161 (Mexico to Brazil). 1916 Lestes forficula - Ris, Arch. Naturg., (A) 82: 62 (Antilles, Brazil). 1919 Lestes forficula - CALVERT, Trans. Am. ent. Soc., 45: 346 (Cuba).

Distribution: Central America, Brazil. helix Ris 1916 Lestes helix Ris, Arch. Naturg., (A) 82: 63. Type male: DEI, Eberswalde; type-locality: Peru. 1943 Lestes helix - SCHMIDT, Beitr. Fauna Perus, 2: 226 (redescription). Distribution: Peru.

115 henshawi

CALVERT

1906 Lestes henshawi CALVERT, Biol. Centr.-Amer., Neur.: 350. Type male: Calvert's Collection; type-locality: Costa Rica. 1916 Lestes henshawi - Ris, Arch. Naturg., (A) 82: 61 (Colombia).

Distribution: Costa Rica, Colombia. ictericus

GERSTAECKER

1869 Lestes ictericus GERSTAECKER, Arch. Naturgesch., 35: 222. Type male: Peringuey's Collection; type-locality: Mombas. 1890 Lestes ictericus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1910 Lestes jacobi MARTIN, Ann. Soc. ent. France, 79: 93. Type male: unknown, perhaps in Mus. Nation. Hist. Nat., Paris; type-locality: Belgian Congo. 1921 Lestes ictericus - Ris, Annls S. Afr. Mus., 18: 270 (South Africa). 1951 Lestes geminata FRASER, Bull. Inst, franc. Afr. noire, 13: 1093. Type female: Ifan (ad Paris Museum); type-locality: Senegal. 1962 Lestes (Xerolestes) geminatus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 96. 1962 Lestes (Xerolestes) pallidus f. ictericus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1962 Lestes (Xerolestes) jacobi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95 (Congo). 1 9 8 4 Lestes (Icterolestes) icterius (as generotype of Icterolestes PINHEY, 1 9 8 4 ) - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 31.

Distribution: West Africa, inaequalis

WALSH

1862 Lestes inaequalis WALSH, Proc. Acad. nat. Sei. Philad., 1862: 385. Types: destroyed; type-locality: Illinois.

Distribution: North America, japonicus

SELYS

1883 Lestes (Lestes) japonica SELYS, Ann. Soc. ent. Belg., 27: 130. Type male: R. Mus., Brussels; type-locality: Japan. 1890 Lestes japonicus - KIRBY, Syn. Cat. Neur.-Odon., London: 162.

Distribution: Japan. macrostigma (EVERSMANN) 1836 Agrion macrostigma EVERSMANN, Bull. Mose., 9: 246. Type male: unknown; type-locality: Russia: Chkalov. 1840 Lestes picteti GENE, in SELYS (nec FONSCOLOMBE, 1838), M o n o g r . Libell. E u r o p . , Paris: 138.

Type male: Mus. Naturk., Berlin; type-locality: Portugal. 1842 Lestes macrostigma - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 249. 1 8 5 0 Lestes macrostigma - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 5 0 (Russia, Sicily, Sardegna). 1 8 5 0 Lestes picteti - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 5 0 [proposed synonymy with macrostigma

(EVERSMANN, 1 8 3 6 ) ] .

1862 Lestes (Lestes) macrostigma -SELYS, Synopsis des Agrionines, 2: 12 (Spain, Hungary, Sicily, Sardinia, Cyprus, South European Russia, Asia Minor). 1 9 0 5 Lestes macrostigma - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 7 (Europa, Sicilia). 1940 Lestes macrostigma - NIELSEN, Mem. Soc. ent. ital., 19: 238 (Sardinia). 1948 Lestes macrostigma - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 149 (Armenia). 1950 Lestes macrostigma - MORERA, Inst. Esp. Ent., Madrid, 76 (Spain). 1961 Lestes macrostigma - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 36 (Kazakhstan).

116 1984 Lestes macrostigma

- STEINMANN, F a u n a Hung., 160: 44 (Hungary).

Distribution: Western Palaearctic region. malabarica

FRASER

1929 Lestes malabarica FRÄSER, J. Bombay nat. Hist. Soc., 33: 847. Type male: British Museum (Nat. Hist.), London: type-locality: India. 1931 Lestes malabarica - FRÄSER, Ree. Indian Mus., 33: 463.

Distribution: India. malaisei

SCHMIDT

1964 Lestes malaisei SCHMIDT, Ent. Tidskr., 85:146. Type male: Schmidt's Collection; type-locality: Burma.

Distribution: Burma, mediorufus

CALVERT

1909 Lestes mediorufus CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 97. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil, minutus

SELYS

1862 Lestes (Lestes) minutus SELYS, Synopsis des Agrionines, 2: 21. Type male: Selys's Collection;

type-locality: Brazil. 1890 Lestes minutus - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: Brazil. nigriceps

FRASER

1924 Lestes nigriceps FRASER, Mem. Dept. Agr. India, 8: 85. Type male: British Museum (Nat. Hist.), London; type-locality: Pusa.

Distribution: India. nodalis

SELYS

1892 Lestes nodalis SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 496. Type male: Mus. civ. Stor. nat. Genova; type-locality: Burma. 1960 Lestes nodalis - LIEFTINCK, Mem. Soc. ent. ital., 38: 232 (redescription).

Distribution: Burma. ochraceus ochraceus

SELYS

1862 Lestes (Lestes) ochracea SELYS, Synopsis des Agrionines, 2: 41. Type male: not given; typelocality: Africa (Cap?). 1890 Lestes ochraceus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1921 Lestes ochraceus - Ris, Annls S. Afr. Mus., 18: 271 (South Africa). 1958 Lestes pallidus f. ochraceus - PINHEY, Stuttg. Beitr. Naturk., 1958 (10): 1 (tropical and subtropical Africa). 1962 Lestes (Xerolestes) pallidus f. ochraceus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1 9 8 4 Lestes ochraceus ochraceus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 3 .

Distribution: South Africa.

117 ochraceus aldabrensis BLACKMAN & PINHEY 1984 Lestes ochraceus aldabrensis BLACKMAN & PINHEY, 1967 - loc. cit. DA VIES & TOBIN, T h e D r a -

gonflies of the World, Utrecht, 1: 31.

Distribution: Seychelles: Aldabra Island. ochraceus unicolor MCLACHLAN 1895 Lestes unicolor MCLACHLAN, Ann. Mag. nat. Hist., 16: 27. Type male: British Museum (Nat. Hist.), London; type-locality: Madagascar. 1917 Lestes unicolor - SJÖSTEDT, Ark. Zool., Uppsala, 11 (13): 10 (Madagascra). 1962 Lestes (Xerolestes) unicolor - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 96. 1984 Lestes ochraceus unicolor - DA VIES & TOBIN. The Dragonflies of the World, Utrecht, 1: 33.

Distribution: Madagascar. pallidus RAMBUR 1842 Lestes pallida RAMBUR (nec NAVAS, 1924), Hist. nat. Ins. Neuropt., Paris: 252. Type female: Mus. Nation. Hist. Nat., Paris; type-locality: Cape of Good Hope. 1862 Lestes (Lestes) pallida - SELYS, Synopsis des Agrionines, 2: 41. 1890 Lestes pallidus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1951 Xerolestes pallidus - FRASER, Ent. News, 62: 69 (as generotype of Xerolestes FRASER, 1951). 1962 Lestes (Xerolestes) pallidus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1984 Lestes pallidus - RAYAH & ZUBEIR, Entomologist's month. Mag., 120: 157 (Sudan).

Distribution: tropical and subtropical Africa. patricius FRASER 1924 Lestes patricia FRASER, Ree. Indian Mus., 26: 486. Type male: British Museum (Nat. Hist.), London; type-locality: West India.

Distribution: West India, paulistus CALVERT 1909 Lestes paulistus CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 99. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil. pertinax LIEFTINCK 1932 Lestes pertinax LIEFTINCK, Nova Guinea, 15 (5): 9. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. pictus HÄGEN 1862 Lestes (Lestes) picta HAGEN, in SELYS, Synopsis des Agrionines, 2: 30. Type male: Hagen's Collection; type-locality: Brazil. 1890 Lestes pictus - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: Brazil. pinheyi FRASER 1955 Lestes pinheyi FRASER, Explor. Pare. nat. Upemba, Miss, de Witte, 38 (1): 10. Type male: British Museum (Nat. Hist.) London; type-locality: Congo.

118 1962 Lestes (Paralestes) pinheyi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94 (Rhodesia, Congo, Nigeria).

Distribution: Rhodesia, Angola, Zambia, Congo Republic, Nigeria, plagiatus

(BURMEISTER)

1839 Agrion plagiatum BURMEISTER, Handb. Ent., 2: 824. Type male: Drege's Collection; type-locality: Cape of Good Hope. 1842 Lestes forceps RAMBUR, Hist. nat. Ins. Neuropt., Paris: 245. Type male: Serville's Collection; type-locality: Cap of Good Hope. 1862 Lestes (Lestes) plagiata - SELYS, Synopsis des Agrionines, 2: 40. 1862 Lestes forceps - SELYS, Synopsis des Agrionines, 2: 40 [proposed synonymy with plagiatus (BURMEISTER, 1 8 3 9 ) ] .

1890 Lestes plagiatus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1898 Lestes obscurus KIRBY, Ann. Mag. nat. Hist., (7) 2: 245. Type male: British Museum (Nat. Hist.), London; type-locality: South Africa. 1898 Lestes plagiatus - CALVERT, Trans. Am. ent. Soc., 25: 46. 1921 Lestes plagiatus - Ris, Annls S. Afr. Mus., 18: 276 (Transvaal, Mashonaland, Natal, Cape). 1962 Lestes (Lestes) plagiatus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 93 (Congo, Angola, Nigeria, South Africa). 1962 Lestes obscurus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 93 [proposed synonymy with plagiatus (BURMESITER, 1 8 3 9 ) ] . 1984 Lestes (Paralestes) plagiatus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33. 1962 Lestes tarryi PINHEY, J. ent. Soc. S. Afr., 25: 20. Type male: Pinhey's Collection; type-locality: Nigeria. 1984 Lestes (Paralestes) plagiatus f. tarryi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 3 .

Distribution: tropical and subtropical Africa, praecellens

LIEFTINCK

1937 Lestes praecellens LIEFTINCK, Treubia 16 (1): 59. Type male: Buitenzorg Museum; type-locality: Java. 1939 Lestes praecellens - LIEFTINCK, Treubia, 17: 50. 1954 Lestes praecellens - LIEFTINCK, Treubia, 22 Suppl.: 25 (Java).

Distribution: Java. praemorsus praemorsus

HÄGEN

1862 Lestes (Lestes) praemorsa HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 36. Type female: Mus. Naturk., Berlin; type-locality: Manila. 1878 Lestes praemorsa - SELYS, Mitt. Zool. Mus. Dresden, 3: 317. 1890 Lestes praemorsus - KIRBY, Syn. Cat. Neur.-Odon., London: 162. 1924 Lestes praemorsa andamanensis FRÄSER, Ree. Indian Mus., 26: 410. Type female: British Museum (Nat. Hist.), London; type-locality: Andamans. 1927 Lestes praemorsus - Ris, Zool. Meded., Leiden, 10 (1): 11 (Sumatra). 1930 Lestes praemorsa - NEEDHAM, Man. Dragonflies of China: 235 (China). 1934 Lestes praemorsus praemorsus - LIEFTINCK, Treubia, 14 (4): 388 (Java). 1939 Lestes praemorsa - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 261 (Philippines). 1951 Paralestes praemorsus - SCHMIDT, Mem. Inst. sei. Madagascar, 6A; 124 (as generotype of Paralestes SCHMIDT, 1951). 1954 Lestes praemorsa praemorsa - LIEFTINCK, Treubia, 22: Suppl.: 26.

Distribution: Philippine Islands, China, India, Bangladesh, Laos, Burma, Sri Lanka, Java, Timor.

119 praemorsus decipiens

KIRBY

1894 Lestes decipiens KIRBY, J. Linn. Soc. 24: 565. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1924 Lestes praemorsa decipiens - FRÄSER, Ree. Indian Mus., 26: 486 (West India). 1949 Lestes praemorsus decipiens - LIEFTINCK, Nova Guinea, (N.S.) 5: 33. 1953 Lestes praemorsa decipiens - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 148 (Malaysia, India, Ceylon, A n d a m a n , Burma, Tonkin, Sumba, Timor, Borneo). 1954 Lestes praemorsa decipiens - LIEFTINCK, Treubia, 22 suppl.: 26 (Siam, Malaya, Sumatra, Borneo, Java). 1955 Lestes praemorsa decipiens - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69 (Ceylon).

Distribution: Southeastern Asia. praemorsus sikkimus

FRASER

1929 Lestes praemorsa sikkima FRÄSER, J. Bombay nat. Hist. Soc., 33: 843. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1966 Paralestes praemorsa sikkima - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 213.

Distribution: India. praevius

LIEFTINCK

1940 Lestes praevius LIEFTINCK, Treubia, 17: 344. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Lestes praevius - LIEFTINCK, Treubia, 22: Suppl.: 26 (Borneo). 1984 Lestes praevius - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33 (E. Indies).

Distribution: Borneo. quadristriatus

CALVERT

1909 Lestes quadristriatus CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 104. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil. quercifolius

SELYS

1878 Lestes quercifolia SELYS, Mitt. zool. Mus. Dresden, 3: 318. Type male: Mus. Tierk., Dresden; type-locality: East Asian Is. 1943 Lestes nigrofasciatus FRÄSER, Proc. R. ent. Soc. London, (B) 12: 113. Type male: British Museum (Nat. Hist.), London; type-locality: Burn. 1949 Lestes quercifolia - LIEFTINCK, N o v a Guinea (N.S.), 5: 35. 1949 Lestes nigrofasciatus - LIEFTINCK, Nova Guinea (N.S.), 5: 35 (proposed synonymy with quercifolius

SELYS, 1 8 7 8 ) .

1984 Lestes quercifolius f. nigrofasciatus

-

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 3 2 .

Distribution: East Asian Islands. radiatus

MARTIN

1910 Lestes radiatus MARTIN, Ann. Soc. ent. France, 79: 94. Type male: unknown; type-locality: Abyssinia. 1962 Lestes radiatus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94. 1984 Lestes pallidus f. radiatus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33.

Distribution: Ethiopia, and Sudan.

120 rectangularis SAY 1839 Lestes rectangularis SAY, J1 Acad. nat. Sei. Philad., 8: 34. Type male: Boston Museum; typelocality: Indiana. 1861 Lestes rectangularis - HAGEN, Syn. Neur. N.Amer., Washington: 66 (Chicago, Indiana, Maryland, Pennsylvania, New Jersey, New York, Georgia, Minnesota). 1862 Lestes (Lestes) rectangularis - SELYS, Synopsis des Agrionines, Add. 2: 22. 1890 Lestes rectangularis - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: U.S.A. regulatus

MARTIN

1910 Lestes regulatus MARTIN, Ann. Soc. ent. France, 79: 90. Type male: unknown; type-locality: Abyssinia. 1962 Lestes (Lestes) regularis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94.

Distribution: Ethiopia, and Somalia. scalaris

CALVERT

1909 Lestes scalaris CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 96. Type male: Mus. Comp. Zool., Cambridge (Mass.); type-locality: Cuba. 1919 Lestes scalaris - CALVERT, Trans. Am. ent. Soc., 45: 347 (Cuba) (type in Instituto Havanna). 1 9 8 4 Lestes ,.scalaris G R U N D L A C H , 1 8 8 8 : 2 1 6 " - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33.

Distribution: Cuba. sigma

CALVERT

1901 Lestes sigma CALVERT, Biol. Centr.-Amer. Neur.: 49. Type male: Calvert's Collection; typelocality: Texas-Mexico.

Distribution: Mexico, U.S.A.: Texas. silvaticus

(SCHMIDT)

1951 Chalcolestes silvaticus SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 129. Type: sex not given; Schmidt's Collection; type-locality: Madagascar. 1962 Lestes (Chalcolestes) silvaticus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94. 1984 Lestes (Pseudochalcolestes) silvaticus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 34.

Distribution: Madagascar. simplex

HÄGEN

1861 Lestes simplex HÄGEN, Syn. Neur. N.Amer., Washington: 68. Type male: not given, perhaps in Mus. Naturk., Berlin or Mus. Comp. Zool., Cambridge (Mass.); type-locality: Mexico. 1862 Lestes (Lestes) simplex - SELYS, Synopsis des Agrionines, Add. 2: 14 (Mexico, Texas). 1890 Lestes simplex - KIRBY, Syn. Cat. Neur.-Odon., London: 160.

Distribution: Mexico, U.S.A.: Texas. simulans

MARTIN

1910 Lestes simulans MARTIN, Ann. Soc. ent. France, 79: 88. Type male: unknown, perhaps in Mus. Nation. Hist, nat., Paris; type-locality: East Africa. 1962 Lestes (Paralestes) simulans - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94 (Zambezi).

Distribution: Zambezi.

121 simulatrix

MCLACHLAN

1895 Lestes simulatrix MCLACHLAN, Ann. Mag. nat. Hist., (6) 6: 25. Type male: British Museum (Nat. Hist.), London; type-locality: Madagascar. 1910 Lestes pruinescens MARTIN, Ann. Soc. ent. France, 79: 87. Type male: Mus. Nation. Hist, nat. Paris; type-locality: Madagascar. 1917 Lestes simulatrix - SJÖSTEDT, Ark. Zool., Uppsala, 11 (13): 8 (redescription). 1951 Paratestes simulatrix - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 124. 1951 Lestes pruinescens - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 124 (proposed synonymy with simulatrix M C L A C H L A N , 1 8 9 5 ) . 1952 Lestes (Paralestes) simulatrix - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94. 1 9 8 4 Lestes ( Malgassolestes) pruinescens - DAVIF.S & TOBIN, The Dragonflies of the World, Utrecht, 1: 33 (as good species).

Distribution: Madagascar. smaragdulus

BUCHECKER

1878 Lestes smaragdula BUCHECKER, Syst. Ent., Odonata, 1: 16. Type: not given; type-locality: Zürich.

Distribution: Suisse. somalicus

FÖRSTER

1906 Lestes somalicus FÖRSTER, Jb. Nassau Ver. nat., 59: 339. Type female: Ann Arbor Museum; type-locality: N.O. Afrika. 1962 Lestes (Xerolestes) pallidus f. somalicus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95.

Distribution: Somalia, Kenya, Uganda, Tanzania. spatulus

FRASER

1946 Lestes spatula FRÄSER, Proc. R. ent. Soc. London, (B) 15: 46. Type male: British Museum (Nat. Hist.), London; type-locality: Argentina. 1948 Lestes spatula - FRASER, Acta Zool. lilloana, 5: 63.

Distribution: Argentina. sponsa

(HANSEMANN)

1823 Agrion sponsa HANSEMANN, Wiede. Zool. Mag., 2 (1): 159. Type male: unknown; type-locality: Germany. 1825 Agrion forcipula CHARPENTIER, Horae Ent., Paris: 6. Type male: Halle Museum; type-locality: Europe. 1839 Agrion picteti FONSCOLOMBE (nec GENE, 1840) ( e m e n d a t i o n ) .

1840 Lestes sponsa - SELYS, Monogr. Libell. europ., Paris: 140. 1840 Agrion (Anapetes) forcipula - CHARPENTIER, Libell. Europ. Lipsiae: 24. 1 8 5 0 Lestes sponsa - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 5 4 (England, Holland, Beige, France, Germany, Poland, Hungary, Russia). 1856 Agrion spectrum KOLENATI, Bull. Mose., 29 (4): 501. Type male: unknown; type-locality: Russia. 1862 Lestes (Lestes) sponsa - SELYS, Synopsis des Agrionines, Add. 2: 17. 1862 Agrion forcipula - SELYS, Synopsis des Agrionines, Add. 2:17 [proposed synonymy with sponsa (HANSEMANN, 1823)].

1862 Agrion spectrum - SELYS, Synopsis des Agrionines, Add. 2: 17 [proposed synonymy with sponsa

(HANSEMANN, 1 8 2 3 ) ] .

1890 Lestes sponsa - KIRBY, Syn. Cat. Neur. Odon., London: 160. 1934 Anapetes forcipula - C O W L E Y , Entomologist, 67: 250 (as generotype of Anapetes 1840). 1940 Lestes sponsa - NIELSEN, Mem. Soc. ent. ital., 19: 238 (Sardinia).

CHARPENTIER,

122 1948 1950 1956 1958 1960 1968 1984

Lestes Lestes Lestes Lestes Lestes Lestes Lestes

sponsa sponsa sponsa sponsa sponsa sponsa sponsa

-

AKRAMOVSKI, Zool. Zborn., Moscow, 5: 149 (Armenia). MORERA, Inst. Esp. Ent., Madrid: 77 (Spain). FRÄSER, R. Ent. Soc. London, 1 (10): 33 (England). ASAHINA - Insecta matsum., 22: 65 (Kurille Islands). HOFFMANN, Archives, 27: 223 (France, Belgium, Holland, Germany). BENEDEK, Reichenbachia, 11 (17): 185 (Mongolia). STEINMANN, Fauna Hung., 160: 41 (Hungary).

Distribution: Central and northern Europe, and palearctical Asia. spumarius

HÄGEN

1862 Lestes (Lestes) spumaria HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 25. Type male: Mus. Naturk., Berlin; type-locality: Porto Rico. 1890 Lestes spumarius - KIRBY, Syn. Cat. Neur.-Odon., London: 161. 1910 Lestes spumarius - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 39. 1919 Lestes spumaria - CALVERT, Trans. Am. ent. Soc., 45: 346 (Cuba).

Distribution: Cuba, Puerto Rico. sternalis

NAVÄS

1924 Lestes pallidus NAVÄS (nec RAMBUR, 1842), Mem. R. Acad. Barcelona, 18 (13): 11. Type male: unknown; type-locality: Colombia. 1930 Lestes sternalis NAVÄS, Bol. Soc. ent. Esp., 13: 21 (new name for Lestes pallidus NAVÄS, 1924, a junior secondary homonymy of Lestes pallidus RAMBUR, 1842). 1950 Lestes confusus FRASER, Mem. Inst, franc. Afr. noire, 10: 107 (new name for Lestes pallidus NAVÄS, 1924, a junior secondary homonymy of Lestes pallidus RAMBUR, 1842) syn. nov.

Distribution: Colombia. stigmaticus

NAVÄS

stat. nov.

1924 Lestes stigmaticus NAVÄS, Mem. R. Acad., Barcelona, 18 (13): 9. Type male: ? Barcelona Museum; type-locality: East Africa. 1962 Lestes (Xerolestes) pallidus f. stigmaticus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1984 Lestes pallidus f. stigmatus (misprint) - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 33.

Distribution: East Africa, striatus

SELYS 1862 Lestes (Lestes) striata SELYS (after HAGEN, Amer. sans description), Synopsis des Agrionines, Add. 2: 25. Type: undesignated, from Venezuela. 1890 Lestes striatus - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: Venezuela. stultus

HÄGEN

1861 Lestes stulta HAGEN, Syn. Neur. N.Amer., Washington: 67. Type: sex not given: Mus. comp. Zool. Cambridge (Mass.) - lost ?; type-locality: California. 1862 Lestes (Lestes) stulta - SELYS, Synopsis des Agrionines, Add. 2: 20 (San Francisco). 1890 Lestes stultus - KIRBY, Syn. Cat. Neur.-Odon., London: 161. 1917 Lestes stultus - KENNEDY, Proc. U.S. natn. mus., 52: 485 (California, Nevada) (redescription).

Distribution: U.S.A. sublatus

HÄGEN

1862 Lestes (Lestes) sublata HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 23. Type female: Mus. Naturk., Berlin; type-locality: Suriname.

123 1890 Lestes sublatus - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: Suriname. temporalis

SELYS

1883 Lestes temporalis locality: Japan. 1905 Lestes temporalis 1916 Lestes temporalis 1973 Lestes temporalis

SELYS, Ann. Soc. ent. Belg., 27: 129. Type male: Selys's Collection; type- JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 808. - Ris, Suppl. ent. Berlin, 5: 11. - BELYSHEV, The Dragonflies of Siberia, 1 (2): 511.

Distribution: Japan. tenuatus

RAMBUR

1842 Lestes tenuatus RAMBUR, Hist. nat. Ins. Neuropt., Paris: 245. Type: sex not given; Guerin & Serville's Collection; type-locality: Martinique. 1862 Lestes (Lestes) tenuata - SELYS, Synopsis des Agrionines, Add. 2: 31. (Martinique, Antilles). 1890 Lestes tenuatus - KIRBY, Syn. Cat. Neur.-Odon., London: 162. 1916 Lestes tenuatus - Ris, Arch. Naturg., (A) 82: 62 (Antilles, Ecuador). 1919 Lestes tenuata - CALVERT, Trans. Am. ent. Soc., 45: 346 (Cuba). 1930 Lestes tenuatus - BYERS, Univ. Florida Publ. Biol. Sei., 1 (1): 165 (U.S.A.).

Distribution: Central America, and Ecuador. tikalus

KORMONDY

1959 Lestes tikalus KORMONDY, Ohio J. Sei., 59: 310. Type male: Kormondy's Collection; typelocality: Guatemala.

Distribution: Guatemala. thoracicus

LAIDLAW

1920 Lestes thoracica LAIDLAW, Ree. Indian Mus., 19: 152. Type male: Nation. Coll. zool. Surv India; type-locality: India.

Distribution: India. tricolor

HOFFMANSEGG

1848 Lestes tricolor HOFFMANSEGG, in ERICHSON, Reisen Brit. Guiana, 3: 585. Type male: Hagen's Collection; type-locality: British Guiana. 1862 Lestes (Lestes) tricolor - SELYS, Synopsis des Agrionines, Add. 2: 29. 1890 Lestes tricolor - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: British Guyana. tridens

MCLACHLAN

1895 Lestes tridens MCLACHLAN, Ann. Mag. nat. Hist., (6) 6: 24. Type male: British Museum (Nat. Hist.), London; type-locality: Delagoa Bay. 1910 Lestes tridens - MARTIN, Ann. Soc. ent. France, 79: 85. 1910 Lestes niger MARTIN, Ann. Soc. ent. France, 79: 88. Type male: lost; tyle-locality: Delagoa Bay. 1917 Lestes tridens - SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 24 (Congo)(description of female). 1951 Paralestes tridens - SCHMIDT, Mem. Inst. Sei. Madagascar, 6A: 124. 1962 Lestes (Paralestes) tridens - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94 (Natal to East Africa, Belgian Congo, Nigeria).

Distribution: tropical and subtropical Africa.

124 umbrinus SELYS 1892 Lestes umbrina SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 497. Type male: Mus. civ. Stor. nat., Genova; type-locality: Burma. 1960 Lestes umbrinus - LIEFTINCK, Mem. Soc. ent. ital., 38: 230 (redescription). 1962 Lestes (Xerolestes) umbrina - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95 (incorrect synonymy with pallidus RAMBUR, 1842).

Distribution: Burma. uncifer KARSCH 1899 Lestes uncifer KARSCH, Ent. Nachr., 25 (24): 381. Type male: Mus. Naturk., Berlin; typelocality: S.-E. Africa. 1921 Lestes uncifer - Ris, Annls S. Afr. Mus., 18: 278. 1951 Paratestes unciver - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 124. 1962 Lestes (Paralestes) uncifer - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 94 (Tanganyika, Kenya, Uganda, Nigeria).

Distribution: southeastern Africa. undulatus SAY 1839 Lestes undulata SAY, J1 Acad. nat. Sei. Philad., 8: 35. Type male: Mus. Naturk., Berlin; typelocality: Chile. 1849 Agrion viridivittatum BLANCHARD, IN SAY, Faun. Chile, 6: 118. Type: unknown; type-locality: Chile. 1850 Lestes vittata SELYS & HÄGEN, Revue Odon. Libell. europ., Paris: 331. Type unknown. 1862 Lestes (Lestes) undulata - SELYS, Synopsis des Agrionines, Add. 2: 27 (Chile, Montevideo, Buenos-Aires). 1890 Lestes undulatus - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: Chile, Argentina, Uruguay. unguiculatus HÄGEN 1861 Lestes unguiculata HAGEN, Syn. Neur. N.Amer., Washington: 70. Type male: Comp. zool. Mus., Cambridge (Mass.); type-locality: Chicago. 1862 Lestes (Lestes) unguiculata - SELYS, Synopsis des Agrionines, Add. 2: 15 (Missouri, New Jersey). 1890 Lestes unguiculatus - KIRBY, Syn. Cat. Neur.-Odon., London: 160. 1910 Lestes unguiculatus - MUTTKOWSKI, Bull. Publ. cult. Mus., Milwaukee, 1 (1): 40.

Distribution: U.S.A. urubamba KENNEDY 1942 Lestes urubamba KENNEDY, Rev. Ent. Rio de Janeiro, 13: 282. Type: unknown; type-locality: Peru.

Distribution: Peru, viduus HÄGEN 1861 Lestes vidua HÄGEN, Syn. Neur. N.Amer., Washington: 69. Type: sex not given; Naturhist. Mus., Wien; type-locality: New Orleans. 1862 Lestes (Lestes) vidua - SELYS, Synopsis des Agrionines, Add. 2: 33. 1890 Lestes vidua - KIRBY, Syn. Cat. Neur.-Odon., London: 162.

Distribution: U.S.A.

125 vigilax HAGEN 1862 Lestes (Lestes) vigilax HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 22. Type male: Comp. Zool. Mus., Cambridge (Mass.); type-locality: New Jersey. 1890 Lestes vigilax - KIRBY, Syn. Cat. Neur.-Odon., London: 161.

Distribution: U.S.A. virens virens (CHARPENTIER) 1825 Agrion virens CHARPENTIER, Hor. Ent.: 8. Type: lost; type-locality: Lusitania. 1 8 5 0 Lestes virens - SELYS & H Ä G E N , Revue Odon. Libell. europ., Paris: 1 5 6 (England, Germany, Poland, Russia, Beige, France, Spain, Albania, Italy, Portugal). 1862 Lestes (Lestes) virens - SELYS, Synopsis des Agrionines, Add. 2: 33 (Europe merid., Asia Minor, Algeria). 1862 Lestes (Lestes) sellata HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 34. Type male: Mus. Naturk., Berlin; type-locality: Syria. 1890 Lestes virens - KIRBY, Syn. Cat. Neur.-Odon., London: 162. 1890 Lestes sellatus - KIRBY, Syn. Cat. Neur.-Odon., London: 162 (Egypt, Syria). 1 9 0 5 Lestes sellata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 0 9 (Syria). 1928 Lestes sellatus - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 21 (Egypt). 1933 Lestes virens - ANDERS, Ent. Tidskr., 54: 120 (Sweden). 1938 Lestes virens - SCHMIDT, Sitz. Akad. Wiss., Wien, (1) 147: 138 (Algeria, Spain, Portugal, Asia Minor). 1940 Lestes virens - NIELSEN, Mem. Soc. ent. ital., 19: 238 (Sardinia). 1948 Lestes virens - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 148 (Armenia). 1950 Lestes virens - MORERA, Inst. Esp. Ent., Madrid: 75 (Spain). 1960 Lestes virens - HOFFMANN, Archives, 97: 221 (N. Africa, C. and S. Europe). 1961 Lestes virens virens - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 37.

Distribution: from Armenia to Algeria. virens marikovskii BELYSHEV 1961 Lestes virens marikovskii BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 37. Type male: Alma-Ata Museum ?; type-locality: Kazakhstan.

Distribution: Kazakhstan, virens vestalis RAMBUR 1842 Lestes vestalis RAMBUR, Hist. nat. Ins. Neuropt., Paris: 250. Type: sex and locality not given. 1 9 5 0 Lestes vestalis - M O R E R A , Inst. Esp. Ent., Madrid: 7 5 [proposed synonymy with virens ( C H A R PENTIER, 1 8 2 5 ) ] .

1978 Lestes virens vestalis - DEVAI, Debr. Deri Mus. evk., 1977: 90. 1984 Lestes virens vestalis - STEINMANN, Fauna Hung., 160: 43 (North America, West, Central & South Europe, Syria, Caucaus).

Distribution: North America, Caucasus to Spain, virgatus (BURMEISTER) 1839 Agrion virgatum BURMEISTER, Handb. Ent., 2: 824. Type male: Halle Museum; type-locality: Good Hope Cap. 1862 Lestes (Lestes) virgata - SELYS, Synopsis des Agrionines, Add. 2: 39. 1890 Lestes virgatus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1898 Lestes virgatus - CALVERT, Trans. Am. ent. Soc., 25: 46. 1906 Lestes rothschildi MARTIN, Bull. Mus. Paris, 1906: 508. Type male: unknown; type-locality: East Africa. 1920 Africalestes virgatus - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Africalestes KENNEDY, 1920).

126 1921 Lestes virgatus - Ris, Annls S. Afr. Mus., 18: 274 (Natal, Mashonaland, Bulawayo). 1958 Lestes virgatus - PINHEY, Stuttg. Beitr. Naturk., 1958 (10): 1 (tropical and subtropical Africa). 1962 Lestes (Lestes) virgatus - PINHEY, Publ.-cult. Comp. Diam. Angola, 59: 94 (Congo, Nigeria).

Distribution: tropical and subtropical Africa. viridis viridis

(VAN DER LINDEN)

1825 Agrion viridis VAN DER LINDEN, Mon. Lib. Eur.: 36: Type male: unknown; type-locality: unknown. 1825 Agrion leucopsalis CHARPENTIER, Horae Ent.: 5. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Silesia. 1840 Lestes viridis - SELYS, Monogr. Libell. europ., Paris: 137 (Europe). 1 8 5 0 Lestes viridis - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 4 8 (Beige, France, Germany, Portugal, Italy, Turkey, Algeria, England). 1862 Lestes (Lestes) viridis - SELYS, Synopsis des Agrionines, Add. 2:13 (Eur. merid., Asia Minor, Algeria). 1890 Lestes viridis - KIRBY, Syn. Cat. Neur.-Odon., London: 160. 1890 Agrion leucopsalis - KIRBY, Syn. Cat. Neur.-Odon., London: 160 [proposed synonymy with viridis (VAN DER L I N D E N , 1 8 2 5 ) ] . 1905 Lestes viridis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 807. 1920 Chalcolestes viridis - KENNEDY, Ohio J. Sei. Columbus, 21: 84 (as generotype of Chalcolestes KENNEDY, 1 9 2 0 ) .

1928 1940 1950 1960

Lestes virdis Lestes viridis Lestes viridis Lestes viridis bourg). 1984 Lestes viridis -

ANDRES, Mem. Soc. R. ent. Egypt, 3 (1): 21 (Egypt). NIELSEN, Mem. Soc. ent. ital., 19: 239 (Sardinia). MORERA, Inst. Esp. Ent., Madrid: 77 (Spain). HOFFMANN, Archives, 27: 222 (Germany, Belgium, France, Holland, LuxemSTEINMANN, F a u n a Hung., 160: 44 (Hungary).

Distribution: Southern and Central Europe, Asia Minor, North Africa. viridis parvidens

ARTOBOLEWSKI

1929 Lestes viridis parvidens ARTOBOLEWSKI, Bull. Soc. nat. Crimee, 11: 141. Type male: unknown; type-locality: Crimea. 1938 Lestes viridis parvidens - SCHMIDT, Sitz. Akad. Wiss., Wien (I) 147: 141 (Syria, Lebanon, Cyprus, Kreta).

Distribution: from Crimea to Cyprus. viridulus

RAMBUR

1842 Lestes viridula RAMBUR, Hist. nat. Ins. Neuropt., Paris: 252. Type: sex not given: Marchal's Collection; type-locality: Bombay. 1862 Lestes (Lestes) viridula - SELYS, Synopsis des Agrionines, Add. 2: 38. 1890 Lestes viridulus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1924 Lestes viridula - FRÄSER, Ree. Indian Mus., 26: 487. 1951 Paralestes viridulus - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 124.

Distribution: India. wahlbergi Ris 1921 Lestes wahlbergi Ris, Annls S. Afr. Mus., 18: 272. Type male: Naturh. Riksmus., Stockholm; type-locality: Caffraria. 1962 Lestes (Xerolestes) pallidus f. wahlbergi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 95. 1984 Lestes pallidus f. wahlbergi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht: 1: 33.

Distribution: South Africa.

127 Genus Orolestes

MCLACHLAN

1895 Orolestes MCLACHLAN, Ann. Mag. nat. Hist., (6) 16: 21. Type species: Orolestes selysi MCLACHLAN, 1895 (original designation). - 1905 Orolestes - JACOBSON & BIANKI, Pryam. lozhnos. Ross.imp.: 811. - 1928 Orolestes - CAMPION & LAIDLAW, Proc. Zool. Soc.London, 1928: 135 (key to species). - 1933 Orolestes - FRÄSER, Ree. Indian Mus., 35: 175 (revision, key to Indian species). - 1954 Orolestes - LIEFTINCK, Treubia, 22 Suppl.: 26.

excelsus

FRASER

1933 Orolestes excelsus FRÄSER, Ree. Indian Mus., 35:181. Type male: British Museum (Nat. Hist.), London; type-locality: Siam.

Distribution: Thailand, motis

BAIJAL & AGARWAL

1955 Orolestes motis BAIJAL & AGARWAL, Agra Univ. J. Res., 4 (2): 467. Type male: Zool. Surv. India; type-locality: India.

Distribution: India, octomaculatus

MARTIN

1903 Orolestes octomaculata MARTIN, Bull. Mus. Paris, 8: 506. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Cambodia.

Distribution: Kambodia. selysi MCLACHLAN 1895 Orolestes selysi MCLACHLAN, Ann. Mag. nat. Hist., (6) 16: 22 (as generotype of Orolestes MCLACHLAN, 1895). Type male: British Museum (Nat. Hist.), London; type-locality: Darjeeling. 1913 Megalestes mirabilis MATSUMURA, Insecta matsum., 1: 50. Type male: Hokkaido Univ.; typelocality: Japan. 1950 Orolestes mirabilis - CHEN, Quart. J. Taiwan Mus., 3: 24. 1950 Orolestes koxingai CHEN, Quart. J. Taiwan Mus., 3: 25. Type male: Hokkaido Univ.; typelocality: Taiwan. - syn. nov. 1961 Orolestes selysi - ASAHINA, Insecta matsum., 24: 57. 1961 Megalestes mirabilis - ASAHINA, Insecta matsum., 24: 57 (proposed synonymy with Selysi MCLACHLAN, 1895).

Distribution: Japan to India. wallacei

(KIRBY)

1889 Lestes wallacei KIRBY, Proc. Zool. Soc. London, 1889: 302. Type male: British Museum (Nat. Hist.), London; type-locality: Sarawak. 1898 Lestes udeana KRÜGER, Stettiner ent. Ztg, 59:127. Type male: unknown, perhaps in Warszawa Museum; type-locality: Sumatra. 1902 Lestes ridleyi LAIDLAW, Proc. Zool. Soc. London, 1902: 92. Type: unknown; type-locality: Perak. 1927 Orolestes udeana - RIS, Zool. Meded., Leiden, 10 (1): 11 (redescription). 1928 Lestes ridleyi -CAMPION & LAIDLAW, Proc. Zool. Soc. London, 1928:134 [proposed synonymy with wallacei (KIRBY, 1889)]. 1935 Orolestes wallacei - LIEFTINCK, Misc. Zool. Sum., 92-93: 5. 1935 Lestes udeana - LIEFTINCK, Misc. Zool. Sum., 92-93: 5 [proposed synonymy with wallacei (KIRBY, 1889)].

Distribution: India, Vietnam, Borneo, Malaysia, Sumatra, Java, Borneo.

128

Genus Platylestes SELYS 1862 Lestes (Platylestes) SELYS, Synopsis des Agrionines, Add. 2: 53. Type species: Lestesplatystyla RAMBUR, 1842 (monotyp). - 1890 Platylestes - KIRBY, Syn. Cat. Neur.-Odon., London: 164. - 1905 Platylestes - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 811. - 1954 Platylestes - LIEFTINCK, Treubia, 22 Suppl.: 27.

heterostylus

LIEFTINCK

1932 Platylestes heterostylus LIEFTINCK, Stylops, 1 (2): 248. Type male: Buitenzorg Museum; typelocality: Java. 1934 Platylestes heterostylus - LIEFTINCK, Treubia, 14 (4): 389 (Java). 1954 Platylestes heterostylus - LIEFTINCK, Treubia, 22 Suppl.: 27 (Malaya, Java, Borneo).

Distribution: Malaysia, Java, Borneo. orientalis

BAIJAL & A G A R W A L

1955 Platylestes orientalis BAIJAL & AGARWAL, Agra Univ. J. Res., 4 (2): 469. Type male: Nation. Coli. Zool. Surv. India, Calcutta; type-locality: India.

Distribution: India. platystylus (RAMBUR) 1842 Lestes platystyla RAMBUR, Hist. nat. Ins. Neuropt., Paris: 254. Type female: Serville's Collection; type-locality: India. 1862 Lestes (Platylestes) platystyla - SELYS, Synopsis des Agrionines, Add. 2: 54 (as generotype o f Platylestes

SELYS, 1 8 6 2 ) .

1890 Platylestes platystylus - KIRBY, Syn. Cat. Neur.-Odon., London: 35. 1984 Platylestes platystyla - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 35.

Distribution: India. Genus Sinhalestes

FRASER

1951 Sinhalestes FRASER, Ent. News, 62: 69. Type species: Lestes (Lestes) designation).

orientalis HÄGEN (original

orientalis (HÄGEN) 1862 Lestes (Lestes) orientalis HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 38. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Lestes orientalis - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1951 Sinhalestes orientalis-FRASER, Ent. News, 62: 69 (as generotype of Sinhalestes FRASER, 1951). 1951 Paralestes orientalis - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 124. 1955 Lestes orientalis - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69.

Distribution: Sri Lanka. Subfamily Sympecmatinae Wings usually closed over back at rest. Dc in forewing shorter than in hind wing. Stigma not elongated. 1984 Sympecmatinae auct. - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36 (with genera Austrolestes, Indolestes, Sympecma).

129 Genus Austrolestes TILLYARD 1913

Austrolestes

Proc. Linn. Soc. N.S. Wales, 37: 4 2 1 . Type species: Agrion cingulatum BURMEISTER, 1 8 3 9 (original designation). - 1 9 2 0 Ceylonolestes K E N N E D Y , Ohio J . Sei. Columbus, 21: 8 4 . Type species: Lestes analis RAMBUR, 1 8 4 2 (original designation). - 1 9 3 3 Ceylonolestes - FRASER, Fauna brit. India, Odon., 1: 63 (key to Indian species). - 1960 Austrolestes - LIEFTINCK, Nova Guinea (N.S.) (Zool.), 1: 130. - 1960 Ceylonolestes - LIEFTINCK, Nova Guinea (N.S.) (Zool.), 1: 1 3 0 (proposed synonymy with Austrolestes TILLYARD, 1 9 1 3 ) . T I L L YARD,

aleison WATSON & MOULDS 1984

Austrolestes aleison WATSON & of the World, Utrecht, 1: 36.

MOULDS, 1 9 7 9

- loc. cit.

DAVIES & TOBIN,

The Dragonflies

Distribution: West Australia. analis (RAMBUR) 1842 Lestes analis RAMBUR, Hist. nat. Ins. Neuropt., Paris: 253. Type female: not given; typelocality: Australia. 1862 Lestes (Lestes) analis - SELYS, Synopsis des Agrionines, Add. 2: 49. 1890 Lestes analis - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1901 Lestes analis - MARTIN, Mem. Soc. zool. France, 14: 244 (Australia). 1920 Ceylonolestes analis -KENNEDY, Ohio J. Sei. Columbus, 21: 84 (as generotype of Ceylonolestes KENNEDY, 1920).

1984 Austrolestes analis -

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1:

36.

Distribution: S. & W. Australia. annulosus (SELYS) 1862 Lestes (Lestes) annulosa SELYS, Synopsis des Agrionines, Add. 2: 48. Type male: Selys's Collection; type-locality: Australia. 1890 Lestes annulosus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1901 Lestes annulosus - MARTIN, Mem. Soc. zool. France, 14: 244 (Victoria, South Australia). 1 9 8 4 Austrolestes annulosus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 6 .

Distribution: S. & W. Australia. aridus (TILLYARD) 1907 Lestes aridus TILLYARD, Proc. Linn. Soc N.S. Wales, 32: 762. Type male: not given, perhaps in Tillyard's Collection; type-locality: Central Australia. 1913 Austrolestes aridus - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 427. 1952 Lestes humphriesi LIEFTINCK, Ent. Ber. Austr., 14: 126. Type male: Lieftinck's Collection; type-locality: South Australia. 1 9 8 4 Austrolestes aridus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 6 . 1984 Lestes humphriesi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36 [proposed synonymy with aridus (TILLYARD, 1907)].

Distribution: Central and South Australia. boninensis ASAHINA 1952 Austrolestes boninensis ASAHINA, Mushi, 23 (6): 49. Type male: Asahina's Collection; typelocality: Chicki Jima. 1962 Lestes boninensis - LIEFTINCK, Insects of Micronesia, 5 (1): 12.

Distribution: Bonin Islands.

130 cheesmanae

KIMMINS

1936 Austrolestes cheesmanae KIMMINS, Ann. Mag. nat. Hist., (10) 18: 69. Type male: British Museum (Nat. Hist.), London; type-locality: Banks Island.

Distribution: West Pacific Islands. cingulatus

(BURMEISTER)

1839 Agrion cingulatum BURMEISTER, Handb. Ent., 2: 823. Type male: Halle Museum; type-locality: Australia. 1862 Lestes (Lestes) cingulata - SELYS, Synopsis des Agrionines, Add. 2: 50. 1890 Lestes cingulatus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1897 Lestes cingulatus - CALVERT, Trans. Am. ent. Soc., 25: 45 (redescription of holotype). 1901 Lestes cingulatus - MARTIN, Mem. Soc. zool. France, 14: 244 (Victoria). 1912 Austrolestes cingulatus - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 425 (as generotype of Austrolestes TILLYARD, 1912).

Distribution: Australia. colensonis

(WHITE)

1846 Agrion colensonis WHITE, Voy. Erebus & Terror, Ins.: 6. Type male: British Museum (Nat. Hist.), London; type-locality: New Zealand. 1962 Lestes (Lestes) colensonis - SELYS, Synopsis des Agrionines, Add. 2: 44 (Sydney). 1890 Lestes colensonis - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1 9 8 4 Austrolestes colensonis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 6 .

Distribution: New Zealand, South Australia. insularis

TILLYARD

1913 Austrolestes insularis TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 425. Type male: Tillyard's Collection; type-locality: Banks Island.

Distribution: Banks Island. io (SELYS) 1862 Lestes (Lestes) io SELYS, Synopsis des Agrionines, Add. 2: 46. Type male: Selys's Collection; type-locality: Australia. 1890 Lestes io - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1901 Lestes io - MARTIN, Mem. Soc. zool. France, 14: 244 (Victoria). 1984 Austrolestes io - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36.

Distribution: South Australia. leda

(SELYS)

1862 Lestes (Lestes) leda SELYS, Synopsis des Agrionines, Add. 2: 47., Type male: Mus. Naturk., Berlin; type-locality: Australia. 1890 Lestes leda - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1901 Lestes leda - MARTIN, Mem. Soc. zool. France, 14: 244 (Victoria). 1906 Lestes leda - TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 409. 1917 Lestes leda - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 28 (Queensland). 1984 Austrolestes leda - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36. Distribution: A u s t r a l i a .

131 minjerriba

WATSON

1 9 8 4 A u s t r o l e s t e s m i n j e r r i b a WATSON, 1 9 7 9 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

1: 36.

Distribution: Queensland. psyche

(HÄGEN)

1862 Lestes (Lestes) psyche HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 45. Type male: Hagen's Collection; type-locality: Australia. 1890 Lestes psyche - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1901 Lestes psyche - MARTIN, Mem. Soc. zool. France, 14: 244. 1984 Austrolestes psyche - DA VIES & TOBIN, The Dragonflies of the World, 1: 36.

Distribution: Australia. Genus Indolestes

FRASER

1922 Indolestes FRASER, Mem. Dept. Agric. India, 7: 57. Type species: Indolestes indicus FRASER, 1922 (original designation). - 1924 Ceylanicolestes FRASER, Ree. Indian Mus., 26: 487. Type species: Lestes (Lestes) gracilis HÄGEN, 1862 (first emandation). - 1931 Ceylanicolestes FRASER, R e e . I n d i a n M u s . , 3 3 : 4 5 0 ( p r o p o s e d s y n o n y m y w i t h Ceylanicolestes

KENNEDY, 1 9 2 0 ) .

- 1951 Tricholestes FRASER, Ent. News, 62: 67. Type species: Lestes risi WEELE, 1909 (original designation). - 1957 Esaulestes FRASER, R. Zool. Soc. N.S. Wales, 1957: 56 (new name for Tricholestes FRASER, 1951, a primary junior homonymy of Tricholestes.

albicauda

(MCLACHLAN)

1895 Lestes albicauda MCLACHLAN, Ann. Mag. nat. Hist., (6) 16: 23. Type male: British Museum (Nat. Hist.), London; type-locality: New Guinea. 1984 Indolestes albicaudus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36.

Distribution: New Guinea. alfurus

LIEFTINCK

I960 Lestes (Indolestes) alfurus LIEFTINCK, Nova Guinea (N.S.) (Zool.), 1: 143. Type male: Lieftinck's Collection; type-locality: Halmahera.

Distribution: New Guinea. alleni (TILLYARD) comb. nov. 1912 Austrolestes alleni TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 425. Type male: Tillyard's Collection; type-locality: Cairns. 1 9 8 4 A u s t r o l e s t e s t i n d a l e i TILLYARD, 1 9 2 5 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 37.

Distribution: North Australia. anomalus

FRASER

1946 Indolestes anomala FRASER, Proc. R. ent. Soc. London, (B) 15: 43. Type female: Fraser's Collection; type-locality: Siam. 1966 Indolestes anomala - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 178 (type male: British Museum (Nat. Hist.), London).

Distribution: Thailand.

132 aruanus

(LIEFTINCK)

1951 Lestes aruanus LIEFTINCK, Amer. Mus. Novit., 1488: 8. Type: unknown; type-locality: Aru Island. 1984 Indolestes aruanus - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36.

Distribution: Aru. assamicus

FRASER

1930 Indolestes assamica Fraser, J. Bombay nat. Hist. Soc., 34: 104. Type female: British Museum (Nat. Hist.), London; type-locality: India.

Distribution: India. bellax

(LIEFTINCK)

comb. nov.

1953 Lestes bellax LIEFTINCK, Verh. naturf. Ges. Basel, 64: 145. Type male: Naturh. Mus., Basel; type-locality: Sumba.

Distribution: Sumba. bilineatus (SELYS) comb. nov. 1862 Lestes ? bilineata SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 498. Type female: Mus. civ. Stor. nat., Genova; type-locality: Burma. 1960 Lestes bilineatus - LIEFTINCK, Mem. Soc. ent. ital., 38: 234 (redescription). 1964 Lestes bilineatus - SCHMIDT, Ent. Tidskr., 85: 146.

Distribution: Burma. coeruleus

FRASER

1924 Indolestes coerulea FRASER, J. Asiat. Soc. Bengal, 19: 460. Type female: British Museum (Nat. Hist.), London; type-locality: Yunnan. 1928 Indolestes coerulea - MORTON, Trans, ent. Soc. London, 76: 111 (description of male).

Distribution: China. cyaneus(SELYS) 1862 Lestes (Lestes) cyanea SELYS, Synopsis des Agrionines, Add. 2: 51. Type male: Selys's Collection; type-locality: India. 1890 Lestes cyaneus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1905 Lestes cyanea - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 809. 1920 Lestes cyanea - LAIDLAW, Ree. Indian Mus., 19: 147. 1922 Indolestes helena FRASER, Mem. Dept. Agric. India, 7: 60. Type male: British Museum (Nat. Hist.), London; type-locality: Kurseong. 1924 Indolestes veronica FRASER, Mem. Dept. Agric. India, 8: 85. Type female: British Museum (Nat. Hist.), London; type-locality: India. 1927 Lestes cyanea - LIEFTINCK, Tijdschr. Ent., 70: 86 (Tibet). 1933 Ceylonolestes cyanea - FRASER, Fauna brit. India, Odon., 1: 74. 1933 Indolestes helena - FRASER, Fauna brit. India, Odon., 1: 74 [proposed synonymy with cyaneus (SELYS, 1862)].

1933 Indolestes veronica- FRASER, Fauna brit. India, Odon., 1: 74 [proposed synonymy with cyaneus (SELYS, 1862)].

1955 Lestes manaliensis SINGH, Agra Univ. J. Res., 4: 172. Type male: Zool. Surv. India; typelocality: Himalaya, syn. nov. 1955 Archibasis sushmae BAIJAL, in SINGH, Agra Univ. J. Res., 4 (Suppl.): 747. Type male: Zool. Surv. I n d i a ; type-locality: H i m a l a y a .

133 1989 Archibasis sushmae - HÄMÄLÄINEN, Odonatologica, 18 (1): 17 [proposed synonymy with cyaneus SELYS, 1862)].

Distribution: India, Tibet, Neapel, Bhutan.

divisus

comb. nov.

(HAGEN)

1862 Lestes (Lestes) divisa HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 44. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Lestes divisus - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1892 Lestes divisa - SELYS, Annali Mus. civ. Stor. nat. Genova, 2 (10): 495. 1955 Lestes divisa - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69.

Distribution: Sri Lanka.

extraneus

(NEEDHAM)

1930 Lestes extranea NEEDHAM, Zool. Sinica, (A) 11: 233. Type female: Peking Museum ?; typelocality: China. 1931 Lestes gracilis extraneus - SCHMIDT, Konowia, 10 (3): 178. 1935 Lestes monteili NAVAS, Mus. Heude Notes Ent. chinoise, 2 (5): 90. Type male: not given, perhaps in Heude Museum; type-locality: China. 1956 Ceylonolestes gracilis extraneus - ASAHINA, Ent. Medd., 27 (4-5); 218. 1962 Indolestes extraneus - CHAO, Acta ent. sinica, Suppl., 11: 27.

Distribution: China.

floresianus

LIEFTINCK

I960 Lestes (Indolestes) floresianus LIEFTINCK, Nova Guinea (N.S.) (Zool.), 1:141. Type: unknown; type-locality: Flores.

Distribution: Flores.

goniocerus

LIEFTINCK

I960 Lestes (Indolestes) goniocerus LIEFTINCK, Nova Guinea (N.S.) (Zool.), 1:154. Type: unknown; type-locality: New Guinea.

Distribution: New Guinea.

gracilis gracilis

(HÄGEN)

1862 Lestes (Lestes) gracilis HAGEN, in SELYS, Synopsis des Agrionines, Add. 2: 43. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Lestes gracilis - KIRBY, Syn. Cat. Neur.-Odon., London: 163. 1916 Lestes gracilis gracilis - Ris, Suppl. ent. Berlin, 5: 14 (key to subspecies) (Ceylon). 1924 Ceylonicolestesgracilis - FRÄSER, Ree. Indian Mus., 26: 487 (as generotype of Ceylonicolestes FRASER, 1924).

1955 Lestes gracilis gracilis - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69. 1984 Indolestes gracilis gracilis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 36.

Distribution: Sri Lanka.

134 gracilis birmanus

(SELYS)

1892 Lestes (Lestes) birmana SELYS, Annali Mus. civ. Stor. nat. Genova. 2 (10): 495. Type male: unknown, perhaps in Mus. civ. Stor. nat., Genova; type-locality: Burma. 1916 Lestes gracilis birmanus - Ris, Suppl. ent. Berlin, 5: 14 (South India). 1924 Ceylonicolestes gracilis birmanus - FRÄSER, Ree. Indian Mus., 2 6 : 487 (West India). 1930 Ceylonolestes davenporti FRÄSER, J. Bombay nat. Hist. Soc., 34:96 (new name for Ceylonolestes gracilis birmanus Ris, 1916 - nec SELYS, 1892). 1966 Lestes (Indolestes) davenporti - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 1 8 : 188. 1984 Indolestes gracilis birmanus - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 36.

- Distribution: Burma, India. gracilis peregrinus (Ris) comb. nov. 1916 Lestes gracilis peregrinus Ris, Suppl. ent. Berlin, 5: 15. Type female: Ris's Collection; typelocality: Japan.

Distribution: Japan. indicus

FRASER

1922 Indolestes indica FRASER, Mem. Dept. Agric. India, 7: 58. Type male: British Museum (Nat. Hist.), London; type-locality: Shillong. 1922 Indolestes buddha FRASER, Mem. Dept. Agric. India, 7: 59. Type male: British Museum (Nat. Hist.), London; type-locality: Assam. 1966 Indolestes indica - KIMMINS, Bull. Brit. Mus.nat. Hist. (Ent.), 1 8 : 197.

Distribution: India. inflatus

(FRASER)

1933 Ceylonolestes inflata FRASER, F a u n a brit. India, Odon., 1: 67. Type male: British Museum (Nat. Hist.), London; type-locality: Burma. 1966 Indolestes inflata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 1 8 : 197. 1984 Indolestes inflatus - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 37.

Distribution: Burma. linsleyi

LIEFTINCK

1960 Lestes (Indolestes) linsleyi LIEFTINCK, , Nova Guinea (N.S.) (Zool.), 1: 148. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. lundquisti

(LIEFTINCK)

comb. nov.

1949 Lestes lundquisti LIEFTINCK, N o v a Guinea, (N.S.) (Zool.), 5: 31. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. luxatus

(LIEFTINCK)

comb. nov.

1932 Lestes luxatus LIEFTINCK, N o v a Guinea, Leiden; type-locality: New Guinea. Distribution: N e w G u i n e a .

15

(5): 11. Type male: Rijksmus. Natuurl., Hist.,

135 lygisticerus (LIEFTINCK) comb. nov. 1932 Lestes lygisticerus LIEFTINCK, Nova Guinea, 15 (5): 13. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, neglectus FRÖHLICH 1984

Indolestes neglectus F R Ö H L I C H 1 9 0 3 , - loc. cit. Utrecht. 1: 37 (locality not given).

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: unknown. obiri WATSON 1984 Indolestes obiri Utrecht, 1: 37.

WATSON,

1979 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: North Australia. pulcherrimus (FRASER) 1924 Ceylanicolestes pulcherrima FRÄSER, Ree. Indian Mus., 26: 487. Type male: British Museum (Nat. Hist.), London; type-locality: South India. 1966 Lestes (Indolestes) pulcherrimus - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 210.

Distribution: South India, risi (WEELE) 1909 Lestes risi WEELE, Nova Guinea, 9: 23. Type male: unknown; type-locality: New Guinea. 1951 Tricholestes risi - FRASER, Ent. News, 62: 67 (as generotype of Tricholestes FRASER, 1951). 1984 Indolestes risi (VAN DER WEELE) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 3 7 .

Distribution: New Guinea. sutteri (LIEFTINCK) comb. nov. 1953 Lestes sutteri LIEFTINCK, Verh. naturf. Ges. Basel, 64: 142. Type male: Naturh. Mus., Basel; type-locality: Sumba.

Distribution: Sumba. tenuissimus (TILLYARD) comb. nov. 1906 Lestes tenuissimus TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 179. Type male: Tillyard's Collection; type-locality: Queensland. 1913 Austrolestes tenuissimus - TILL YARD, Proc. Linn. Soc. N.S. Wales, 37: 426.

Distribution: Australia. vitiensis (TILLYARD) 1923 Austrolestes vitiensis TILLYARD, Trans, ent. Soc. London, 1923: 309. Type male: Cawthron Inst. Coll. type-locality: Fiji. 1984 Indolestes vitiensis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 37.

Distribution: Fiji. Genus Sympecma BURMEISTER 1839 Sympecma BURMEISTER, Handb. Ent., 2: 824 undescribed name. - 1940 Agrion (Sympycna) CHARPENTIER, Libell. europ. Lipsiae: 1 9 . Type species: Agrion phallatum CHARPENTIER, 1 8 2 5 (monotypy). - 1840 Sympecma - SELYS, Mon. Lib. eur.: 144. Type species: Agrion fusca VAN

136 (monotype). - 1 9 0 5 Sympycna - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 810 (key to species). - 1954 Sympecma - LONGFIELD, Entomologist's month. Mag., 90: 1 4 5 .

DER L I N D E N , 1 8 2 3

annulata annulata

SELYS

1887 Sympecma annulata SELYS, Ann. Soc. ent. Belg., 31: 43. Type male: R. Mus., Brussels; typelocality: Turkey: Anatolia. 1848 Sympecma annulata - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 145 (Armenia). 1984 Sympecma annulata - STEINMANN, F a u n a Hung., 160: 39 (Caucasus).

Distribution: Caucasus, Kashmir. annulata braueri 1905 Sympycna Inst. Mus. 1948 Sympycna 1961 Sympycna

(JACOBSON & BIANKI)

braueri JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 811. Type male: Zool. Acad. Sei., St. Petersburg; type-locality: Russia. annulata braueri - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 145 (Armenia). braueri - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 38 (Kazakhstan).

Distribution: Russia to Kazakhstan. annulata gobica 1900 1905 1961 1984

Sympycna Sympycna Sympycna Sympecma

FÖRSTER

annulata race gobica FÖRSTER, Wiener ent. Z., 19: 258. Male from Gobi. gobica - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 811. gobica - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 38 (Kazakhstan). annulata gobica - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 37.

Distribution: West and Central Asia. fuscum

(VAN DER LINDEN)

1823 Agrion fuscum VAN DER LINDEN, Opusc. Scient., 4: 102. Type male: unknown; type-locality: Silesia. 1825 Agrion phallatum CHARPENTIER, Hor. Ent., Paris: 10. Type female: Mus. Nation. Hist. Nat., Paris; type-locality: Europe. 1840 Sympecma fusca - SELYS, Monogr. Libell. europ., Paris: 145 (Beige, Germany, France, Italy) (as generotype of Sympecma BURMEISTER, 1839). 1840 Agrion (Sympycna) phallata - CHARPENTIER, Libell. Europ., Lipsiae: 24. 1862 Lestes (Sympycna) fusca - SELYS, Synopsis des Agrionines, Add. 2: 53 (Europe, Asia Minor, North Africa). 1850 Lestes fusca - SELYS & H Ä G E N , Revue des Odonates, Paris: 161 (Europe, Algeria, Egypt). 1898 Agrion phallatum - CALVERT, Trans. Am. ent. Soc., 25: 45 (Hungary, Germany) (description of holotype). 1927 Sympecma aragoniensis NAVÄS, Rev. Acad. Ci. Zaragoza, 10: 89. Type male: Barcelona Museum; type-locality: Spain. 1928 Sympecma fusca - ANDERS, Mem. Soc. R. ent. Egpyt, 3 (1): 21 (Egypt). 1940 Sympecma fusca - NIELSEN, Mem. Soc. ent. ital., 19: 237 (Sardinia). 1948 Sympycna fusca - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 144 (Armenia). 1956 Sympecma fusca - ADAMOVIC, Bull. Mus. Hist. nat. Serbe, (Β) 8: 103 (Serbia). 1960 Sympecma fusca - HOFFMANN, Archives, 27: 221 (Germany, Belgium, France, Holland, Luxembourg). 1961 Sympecma fusca - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 38 (Kazakhstan). 1984 Sympecma fusca - STEINMANN, F a u n a Hung., 160: 38 (Hungary).

Distribution: from Turkestan to South and Central Europe.

137 paedisca paedisca

(EVERSMANN)

1836 Agrion paedisca EVERSMANN, Bull. Mose., 9: 247. Type male: unknwon, perhaps undesignated in Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: South Russia. 1880 Sympycna paedisca - BRAUER, Verh. Zool.-Bot. Ges. Wien., 30: 231. 1930 Sympycna paedisca - FUDAKOWSKI, Fragm. faun. Mus. zool. polon., 1: 193 (Poland). 1944 Sympecma paedisca f. kashmirensis ANDER, Förh. k. Fysion. Sällsk. Lund, 14 (13): 175. Male from Kashmir. 1 9 8 4 Sympecma paedisca - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 8 [proposed synonymy with annulata braueri (JACOBSON & BIANKI, 1 9 0 5 ) ] .

Distribution: from Japan to England. paedisca striata

ST. QUENTIN

1963 Sympecma paedisca striata ST. QUENTIN, Ann. naturh. Mus., Wien, 66:381. Type male: Naturh. Mus., Wien; type-locality: Central Europe. 1 9 8 4 Sympecma paedisca striata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 3 8 [proposed synonymy with annulata braueri (JACOBSON & BIANKI, 1 9 0 5 ) ] .

Distribution: Central Europe.

Family Lestoideidae Small brown and yellow species. Dc a regular rectangle. Reduced IA & CuP; recessed IRiii and Riv; intercalated veins present. Larva with saccoid gills and quadrate labial mask. 1938 Lestoideidae TILLYARD & subfamily Lestoideinae).

FRASER,

Austr. Zool., 9: 157 (in superfamily Lestoidea, and with

Subfamily Lestoideinae 1919 Lestoidinae MUNZ, Mem. Amer. Ent. Soc., 3: 17 (for genus Lestoidea). - 1938 Lestoideinae - TILL YARD & FRASER, Austr. Zool., 9: 157 (with single genus Lestoidea). - 1957 Lestoideinae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 63.

Genus Lestoidea

TILLYARD

1913 Lestoidea TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 428. Type species: Lestoidea conjuncta TILLYARD, 1913 (original designation). - 1919 Lestoidea - MUNZ, Mem. Amer. Ent. Soc., 3: 17. - 1938 Lestoidea - TILLYARD & FRASER, Austr. Zool., 9: 157. - 1957 Lestoidea - FRASER, R. Zool. Soc. N.S. Wales, 1957: 63.

barbarae

WATSON

1984 Lestoidea barbarae Utrecht, 1: 38.

WATSON,

1967 - loc. cit.

DAVIES & TOBIN,

The Dragoflies of the World,

Distribution: Queensland. conjuncta

TILLYARD

1913 Lestoidea conjuncta TILLYARD, Proc. Linn. Soc. N.S. Wales, 37:428 (as generotype of Lestoidea TILLYARD, 1913). Type male: Tillyard's Collection; type-locality: Queensland.

Distribution: Australia.

138 Family Megapodagrionidae Small to large Zygoptera. Usually only 2 antenodals. Stigma usually not braced; IRiii & Riv originate near subnodus; intercalated veins usually present. Nodus one third to one fourth of wing length from base. Proximal end of stigma acute (radius embraces). Dc long with costal side about twice length of lower side. Male genital hamules quadrate. 1892 Megapodagrionidae SELYS, Ann. Soc. ent. Belg., 36: 86. - 1926 Megapodagriidae TILLYARD, Insects of Austr. N. Zeal.: 76 - 1938 Megapodagriidae - TILLYARD & FRASER, Austr. Zool., 9: 159 (with subfamilies Thaumatoneuridae, Megapodagriinae). - 1955 Megapodagriidae FRÄSER, Proc. R. ent. Soc. London, (B) 24: 139 (revision). - 1962 Megapodagriidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 96 (key to subfamilies Megapodagriidae, Coryphagriinae). - 1984 Megapodagriidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 38 (with subfamilies Argiolestinae, Coryphagrioninae, Hypolestinae, Megapodagriinae, Philosininae, Thaumatoneurinae). - 1959 Philogenini, Heteragrionini, Austroargiolestini RACENIS, Acta biol. venezuel., 2: 339-340.

Subfamily Argiolestinae Medium sized. Petiolation to posterior Ac. IRiii arising near subnodus and Riv 1-2 cells proximal to it. Stigma often with forward angle acute. 1957 Argiolestinae FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 60 (with genera Paraphlebia, Rhinagrion, Podopteryx, Podolestes, Argiolestes, Nesolestes, Philogenia, Allolestes, Dimeragrion, Heteropodagrion, Protolestes, Neurolestes, Neuragrion, Trineuragrion, Heteragrion, Mesagrion, Burmargiolestes, Agriomorpha, Podopteryx, Caledarargiolestes, Celebargiolestes). - 1962 Argiolestinae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 97 (proposed synonymy with Megapodagriinae). -1984 A rgiolestinae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 38 (as valid subfamily).

Genus Agnophilogenia

KENNEDY

1941 Agnophilogenia KENNEDY, Ann. ent. Soc. Amer., 34: 490. Type species: Agnophilogenia monotis KENNEDY, 1941 (original designation).

monotis

KENNEDY

1941 Agnophilogenia monotis KENNEDY, Ann. ent. Soc. Amer., 34: 492 (as generotype of Agnophilogenia KENNEDY, 1941). Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador.

Genus Agriomorpha MAY 1933 Agriomorpha MAY, Senckenbergiana, 15: 341. Type species: Agriomorpha fusca MAY, 1933 (original designation).

fusca MAY 1933 Agriomorpha fusca MAY, Senckenbergiana, 15: 342 (as generotype of Agriomorpha MAY, 1933). Type male: Senckenberg Museum: type-locality: China.

139 1965 Agriomorpha fusca - ASAHINA, Kontyü, 33 (4): 497 (Hongkong).

Distribution: China, Hong Kong. Genus Allolestes

SELYS

1869 Allolestes SELYS, Ann. Soc. ent. Belg., 12: 97. Type species: Allolestes maclachlani SELYS, 1869 (monotypy). - 1890 Allolestes-KIRBY, Syn. Cat. Neur.-Odon., London: 125. - 1919 Allolestes - MUNZ, Mem. Am. ent. Soc., 3 (1): 52. - 1962 Allolestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 100.

maclachlani

SELYS

1869 Allolestes maclachlani SELYS, Ann. Soc. ent. Belg., 12: 97 (as generotype of Allolestes SELYS, 1869). Type female: Selys's Collection; type-locality: Seychelles. 1896 Allolestes maclachlani - MARTIN, Mem. Soc. Zool. France, 9: 107 (description of male). 1896 Allolestes nigra MARTIN, Mem. Soc. Zool. France, 9:107. Type male: lost; type-locality: Seychelles. 1962 Allolestes maclachlani - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 100. 1962 Allolestes nigra - PINHEY, Publ. cult. Comp. Diam. Angola, 9: 100 (as valid taxa).

Distribution: Seychelles. Genus Amanipodagrion

PINHEY

1962 Amanipodagrion PINHEY, J . ent. Soc. S.Afr., 25: 22. Type species: Amanipodagrion HEY, 1962 (original designation).

gilliesi 1962

gilliesi P I N -

PINHEY

Amanipodagrion gilliesi PINHEY, J . ent. Soc. S. Afr., 25: 2 2 (as generotype of PINHEY, 1 9 6 2 ) . Type male: Pinhey's Collection; type-locality: Tanganyika.

Amanipodagrion

Distribution: Tanzania. Genus Archeopodagrion

KENNEDY

1939 Archeopodagrion KENNEDY, Ann. ent. Soc. Amer., 32: 32. Type species: Archeopodagrion corne KENNEDY, 1939 (original designation).

bicorne

bi-

KENNEDY

1939 Archeopodagrion bicorne KENNEDY, Ann. ent. Soc. Amer., 32: 33 (as generotype of Archeopodagron KENNEDY, 1939). Type: unknown; type-locality: Ecuador.

Distribution: Ecuador. bilobata

KENNEDY

1946 Archeopodagrion bilobata KENNEDY, Ann. ent. Soc. Amer., 39: 171. Type male: Kennedy's Collection; type-locality: Ecuador. 1984 Archeopodagrion bilobatum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 38.

Distribution: Ecuador. Genus Argiolestes 1862

SELYS

Argiolestes SELYS, Synopsis des Agrionines, Add. 3 : 3 6 . Type species: Agrion australis G U E R I N , 1 8 3 0 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 2 5 ) . - 1 9 0 0 Argiolestes - FÖRSTER, Termeszetr. Füz., Budapest, 23:104. - 1900 Wahnesia FÖRSTER, Termeszetr. Füz., Budapest, 23:105. Type species: Wahnesia kirbyi FÖRSTER, 1900 (original designation). - 1913

140 Argiolestes - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 420 (key to Australian species). 1913 Metagrion CALVERT, Proc. Acad. nat. Sei. Philad., 65: 261. Type species: Argiolestes postnodalis SELYS, 1878 (original designation). - 1925 Archiargiolestes KENNEDY, Bull. Mus. Comp. Zool., 67: 291. Type species: Archiargiolestes pusillissimus KENNEDY, 1925 (original designation). - 1925 Austroargiolestes KENNEDY, Bull. Mus. Comp. Zool., 67: 294. Type species: Argiolestes icteromelas SELYS, 1862 (original designation). - 1926 Risiolestes FRASER, Treubia, 8 ( 3 - 4 ) : 486 (new name new status for Argiolestes (Argiolestes) SELYS, 1862 (partim). Type species: Argiolestes icteromelas SELYS, 1862 (original designation). - 1926 Wahnesia FRASER, Treubia, 8 ( 3 - 4 ) : 486 (proposed synonymy with Argiolestes SELYS, 1862). - 1932 Wahnesia - LIEFTINCK, N o v a Guinea, 1 5 (5): 215 (proposed synonymy with Argiolestes SELYS, 1862). - 1932 Argiolestes - LIEFTINCK, Nova Guinea, 15 (5): 193 (revision, key to species). - 1935 Argiolestes - LIEFTINCK, N o v a Guinea, 17: 211 (revision, key to species). - 1956 Argiolestes - LIEFTINCK, Nova Guinea (N.S.), 7: 64 (revision, key to species). - 1986 Austroargiolestes - THEISCHINGER & O ' F A R E L L , Odonatologica, 1 5 (4): 391 (as good genus).

alfurus

LIEFTINCK

1956 Argiolestes alfurus LIEFTINCK, Nova Guinea (N.S.), 7: 72. Type male: Tillyard's Collection; type-locality: Batjan Island.

Distribution: Batjan Island. alpinus

TILLYARD

1913 Argiolestes alpinus TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 417. Type male: Tillyard's Collection; type-locality: New South Wales.

Distribution: Australia. amabilis

FÖRSTER

1899 Argiolestes amabilis FÖRSTER, Ann. Soc. ent. Belg., 43: 71. Type male: Mus. Naturk., Berlin; type-locality: New South Wales. 1913 Argiolestes amabilis - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 415 (description of male).

Distribution: Australia. amphistylus

LIEFTINCK

1949 Argiolestes amphistylus LIEFTINCK, Nova Guinea (N.S.), 5:41. Type male: Tillyard's Collection; type-locality: New Guinea.

Distribution: New Guinea. annulipes

LIEFTINCK

1956 Argiolestes annulipes LIEFTINCK, N o v a Guinea (N.S.), 7:102. Type male: Tillyard's Collection; type-locality: New Guinea.

Distribution: New Guinea. armeniacus

LIEFTINCK

1956 Argiolestes armeniacus LIEFTINCK, N o v a Guinea (N.S.), 7: 90. Type male: Tillyard's Collection; type-locality: d'Entrecasteaux Island.

Distribution: d'Entrecasteaux Island. aulicus

LIEFTINCK

1949 Argiolestes aulicus LIEFTINCK, Nova Guinea (N.S.), 5: 43. Type male: Lieftinck's Collection; type-locality: New Guinea.

Distribution: N e w G u i n e a .

141 aurentiacus Ris 1898 Argiolestes aurantiaca Ris, Ent. Nachr., 24: 324. Type male: Ris's Collection; type-locality: Bismarck Archipelago. 1919 Argiolestes aurantiacus - CAMPION, Entomologist, 52: 249 (New Britain). 1949 Argiolestes aurantiacus - LIEFTINCK, Treubia, 20 (2): 335.

Distribution: Bismarck Archipelago. aureus

TILL YARD

1906 Argiolestes aureus TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 178. Type male: not given; type-locality: Queensland. 1913 Argiolestes aureus - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 415.

Distribution: Australia. australis

(GUERIN)

1830 Agrion australis GUERIN, Voy. Coq. Zool., (2) 2: 196. Type female: R. Mus. Brussels; typelocality: Offack Island. 1842 Argia australis - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 256. 1862 Argiolestes australis - SELYS, Synopsis des Agrionines, Add. 3: 37. 1890 Argiolestes australis - KIRBY, Syn. Cat. Neur.-Odon., London: 126 (as generotype of Argiolestes

SELYS, 1 8 6 2 ) .

1935 Argiolestes australis - LIEFTINCK, Nova Guinea, 17: 218 (redescription). 1938 Argiolestes australis - LIEFTINCK, Nova Guinea (N.S.), 2: 70 (redescription).

Distribution: Oceania, calcaris

FRÄSER

1958 Argiolestes calcaris FRASER, Proc. R. Zool. Soc. N.S. Wales, Collection; type-locality: New South Wales.

1956-57:

70. Type male: Fraser's

Distribution: Australia. chrysoides

TILLYARD

1914 Argiolestes chrysoides TILLYARD, Proc. Linn. Soc. N.S. Wales, 38: 237. Type male: Tillyard's Collection; type-locality: Queensland. 1917 Argiolestes chrysoides - TILLYARD, Proc. Linn. Soc. N.S. Wales, 42: 475. 1 9 8 4 Argiolestes chrysoides „ T I L L Y A R D , 1 9 1 3 " - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 39.

Distribution: Australia. coartans

LIEFTINCK

1956 Argiolestes coartans LIEFTINCK, Nova Guinea (N.S.), 7: 81. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Waigeu Island.

Distribution: Waigeu. connectens

LIEFTINCK

1956 Argiolestes connectens LIEFTINCK, Nova Guinea (N.S.), 7: 79. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea.

142 convergens

LIEFTINCK

1949 Argiolestes convergens LIEFTINCK, N o v a G u i n e a (N.S.), 5: 47. Type male: R i j k s m u s . N a t u u r l . Hist., Leiden; type-locality: N e w G u i n e a .

Distribution: New Guinea. ephippiatus

LIEFTINCK

1956 Argiolestes ephippiatus LIEFTINCK, N o v a G u i n e a (N.S.), 7: 97. Type male: R i j k s m u s . N a t u u r l . , Hist., Leiden; type-locality: N e w G u i n e a .

Distribution: New Guinea. esuriensis

LIEFTINCK

1956 Argiolestes esuriensis LIEFTINCK, N o v a G u i n e a (N.S.), 7: 93. Type male: R i j k s m u s . N a t u u r l . Hist., Leiden; type-locality: N e w G u i n e a .

Distribution: New Guinea. fontanus

TILL YARD

1913 Argiolestes fontanus TILLYARD, Proc. Linn. Soc. N . S . Wales, 37: 419. Type male: Tillyard's Collection; type-locality: N e w S o u t h Wales.

Distribution: Australia. fontinalis

LIEFTINCK

1956 Argiolestes fontinalis LIEFTINCK, N o v a G u i n e a , (N.S.), 7: 84. Type male: R i j k s m u s . N a t u u r l . Hist., Leiden; type-locality: N e w G u i n e a .

Distribution: New Guinea. griseus griseus

HÄGEN

1862 Argiolestes grisea HAGEN, in SELYS, Synopsis des Agrionines, A d d . 3: 38. Type male: H a g e n ' s Collection; type-locality: A u s t r a l i a . 1901 Argiolestes grisea - MARTIN, M e m . Soc. zool. F r a n c e , 14: 244 (Victoria). 1913 Argiolestes griseus - TILLYARD, P r o c . Linn. Soc. N . S . Wales, 37: 410 (redescription). 1913 Argiolestes griseus race intermedius TILLYARD, P r o c . Linn. Soc. N . S . Wales, 37: 412. M a l e f r o m N e w S o u t h Wales. 1913 Argiolestes griseus race eboracus TILLYARD, Proc. Linn. Soc. N . S . Wales, 37: 413. M a l e f r o m N e w S o u t h Wales. 1913 Argiolestes griseus race tenuis TILLYARD, P r o c . Linn. Soc. N . S . Wales, 37: 413. M a l e f r o m N e w S o u t h Wales. 1913 Argiolestes griseus race albescens TILLYARD, P r o c . Linn. Soc. N . S . Wales, 37: 414. M a l e f r o m Queensland.

Distribution: Australia. griseus subgriseus

FRÄSER

1 9 8 4 A r g i o l e s t e s griseus subgriseus FRASER, 1 9 5 9 - loc. cit. D A VIES & TOBIN, T h e D r a g o n f l i e s o f

the World, U t r e c h t , 1: 40.

Distribution: East Australia. icteromelas

SELYS

1862 Argiolestes icteromelas SELYS, Synopsis des Agrionines, A d d . 3: 38. Type male: Selys's Collection; type-locality: A u s t r a l i a . 1901 Argiolestes icteromelas - MARTIN, M e m . Soc. zool. F r a n c e , 14: 244 (Victoria, N . S . Wales). 1913 Argiolestes icteromelas - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 410.

143 1913 Argiolestes icteromelas f. nobilis TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 410. Male from New South Wales. 1917 Argiolestes icteromelas - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 26 (Queensland). 1925 Austroargiolestes incteromelas - KENNEDY, Bull. Mus. Comp. Zool., 67: 294 (as generotype o f Austroargiolestes

KENNEDY, 1 9 2 5 ) .

1926 Risiolestes icteromelas - FRASER, Treubia, 8 ( 3 - 4 ) : 486 (as generotype of Risiolestes FRASER, 1926). 1 9 8 6 Argiolestes

icteromelas

- THEISCHINGER & O'FARRELL, O d o n a t o l o g i c a , 15 (4): 3 8 7 .

Distribution: Australia. kirbyi

LIEFTINCK

1900 Wahnesia kirbyi FÖRSTER (nom.nud.), Termeszetr. Füz., Budapest, 23: 103 (without description, as generotype of Wahnesia FÖRSTER, 1900). 1915 Wahnesia kirbyi - SCHMIDT, Zool. Jb., 39 (1): 147. 1926 Argiolestes kirbyi - FRASER, Treubia, 8 ( 3 - 4 ) : 490 (New Guinea). 1935 Argiolestes kirbyi LIEFTINCK (as spec.nov.), Nova Guinea, 17: 228. Type male: Termeszett. Müs., Budapest; type-locality: New Guinea.

Distribution: New Guinea. lamprostomus

LIEFTINCK

1949 Argiolestes lamprostomus LIEFTINCK, Nova Guinea (N.S.), 5: 49. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. luteipes

LIEFTINCK

1956 Argiolestes luteipes LIEFTINCK, Nova Guinea (N.S.), 7: 94. Type male: Rijkmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. macrostylis Ris 1913 Argiolestes macrostylis Ris, Nova Guinea, 9: 475. Type male: Mus. Naturk., Berlin; typelocality: New Guinea.

Distribution: New Guinea. metallicus

SJÖSTEDT

1917 Argiolestes metallicus SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 24. Type male: N a t u r h . Riksmus., Stockholm; type-locality: Queensland.

Distribution: Australia. microstigma

LIEFTINCK

1956 Argiolestes microstigma LIEFTINCK, Nova Guinea (N.S.), 7:99. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. minimus

TILL YARD

1908 Argiolestes minimus TILLYARD, Proc. Linn. Soc. N.S. Wales, 32: 735. Type male: Tillyard's Collection; type-locality: Australia. 1913 Argiolestes minimus - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 415.

Distribution: Australia.

144

montivagans

FÖRSTER

1900 Argiolestes montivagans FÖRSTER, Termeszetr. Füz., Budapest, 23: 103. Type male: Ann Arbor Museum; type-locality: New Guinea. 1938 Argiolestes montivagans - LIEFTINCK, Nova Guinea (N.S.), 2: 77 (redescription).

Distribution: New Guinea. obiensis

LIEFTINCK

1956 Argiolestes obiensis LIEFTINCK, Nova Guinea (N.S.), 7: 79. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Moluccas.

Distribution: Malaysia. ochraceus

(MONTROUSIER)

1864 Sympecma ochracea MONTROUSIER, Ann.Soc. Lyon, 11: 247. Type male: unknown, perhaps in Mus. Nation. Hist. Nat., Paris; type-locality: New Caledonia. 1890 Sympycna ochracea - KIRBY, Syn. Cat. Neur.-Odon., London: 164. 1916 Argiolestes rouxi Ris, Nova Caledonia, Zool., 2: 60. Type male: unknown, perhaps undesignated in Mus. Naturk., Berlin; type-locality: New Caledonia.

Distribution: New Caledonia. ochrostomus

LIEFTINCK

1949 Argiolestes ochrostomus LIEFTINCK, Nova Guinea (N.S.), 5: 48. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Waigeoe Island.

Distribution: Waigeoe. ornatus ornatus

SELYS

1878 Argiolestes ornata SELYS, Mitt. Zool. Mus. Dresden, 3: 321. Type male: not given; type-locality: New Guinea. 1 8 7 8 Argiolestes obscura SELYS (nec M A R T I N , 1 9 0 9 ) , Mitt. Zool. Mus. Dresden, 3: 3 2 1 . Type female: not given; type-locality: New Guinea. 1925 Wahnesia ornata - KENNEDY, Bull. Mus. Comp. Zool., 67 (7): 301. 1935 Argiolestes ornatus ornatus - LIEFTINCK, Nova Guinea 17: 223 (redescription).

Distribution: New Guinea. ornatus subornatus

LIEFTINCK

1935 Argiolestes ornatus subornatus LIEFTINCK, Nova Guinea 17: 224. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. pallidistylus

SELYS

1878 Argiolestes pallidistyla SELYS, Mitt. Zool. Mus. Dresden, 3: 320. Type male: not given; typelocality: New Guinea. 1935 Argiolestes pallidistyla - LIEFTINCK, Nova Guinea, 17: 220 (redescription).

Distribution: New Guinea. parvulus

WATSON

1984 Argiolestes parvulus Utrecht, 1: 40.

WATSON,

1977 - loc. cit.

Distribution: Southwest Australia.

DAVIES & TOBIN,

The Dragonflies of the World,

145 pectitus

LIEFTINCK

1949 Argiolestes pectitus LIEFTINCK, Nova Guinea (N.S.), 5: 53. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, postnodalis

SELYS

1878 Argiolestes postnodalis SELYS, Mitt. Zool. Mus. Dresden, 3: 319. Type male: Selys's Collection; type-locality: New Guinea. 1913 Metagrion postnodalis - CALVERT, Proc. Acad. nat. Sei. Philad., 65: 261 (as generotype of Metagrion CALVERT, 1 9 1 3 ) . 1 9 3 5 Argiolestes postnodalis - LIEFTINCK, Nova Guinea, 1 7 : 2 3 1 (redescription).

Distribution: New Guinea. prothoracalis

LIEFTINCK

1956 Argiolestes prothoracalis LIEFTINCK, Nova Guinea (N.S.), 7: 98. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, pusillissimus

(KENNEDY)

1925 Archiargiolestes pusillissimus K E N N E D Y , Bull. Mus. Comp. Zool., 67: 292. Type male: Comp. Zool. Mus. Cambridge (Mass.); type-locality: West Australia (as generotype of Archiargiolestes K E N N E D Y , 1925). 1984 Argiolestes pusillissimus - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 40.

Distribution: West Australia. pusillus

TILL YARD

1908 Argiolestes pusillus T I L L Y A R D , Proc. Linn. Soc. N.S. Wales, 32: 736. Type male: Tillyard's Collection; type-locality: S. & W. Australia.

Distribution: Australia. saltator

LIEFTINCK

1956 Argiolestes saltator LIEFTINCK, Nova Guinea (N.S.) 7: 105. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. salutarius

LIEFTINCK

1956 Argiolestes salutarius LIEFTINCK, N o v a Guinea (N.S.) 7: 101. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, sidonius

MARTIN

1900 Wahnesia sidonia - FÖRSTER, Termeszetr. Füz., Budapest, 23: 103 (without description, as generotype of Wahnesia FÖRSTER, 1 9 0 0 ) . 1 9 0 8 Argiolestes sidonia M A R T I N , Boll. Soc. ent. Ital., 40: 1 9 5 . Type male: Mus. civ. Stor. nat., Genova; type-locality: New Guinea. 1925 Wahnesia sidonia - K E N N E D Y , Bull. Mus. Comp. Zool. 6 7 (7): 302. 1935 Argiolestes sidonia - LIEFTINCK, Nova Guinea, 1 7 : 233 (redescription). 1938 Argiolestes sidonia - LIEFTINCK, Nova Guinea (N.S.) 2 : 75 (redescription of holotype).

Distribution: New Guinea.

146 simplex

LIEFTINCK

1949 Argiolestes simplex LIEFTINCK, N o v a Guinea (N.S.), 5: 51. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. sponsus

LIEFTINCK

1956 Argiolestes sponsus LIEFTINCK, Nova Guinea (N.S.) 7: 86. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. tenuispinus

LIEFTINCK

1908 Argiolestes obscurus M A R T I N (nec SELYS, 1878), Boll. Soc. ent. Ital., 40: 202. Type male: Mus. civ. Stor. nat. Genova; type-locality: New Guinea. 1 9 3 8 Argiolestes tenuispinus LIEFTINCK, N o v a Guinea (N.S.) 2 : 7 3 (new name for obscurus M A R T I N , 1908).

Distribution: New Guinea. tristis

LIEFTINCK

1935 Argiolestes tristis LIEFTINCK, N o v a Guinea, 17: 229. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Bornargiolestes

KIMMINS

1936 Bornargiolestes KIMMINS, J. Fed. Malay. States Mus., 13: 86. Type species: Bornargiolestes nigra KIMMINS, 1936 (original designation). - 1954 Bornargiolestes - LIEFTINCK, Treubia, 22 Suppl.: 27.

nigra

KIMMINS

Bornargiolestes nigra KIMMINS, J . Fed. Malay. States, Mus., 13: 8 6 (as generotype of Bornargiolestes KIMMINS, 1 9 3 6 ) . Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Bornargiolestes nigra - LIEFTINCK, Treubia, 22 Suppl.: 27. 1936

Distribution: Borneo. Genus Burmargiolestes

KENNEDY

1925 Burmargiolestes KENNEDY, Bull. Mus. Comp. Zool., 67: 296. Type species: Argiolestes nothorax Selys, 1892 (original designation).

laidlawi

mela-

LIEFTINCK

Argiolestes melanothorax L A I D L A W (nec SELYS, 1 8 9 2 ) , Ree. Indian Mus., 13: 3 2 3 . Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: India. 1960 Burmargiolestes laidlawi LIEFTINCK, Mem. Soc. ent. Ital., 38: 236 (new name for melanothorax LAIDLAW, 1917, a junior secondary homonymy of Argiolestes melanothorax SELYS, 1892). 1917

Distribution: India. melanothorax 1892

(SELYS)

Argiolestes melanothorax SELYS (nec L A I D L A W , 1 9 1 7 ) , Ann. Mus. civ. Stor. nat. Genova, 10: 500. Type male: Mus. civ. Stor. nat., Genova; type-locality: Burma.

(2)

147 1925 Burmargiolestes melanothorax - KENNEDY, Bull. Mus. Comp. Zool., 67: 296 (as generotype o f Burmargiolestes

KENNEDY, 1925).

1933 Burmargiolestes flaviceps FRÄSER, J. Siam. Soc. nat. Hist., Suppl., 9: 120. Type male: British Museum (Nat. Hist.), London; type-locality: Laos. 1966 Burmargiolestes flaviceps - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 191 [proposed synonymy with melanothorax (SELYS, 1892)].

Distribution: Burma, Laos. Genus Caledargiolestes

KENNEDY

1925 Caledargiolestes KENNEDY, Bull. Mus. Comp. Zool., 67: 293. Type species: Argiolestes miseries Ris, 1916 (original designation). - 1975 Caledargiolestes - LIEFTINCK, Cah. Off. Rech. sei. tech. Hydrob., 9 (3): 130. - 1984 Caledargiolestes - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 40.

janiceae

LIEFTINCK

1975 Caledargiolestes janiceae LIEFTINCK, Cah. Off. Reh. sei. tech. Hydrob., 9 (3): 131. Type male: Lieftinck's Collection; type-locality: New Caledonia.

Distribution: New Caledonia. uniseries (Ris) 1916 Argiolestes uniseries Ris, Nova Caledonia, Zool., 2: 62. Type male: Mus. Naturk., Berlin; type-locality: New Caledonia. 1925 Caledargiolestes uniseries - KENNEDY, Bull. Mus. Comp. Zool., 67: 293 (as generotype of Caledargiolestes

KENNEDY 1925).

Distribution: New Caledonia. Genus Caledopteryx

KENNEDY

1925 Caledopteryx KENNEDY, Bull. Mus. Comp. Zool., 67: 295. Type species: Argiolestes sarasini Ris, 1916 (original designation).

maculata

WINSTANLEY & DAVIES

1984 Caledopteryx maculata WINSTANLEY & DAVIES, 1982 - loc. cit. DAVIES & TOBIN, T h e D r a -

gonflies of the World, Utrecht, 1: 40.

Distribution: New Caledonia. sarasini (Ris) 1916 Argiolestes sarasini Ris, Nova Caledonia, Zool., 2: 58. Type male: Mus. Naturk., Berlin; typelocality: New Caledonia. 1925 Caledopteryx sarasini - KENNEDY, Bull. Mus. Comp. Zool., 67: 295 (as generotype of Caledopteryx

KENNEDY, 1925).

Distribution: New Caledonia. Genus Calilestes

FRASER

1926 Calilestes FRASER, Treubia, 8 (3-4): 487. Type species: Calilestes pallidistigma FRASER, 1926 (original designation).

148 pallidistigma

FRÄSER

1926 Calilestes pallidistigma FRÄSER, Treubia, 8 ( 3 - 4 ) : 489 (as generotype of Calilestes 1926). Type male: British Museum (Nat. Hist.), London; type-locality: Tonkin.

FRÄSER,

Distribution: Vietnam. Genus Celebargiolestes

KENNEDY

1925 Celebargiolestes KENNEDY, Bull. Mus. Comp. Zool., 67: 295. Type species: Argiolestes cincta SELYS, 1 8 8 6 (original designation). - 1 9 8 4 Celebargiolestes - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 41.

cinctus

(SELYS)

1886 Argiolestes cincta SELYS, Mem. Cour. Acad. Belg., 38: 86. Type male: Selys's Collection; typelocality: Celebes. 1925 Celebargiolestes cincta - KENNEDY, Bull. Mus. Comp. Zool., 67: 296 (as generotype of Celebargiolestes KENNEDY, 1825). 1 9 2 6 Argiolestes karnyi FRÄSER, Treubia, 8 ( 3 - 4 ) : 4 8 6 . Type male: not given; type-locality: Java. 1984 Argiolestes karnyi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 41 [proposed synonymy with cinctus (SELYS, 1886)].

Distribution: Celebes, Java. Genus Dimeragrion

CALVERT

1913 Dimeragrion CALVERT, Proc. Acad. Philad., 65: 265. Type species: Dimeragrion CALVERT, 1913 (original designation).

percubitale

percubitale

CALVERT

1913 Dimeragrion percubitale CALVERT, Proc. Acad. Philad., 65: 265 (as generotype of Dimeragrion CALVERT, 1913). Type male: Acad. nat. Sei., Philadelphia; type-locality: British Guyana.

Distribution: British Guyana. secundum

NEEDHAM

1933 Dimeragrion secundum NEEDHAM, Amer. Mus. Novit., 664: 5. Type: unknown; type-locality: Venezuela.

Distribution: Venezuela. Genus Heteragrion

SELYS

1862 Heteragrion SELYS, Synopsis des Agrionines, Add. 3: 17. Type species: Heteragrion flavovittatum SELYS, 1862 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 123). - 1916 Heteragrion - Ris, Arch. Naturg., (A) 82: 85. - 1987 Heteragrion - DE MARMELS, Odonatologica, 16 (3): 225.

aequatoriale

SELYS

1886 Heteragrion aequatoriale SELYS, Mem. Cour. Acad. Belg., 38: 63. Type male: lost; type-locality: Bogota. 1916 Heteragrion aequatoriale - Ris, Arch. Naturg., (A) 82: 85 (Peru). 1943 Heteragrion aequatoriale - SCHMIDT, Beitr. F a u n a Perus, 2: 228 (Colombia, Ecuador, Peru).

Distribution: South America.

149 albifrons Ris 1916 Heteragrion albifrons Ris, Arch. Naturg., (A) 82: 89. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Costa Rica.

Distribution: Costa Rica. alienum

WILLIAMSON

1919 Heteragrion alienum WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Michigan, 68: 33. Type male: Williamson's Collection; type-locality: Central America.

Distribution: Central America. angustipenne

SELYS

1886 Heteragrion angustipenne locality: Ecuador. 1909 Heteragrion angustipenne 1919 Heteragrion angustipenne 1943 Heteragrion angustipenne

SELYS, Mem. Cour., Acad. Belg., 38: 64. Type male: unknown; type- CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 108 (Peru). - WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Michigan, 68: 13. - SCHMIDT, Beitr. Fauna Perus, 2: 228.

Distribution: Ecuador, Peru. aurantiacum

SELYS

1862 Heteragrion aurantiacum SELYS, Synopsis des Agrionines, Add. 3: 23. Type male: Selys's Collection; type-locality: Brazil. 1862 Heteragrion cinnamomeum HEYER, in SELYS, Synopsis des Agrionines, Add. 3: 24. Type male: Selys's Collection; type-locality: Bahia. 1886 Heteragrion amazonicum SELYS, Mem. Cour., Acad. Belg., 38: 60. Type female: R. Mus. Brussels; type-locality: Amazonas. 1890 Heteragrion aurantiacum - KIRBY, Syn. Cat. Neur.-Odon., London: 124. 1890 Heteragrion cinnamomeum - KIRBY, Syn. Cat. Neur.-Odon., London: 124 (proposed synonymy w i t h aurantiacum

SELYS, 1 8 6 2 ) .

1890 Heteragrion amazonicum -KIRBY, Syn. Cat. Neur.-Odon., London: 124 (proposed synonymy with aurantiacum SELYS, 1862). 1984 Heteragrion cinnamomeum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 41 (incorrect synonymy with ochraceum HÄGEN, 1862).

Distribution: Brazil. azulum

DUNKLE

1989 Heteragrion azulum DUNKLE, Odonatologica, 18 (2): 195. Type male: Florida State Coll., Gainesville; type-locality: Mexico.

Distribution: Mexico. beschki

SELYS

1862 Heteragrion beschkii SELYS, Synopsis des Agrionines, Add. 3: 25. Type male: Beschk's Collection; type-locality: Brazil.

Distribution: Brazil. calendulum

WILLIAMSON

1919 Heteragrion calendulum WILLIAMSON, Occ. Pap. Mus. Zool. Univ. Michigan, 68: 36. Type male: Williamson's Collection; type-locality: Colombia.

Distribution: Colombia.

150 chrysops

HAGEN

1862 Heteragrion chrysops HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 26. Type male: H a gen's Collection; type-locality: Venezuela. 1907 Heteragrion chrysops - CALVERT, Biol. Centr.-Amer. Neur.: 357 (Costa Rica, Panama). 1916 Heteragrion chrysops - Ris, Arch. Naturg., (A) 82: 88 (Panama).

Distribution: Central America, Venezuela. consors

HÄGEN

1862 Heteragrion consors HAGEN, in SELYS, Synopsis des Agrionines, Add. 3:25. Type male: Hagen's Collection; type-locality: Bahia.

Distribution: Brazil. dorsale

SELYS

1862 Heteragrion dorsale SELYS, Synopsis des Agrionines, Add. 3: 21. Type male: Selys's Collection; type-locality: Brazil.

Distribution: Brazil. eboratum

DONNELLY

1 9 8 4 H e t e r a g r i o n e b o r a t u m DONNELLY, 1 9 6 5 - l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e

World, Utrecht, 1: 41.

Distribution: Guatemala. erythrogastrum

SELYS

1886 Heteragrion erythrogastrum SELYS, Mem. Cour., Acad. Belg., 38: 61. Type male: Selys's Collection; type-locality: Panama. 1907 Heteragrion erythrogastrum - CALVERT, Biol. Centr.-Amer., Neur.: 62 (Costa Rica, Panama). 1916 Heteragrion erythrogastrum - Ris, Arch. Naturg., (A) 82: 88 (Panama).

Distribution: Central America. flavidorsum

CALVERT

1909 Heteragrion flavidorsum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 111. Type male: Calvert's Collection; type-locality: Bolivia.

Distribution: Bolivia. flavovittatum

SELYS

1862 Heteragrion flavovittatum SELYS, Synopsis des Agrionines, Add. 3:18. Type male: Selys's Collection; type-locality: Brazil. 1890 Heteragrion flavovittatum - KIRBY, Syn. Cat. Neur.-Odon., London: 123 (as generotype of Heteragrion SELYS, 1862).

Distribution: Brazil. ictericum

WILLIAMSON

1919 Heteragrion ictericum WILLIAMSON, Occ. Pap. Mus. Zool. Univ. Michigan, 68: 17. Type male: Williamson's Collection; type-locality: British Guyana.

Distribution: British Guyana.

151 icterops

SELYS

1862 Heteragrion icterops SELYS, Synopsis des Agrionines, Add. 3: 27. Type male: Selys's Collection; type-locality: Santarem. 1886 Heteragrion icterops - SELYS, Bull. Acad. r. Belg., (3) 38: 65. 1919 Heteragrion icterops - WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Michigan, 68: 51. 1943 Heteragrion icterops - SCHMIDT, Beitr. Fauna Perus, 2: 228 (Ecuador, Venezuela, Peru).

Distribution: Ecuador, Venezuela, Peru. inca

CALVERT

1901 Heteragrion inca HIGGINS, Proc. Acad. nat. Sei. Philad., 53: 137 (without description). 1909 Heteragrion inca CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 108. Type male: Carnegie Museum; type-locality: Iquitos. 1943 Heteragrion inca - SCHMIDT, Beitr. Fauna Perus, 2: 228.

Distribution: Peru. macilentum

HÄGEN

1862 Heteragrion macilentum HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 24. Type male: Naturhist. Mus., Wien; type-locality: Venezuela.

Distribution: Venezuela. majus

SELYS

1886 Heteragrion majus SELYS, Mem. Cour., Acad. Belg., 38: 62. Type male: unknown; type-locality: Chiriqui. 1916 Heteragrion majus - Ris, Arch. Naturg., (A) 82: 86 (Peru) (redescription). 1943 Heteragrion majus - SCHMIDT, Beitr. Fauna Perus, 2: 228.

Distribution: Peru. melanurum

WILLIAMSON

1919 Heteragrion melanurum WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Michigan, 68: 20. Type male: Williamson's Collection; type-locality: British Guyana.

Distribution: British Guyana, mitratum

WILLIAMSON

1919 Heteragrion mitratum WILLIAMSON, Occ. Pap. Mus. Zool. Univ. Michigan, 68: 39. Type male: Williamson's Collection; type-locality: Colombia.

Distribution: Colombia, obsoletum

SELYS

1886 Heteragrion obsoletum SELYS, Mem. Cour. Acad. Belg., 38: 57. Type male: unknown; typelocality: Brazil.

Distribution: Brazil. ochraceum HÄGEN 1862 Heteragrion ochraceum HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 22. Type male: Burmeister's Collection; type-locality: Brazil.

Distribution: Brazil.

152 ovatum SELYS 1862 Heteragrion ovatum SELYS, Synopsis des Agrionines, Add. 3: 19. Type male: Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Brazil.

Distribution: Brazil. pemon

DE MARMELS

1987 Heteragrion pemon DE M A R M E L S , Odonatologica, 16 (3): 231. Type male: de Marmels's Collection; type-locality: Venezuela.

Distribution: Venezuela. peregrinum WILLIAMSON 1919

Heteragrion peregrinum WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Michigan, male: Williamson's Collection; type-locality: Colombia.

68: 51.

Type

Distribution: Colombia. romani SJÖSTEDT Heteragrion romani SJÖSTEDT, Ark. Zool., Uppsala, 1 1 ( 1 5 ) : 8 . Type male: Naturh. Riksmus., Stockholm; type-locality: Amazonas. 1987 Heteragrion romani - DE MARMELS, Odonatologica, 1 6 (3): 227.

1917

Distribution: Brazil. silvarum 1917

SJÖSTEDT

Heteragrion silvarum SJÖSTEDT, Ark. Zool., Uppsala, Stockholm; type-locality: Amazonas.

11 (15): 5.

Type male: Naturh. Riksmus.,

Distribution: Brazil. simulatum WILLIAMSON 1919 Heteragrion simulatum WILLIAMSON, Occ. Pap. Mus. Zool. Univ. Michigan, 68: 47. Type male: Williamson's Collection; type-locality: British Guyana.

Distribution: British Guyana. speciosum

SJÖSTEDT

1917 Heteragrion speciosum SJÖSTEDT, Ark. Zool., Uppsala, 11( 15): 7. Type male: Naturh. Riksmus., Stockholm; type-locality: Peru. 1987 Heteragrion speciosum - DE M A R M E L S , Odonatologica, 1 6 (3): 231.

Distribution: Peru. triangulare HÄGEN 1862 Heteragrion triangulare H A G E N , in SELYS, Synopsis des Agrionines, Add. 3: 20. Type female: Naturh. Mus. Wien; type-locality: Brazil.

Distribution: Brazil. tricellulare CALVERT 1901

Heteragrion tricellulare CALVERT, Biol.-Centr. Amer. Neuropt.: Museum; type-locality: Guatemala.

Distribution: Guatemala.

63.

Type male: Pittsburgh

153 Genus Heteropodagrion

SELYS

1885 Heteropodagrion SELYS, C.R. SOC. Ent. Belg., 29: 144. Type species: Heteropodagrion nipes SELYS, 1885 (monotypy).

sanguinipes

sangui-

SELYS

1885 Heteropodagrion sanguinipes SELYS, C.R. Soc. Ent. Belg., 29: 144 (as generotype of Heteropodagrion SELYS, 1885). Type male: R. Mus., Brussels; type-locality: Quito.

Distribution: Ecuador. superbum Ris 1916 Heteropodagrion superbum Ris, Arch. Naturg., (A) 82: 89. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Colombia.

Distribution: Colombia. Genus Mesagrion

SELYS

1885 Mesagrion SELYS, C.R. Soc. Ent. Belg., 29: 144. Type species: Mesagrion leucorhinum SELYS, 1885 (monotypy).

leucorhinum

SELYS

1885 Mesagrion leucorhinum SELYS, C.R. Soc. Ent. Belg., 29: 144 (as generotype of SELYS, 1885). Type male: unknown; type-locality: Bogota. 1916 Mesagrion leucorhinum - Ris, Arch. Naturg., (A) 82: 90.

Mesagrion

Distribution: Colombia. Genus Nesolestes

SELYS

1891 Podagrion (Nesolestes) SELYS, C.R. Soc. Ent. Belg., 35: 399. Type species: Podagrion (Nesolestes) alboterminata SELYS, 1891 (monotypy). - 1951 Nesolestes - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 132 (key to Malagascan species). - 1955 Nesolestes - FRÄSER, Nat. malgache, 7: 45 (key to Malagascan species). - 1962 Nesolestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 98.

albicauda

FRÄSER

1952 Nesolestes albicauda FRASER, Mem. Inst. sei. Madagascar, (Ε) 1:137. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Nesolestes albicauda - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 98.

Distribution: Madagascar. albicolor

FRASER

1955 Nesolestes albicolor FRASER, Nat. malgache, 7 (1): 50. Type male: Inst. Rech. Sei. Madagascar; type-locality: Madagascar. 1962 Nesolestes albicolor - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 98.

Distribution: Madagascar. alboterminatus 1891 Podagrion Nesolestes 1919 Nesolestes 1962 Nesolestes

SELYS

(Nesolestes) alboterminata SELYS, C.R. Ent. Soc. Belg., 35: 400 (as generotype of SELYS, 1891). Type male: R. Mus., Brussels; type-locality: Madagascar. alboterminata - MUNZ, Mem. Am. ent. Soc., 3 (1): 3. alboterminata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 98.

Distribution: Madagascar.

154 angydna

SCHMIDT

1951 Nesolestes angydna SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 142. Type male: Schmidt's Collection; type-locality: Madagascar. 1 9 4 4 Nesolestes angydna - SCHMIDT, Dtsch. ent. Z . , 1 9 4 4 ( 1 - 2 ) : 7 6 (as invalid taxa). 1962 Nesolestes angydna - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 98.

Distribution: Madagascar. drocerus

FRASER

1951 Nesolestes drocera FRASER, Mem. Inst. sei. Madagascar, 6A: 415. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Nesolestes drocera - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 98.

Distribution: Madagascar. elizabethae

LIEFTINCK

1965 Nesolestes elizabethae LIEFTINCK, Verh. Naturf. Ges. Basel, 76: 243. Type male: Naturh. Mus., Basel; type-locality: Madagascar.

Distribution: Madagascar, forficuloides

FRASER 1955 Nesolestes forficuloides FRASER, Nat. malgache, 7: 51. Type male: Inst. Rech. Sei. Madagasgar; type-locality: Madagascar. 1962 Nesolestes forficuloides - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

Distribution: Madagascar. mariae 1984

AGUESSE

Nesolestes mariae Utrecht, 1: 42.

AGUESSE, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Madagascar. martini

SCHMIDT

1951 Nesolestes martini SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 138. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Nesolestes martini - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

Distribution: Madagascar. nigeriensis 1984

GAMBLES

Nesolestes nigeriensis Utrecht, 1: 42.

GAMBLES, 1 9 7 0

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Nigeria, pauliani FRASER 1951 Nesolestes pauliani

FRASER, Mem. Inst. sei. Madagascar, 6A: 413. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Nesolestes pauliani - PINHEY, Pub. cult. Comp.Diam. Angola, 59: 99.

Distribution: Madagascar.

155 pulverulans

LIEFTINCK

1 9 8 4 N e s o l e s t e s pulverulans LIEFTINCK, 1 9 6 5 - l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e

World, Utrecht, 1: 42.

Distribution: Madagascar. radama

LIEFTINCK

1984 Nesolestes radama LIEFTINCK, 1965 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 42.

Distribution: Madagascar. ranavalona

SCHMIDT

1951 Nesolestes ranavalona SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 136. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Nesolestes ranavalona - PINHEY, Publ. cult. comp. Diam. Angola, 59: 99.

Distribution: Madagascar. robustus

AGUESSE

1984 Nesolestes robustus AGUESSE, 1968 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 42.

Distribution: Madagascar. rubristigma

MARTIN

1903 Nesolestes rubristigma MARTIN, Bull. Mus. Paris, 8: 510. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Nesolestes rubristigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

Distribution: Madagascar. tuberculicollis

SCHMIDT

1949 Nesolestes tuberculicollis SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 3A: 11. Type female: Schmidt's Collection; type-locality: Madagascar. 1962 Nesolestes tuberculicollis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

Distribution: Madagascar. Genus Neuragrion

KARSCH

1891 Neuragrion KARSCH, SOC. Ent., 6: 105. Type species: Neuragrion mysticum KARSCH, 1891 (original designation).

mysticum

KARSCH

1891 Neuragrion mysticum KARSCH, SOC. Ent., 6: 105 (as generotype of Neuragrion KARSCH, 1891). Type male: unknown, perhaps in Mus. Naturk., Berlin; type-locality: Ecuador.

Distribution: Ecuador. Genus Neurolestes

SELYS

1882 Neurolestes SELYS, Congr. Sei., 10: 668. Type species: Neurolestes trinervis SELYS, 1885 (emendation). - 1885 Neurolestes -SELYS, C.R. SOC. Ent. Belg., 29: 144. - 1890 Neurolestes - KIRBY,

156 Syn. Cat. Neur.-Odon., London: 124. - 1962 Neurolestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

trinervis

SELYS

1885 Neurolestes trinervis SELYS, C.R. Soc. Ent. Belg., 2 9 : 144 (as generotype of Neurolestes 1882). Type male: British Museum (Nat. Hist.), London; type-locality: Cameroon. 1919 Neurolestes trinervis - MUNZ, Mem. Am. ent. Soc., 3 (1): 10. 1962 Neurolestes trinervis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

SELYS,

Distribution: Cameroon. Genus Oxystigma 1919

WILLIAMSON

Oxystigma WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Mich., 68: 5 8 . Type species: Oxystigma cyanofrons WILLIAMSON, 1 9 1 9 (original designation). - 1 9 3 4 Oxystigma - C O W L E Y , Entomologist's month. Mag., 70: 245.

cyanofrons

WILLIAMSON

1919 Oxystigma cyanofrons WILLIAMSON, Occ. Pap. Mus. Zool. Univ. Mich., 68: 58 (as generotype of Oxystigma WILLIAMSON 1919). Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. petiolatum

(SELYS)

1862 Heteragrion petiolatum SELYS, Synopsis des Agrionines, Add. 3: 28. Type male: R. Mus., Brussels; type-locality: Santarem. 1984 Oxystigma petiolatum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:42 (Brazil).

Distribution: Brazil. williamsoni 1984

GEIJSKES

Oxystigma williamsoni World, Utrecht, 1: 42.

GEIJSKES, 1 9 7 6

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Suriname. Genus Paraphlebia

SELYS

1861 Paraphlebia SELYS, in HÄGEN, Syn. Neur. N. Amer., Washington: 71. Type species: Paraphlebia zoe H Ä G E N , 1 8 6 1 (original designation). - 1 9 6 2 Paraphlebia - SELYS, Synopsis des Agrionines, Add. 3 : 6 (revision). - 1 9 8 4 Paraphleoia (misprint) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 42.

abrogata

CALVERT

1907 Paraphlebia abrogata CALVERT, Biol. Centr.-Amer., Neuropt.: 355. Type male: Calvert's Collection; type-locality: Guatemala.

Distribution: Guatemala. duodecima

CALVERT

1901 Paraphlebia duodecima CALVERT, Biol. Centr.-Amer., Neuropt.: 60. Type male: Calvert's Collection; type-locality: Guatemala.

Distribution: Guatemala.

157 hyalina BRAUER 1871 Paraphlebia hyalina B R A U E R , Verh. Zool.-Bot. Ges. Wien, Mus., Wien; type-locality: Mexico.

21:

105. Type male: Naturhist.

Distribution: Mexico. quinta CALVERT 1901 Paraphlebia quinta C A L V E R T , Biol. Centr.-Amer., Neuropt.: 60. Type male: Calvert's Collection; type-locality: Guatemala.

Distribution: Guatemala. z o e SELYS 1861 Paraphlebia zoa SELYS, in H A G E N , Syn. Neur. N. Amer., Washington: 72 (as generotype of Paraphlebia SELYS, 1861) (without description). Type: unknown. 1 8 6 2 Paraphlebia zoa - SELYS, Synopsis des Agrionines, Add. 3 : 7 (description of male, specimen from Selys's Collection).

Distribution: Mexico. Genus Philogenia SELYS 1862 Philogenia SELYS, Bull. Acad. r. Belg., (2) 14: 10. Type species: Philogenia margarita SELYS, 1862 (designated by K I R B Y , 1890, Syn. Cat. Neur.-Odon., London: 122). - 1890 Philogenia - K I R B Y , Syn. Cat. Neur.-Odon., London: 122. - 1916 Philogenia - Ris, Arch. Naturg., (A) 8 2 : 77 (key to species). - 1943 Philogenia - SCHMIDT, Beitr. Fauna Perus, 2 : 227. - 1986 Philogenia - D U N K L E , Odonatologica, 1 5 (1): 43. - 1988 Philogenia - B I C K & B I C K , Odonatologica, 17 (1): 14 (key to species).

angusti CALVERT 1924 Philogenia angusti CALVERT, Trans. Am. ent. Soc., 50: 42. Type male: Calvert's Collection; type-locality: Panama.

Distribution: Panama. berenice

HIGGINS

1901 Philogenia berenice H I G G I N S , Proc. Acad. nat. Sei. Philad., 53: 136. Type: unknown; typelocality: Peru. 1943 Philogenia berenice - S C H M I D T , Beitr. Fauna Perus, 2 : 227.

Distribution: Peru. boliviana BICK & BICK 1988 Philogenia boliviana cality: Bolivia.

BICK & BICK,

Odonatologica,

17

(1): 16. Type male: unknown; type-lo-

Distribution: Bolivia. buenavista BICK & BICK 1988 Philogenia buenavista locality: Bolivia.

BICK & B I C K ,

Distribution: Bolivia.

Odonatalogica,

17

(1): 27. Type male: unknown; type-

158 carrillica

CALVERT

1907 Philogenia carrillica CALVERT, Biol. Centr. Amer., Neuropt.: 356. Type male: Calvert's Collection; type-locality: Costa Rica. 1916 Philogenia carrillica - Ris, Arch. Naturg., (A) 82: 81.

Distribution: Costa Rica. cassandra

HÄGEN

1862 Philogenia cassandra HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 10. Type male: Naturhist. Mus., Wien; type-locality: Venezuela. 1943 Philogenia cassandra - SCHMIDT, Beitr. Fauna Perus, 2: 227 (Peru).

Distribution: Venezuela, Peru. championi

CALVERT

1901 Philogenia championi CALVERT, Biol. Centr.-Amer., Neuropt.: 61. Type male: Calvert's Collection; type-locality: Panama. 1988 Philogenia championi - BICK & BICK, Odonatologica, 17 (1): 117.

Distribution: Panama. cristalina

CALVERT

1924 Philogenia cristalina CALVERT, Trans. Am. ent. Soc., 50: 35. Type male: Calvert's Collection; type-locality: Colombia.

Distribution: Colombia. ebona

DUNKLE

1986 Philogenia ebona DUNKLE, Odonatologica, 15 (1): 48. Type male: Dunkle's Collection; typelocality: Colombia.

Distribution: Colombia, elisabeta

CALVERT 1924 Philogenia elisabeta CALVERT, Trans. Am. ent. Soc., 50: 29. Type male: Calvert's Collection; type-locality: Peru. 1943 Philogenia elisabeta - SCHMIDT, Beitr. Fauna Perus, 2: 227.

Distribution: Peru. expansa

CALVERT

1924 Philogenia expansa CALVERT, Trans. Am. ent. Soc., 50: 50. Type female: Calvert's Collection; type-locality: Costa Rica.

Distribution: Costa Rica, ferox

RACENIS

1959 Philogenia ferox RACENIS, Acta biol. venezuel., 2: 353. Type male: Racenis's Collection; typelocality: Venezuela.

Distribution: Venezuela. helena

HÄGEN

1879 Philogenia helena HAGEN, Stettiner ent. Ztg., 30: 261. Type male: Hagen's Collection; typelocality: Bogota. 1916 Philogenia helena - Ris, Arch. Naturg., (A) 82: 78 (Colombia, Peru).

159 1943 Philogenia helena - SCHMIDT, Beitr. Fauna Perus, 2: 227 (Peru).

Distribution: Colombia, Peru. lankesteri

CALVERT

1924 Philogenia lankesteri CALVERT, Trans. Am. ent. Soc., 50: 51. Type male: Calvert's Collection; type-locality: Costa Rica.

Distribution: Costa Rica. leonora

WESTFALL & CUMMING

1956 Philogenia leonora WESTFALL & CUMMING, Bull. Fla State Mus., 1: 244. Type male: British Museum (Nat. Hist.), London; type-locality: Panama.

Distribution: Panama. macuma

DUNKLE

1986 Philogenia macuma DUNKLE, Odonatologica, 15 (1): 45. Type male: unknown; type-locality: Ecuador.

Distribution: Ecuador. mangosisa

BICK & BICK

1988 Philogenia mangosisa BICK & BICK, Odonatologica, 17 (1): 24. Type male: Bick & Bick's Collection; type-locality: Ecuador.

Distribution: Ecuador. margarita

SELYS

1862 Philogenia margarita SELYS, Synopsis des Agrionines, Add. 3: 9. Type female: Selys's Collection; type-locality: Amazonas, (as generotype of Philogenia SELYS, 1862).

Distribution: Brazil. minteri

DUNKLE

1986 Philogenia minteri DUNKLE, Odonatologica, 15 (1): 43. Type male: unknown; type-locality: Ecuador.

Distribution: Ecuador. peruviana

BICK & BICK

1988 Philogenia peruviana BICK & BICK, Odonatologica, 17 (1): 23. Type male: Bick & Bick's Collection; type-locality: Peru.

Distribution: Peru. polyxena

CALVERT

1924 Philogenia polyxena CALVERT, Trans. Am. ent. Soc., 50: 27. Type male: Calvert's Collection; type-locality: Venezuela.

Distribution: Venezuela. raphaella

SELYS

1886 Philogenia raphaella SELYS, Mem. Cour. Acad. Belg., 38: 37. Type male: R. Mus., Brussels; type-locality: Bogota.

Distribution: Colombia.

160 schmidti Ris 1916 Philogenia schmidti Ris, Arch. Naturg., (A) 82: 79. Type male: not given; type-locality: Bolivia.

Distribution: Bolivia, silvarum Ris 1916 Philogenia silvarum Ris, Arch. Naturg., (A) 82: 80. Type male: not given; type-locality: Peru. 1943 Philogenia silvarum - SCHMIDT, Beitr. Fauna Perus, 2: 227 (Peru).

Distribution: Peru. sucra DUNKLE 1986 Philogenia sucra DUNKLE, Odonatologica, 15 (1): 47. Type male: Dunkle's Collection; typelocality: Colombia.

Distribution: Colombia. terraba CALVERT 1907 Philogenia terraba CALVERT, Biol. Centr.-Amer. Neuropt.: 356. Type male: Calvert's Collection; type-locality: Costa Rica.

Distribution: Costa Rica, tinlandica BICK & BICK 1988 Philogenia tinlandica BICK & B I C K , Odonatologica, 17 (1): 18. Type male: Bick & Bick's Collection; type-locality: Ecuador.

Distribution: Ecuador. umbrosa Ris 1916 Philogenia umbrosa Ris, Arch. Naturg., (A) 82: 78. Type male: not given; type-locality: Peru. 1943 Philogenia umbrosa - SCHMIDT, Beitr. Fauna Perus, 2: 277 (Peru).

Distribution: Peru. zeteki WESTFALL & CUMMING 1956

Philogenia zeteki WESTFALL & C U M M I N G , Bull. Fla State Mus., 1: Museum (Nat. Hist.), London; type-locality: Panama.

241.

Type male: British

Distribution: Panama. Genus Podolestes SELYS 1862 Podolestes Selys, Synopsis des Agrionines, Add. 3: 39. Type species: Podolestes orientalis SELYS, 1862 (monotypy). - 1950 Podolestes - LIEFTINCK, Zool. Meded., Leiden, 31: 39 (key to Malaysian species). - 1954 Podolestes - LIEFTINCK, Treubia, 22 Suppl.: 28.

atomarius

LIEFTINCK

1950 Podolestes atomarius LIEFTINCK, Zool. Meded., Leiden, 31: 41. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Borneo. 1954 Podolestes atomarius - LIEFTINCK, Treubia, 22 Suppl.: 28.

Distribution: Borneo. buwaldai

LIEFTINCK

1940 Podolestes buwaldai LIEFTINCK, Treubia, 17: 347. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Sumatra.

161 1950 Podolestes buwaldai1954 Podolestes buwaldai -

LIEFTINCK, LIEFTINCK,

Zool. Meded., Leiden, 3 1 : 40. Treubia, 2 2 Suppl.: 28 (Sumatra).

Distribution: Sumatra, chrysopus

SELYS

1889 Podolestes Stor. nat., 1924 Podolestes 1935 Podolestes 1954 Podolestes

chrysopus SELYS, Ann.MUS. civ. Stor. nat., Genova, 27: 479. Type male: Mus. civ. Genova; type-locality: Labuan. chrysopus - L A I D L A W , J . Mal. Br. R . Asiat. Soc., 2 : 302. chrysopus - LIEFTINCK, Treubia, 1 5 (2): 181 (redescription) (Borneo). chrysopus - LIEFTINCK, Treubia, 2 2 Suppl.: 28.

Distribution: Malaysia: Labuan Island, and Borneo. coomansi

LIEFTINCK

1941 Podolestes coomansi LIEFTINCK, Treubia, 17: 348. Type male: Buitenzorg Museum; type-locality: Sumatra. 1950 Podolestes coomansi - LIEFTINCK, Zool. Meded., Leiden, 3 1 : 40. 1954 Podolestes coomansi - LIEFTINCK, Treubia, 2 2 Suppl.: 28.

Distribution: Sumatra. furcifer

LIEFTINCK

1950 Podolestes furcifer LIEFTINCK, Zool. Meded., Leiden, 31: 44. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Borneo. 1954 Podolestes furcifer - LIEFTINCK, Treubia, 2 2 Suppl.: 29.

Distribution: Borneo. harrissoni

LIEFTINCK

1953 Podolestes harrissoni LIEFTINCK, Treubia, 2 2 : 233. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Borneo. 1 9 5 4 Podolestes harrissoni - LIEFTINCK, Treubia, 2 2 Suppl.: 2 9 .

Distribution: Borneo. orientalis

SELYS

1862 Podolestes orientalis SELYS, Synopsis des Agrionines, Add. 3:40. Type male: Selys's Collection; type-locality: Malacca (as generotype of Podolestes SELYS, 1862). 1927 Podolestes orientalis - Ris, Zool. Meded., Leiden, 10 (1): 15 (redescription) (Sumatra). 1935 Podolestes orientalis - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 6 (Sumatra). 1935 Podolestes orientalis - LIEFTINCK, Treubia, 15 (2): 177 (Sumatra, Borneo, Malaya) (redescription). 1 9 5 4 Podolestes orientalis - LIEFTINCK, Treubia, 2 2 Suppl.: 2 9 (Malaya, Sumatra, Borneo).

Distribution: Malaysia, Sumatra, Borneo. Genus Podopteryx

SELYS

1871 Podopteryx SELYS, Trans, ent. Soc. London, 1871: 415. Type species: Podopteryx roseonotata SELYS, 1871 (monotypy). - 1935 Podopteryx - LIEFTINCK, Nova Guinea, 17: 206 (revision).

casuarina

LIEFTINCK

1949 Podopteryx casuarina LIEFTINCK, Nova Guinea, (N.S.) 5: 39. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea.

162 roseonotata

SELYS

1871 Podopteryx roseonotata SELYS, Trans, ent. Soc. London, 1871: 415 (as generotype of Podopteryx SELYS, 1871). Type male: British Museum (Nat. Hist.), London; type-locality: Aru. 1878 Podopteryx roseonotata - SELYS, Mitt. Zool. Mus. Dresden, 3: 319. 1901 Podopteryx roseonotata - MARTIN, Mem. Soc. Zool. France, 14: 244 (New South Wales, Queensland). 1917 Podopteryx roseonotata - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 24 (Queensland). 1935 Podopteryx roseonotata - LIEFTINCK, Nova Guinea, 17: 207.

Distribution: Australia, Aru. selysi

(FÖRSTER)

1899 Agriolestes selysi FÖRSTER, Ann. Soc. ent. Belg., 43: 70. Type male: unknown; type-locality: New Guinea. 1925 Podopteryx selysi - KENNEDY, Bull. Comp. Zool. Harv. Coll., 67: 294. 1935 Podopteryx selysi - LIEFTINCK, Nova Guinea, 17: 208 (redescription).

Distribution: New Guinea, Australia. Genus Protolestes

FÖRSTER

1899 Protolestes FÖRSTER, Ent. Nachr., 25:187. Type species: Protolestesfickei FÖRSTER, 1899 (monotypy). - 1 9 1 3 Protolestes - CALVERT, Trans. Congr. Ent., 2: 260. - 1 9 6 2 Protolestes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 99.

fickei

FÖRSTER

1899 Protolestes fickei FÖRSTER, Ent. Nachr., 25:188 (as generotype of Protolestes FÖRSTER, 1899). Type female: Ann Arbor Museum; type-locality: Madagascar. 1962 Protolestes fickei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 100.

Distribution: Madagascar. furcatus 1984

AGUESSE

Protolestes furcatus AGUESSE, Utrecht, 1: 44.

1967

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Madagascar. kerckhoffae

SCHMIDT

1949 Protolestes kerckhoffae SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 3A: 14. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Protolestes kerckhoffae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 100.

Distribution: Madagascar. leonorae

FRASER

1949 Protolestes leonorae FRASER, Mem. Inst. sei. Madagascar, 3A: 15. Type female: Fraser's Collection; type-locality: Madagascar. 1952 Protolestes leonorae - FRASER, Mem. Inst. sei. Madagascar, (Ε) 1: 137 (description of male).

Distribution: Madagascar. milloti

FRASER

1949 Protolestes milloti FRASER, Mem. Inst. sei. Madgascar, 3A: 13. Type male: Inst. Rech. Sei., Madagascar; type-locality: Madagascar.

163 1962 Protolestes milloti-

PINHEY, Publ. cult. Comp. Diam. Angola, 59: 100.

Distribution: Madagascar. proselytus

LIEFTINCK

1 9 8 4 P r o t o l e s t e s p r o s e l y t u s LIEFTINCK, 1 9 6 5 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e

World, Utrecht, 1: 44.

Distribution: Madagascar. rufescens

AGUESSE

1984 Protolestes rufescens AGUESSE, 1967 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 44.

Distribution: Madagascar. simonei

AGUESSE

1984 Protolestes simonei AGUESSE, 1967 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 44.

Distribution: Madagascar. Genus Rhinagrion

CALVERT

1862 Amphilestes SELYS, Synopsis des Agrionines, Add. 3: 40. Type species: Amphilestes macrocephale SELYS, 1862 (monotypy). - 1913 Rhinagrion CALVERT, Proc. Acad. Philad., 65: 258 (new name for Amphilestes SELYS, 1862, a junior primary homonymy of Amphilestes OWEN, 1859 in Mammalia). - 1954 Rhinagrion - LIEFTINCK, Treubia, 22 Suppl.: 29.

borneensis

(SELYS)

1886 Amphilestes macrocephaa race borneensis SELYS, Mem. Cour. Acad. Belg., 38: 93. Male from Lebuan (Borneo). 1920 Rhinagrion borneensis - LAIDLAW, Proc. Zool. Soc. London, 1920: 3322. 1954 Rhinagrion borneensis - LIEFTINCK, Treubia, 22 Suppl.: 29.

Distribution: Borneo. elopurae

(MCLACHLAN)

1886 Amphilestes elopurae MCLACHLAN, in SELYS, Mem. Cour., Acad. Belg., 38: 221. Type male: British Museum (Nat. Hist.), London; type-locality: North Borneo. 1924 Rhinagrion elopurae - LAIDLAW, J. Mal. Br. R. Asiat. Soc., 2: 203. 1954 Rhinagrion elopurae - LIEFTINCK, Treubia, 22: Suppl.: 30 (Borneo).

Distribution: Borneo. macrocephalum

(SELYS)

1862 Amphilestes macrocephala SELYS, Synopsis des Agrionines, Add. 3: 41 (as generotype of Amphilestes SELYS, 1862). Type male: Selys's Collection; type-locality: Malacca. 1913 Rhinagrion macrocephala - CALVERT, Proc. Acad. Philad., 65: 258 (as generotype of Rhinagrion CALVERT, 1 9 1 3 ) .

1919 Rhinagrion macrocephala - MUNZ, Mem. Amer. ent. Soc., 3: 9. 1935 Rhinagrion macrocephalum - LIEFTINCK, Misc. Zool. Sum., 92-93: 6 (Sumatra). 1954 Rhinagrion macrocephalum - LIEFTINCK, Treubia, 22 Suppl.: 30 (Malaya, Sumatra).

Distribution: Malaysia, Sumatra.

164 mima

(KARSCH)

1891 Amphilestes mima KARSCH, Ent. Nachr., 17 (16): 242. Type male: Mus. Naturk., Berlin; typelocality: Sumatra. 1902 Amphilestes mima - LAIDLAW, Proc. Zool. Soc. London, 1902: 382 (description of female). 1924 Rhinagrion mima - LAIDLAW, J. Mal. Br. R. Asiatic Soc., 2: 302. 1935 Rhinagrion mima - LIEFTINCK, Misc. Zool. Sum., 92-93: 6. 1954 Rhinagrion mima - LIEFTINCK, Treubia, 22 Suppl.: 30 (Siam, Malaya, Sumatra).

Distribution: from Thailand to Sumatra. philippinum

(SELYS)

1882 Amphilestes philippina SELYS, Ann. Soc. Esp. nat. Hist., 11:21. Type male: Semper's Collection; type-locality: Bohol (Philippines). 1939 Rhinagrion philippinum - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 266 (redescription).

Distribution: Philippine Islands. tricolor

(KRÜGER)

1898 Amphilestes tricolor KRÜGER, Stettiner ent. Ztg, 59: 136. Type male: Mus. Naturk., Berlin; type-locality: Java. 1924 Rhinagrion tricolor - LAIDLAW, J. Mal. Br. R. Asiat Soc., 2: 303. 1934 Rhinagrion tricolor - LIEFTINCK, Treubia, 14 (4): 389 (type in Stettiner Museum). 1954 Rhinagrion tricolor - LIEFTINCK, Treubia, 22 Suppl.: 31 (Java).

Distribution: Java. viridatum

FRÄSER

1938 Rhinagrion viridatum FRASER, Proc. R. ent. Soc. London, (B) 7: 198. Type male: British Museum (Nat. Hist.), London; type-locality: Siam. 1954 Rhinagrion viridatum - LIEFTINCK, Treubia, 22 Suppl.: 30 [incorrect synonymy with mima (KARSCH, 1891)].

1966 Rhinagrion viridatum - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 218 (as valid taxa).

Distribution: Thailand.

Genus Sciotropis

RACENIS

1959 Sciotropis RACENIS, Acta biol. venezuel., 2: 356. Type species: Sciotropis cyclanthorum 1959 (original designation).

cyclanthorum

RACENIS,

RACENIS

1959 Sciotropis cyclanthorum RACENIS, Acta biol. venezuel., 2: 357 (as generotype of Sciotropis RACENIS, 1959). Type male: Recenis's Collection; type-locality: Venezuela.

Distribution: Venezuela.

Genus Trineuragrion Ris 1915 Trineuragrion Ris, Nova Caledonia, Zool., 2: 63. Type species: Trineuragrion percostale Ris, 1915 (original

designation).

165 percostale Ris 1915 Trineuragrion percostale Ris, Nova Caledonia, Zool., 2: 63 (as generotype of Trineuragrion Ris, 1915). Type male: Mus. Naturk., Berlin; type-locality: New Caledonia.

Distribution: New Caledonia. Subfamily Coryphagrioninae Very large. Clear wings, petiolate to about Ac. Anal vein and CuP well developed. Nodus over quarter length from wing base. Stigma rhomboidal, well braced. Two antenodals, Ac nearer 2nd than 1st antenodal, arc at 2nd. Dc trapezoid with lower distal angle acute. Postnodals 28 to 40. Supplementary sectors poorly developed, only short intercalaries from or beyond level of stigma. R2 rather sharply curved below stigma and almost angulated. 1962 Coryphagrioninae PINHEY, Publ. cult. Comp. Diam. Angola, 59: 101 (for single genus: Coryphagriori).

Genus Coryphagrion

MORTON

1924 Coryphagrion MORTON, Entomologist, 57: 217. Type species: Coryphagrion grandis MORTON, 1924 (monotypy). - 1962 Coryphagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 102.

grandis

MORTON

1924 Coryphagrion grandis MORTON, Entomologist, 57: 218. (as generotype of Coryphagrion MORTON, 1924). Type male: British Museum (Nat. Hist.), London; type-locality: East Africa. 1955 Coryphagrion grandis - FRASER, Entomologist's month. Mag., 88: 139. 1962 Coryphagrion grandis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 102.

Distribution: Tanzania. Subfamily Hypolestinae IRiii and Riv recessed; no accessory antenodals; postnodals aligned. Dc with slightly acute lower distal angle. 1938 Hypolestinae TILLYARD & FRASER, Austral. Zool., 9: 156 (in family Lestidae) (with genera: Hypolestes, Pseudolestes). - 1984 Hypolestinae - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 44 (in family Megapodagrionidae).

Genus Hypolestes

GUNDLACH

1888 Hypolestes GUNDLACH, Contr. Fauna cubana, Ent., 2: 216. Type species: Hypolestes trinitatis GUNDLACH, 1888 (monotypy). - 1891 Ortholestes CALVERT, Ent. News, 2: 199. Type species: Ortholestes clara CALVERT, 1891 (monotypy). - 1919 Lestes (Hypolestes) - CALVERT, Trans. Am. ent. Soc., 45: 347. - 1919 Ortholestes CALVERT, Trans. Am. ent. Soc., 45: 347 (proposed synonymy with Hypolestes GUNDLACH, 1888). - 1984 Hypolestes - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 44.

clara

(CALVERT)

1891 Ortholestes clara CALVERT, Ent. News, 2: 199 (as generotype of Ortholestes CALVERT, 1891). Type male: Calvert's Collection; type-locality: Jamaica.

166 1919 Lestes (Hypolestes) clara - CALVERT, Trans. Am. ent. Soc., 45: 347. 1984 Hypolestes clara - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 44.

Distribution: Jamaica. trinitatis

GUNDLACH

1888 Hypolestes trinitatis GUNDLACH, Contr. Fauna cubana, Ent., 2: 219 (as generotype of Hypolestes GUNDLACH, 1888). Type female: Instituto at Havana; type-locality: Cuba. 1891 Ortholestes abbotti CALVERT, Proc. Acad. Philad. nat. Hist., 5: 382. Type male: Amer. Ent. Soc. Coll.; type-locality: Haiti. 1919 Lestes (Hypolestes) trinitatis - CALVERT, Trans. Am. ent. Soc., 45: 347 (Trinidad, Cuba, Haiti). 1919 Ortholestes abbotti - CALVERT, Trans. Am. ent. Soc., 45: 347 (proposed synonymy with trinitatis

GUNDLACH, 1 8 8 8 ) .

1984 Hypolestes trinitatis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 44.

Distribution: Cuba, Haiti, Trinidad. Subfamily Megapodagriinae Medium sized. Longs legs. Wings petiolate to cubital cross vein; postnodals aligned; Dc with lower distal angle acute. IRiii arising at nodus, Riv 1 - 2 cells proximal to it. Anal vein arising level to proximal third of Dc. Male superior anal appendages forcipate. 1895 Megapodagriinae MCLACHLAN, Ann. Mag. nat. Hist., (6) 16: 19. Type-genus: Megapodagrion SELYS, 1885. - 1919 Megapodagrioninae - MUNZ, Mem. Amer. ent. Soc., 3: 50 (key to genera Thaumatoneura, Rhipidolestes, Podopteryx, Rhinagrion, Trineuragion, Argiolestes, Mesopodagrion, Megapodagrion, Allopodagrion, Argiolestes, Philogenia, Podolestes, Neurolestes, Nesolestes, Synlestes, Paraphlebia, Wahnesia, Metagrion, Chlorolestes, Allolestes, Neuragrion, Dimeragrion, Heteropodagrion, Mesagrion, Protolestes, Heteragrion, Chorismagrion, Perilestes). - 1938 Megapodagriinae - TILLYARD & FRASER, Austr. Zool., 9:160 (with genera Neurolestes, Neuragrion, Trineuragrion, Rhinagrion, Rhipidolestes, Podopteryx, Argiolestes, Podolestes, Megapodagrion, Mesopodagrion, Nesolestes, Allopodagrion, Philogenia, Paraphlebia, Allolestes, Dimeragrion, Heteropodagrion, Protolestes, Heteragrion, Mesagrion, Burmargiolestes, Caledargiolestes, Celebarargiolestes, Agriomorpha, Lestomima). - 1905 Podagrioninae JACOBSON & BIANKI, Pryam lozhnos. Ross, imp.: 836 (for Mesopodagrion, Podagrion). - 1959 Tatocnemidinae RACENIS, Acta biol. venezuel., 2: 339 (for Tatocnemis). - 1962 Megapodagriinae PINHEY, Publ. cult. Comp. Diam. Angola, 59: 97 (key to genera Neurolestes, Protolestes, Allolestes, Tatocnemis, Nesolestes). - 1984 Megapodagrioninae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 45 (with genera Arrhenocnemis, Megapodagrion, Mesopodagrion, Tatocnemis).

Genus Arrhenocnemis

LIEFTINCK

1933 Arrhenocnemis LIEFTINCK, Nova Guinea, 17:4. Type species: Arrhenocnemis sinuatipennis LIEFTINCK. 1933 (original designation).

amphidactylis

LIEFTINCK

1949 Arrhenocnemis amphidactylis LIEFTINCK, Nova Guinea (N. S.), 5: 92. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. sinuatipennis

LIEFTINCK

1933 Arrhenocnemis sinuatipennis LIEFTINCK, Nova Guinea, 17: 6 (as generotype of Arrhenocnemis LIEFTINCK, 1933). Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

167 1949 Arrhenocnemis sinuatipennis - LIEFTINCK, Nova Guinea (N. S.), 5: 91 (redescription).

Distribution: New Guinea.

Genus Megapodagrion

SELYS

1862 Podagrion SELYS, Synopsis des Agrionines, Add. 3: 11. Type species: Podagrion megalopus SELYS, 1862 (original d e s i g n a t i o n ) . - 1885 Megapodagrion SELYS, C . R . E n t . Belg., 29: 142.

Type species: Podagrion megalopus SELYS, 1862 (designated by KIRBY, 1890, Syn. Cat. Neur.Odon., London: 122). - 1890 Podagrion - KIRBY, Syn. Cat. Neur.-Odon., London: 122 (proposed synonymy with Megapodagrion SELYS, 1885). - 1910 Allopodagrion FÖRSTER, Wien, ent. Ztg., 29: 54. Type species: Podagrion contortum HÄGEN, 1862 (monotypy). - 1916 Megapodagrion- Ris, Arch. Naturg., (A) 82: 81. - 1959 Megapodagrion (Allopodagrion) - RACENIS, Acta biol. venezuel., 2: 346.

arachne

RACENIS

1959 Megapodagrion (Megapodagrion) arachne RACENIS, Acta biol. Venezuela., 2: 342. Type male: Racenis's Collection; type-locality: Venezuela.

Distribution: Venezuela. contortum

(HÄGEN)

1862 Podagrion contortum HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 13. Type male: Halle Museum; type-locality: Brazil. 1886 Megapodagrion contortum - SELYS, Mem. Cour., Acad. Belg., 38: 41. 1910 Allopodagrion contortum - FÖRSTER, Wien. ent. Ztg., 29: 54 (as generotype of Allopodagrion FÖRSTER, 1910.

Distribution: Brazil. curtum

SELYS

1886 Megapodagrion curtum SELYS, Mem. Cour. Acad. Belg., 38: 46. Type male: R. Mus., Brussels; type-locality: Ecuador.

Distribution: Ecuador. erinys Ris 1913 Megapodagrion erinys Ris, Mem. Soc. ent. Belg., 22: 60. Type male: unknown; type-locality: Argentina.

Distribution: Argentina. lepidum

RACENIS

1959 Megapodagrion (Allopodagrion) lepidum RACENIS, Acta biol. venezuel., 2: 346. Type: unknown; type-locality: Venezuela.

Distribution: Venezuela. macropus

(SELYS)

1862 Podagrion macropus SELYS, Synopsis des Agrionines, Add. 3:14. Type male: Selys's Collection; type-locality: Venezuela. 1886 Megapodagrion macropus - SELYS, Mem. Cour., Acad. Belg., 38: 45. 1916 Megapodagrion macropus - Ris, Arch. Naturg., (A) 82: 83 (Bolivia). 1948 Megapodagrion macropus - FRASER, Acta Zool. lilloana, 5: 61 (redescription).

Distribution: Venezuela, Bolivia.

168 megalopus

(SELYS)

1862 Podagrion megalopus SELYS, Synopsis des Agrionines, Add. 3: 12 (as generotype of Podagrion SELYS, 1862). Type male: Dale & Selys's Collection; type-locality: Para. 1885 Megapodagrion megalopus - SELYS, C.R. Soc. Ent. Belg., 2 9 : 142. 1 8 8 6 Megapodagrion megalopus - SELYS, Mem. Cour., Acad. Belg., 3 8 : 4 1 . 1890 Megapodagrion megalopus - KIRBY, Syn. Cat. Neur.-Odon. London: 122 (as generotype of Megapodagrion SELYS, 1885). 1946 Megapodagrion megalopus - FRASER, Trans. R. ent. Soc. London, 9 7 : 465 (Peru).

Distribution: Brazil, Peru. mercenarium (Hägen) 1869 Podagrion mercenarium Bogota. 1886

Megapodagrion

HAGEN,

mercenarium

Stettiner ent. Ztg., 30:261. Type male: unknown; type-locality:

- SELYS,

Mem. Cour. Acad. Belg.,

38: 47.

Distribution: Colombia. nebulosum

SELYS

1886 Megapodagrion nebulosum SELYS, Mem. Cour. Acad. Belg., 3 8 : 43. Type male: Selys's Collection; type-locality: Peru. 1890 Megapodagrion nebulosum - KIRBY, Syn. Cat. Neur.-Odon., London: 122. 1916 Megapodagrion nebulosum - Ris, Arch. Naturg., (A) 82: 84 (Peru, Bolivia). 1943 Megapodagrion nebulosum - SCHMIDT, Beitr. F a u n a Perus, 2: 227 (Peru, Bolivia).

Distribution: Peru, Bolivia. oscillans

(SELYS)

1862 Podagrion oscillans SELYS, Synopsis des Agrionines, Add. 3:14. Type male: R. Mus., Brussels; type-locality: Bogota. 1886 Megapodagrion oscillans - SELYS, Mem. Cour. Acad. Belg., 3 8 : 42.

Distribution: Colombia. setigerum

SELYS

1886 Megapodagrion setigerum SELYS, Mem. Cour. Acad. Belg., 3 8 : 42. Type male: Selys's Collection; type-locality: Ecuador. 1916 Megapodagrion setigerum - Ris, Arch. Naturg., (A) 82: 227 (Peru). 1943 Megapodagrion setigerum - SCHMIDT, Beitr. F a u n a Perus, 2: 227 (Peru). 1946 Megapodagrion setigerum - FRASER, Trans. R. ent. soc. London, 9 7 : 466 (Peru).

Distribution: Ecuador, Peru. temporale

(SELYS)

1862 Podagrion temporale

SELYS,

type-locality: Bogota. 1886 Megapodagrion temporale

Synopsis des Agrionines, Add. 3:16. Type male: Selys's Collection;

- SELYS,

Mem. Cour. Acad. Belg.,

38:

45.

Distribution: Colombia. venale

(HÄGEN)

1862 Podagrion venale H A G E N , in SELYS, Synopsis des Agrionines, Add. 3: 15. Type male: Hagen's Collection; type-locality: Venezuela. 1886 Megapodagrion venale - SELYS, Mem. Cour. Acad. Belg., 3 8 : 44. Distribution: V e n e z u e l a .

169 Genus Mesopodagrion

MCLACHLAN

1896 Mesopodagrion MCLACHLAN, Ann. Mag. nat. Hist., (6) 17: 372. Type species: tibetanum

MCLACHLAN,

1896 (monotypy). -

1 9 0 5 Mesopodagrion

-

Mesopodagrion

JACOBSON &

BIANKI,

Pryam. lozhnos. Ross, imp.: 836.

tibetanum

MCLACHLAN

1896 Mesopodagrion tibetanum MCLACHLAN, Ann. Mag. nat. Hist., (6) 17: 372 (as generotype of Mesopodagrion MCLACHLAN, 1896). Type male: not given, perhaps in British Museum (Nat. Hist.), London; type-locality: Tibet. 1905 Mesopodagrion tibetanum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 836 (China) (in subfamily Podagrioninae). 1928 Mesopodagrion tibetanum - MORTON, Trans, ent. Soc. London, 76: 112 (Yunnan). 1948 Mesopodagrion tibetanum - LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 7 (Burma). 1955 Mesopodagrion tibetanum - ASAHINA, Ent. Medd., 27 (3): 130 (Central China). 1964 Mesopodagrion tibetanum - SCHMIDT, Ent. Tidschr., 85: 147 (Burma).

Distribution: China to Burma. yachowensis

CHAO

1953 Mesopodagrion yechowensis CHAO, Acta ent. sinica, 3: 530. Type male: Peking Museum; typelocality: S.W. China.

Distribution: China. Genus Tatocnemis

KIRBY

1889 Tatocnemis KIRBY, Proc. Zool. Soc. London, 1889: 300. Type species: Tatocnemis malgassica KIRBY, 1889 (monotypy). - 1891 Prionocnemis (Nesocnemis) SELYS, C.R. Ent. Belg., 35: 401. Type species: Prionocnemis (Nesocnemis) sinuatipennis SELYS, 1891 (original designation). 1951 Tatocnemis- SCHMIDT, Mem. Inst. sei. Madagascar, 6Α: 155 (key to Madagascan species). - 1961 Tatocnemis - FRÄSER, Nat. malgache, 12: 65 (key to species). - 1962 Tatocnemis PINHEY, Publ. cult. Comp. Diam. Angola, 59:101. - 1962 Prionocnemis (Nesocnemis) - PINHEY, Publ. cult. C o m p . Diam. Angola, 59: 101 (proposed synonymy with Tatocnemis KIRBY, 1889).

crenulatipennis

FRÄSER

1952 Tatocnemis crenulatipennis FRÄSER, Mem. Inst. sei. Madagascar, (Ε) 1: 135. Type male: Mus. Nation. Hist, nat., Paris; type-locality: Madagascar. 1962 Tatocnemis crenulatipennis - PINHEY, Publ. cult. comp. Diam. Angola, 59: 101.

Distribution: Madagascar. denticularis

AGUESSE

1 9 8 4 T a t o c n e m i s d e n t i c u l a r i s AGUESSE, 1 9 6 8 -

l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f

the

World, Utrecht, 1: 45.

Distribution: Madagascar. emarginatipes

FRASER

1960 Tatocnemis emarginatipes FRASER, Nat. malgache, 12: 68. Type male: Fraser's Collection; typelocality: Madagascar. 1962 Tatocnemis emarginatipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 101.

Distribution: Madagascar.

170 malgassica

KIRBY

1889 Tatocnemis malgassica KIRBY, Proc. Zool. Soc. London, 1889: 302 (as cnemis KIRBY, 1889). Type male: British Museum (Nat. Hist.), London; gascar. 1951 Tatocnemis malgassica - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 157 with sinuatipennis (SELYS, 1891)]. 1961 Tatocnemis malgassica - PINHEY, Publ. cult. Comp. Diam. Angola, 59:

generotype of Tatotype-locality: M a d a [incorrect synonymy 101.

Distribution: Madagascar. mellisi

SCHMIDT

1951 Tatocnemis mellisi SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 156. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Tatocnemis mellisi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 101.

Distribution: Madagascar. micromalgassica 1984

AGUESSE

Tatocnemis micromalgassica the World, Utrecht, 1: 45.

AGUESSE, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of

Distribution: Madagascar. olsufieffi

SCHMIDT

1951 Tatocnemis olsufieffi SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 164. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Tatocnemis olsufieffi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 101.

Distribution: Madagascar. robinsoni

SCHMIDT

1951 Tatocnemis robinsoni SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 165. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Tatocnemis robinsoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 101.

Distribution: Madagascar. sinuatipennis

(SELYS)

1891 Prionocnemis (Nesocnemis) sinuatipennis SELYS, C.R. Ent. Belg., 35: 402 (as generotype of Nesocnemis Selys, 1891). Type: unknown; type-locality: Madagascar. 1951 Tatocnemis sinuatipennis - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 157. 1962 Tatocnemis sinuatipennis - PINHEY, Publ.cult. Comp. Diam. Angola, 59: 101.

Distribution: Madagascar.

Subfamily Philosininae Large, primitive (?) form. IRiii and Riv arise nearer to subnodus than to arc. Penis with slender, doubly curved recurrent lobes. CuP short. 1984 Philosininae auct. - loc. cit. Type-genus: Philosina

DAVIES & TOBIN,

Ris, 1917.

The Dragonflies of the World, Utrecht, 1: 45.

171 Genus Philosina Ris 1917 Philosina Ris, Ent. Tijdschr., 60: 185. Type species: Philosina designation).

buchi Ris, 1917 (original

buchi Ris 1917 Philosina buchi Ris, Ent. Tijdschr., 60: 189 (as generotype of Philosina Ris, 1917). Type male: Univ. zool. Mus., Kebenhavn; type-locality: China.

Distribution: China. Subfamily Thaumatoneurinae Large waterfall dweller. IRiii and Riv not recessed. Nodus close to wing base; extra antenodals; postnodals not aligned. Many extra cross veins and intercalaries, anal vein well developed. Larva with saccoid gills, quadrate mask. 1938 Thaumatoneurinae TILLYARD & FRASER, Austr. Zool., 9: 160 (in family Megapodagriidae) (for genus Thaumatoneura). - 1957 Thaumatoneurinae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 54 (with genera Thaumatoneura, Dysagrion, Phenacolestes). - 1984 Thaumatoneurinae - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46 (with genus Thaumatoneura).

Genus Thaumatoneura

MCLACHLAN

1897 Thaumatoneura MCLACHLAN, Ent. Mag., 33: 130. Type species: Thaumatoneura inopinata MCLACHLAN, 1897 (monotypy). - 1900 Thaumatoneura - MCLACHLAN, Ent. Mag., 36: 189. - 1916 Thaumatoneura - Ris, Arch. Naturg., (A) 82: 75.

inopinata

MCLACHLAN

1897 Thaumatoneura inopinata MCLACHLAN, Ent. Mag., 33: 131 (as generotype of Thaumatoneura MCLACHLAN, 1897). Type male: British Museum (Nat. Hist.), London; type-locality: Costa Rica. 1897 Thaumatoneura inopinata - in Zoological Record, 1897: 288 type-locality: China or Japan?). 1900 Thaumatoneura inopinata - MCLACHLAN, Ent. Mag., 36: 189 (habitat P a n a m a not Asia).

Distribution: Central America. pellucida

CALVERT

1904 Thaumatoneura pallucida CALVERT, Ent. News, 15: 216. Type: unknown; type-locality: Costa Rica. 1984 Thaumatoneura inopinata f. pellucida - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Costa Rica. Family Perilestidae Small slender species with long abdomen. Arc distal to 2nd antenodal; Riv arising from subnodus; IRiii arising 3 - 5 cells distal to nodus; basal part of radius thickened; Dc impinges on posterior border of wing; no oblique cross vein between Riii and IRiii. 1938 Perilestidae TILLYARD & FRASER, Austr. Zool., 9: 155. - 1957 Perilestidae - FRASER, Ent. News, 62: 62. - 1962 Perilestidae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 89. - 1977 Perilestidae - BELYSHEV & KHARITOV, N a u k a , Novosibirsk: 270 (revision, key to Boreal genera). - 1984 Perilestidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

172

Genus Nubiolestes 1943

FRÄSER

Eolestes SCHMIDT, Mitt. Dtsch. ent. Ges., 11: 1 0 2 . Type species: Eolestes diotima SCHMIDT, 1 9 4 3 (original designation). - 1 9 4 4 Eolestes FRÄSER ( S C H M I D T MS), Proc. R. ent. Soc. London, (B) 13: 65 (as gen.nov.). - 1945 Nubiolestes FRÄSER, Entomologist month. Mag., 81: 9 (new name for Eolestes FRÄSER, 1 9 4 4 (nec SCHMIDT, 1 9 4 3 ) , (a junior primary homonymy of Eolestes COCKERELL, 1 9 4 0 ) . - 1 9 6 2 Nubiolestes - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 8 9 . 1 9 8 4 Camerunolestes SCHMIDT, 1 9 5 8 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 4 6 (as syn. with. Nubiolestes FRÄSER, 1 9 4 5 ) .

diotima

(SCHMIDT)

1943 Eolestes diotima SCHMIDT, Mitt. Dtsch. ent. Ges., 11: 107 (as generotype of Eolestes SCHMIDT, 1943). Type male: H a m b u r g Museum; type-locality: Cameroon. 1944 Eolestes diotima - FRÄSER, Proc. R. ent. Soc. London, (B) 13: 66. 1945 Nubiolestes diotima - FRÄSER, Entomologist month. Mag., 81: 9 (as generotype of Nubiolestes FRÄSER, 1 9 4 5 ) .

1962 Nubiolestes

diotima - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 89.

Distribution: Cameroon. Genus Perilestes

HAGEN

1862 Perilestes HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 29. Type species: fragilis, H A G E N , 1862 (monotypy).

attenuatus

Perilestes

SELYS

1886 Perilestes attenuatus SELYS, Mem. Cour. Acad. Belg., 38: 68. Type male: R. Mus., Brussels; type-locality: Santarem.

Distribution: Brazil. bispinus

KIMMINS

1958 Perilestes bispinus KIMMINS, Bull. Brit. Mus. (Nat. Hist.), Ent., 7: 353. Type male: British Museum (Nat. Hist.), London; type-locality: Brazil.

Distribution: Brazil. fragilis

HAGEN

1862 Perilestes fragilis HAGEN, in SELYS, Synopsis des Agrionines, Add. 3: 30 (as generotype of Perilestes HÄGEN, 1862). Type female: Halle Museum; type-locality: Brazil. 1924 Perilestes fragilis - WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 27 (redescription).

Distribution: Brazil. gracillimus

KENNEDY

1941 Perilestes gracillimus KENNEDY, Ann. ent. Soc. Amer., 34: 669. Type male: Kennedy's Collection; type-locality: Peru.

Distribution: Peru. kahli

WILLIAMSON & WILLIAMSON

1924 Perilestes kahli WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 24. Type male: A n n A r b o r Museum; type-locality: Rio N a p o . 1937 Perilestes kahli - KENNEDY, Ann. Ent. Soc. Amer., 30: 423.

Distribution: Brazil.

173 minor

WILLIAMSON & WILLIAMSON

1924 Perilestes minor WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 34. Type male: Ann A r b o r Museum; type-locality: Brazil.

Distribution: Brazil. solutus

WILLIAMSON & WILLIAMSON

1924 Perilestes solutus WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 30. Type male: Ann Arbor Museum; type-locality: Brazil.

Distribution: Brazil. Genus Perissolestes

KENNEDY

1941 Perissolestes KENNEDY, Ann. ent. Soc. Amer., 34: 660. Type species: Perilestes remotus WILLIAMSON & WILLIAMSON, 1924 (original designation).

aculeatus

(KENNEDY)

1941 Perilestes aculeatus KENNEDY, Ann. ent. Soc., Amer., 43: 684. Type male: A n n Arbor Museum; type-locality: Brazil. 1984 Perissolestes aculeatus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Brazil. castor

(KENNEDY)

1937 Perilestes castor KENNEDY, Ann. ent. Soc. Amer., 30: 411. Type male: A n n A r b o r Museum; type-locality: Ecuador. 1984 Perisolestes castor - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Ecuador. cornutus

(SELYS)

1886 Perilestes cornutus SELYS, Bull. Acad. r. Belg., (3) 38: 68. Type male: Selys's Collection; typelocality: Amazons. 1943 Perilestes cornutus - SCHMIDT, Beitr. F a u n a Perus, 2: 225. 1984 Perissolestes cornutus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Brazil, Peru. guianensis

(WILLIAMSON & WILLIAMSON)

1924 Perilestes guianensis WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 12. Type male: A n n A r b o r Museum; type-locality: Guyana. 1984 Perissolestes guianensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Guyana. klugi

KENNEDY

1941 Perissolestes klugi KENNEDY, Ann. ent. Soc. Amer., 34: 673. Type male: Ann. A r b o r Museum; type-locality: Peru.

Distribution: Peru. magdalenae

(WILLIAMSON & WILLIAMSON)

1924 Perilestes magdalenae WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 17. Type male: A n n A r b o r Museum; type-locality: Colombia. 1984 Perissolestes magdalenae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Colombia.

174 paprzyckii

KENNEDY

1941 Perissolestes paprzyckii KENNEDY, Ann. ent. Soc. Amer., 34: 852. Type male: Michigan Univ.; type-locality: Peru.

Distribution: Peru. pollux

(KENNEDY)

1937 Perilestes pollux KENNEDY, Ann. ent. Soc. Amer., 30: 418. Type male: Michigan Univ.; typelocality: Ecuador. 1984 Perissolestes pollux - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46.

Distribution: Ecuador. remotus

(WILLIAMSON & WILLIAMSON)

1924 Perilestes remotus WILLIAMSON & WILLIAMSON, Univ. Mich. Mus. Zool. Misc. Publ., 14: 14. Type male: A n n A r b o r Museum; type-locality: Canal Zone. 1941 Perissolestes remotus - KENNEDY, Ann. ent. Soc. Amer., 34: 660 (as generotype of Perissolestes KENNEDY, 1 9 4 1 ) .

Distribution: Panama, Colombia. remus

KENNEDY

1941 Perissolestes remus KENNEDY, Ann. ent. Soc. Amer., 34: 682. Type male: Michigan Univ.; type-locality: Ecuador.

Distribution: Ecuador. romulus

KENNEDY

1941 Perissolestes romulus KENNEDY, A n n . ent. Soc. Amer., 34: 676. Type male: Michigan Univ.; type-locality: Peru.

Distribution: Peru.

Family Pseudolestidae IRiii and Riv recessed (incompletely in Rhipidolestes). No accessory antenodals; postnodals aligned (not in Pseudolestes, which has hind wing short and metallic). Dc with slightly acute lower distal angle. 1938 Pseudolestidae TILLYARD & FRASER, Austr. Zool., 9: 150. - 1977 Pseudolestidae - BELYSHEV & KHARITONOV, N a u k a , Novosibirsk: 273 (key to genera). - 1984 Pseudolestidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 46 (with genera Lestomima, Pseudolestes, Rhipidolestes). - 1 9 5 7 Pseudolestinae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957:62. Type-genus: Pseudolestes KIRBY, 1900 (with genera Pseudolestes, Rimanella, Hypolestes, Lestomima, Rhipidolestes, Pseudolestes).

Genus Lestomima MAY 1933 Lestomima MAY, Senckenbergiana, 15: 343. Type species: Lestomima (original designation).

flavostigma

MAY, 1933

flavostigma MAY 1933 Lestomima flavostigma MAY, Senckenbergiana, 15: 345 (as generotype of Lestomima 1933). Type: unknown; type-locality: China.

Distribution: China.

MAY,

175 Genus Pseudolestes

KIRBY

1900 Pseudolestes KIRBY, A n n . Mag. nat. Hist., (3) 5: 537. Type species: Pseudolestes KIRBY, 1 9 0 0 ( o r i g i n a l d e s i g n a t i o n ) . - 1 9 0 5 Pseudolestes

mirabilis

- JACOBSON & BIANKI, P r y a m . l o z h n o s .

Ross, imp.: 812.

mirabilis

KIRBY

1900 Pseudolestes mirabilis KIRBY, Ann. Mag. nat. Hist., (3) 5: 538 (as generotype of Pseudolestes KIRBY, 1900). Type male: British Museum (Nat. Hist.), London; type-locality: Hainan. 1905 Pseudolestes mirabilis - JACOBSON & BIANKI, Pyram. lozhnos. Ross, imp.: 812.

Distribution: Hainan. Genus Rhipidolestes Ris 1912 Rhipidolestes Ris, Suppl. ent. Berlin, 1: 57. Type species: Rhipidolestes aculeatus Ris, 1912 (monotypy). - 1928 Taolestes NEEDHAM, Bull. Pekin Soc. nat. Hist., 2: 12. Type species: Taolestes nectans NEEDHAM, 1928 (original designation). - 1 9 3 1 Rhipidolestes - SCHMIDT, Konowia, 10 (3): 178 (key to Chinese species).

aculeatus aculeatus Ris 1912 Rhipidolestes aculeata Ris, Suppl. ent. Berlin, 1: 59 (as generotype of Rhipidolestes Type male: Ris's Collection; type-locality: Formosa.

Ris, 1912).

Distribution: Taiwan, aculeatus yakusimensis

ASAHINA

1951 Rhipidolestes aculeata yakusimensis ASAHINA, Kontyü, 19:15. Type male: Asahina's Collection; type-locality: Ryukyu.

Distribution: Ryukyu Island. apicatus

NAVÄS

1934 Rhipidolestes apicatus NAVÄS, Mus. Heude Notes ent. chinoise, 2 (1): 4. Type male: not given; type-locality: China. 1 9 5 6 Rhipidolestes

apicatus

- ASAHINA, E n t . M e d d . , 2 7 ( 4 - 5 ) : 2 1 1 .

Distribution: China. jucundus

LIEFTINCK

1948 Rhipidolestes jucundus LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 9. Type male: Naturhist. Riksmus., Stockholm; type-locality: Burma. 1984 Rhipidolestes jucundus - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:47 (China).

Distribution: Burma, China, malaisei

LIEFTINCK

1948 Rhipidolestes malaisei LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 7. Type male: Naturhist. Riksmus., Stockholm; type-locality: Burma.

Distribution: Burma. nectans

(NEEDHAM)

1928 Taolestes nectans NEEDHAM, Bull. Pekin Soc. nat. Hist., 2: 12 (as generotype of Taolestes NEEDHAM, 1928). Type female: Cornell University; type-locality: Hangchow. 1931 Rhipidolestes bidens SCHMIDT, Konowia, 10 (3): 180. Type male: Tsai-Tsche-kiang's Collection; type-locality: China.

176 1956 Rhipidolestes nectans - ASAHINA, Ent. Medd., 27 (4-5): 207. 1956 Rhipidolestes bidens - ASAHINA, Ent. Medd., 27 (4-5): 207 [proposed synonymy with nectans (NEEDHAM, 1928)].

Distribution: Central China. okinawanus ASAHINA 1951 Rhipidolestes okinawana ASAHINA, Kontyü, 19: 15. Type male: Hokkaido University; typelocality: Okinawa. 1961 Rhipidolestes okinawanus - ASAHINA, Insecta matsum., 24: 58. 1984 Rhipidolestes apicalis MATSUMURA, 1913 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 47.

Distribution: Ryukyu Islands. rubripes (NAVAS) 1936 Taolestes rubripes NAVÄS, MUS. Heude Notes ent. chinoise, 3 (4): 44. Type male: not given; type-locality: China. 1984 Rhipidolestes rubripes - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 47.

Distribution: China. t r u n c a t i d e n s SCHMIDT 1931 Rhipidolestes truncatidens SCHMIDT, Konowia, 10 (3): 181. Type male: Mus. Naturk., Berlin; type-locality: Kanton.

Distribution: China. Superfamily Hemiphlebioidea Small Zygoptera, wings with 2 antenodals, postnodals not in line with the cross vein below them. Anal vein separate from wing border from its base. Dc of fore wing open at base; arc distal to second antenodal. 1938 Hemiphlebioidea TILLYARD & FRASER, A u s t r . Z o o l . , 9: 150. - 1984 Hemiphlebioidea

- DA VIES

& TOBIN, The Dragonflies of the World, Utrecht, 1: 48.

Family Hemiphlebiidae Small and slender, metallic green. Wings broadening rather than petiolate; gizzard with likely archaic features. 1938 Hemiphlebiidae TILLYARD & FRASER, Austr. Zool., 9: 150 (with single genus Hemiphlebia SELYS, 1869). - 1957 Hemiphlebiidae

- FRASER, R . Z o o l . S o c . N . S . Wales, 1957: 51. - 1984

Hemiphlebiidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 48.

Genus Hemiphlebia SELYS 1869 Hemiphlebia SELYS. C. R. Soc. Ent. Belg., 11: 129. Type species: Hemiphlebia mirabilis SELYS, 1869 (monotypy). - 1877 Hemiphlebia - SELYS, Bull. Acad. r. Belg., (2) 43: 156. - 1938 Hemiphlebia

- TILLYARD & FRASER, A u s t r . Z o o l . , 9: 150. - 1957 Hemiphlebia

FRASER, R . Z o o l .

Soc. N.S. Wales, 1957: 51. mirabilis SELYS 1869 Hemiphlebia mirabilis SELYS, C. R. Soc. Ent. Belg., 11: 122 (as generotype of Hemiphlebia SELYS, 1869). Type male: unknown; type-locality: Queensland. 1877 Hemiphlebia

mirabilis

- SELYS, Bull. A c a d . r. Belg., (2) 43: 158 ( r e d e s c r i p t i o n ) .

177 1901 1913 1955 1984

Hemiphlebia Hemiphlebia Hemiphlebia Hemiphlebia Australia).

mirabilis mirabilis mirabilis mirabilis

-

MARTIN, Mem. Soc. zool. France, 14: 248 (Victoria, Queensland). TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 463 (redescription). FRASER, Entomologist's month. Mag., 91: 110 (redescription). DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 48 (S.E.

Distribution: Australia. Superfamily Coenagrionoidea Wings petiolate, usually hyaline and with 2 antenodals; arc approx. in line with distal primary antenodals; postnodals usually in line with cross veins below them; Dc closed at base and not crossed; anal vein fused at base with posterior border of wing; IRiii & Riv arise nearer to nodus than to arc. First thoracic suture reaching only to spiracle. Male genital hamules quadrate. Larve with 3 leaf-like caudal appendages. 1921 Agrionidae Ris, Annls S. Afr. Mus., 18: 266 (with key to South African genera Lestes, Chlorolestes, Metacnemis, Allocnemis, Chlorocnemis, Disparoneura, Pseudagrion, Ceriagrion, Agriocnemis, Enallagma, Ischnura). - 1926 Coenagrioidea TILL YARD, Ins. Austr. and New Zeal.: 80. - 1957 Coenagrioidea - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 39 (with families Permagriidae, Pseudostigmatidae, Megapodagriidae, Platystictidae, Protoneuridae, Platycnemididae, Coenagriidae) (key to families). - 1938 Coenagrioidea - TILLYARD & FRASER, Austr. Zool., 9: 152 (key to families: Permagriidae, Synlestidae, Lestidae, Lestoideidae, Pseudostigmatidae, Megapodagriidae, Platystictidae, Protoneuridae, Platycnemididae, Coenagriidae). 1959 Coenagrioidea - RACENIS, Senckenb. biol., 40: 55. - 1962 Coenagrioidea - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 96 (key to African families: Megapodagriidae, Protoneuridae, Platycnemididae, Coenagriidae). - 1965 Agrionoidea BENEDEK, Folia ent.hung., 18: 408 (new n a m e f o r C o e n a g r i o i d e a TILLYARD, 1938). - 1984 Coenagrioidea

- DAVIES & TOBIN, T h e D r a -

gonflies of the World, Utrecht, 1: 49 (with families Coenagrionidae, Isostictidae, Platycnemididae, Platystictidae, Protoneuridae, Pseudostigmatidae).

Family Coenagrionidae Generally small Zygoptera. Anal vein and CuP present, arc at level of 2nd antenodal; Dc short, trapezoidal with distal angle sharply acute. MA and IRiii zig-zagged. 1815 Agrionidae LEACH in BREWSTER, Edinburgh encycl., 9 (1): 176 (emendation). - 1840 Heteropteroides SELYS, Monogr. Lib. Eur. - 1876 Rhomboides BUCHECKER, Syst. Ent. Odonata, 1: 3 - 1890 Coenagrionidae KIRBY, Syn. Cat. Neur.-Odon. London: 119 (new name for Agrionidae LEACH, 1815). - 1905 Agrionidae - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 812 (with subfamilies Agrioninae, Agriocneminae, Platycneminae, Podagrioninae, Protoneurinae). - 1910 Coenagrionidae - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 34 (with subfamilies Lestinae, Pseudostigmatinae, Coenagrioninae). - 1926 Coenagrionidae - TILLYARD, Ins. Austr. and New Zeal.: 80. - 1938 Coenagrionidae - TILLYARD & FRASER, Austr. Zool., 9: 165 (with subfamilies Agriinae, Coenagriinae). - 1948 Puellidae AKRAMOVSKI, Zool. Zborn., Moscow, 5: 150 (new name for Coenagrionidae). - 1957 Coenagriidae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 47 (with subfamilies Amphicneminae, Pseudagriinae, Coenagriinae, Ischnuriinae, Agriocneminae, Agriinae). - 1962 Coenagriidae - LIEFTINCK, Insecta of Micronesia, 5 (1): 16 (key to Micronesian genera). - 1962 Coenagriidea - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 115. - 1984 Coenagrionidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 49 (with subfamilies Agriocneminae, Agriinae, Coenagrioninae, Ischnurinae, Leptobasinae, Pseudagrioninae).

178 Subfamily Agriocnemidinae Very small forms, petiolation stops short of Ac. Arc widely distal to distal antenodal. Pterostigma often differs between fore and hind wings. No vulvar spine. 1957 Agriocneminae FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 49. Type-genus: Agriocnemis SELYS, 1877 (with genera Agriocnemis, Argiocnemis, Mortonagrion, Ceratura). - 1977 Agriocneminae - BELYSHEV & KHARITONOV, Nauka, Novosibirsk: 229 (revision, key to Boreal genera). 1984 Agriocnemis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 49 (with genera Agriocnemis, Argiocnemis, Austrocnemis, Coenagriocnemis, Mortonagrion).

Genus Agriocnemis

SELYS

1877 Argiocnemis (Agriocnemis) SELYS, Bull. Acad. r. Belg., (2) 43: 142. Type species: Agrion pygmaeum RAMBUR, 1842 (monotypy). - 1890 Agriocnemis - KIRBY, Syn. Cat. Neur.-Odon. Lond o n : 158 (as b y SELYS, POLLEN & VAN DAM, F a u n e M a d . , Ins.: 2 4 , 1 8 6 9 ) . - 1905

Agriocnemis

- JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 831 (key to species). - 1919 Agriocnemis - LAIDLAW, Ree. Indian Mus., 16: 178 (key to species). - 1920 Argyrocnemis KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type species: Agriocnemis argentea TILLYARD, 1906 (original designation). - 1921 Agriocnemis - Ris, Annls S. Afr. Mus., 18: 335. - 1923 Agriocnemis - TILLYARD, Trans, ent. Soc. London, 1923: 335 (key to species). - 1931 Agriocnemis - CHUJO, Trans, nat. Hist. Soc. Formosa, 21 (112): 32. - 1932 Agriocnemis - LIEFTINCK, Nova Guinea, 15 (5): 103. - 1951 Agriocnemis - SCHMIDT, Arq. Mus. Bocage, 20:131 (key to African species). - 1962 Agriocnemis - LIEFTINCK, Insects of Micronesia, 5 (1): 41. - 1962 Agriocnemis- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 138. - 1974 Agriocnemis - PINHEY, Occ. Pap. natn. Mus. Rhod. (Nat. Sei.), 5 (4): 194 (key to African species).

aderces

LIEFTINCK

1932 Agriocnemis aderces LIEFTINCK, Nova Guinea, 15 (5): 114. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. aligulae

PINHEY

1974 Agriocnemis aligulae PINHEY, OCC. Pap. natn. Mus. Rhod. (Nat.Sei.), 5 (4): 235. Type male: Pinhey's Collection; type-locality: Zaire.

Distribution: Zaire, angolensis angolensis

LONGFIELD

1947 Agriocnemis angolensis LONGFIELD, Arq. Mus. Bocage, 16: 15. Type male: British Museum (Nat. Hist.), London; type-locality: Angola. 1962 Agriocnemis angolensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 139. 1963 Agriocnemis angolensis - BALINSKY, J. ent. Soc. S. Afr., 26: 252 (redescription). 1974 Agriocnemis angolensis angolensis - PINHEY, Occ. Pap. natn. Mus. Rhod. (Nat.Sei.), 5 (4): 186.

Distribution: Angola. angolensis spatulae

PINHEY

1974 Agriocnemis angolensis spatulae PINHEY, OCC. Pap. natn. Mus. Rhod. (Nat.Sci.), 5 (4): 256. Type male: Pinhey's Collection; type-locality: Zambia.

Distribution: Zambia.

179

angustirami

PINHEY

1974 Agriocnemis angustirami PINHEY, OCC. Pap. natn. Mus. R h o d . (Nat.Sei.), 5 (4): 274. Type male: Pinhey's Collection; type-locality: Liberia.

Distribution: Liberia.

argentea

TILLYARD

1906 Agriocnemis argentea TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 192. Type male: not given; type-locality: Queensland. 1913 Agriocnemis argentea - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 461. 1920 Argyrocnemis argentea - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Argyrocnemis

KENNEDY,

1920).

Distribution: Australia.

australis

SELYS

1877 Argiocnemis (Agriocnemis) australis SELYS, Bull. Acad. r. Belg., (2) 43: 155. Type female: McLachlan's Collection; type-locality: Queensland. 1890 Agriocnemis australis - KIRBY, Syn. Cat. Neur.-Odon. London: 159. 1901 Agriocnemis australis - MARTIN, Mem. Soc. zool. France, 14: 248.

Distribution: Australia.

binocellata

FRASER

1922 Agriocnemis binocellata FRÄSER, J. nat. Hist. Soc. Siam, 4: 233. Type male: unknown; typelocality: Bangkok.

Distribution: Thailand.

carmelita

SELYS

1877 Argiocnemis (Agriocnemis) carmelita SELYS, Bull. Acad. r. Belg., (2) 43: 155. Type female: Selys's Collection; type-locality: A n n a m . 1890 Agriocnemis carmelita - KIRBY, Syn. Cat. Neur.-Odon. London: 159.

Distribution: Annam.

clauseni

FRASER

1923 Agriocnemis clauseni FRASER, Mem. Dept. Agr. India, 7: 53. Type male: British Museum (Nat. Hist.), London; type-locality: Assam.

Distribution: Southeast Asia.

corbeti

KUMAR

&

PRASAD

1978 Agriocnemis corbeti K U M A R & PRASAD, J. Bombay nat. Hist. Soc. Nation. Coll. zool. Surv. India, Calcutta; type-locality: India.

Distribution: India.

75

(1): 174. Type male:

180 dabreui

FRÄSER

1919 Agriocnemis d'abreui FRÄSER, Ree. Indian Mus., 16: 454. Type female: Pusa Collection; typelocality: India. 1931 Agriocnemis dabreui - LAIDLAW, J. Fed. Mai. States Mus., 16: 201 (Pahang). 1948 Agriocnemis dabreui - LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 16 (Tanasserim, India, Assam, Siam, Malay Peninsula). 1954 Agriocnemis dabreui - LIEFTINCK, Treubia, 22 Suppl.: 71 (Siam, Malaya).

Distribution: Southeast Asia. dobsoni

FRÄSER

1954 Agriocnemis dobsoni FRÄSER, Ann. Mag. nat. Hist., (12) 7: 147. Type male: British Museum (Nat. Hist.), London; type-locality: Queensland.

Distribution: Australia, exilis

SELYS 1872 Argiocnemis (Agriocnemis) exilis SELYS, Revue Zool., (2) 23:181. Type series in Mus. Brussels, but type not designated; type-locality: Madagascar. 1877 Argiocnemis (Agriocnemis) exilis - SELYS, Bull. Acad. r. Belg., (2) 43: 154 (Zanzibar, Madagascar). 1890 Agriocnemis exilis - KIRBY, Syn. Cat. Neur.-Odon., London: 159. 1921 Agriocnemis exilis - Ris, Annls S. Afr. Mus., 18: 335 (Beira). 1962 Agriocnemis exilis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 139 (Continental Africa, Madagascar, Mauritius, Reunion). 1963 Agriocnemis pinheyi BALINSKY, J. ent. Soc. S. Afr., 26: 247. Type: unknown; type-locality: Transvaal. 1974 Agriocnemis exilis - PINHEY, Occ. Pap. natn. Mus. Rhod. (Nat.Sci.), 5 (4): 185. 1984 Agriocnemis pinheyi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 49 (as syn. w i t h exilis

SELYS, 1 8 7 2 ) .

1988 Agriocnemis exilis - JACQUEMIN, Notul. odonatol., 1 (1): 14 (Mascarene Island).

Distribution: continental Africa, Madagascar, Mauritius, Reunion. exsudans

SELYS

1877 Argiocnemis (Agriocnemis) exsudans SELYS, Bull. Acad. r. Belg., (2) 43:148. Type male: Selys's Collection; type-locality: New Caledonia. 1890 Agriocnemis exsudans - KIRBY, Syn. Cat.Neur.-Odon., London: 158. 1913 Agriocnemis exsudans - TILL YARD, Proc. Linn. Soc. N.S. Wales, 37: 461 (New Hebrides). 1923 Agriocnemis exsudans - TILLYARD, Trans, ent. Soc. London, 1923: 335 (New Hebrides, Fiji, New Caledonia).

Distribution: New Caledonia, New Hebrides, Fiji. falcifera falcifera

PINHEY

1958 Agriocnemis falcifera PINHEY, J. ent. Soc. S. Afr., 22: 465. Type male: Transvaal Museum; type-locality: Natal. 1962 Agriocnemis falcifera - PINHEY, Publ. cult. Comp. Diam Angola, 59: 139 (Zululand, Natal). 1974 Agriocnemis falcifera falcifera - PINHEY, OCC. Pap. natn. Mus. Rhod. (Nat.Sci.) 5 (4): 213.

Distribution: South Africa. falcifera transvaalica

PINHEY

1974 Agriocnemis falcifera transvaalica PINHEY, OCC. Pap. natn. Mus. Rhod. (Nat.Sci.) 5 (4): 213. Type male: Pinhey's Collection; type-locality: Transvaal.

Distribution: South Africa.

181 femina femina

(BRAUER)

1868 Agrion (Ischnura) femina BRAUER, Abh. Zool.-Bot. Ges. Wien, 18: 554. Type male: Mus. Naturk., Berlin; type-locality: Luzon. 1877 Argiocnemis (Agriocnemis) incisa HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 43: 149. Type male: Mus. Naturk., Berlin; type-locality: Nicobar. 1877 Argiocnemis (Agriocnemis) incisa race pulverulans SELYS, Bull. Acad. r. Belg., (2) 43: 150. Male from Celebes. 1877 Argiocnemis (Agriocnemis) materna HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 43: 151. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Sumatra. 1890 Agriocnemis femina - KIRBY, Syn. Cat. Neur.-Odon. London: 158. 1890 Argiocnemis (Agriocnemis) incisa - KIRBY, Syn. Cat. Neur.-Odon. London: 158 [proposed synonymy with femina femina ( B R A U E R , 1 8 6 8 ) ] . 1890 Agriocnemis materna - Kirby, Syn. Cat. Neur.-Odon. London: 159. 1916 Agriocnemis femina - Ris, Suppl. ent. Berlin, 5: 21. (Formosa, China, Borneo, Java, Philippines, Lombok). 1916 Argiocnemis (Agriocnemis) materna - Ris, Suppl. ent. Berlin, 5: 21 [proposed synonymy with femina femina (BRAUER, 1868)]. 1927 Agriocnemis femina - Ris, Zool. Meded., Leiden, 10 (1): 22 (Sumatra). 1931 Agriocnemis femina - CHUJO, Trans, nat. Hist. Soc. Formosa, 21 (112): 36. 1934 Agriocnemis femina - SCHMIDT, Arch. Hydrob. Suppl., 13: 339 (Java, Sumatra, Bali). 1934 Agriocnemis femina - LIEFTINCK, Treubia, 14 (4): 401. 1936 Agriocnemis femina - LIEFTINCK, Revue suisse Zool., 43 (5): 136 (Flores, Timor). 1939 Agriocnemis femina - NEEDHAM & GYGER, Philipp J. Sei., 70 (3): 278. 1954 Agriocnemis femina - LIEFTINCK, Treubia, 22 Suppl.: 72 (Siam, Krakatau, Sumatra, Java, Borneo, Bali, Mentawei Island). 1955 Agriocnemis femina - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 77 (Ceylon). 1962 Agriocnemis femina femina - LIEFTINCK, Insects of Micronesia, 5 (1): 43 (from India and Ceylon to the Bismarck and North Australia).

Distribution: Oriental region, femina oryzae

LIEFTINCK

1962 Agriocnemis femina oryzae LIEFTINCK, Insects of Micronesia, 5 (1): 44. Type male: US natn. Mus., Washington; type-locality: Okinawa.

Distribution: Taiwan, Hainan, East China, Ryukyu Island, forcipata LE ROI Agriocnemis forcipata L E R O I (nec SJÖSTEDT, 1 9 1 7 ) , Ergeb. zweit. Deutsch. Zentr. Afr. Exp., 1: 341. Type male: Bonn Museum; type-locality: Bahr el Ghazal. 1917 Agriocnemis forcipata SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 22. Type male: Naturh. Rijksmus., Stockholm; type-locality: Congo. 1 9 2 4 Agriocnemis flavilabris C A M P I O N , Entomologist, 5 7 : 1 8 (new name for forcipata SJÖSTEDT, 1917, a junior secondary homonymy of forcipata LE ROI, 1915). 1962 Agriocnemis forcipata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 139. 1962 Agriocnemis flavilabris - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 139 (proposed synonymy with forcipata L E R O I , 1 9 1 5 ) . 1915

Distribution: Central Africa, gratiosa

GERSTAECKER

1891 Agriocnemis gratiosa GERSTAECKER, Jb. Hamb., 9: 190. Type: unknown (lost); type-locality: Zanzibar. 1902 Agriocnemis consimilis GRUNBERG, Senck. Ges. Naturf. Frde. Berlin, 1902: 230. Type male: Mus. Naturk., Berlin; type-locality: East Africa.

182 1958 Agriocnemis gratiosa - P I N H E Y , Stuttg. Beitr. Naturk., 1 9 5 8 (10): 2 (Tanganyika, Belgian Congo, N. Rhodesia, Natal, Madagascar). 1962 Agriocnemis gratiosa - P I N H E Y , Publ. cult. Comp. Diam. Angola, 5 9 : 139 (Belgian Congo, Tanganyika, Zanzibar, Kenya N. Rhodesia, Natal, Madagascar).

Distribution: tropical Africa, Madagascar. interrupta

FRASER

1927 Agriocnemis interrupta F R A S E R , Ins. Samoa, 7: 22. Type male: British Museum (Nat. Hist.), London; type-locality: Samoa. 1953 Agriocnemis interrupta-FRASER, Proc. R. ent. Soc. London, (B) 22:125 (description of female).

Distribution: Samoa. inversa

KARSCH

Agriocnemis inversa K A R S C H , Ent. Nachr., 2 5 ( 2 4 ) : 3 8 0 . Type male: Mus. Naturk., Berlin; type-locality: Victoria-Nyassa. 1962 Agriocnemis inversa - P I N H E Y , Publ. cult. Comp. Diam. Angola, 5 9 : 139 (Belgian Congo, U g a n d a , Sudan). 1899

Distribution: tropical Africa. kevalensis 1984

PETERS

Agriocnemis kevalensis Utrecht, 1: 50.

PETERS, 1 9 8 1

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

D A V I E S & TOBIN,

The Dragonflies of the World,

Distribution: India. kunjina

WATSON

1984 Agriocnemis kunjina Utrecht, 1: 50.

WATSON,

1969 - loc. cit.

Distribution: N.W. Australia. lacteola

SELYS

1877 Argiocnemis Agriocnemis 1890 Agriocnemis 1 9 0 5 Agriocnemis 1919 Agriocnemis 1965 Agriocnemis

(Agriocnemis) lacteola SELYS, Bull. Acad. r. Belg., ( 2 ) 43: 144 (as generotype of SELYS, 1877). Type male: Selys's collection; type-locality: Bengal. lacteola - K I R B Y , Syn. Cat. Neur.-Odon. London: 158. lacteola - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 3 1 (India). lacteola - L A I D L A W , Ree. Indian Mus., 16: 178 (India). lacteola - A S A H I N A , Kontyu, 33 (4): 494 (Hong-Kong).

Distribution: India, Hong Kong. luteola

NAVÄS

1936 Agriocnemis luteola N A V Ä S , M U S . Heude Notes ent. chinoise, 3 (4): 46. Type male: not given, perhaps in Heude Museum; type-locality: China.

Distribution: China. maclachlani

SELYS

1877 Argiocnemis (Agriocnemis) maclachlani SELYS, Bull. Acad. r. Belg., (2) 43: 152. Type male: lost; type-locality: G a b o n . 1890 Agriocnemis maclachlani - K I R B Y , Syn. Cat. Neur.-Odon. London: 159. 1928 Agriocnemis maclachlani - F R A S E R , Trans. R. ent. Soc. London, 76: 125 (type from Senegal) (Uganda). 1947 Agriocnemis

maclachlani

— FRASER, Trans. R. ent. Soc. London, 98: 2 4 (Ivory Coast).

183 1962 Agriocnemis maclachlani - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 139 (Senegal, Ivory Coast, Gabon, Portuguese Guinea, French Guinea, Angola, Belgian Congo, Uganda, Dahomey, Nigeria).

Distribution: tropical Africa, merina

LIEFTINCK

1984 Agriocnemis merina LIEFTINCK, 1965 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 50.

Distribution: Madagascar. minima

SELYS

1877 Argiocnemis (Agriocnemis) minima SELYS, Bull. Acad. r. Belg., (2) 43: 145. Type male: R. Mus., Brussels; type-locality: Java. 1890 Agriocnemis minima - KIRBY, Syn. Cat. Neur.-Odon., London: 158. 1898 Agriocnemis minima - KRÜGER, Stettiner ent. Ztg, 59: 126 (Penang). 1930 Agriocnemis minima - KIEFTINCK, Treubia, 12: 151 (redescription of holotype) (Java). 1934 Agriocnemis minima - LIEFTINCK, Treubia, 14 (4): 401 (Java). 1954 Agriocnemis minima - TREUBIA, 22 Suppl.: 72 Java, Borneo, Sumatra, Penang).

Distribution: Java, Borneo, Sumatra, Malaysia. nana

LAIDLAW

1914 Agriocnemis nana LAIDLAW, Ree. Indian Mus., 8: 348. Type male: Zool. Surv. India; typelocality: Upper Burma. 1923 Agriocnemis naia FRÄSER, J. Bombay nat. Hist. Soc., 29: 747. Type male: British Museum (Nat. Hist.), London; type-locality: sine patria. 1930 Agriocnemis ? nana - LIEFTINCK, Treubia, 12: 153 (redescription) (Malacca). 1933 Agriocnemis naia - FRASER, Fauna brit. India, Odon., 1: 381 (Burma). 1948 Agriocnemis nana - LIEFTINCK, Ark. Zool., Uppsala, 41A (10): 16 (Burma). 1954 Agriocnemis naia - LIEFTINCK, Treubia, 22 Suppl.: 73. 1954 Agriocnemis nana - LIEFTINCK, Treubia, 22 Suppl.: 73 (Siam, Malaya). 1964 Agriocnemis nana - SCHMIDT, Ent. Tidschr., 35: 149. 1966 Agriocnemis nana - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 206. 1966 Agriocnemis naia - KIMMINS - Bull. Brit. Mus. nat. Hist., (Ent.) 18: 206 (proposed synonymy w i t h nana LAIDLAW, 1914).

Distribution: Southeast Asia. palaeforma

PINHEY

1958 Agriocnemis palaeforma PINHEY, J. ent. Soc. S. Afr., 22: 465. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1962 Agriocnemis palaeforma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140.

Distribution: Uganda. pieli

NAVÄS

1933 Agriocnemis pieli NAVÄS, Mus. Heude Notes ent. chinoise, 1 (9): 2. Type: not given, perhaps in Heude Mus.; type-locality: China.

Distribution: China. pieris

LAIDLAW

1919 Agriocnemis pieris LAIDLAW, Ree. Indian Mus., 16: 179. Type male: Zool. Surv., India; typelocality: India.

184 1924 Agriocnemis pieris - FRÄSER, Ree. Indian Mus., 26: 490 (West India).

Distribution: India. pygmaea

(RAMBUR)

1842 Agrion pygmaeum RAMBUR, Hist. nat. Ins. Nevropt., Paris: 278. Type female: lost; type-locality: India. 1877 Argiocnemis (Agriocnemis) pygmaea- SELYS, Bull. Acad. r. Belg., (2) 43: 146 (Ceylon, India, Java). 1882 Agriocnemis velaris SELYS, Ann. Soc. Esp., 11: 31. Type male: Selys's Collection; type-locality: Philippines. 1890 Agriocnemis pygmaea - KIRBY, Syn. Cat. Neur.-Odon., London: 158. 1 9 0 5 Agriocnemis pygmaea - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 1 . 1916 Agriocnemis pygmaea - Ris. Suppl. ent. Berlin, 5: 20 (Ceylon, Formosa, New Guinea, Philippines, Seychelles). 1917 Agriocnemis pygmaea - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 34 (Queensland). 1924 Agriocnemis pygmaea - FRÄSER, Ree. Indian Mus., 26: 490 (West India). 1 9 3 1 Agriocnemis pygmaea - C H U J O , Trans, nat. Hist. Soc. Formosa, 21 ( 1 1 2 ) : 3 2 (Formosa). 1934 Agriocnemis pygmaea - LIEFTINCK, Treubia, 14 (4): 402 (India to Australia, Java). 1 9 3 9 Agriocnemis velaris - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 ( 3 ) : 2 7 8 (Luzon). 1949 Agriocnemis pygmaea - LIEFTINCK, Treubia, 20 (2): 348 (Solomon Islands). 1954 Agriocnemis pygmaea - LIEFTINCK, Treubia, 22 Suppl.: 73 (Malaya, Sumatra, Java, Bali). 1955 Agriocnemis pygmaea - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 77 (Ceylon). 1962 Agriocnemis pygmaea - PINHEY, Publ. cult. C o m p . Diam. Angola, 59: 140 (Seychelles, S. and E. Asia to Australia).

Distribution: Oriental region, and North Australia. ruberrima ruberrima 1961

1963

BALINSKY

Agriocnemis ruberrima BALINSKY, J . ent. Soc. S. Afr., 2 4 : 7 4 . Type: unknown; type-locality: Natal. Agriocnemis ruberrima ruberrima - BALISKY, J . ent. Soc. S. Afr., 2 6 : 2 4 9 .

Distribution: Natal. ruberrima albifrons

BALINSKY

1963 Agriocnemis ruberrima albifrons BALINSKY, J. ent. Soc. S. Afr., 26: 249. Type male: Transvaal Museum; type-locality: Bechuanaland.

Distribution: South Africa, rubricauda

T I L L YARD

1913 Agriocnemis rubricauda TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 459. Type male: Tillyard's Collection; type-locality: Queensland. 1917 Agriocnemis thoracalis SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 35. Type male: N a t u r h . Riksmus., Stockholm; type-locality: Queensland.

Distribution: Australia. salomonis

LIEFTINCK

1949 Agriocnemis salomonis LIEFTINCK, Treubia, 20 (2): 349. Type male: Rijksmus. natuurl. Hist. Nat., Leiden; type-locality: G a n o n g a Island.

Distribution: Ganongga Island, sania

NIELSEN 1958 Agriocnemis sania NIELSEN, Riv. Biol, colon., 16: 33. Type: unknown; type-locality: Libya.

185 1974 Agriocnemis pygmaea sania - PINHEY, OCC. Pap. natn. Mus. Rhod. (Nat.Sei.) 5 (4): 225 (redescription). 1984 Agriocnemis pygmaea sania - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 50.

Distribution: Libya. splendidissima

LAIDLAW

1919 Agriocnemis splendidissima LAIDLAW, Ree. Indian Mus., 16:180. Type male: Zool. Surv. India; type-locality: India. 1924 Agriocnemis splendidissima - FRÄSER, Ree. Indian Mus., 26: 490 (West India). 1931 Agriocnemis splendidissima - FRÄSER, Ree. Indian Mus., 33: 464 (S. Malabar).

Distribution: India. trilobata

SJÖSTEDT

1917 Agriocnemis trilobatus SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 36. Type female: Naturh. Riksmus., Stockholm; type-locality: Cap York.

Distribution: Australia. victoria

FRASER

1928 Agriocnemis victoria FRASER, Trans, ent. soc. London, 76: 123. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1962 Agriocnemis victoria - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140 (Uganda, Nigeria, Sierra Leone).

Distribution: Uganda, Central African Republic, Chad, Togo, Nigeria, Sierra Leone. vitiensis

TILLYARD

1923 Agriocnemis vitiensis TILLYARD, Trans, ent. Soc. London, 1923: 337. Type male: Cawthron Inst. Coll.; type-locality: Fiji. 1984 Agriocnemis vitiensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:51 (incorrect synonymy with exsudans SELYS, 1877).

Distribution: Fiji. zerafica LE ROI 1915 Agriocnemis zerafica LE ROI, Ergeb. zweit. Deutsch. Zentr.-Afr. Exp., 1: 343. Type female: Bonn Museum; type-locality: Bahr el Zeraf. 1924 Agriocnemis ebner! Ris, in Alfken, Denkschr. Akad. Wiss. Wien, 99: 277. Type male: Naturh. Mus., Wien; type-locality: Sudan. 1947 Agriocnemis ebneri- FRASER, Trans. R. ent. Soc. London, 98: 24 (redescription) (Ivory Coast). 1962 Agriocnemis zerafica - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140 (Uganda, Sudan, Angola, Ivory Coast, Nigeria). 1962 Agriocnemis ebneri- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140 (proposed synonymy with zerafica LE ROI, 1915).

Distribution: tropical Africa. Genus Argiocnemis

SELYS

1877 Argiocnemis (Argiocnemis) SELYS, Bull. Acad. r. Belg., (2) 43: 135. Type species: Argiocnemis rubeola SELYS, 1877 (designated by Kirby, 1890, Syn. Cat. Neur.-Odon., London: 158). 1890 Argiocnemis KIRBY, Syn. Cat. Neur.-Odon., London: 158. - 1954 Argiocnemis - LIEFTINCK, Treubia, 22Suppl.: 68 [type-species: Argiocnemis (Argiocnemis) rubescens SELYS, 1877].

186 aborense

LAIDLAW

1914 Argiocnemis aborense LAIDLAW, Ree. Indian Mus., 8: 347. Type male: Zool. Surv. India; type-locality: Abor. 1933 Argiocnemis aborense - FRÄSER, J. Bombay nat. Hist. Soc., 36: 467 (description of female).

Distribution: Abor. ensifera

LIEFTINCK

1932 Argiocnemis ensifera LIEFTINCK, Nova Guinea, 15 (5): 105. Type female: Rijksmus. Natuurl. nat. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, rubescens rubescens

SELYS

1877 Argiocnemis (Argiocnemis) rubescens - KIRBY, Syn. Cat. Neur.-Odon., London: 158. 1901 Argiocnemis rubescens - MARTIN, Mem. Soc. zool. France, 14: 248 (Queensland). 1913 Argiocnemis rubescens - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 454 (Queensland, New South Wales). 1917 Argiocnemis rubescens - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 34. 1927 Argiocnemis rubescens - Ris, Zool. Meded., Leiden, 10 (1): 22 (Sumba). 1934 Argiocnemis rubescens - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 339 (Sumatra). 1934 Argiocnemis rubescens - LIEFTINCK, Treubia, 14 (4): 399 (Java). 1954 Argiocnemis rubescens rubescens - LIEFTINCK, Treubia, 22 Suppl.: 68 (Queensland).

Distribution: Australia, Sumatra, Java, Sumba. rubescens intermedia

SELYS

1877 Argiocnemis (Argiocnemis) rubeola var. or race intermedia SELYS, Bull. Acad. r. Belg., (2) 43: 138. Male from Luzon. 1890 Argiocnemis intermedia - KIRBY, Syn. Cat. Neur.-Odon. London: 158. 1984 Argiocnemis rubescens intermedia - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 51.

Distribution: Philippine Islands, rubescens lunulata

SELYS

1877 Argiocnemis (Argiocnemis) lunulata SELYS, Bull. Acad. r. Belg., (2) 43: 139. Type male: not given; type-locality: Celebes. 1890 Argiocnemis lunulata - KIRBY, Syn. Cat. Neur.-Odon., London: 158 (Java, Celebes). 1939 Argiocnemis lunulata - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 278 (Luzon). 1984 Argiocnemis rubescens lunulata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:51.

Distribution: Philippine rubescens nigricans SELYS Islands, Java, Celebes. 1877 Argiocnemis (Argiocnemis) nigricans SELYS, Bull. Acad. r. Belg., (2) 43:140. Type male: Selys's Collection; type-locality: New Guinea. 1890 Argiocnemis nigricans - KIRBY, Syn. Cat. Neur.-Odon, London: 158. 1954 Argiocnemis (Argiocnemis) nigricans SELYS - Lieftinck, Treubia, 22 Suppl.: 68 (incorrect syno n y m y w i t h rubeola

SELYS, 1877).

1984 Argiocnemis rubescens nigricans - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 51.

Distribution: New Guinea.

187 rubescens obscura

LAIDLAW

1914 Argiocnemis obscura LAIDLAW, Ree. Indian Mus., 8: 346. Type male: Zool. Surv. India; typelocality: Assam. 1984 Argiocnemis rubescens obscura- DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:51.

Distribution: Assam. rubescens rubeola

SELYS

1877 Argiocnemis (Argiocnemis) rubeola SELYS, Bull. Acad. r. Belg., (2) 43: 137 (as generotype of Argiocnemis SELYS, 1877). Type male: Wallace's Collection; type-locality: Malaysia. 1890 Argiocnemis rubeola - KIRBY, Syn. Cat. Neur.-Odon., London: 158 (Celebes). 1898 Argiocnemis sumatrana KRÜGER, Stettiner ent. Ztg., 59: 290. Type male: originally in Stettin Museum; type-locality: Sumatra. 1939 Argiocnemis rubeola - NEEDHAM & GYGER, Philipp J. Sei., 20 (3): 276 (redescription) (Luzon). 1954 Argiocnemis rubescens rubeola - LIEFTINCK, Treubia, 22 Suppl.: 68.

Distribution: Malaysia, Celebes, Sumatra, Luzon. solitaria

(SELYS)

1872 Agriocnemis solitaria SELYS, Revue Zool., (2) 23: 180. Type female: R. Mus. Brussles; typelocality: Rodriguez Island. 1877 Argiocnemis (Argiocnemis) solitaria - SELYS, Bull. Acad. r. Belg., (2) 43: 141. 1962 Argiocnemis solitaria - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140.

Distribution: Rodriguez Island. umbargae

PINHEY

1970 Argiocnemis umbargae PINHEY, Arnoldia, Rhodesia, 4 (37): 1. Type male: Pinhey's Collection; type-localicity: Cameroon.

Distribution: Cameroon. Genus Austrocnemis

TILL YARD

1913 Austrocnemis TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 456. Type species: Austrocnemis hyacinthus TILLYARD, 1913 (original designation).

hyacinthus

TILLYARD

1913 Austrocnemis hyacinthus TILLYARD, Proc. Linn. Soc. N.S. Wales, 37:456. Type male: Tillyard's Collection; type-locality: New South Wales, (as generotype of Austrocnemis TILLYARD, 1913).

Distribution: Australia, meccullochi

TILLYARD 1926 Austrocnemis meccullochi TILLYARD, Ree. Austr. Mus., 15:160. Type: unknown; type-locality: Papua.

Distribution: New Guinea, splendida

(MARTIN)

1901 Agriocnemis splendida MARTIN, Mem. Soc. zool. France, 14: 247. Type male: Martin's Collection; type-locality: Victoria. 1913 Austrocnemis splendida - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 457.

Distribution: Australia.

188 Genus Coenagriocnemis

FRÄSER

1949 Coenagriocnemis FRÄSER, Trans. R. ent. Soc. London, 100: 140. Type species: Agriocnemis rufipes R A M B U R , 1 8 4 2 (original designation). - 1 9 4 9 Coenargiocnemis FRÄSER, Trans. R . ent. Soc. London, 1 0 0 : 1 4 3 . Type species: Agrion insulare SELYS, 1 8 7 2 (original designation). 1962 Coenagriocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 136. - 1962 Coenagriocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 131. - 1962 Coenagriocnemis PINHEY, Proc. R. ent. Soc. London, (B) 31: 117 (proposed synonymy with Coenagriocnemis FRÄSER, 1 9 4 9 ) .

insulare

(SELYS)

1872 Agrion (?) insulare, SELYS, Revue Zool., 2 (23): 179. Type female: Selys's Collection; typelocality: Mauritius. 1890 Coenagrion insulare - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1949 Coenargiocnemis insulare - FRÄSER, Trans. R. ent. Soc. London, 100: 143 (as generotype of Coenargiocnemis FRÄSER, 1949). 1962 Coenagriocnemis insulare - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 131.

Distribution: Mauritius. ramburi

FRÄSER

1950 Coenagriocnemis ramburi FRÄSER, Proc. R. ent. Soc. London (B) 19: 70. Type male: British Museum (Nat. Hist.), London; type-locality: Mauritius. 1962 Coenagriocnemis ramburi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 136. 1966 Agriocnemis ramburi - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 211.

Distribution: Mauritius. reuniensis

FRÄSER

1957 Coenagriocnemis reuniensis FRÄSER, Mem. Inst. sei. Madagascar, (Ε) 8: 17. Type male: Inst. Rech. Sei. Madagascar; type-locality: Reunion. 1962 Coenagriocnemis reuniensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 132.

Distribution: Reunion. rufipes

(RAMBUR)

1842 Agrion rufipes RAMBUR, Hist. nat. Ins. Neuropt., Paris: 258. Type male: Guerin's Collection; type-locality: Mauritius. 1872 Agrion (?) mauritianum SELYS, Revue Zool., (2) 23: 180. Type male: unknown; type-locality: Mauritius. 1876 Agrion (Agrion) rufipes - SELYS, Synopsis des Agrionines, Add. 5: 180. 1890 Agriocnemis rufipes - KIRBY, Syn. Cat. Neur.-Odon., London: 159. 1949 Coenagriocnemis rufipes - FRASER, Trans. R. ent. Soc. London, 100: 140 (as generotype of Coenagriocnemis

1962 Coenagriocnemis

SELYS, 1 8 7 7 ) .

rufipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 136.

Distribution: Mauritius. Genus Mortonagrion

FRASER

1920 Mortonagrion FRASER, J. Bombay nat. Hist. Soc., 27: 147. Type species: Mortonagrion varralli FRASER, 1920 (original designation). - 1922 Indagrion FRASER, Mem. Dept. Agric. India, 7: 50. Type species: Indagrion gautama FRASER, 1922 (original designation). - 1930 Mortonagrion - RIS, Ark. Zool., Uppsala, 21A (31): 1 (key to genera of Agriocnemininae). - 1954 Mortonagrion - LIEFTINCK, Treubia, 22 Suppl.: 70. - 1962 Mortonagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140. - 1962 Indagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 140 ( p r o p o s e d s y n o n y m y w i t h Mortonagrion

FRASER, 1 9 2 0 ) .

189 alcyone

(LAIDLAW)

1931 Agriocnemis alcyone LAIDLAW, J. Fed. Mai. States Mus., 16: 249. Type male: Laidlaw's Collection; type-locality: Borneo. 1954 Agriocnemis alcyone - LIEFTINCK, Treubia, 22 Suppl.: 71 (Borneo). 1984 Mortonagrion alcyone - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 52.

Distribution: Borneo. amoenum (Ris) 1915 Agriocnemis amoena Ris, cality: Sumatra. 1930 Mortonagrion amoenum 1935 Mortonagrion amoenum 1948 Mortonagrion amoenum 1954 Mortonagrion amoenum -

Tijdschr. Ent., 58: 10. Type male: Jacobson's Collection; type-loRis, Ark. Zool., Uppsala, 21A (31): 12. LIEFTINCK, Misc. Zool. Sum., 92-93: 12. LIEFTINCK, Treubia, 19: 285. LIEFTINCK, Treubia, 22 Suppl.: 70 (Java, Sumatra, Borneo).

Distribution: Java, Sumatra, Borneo. appendiculatum

LIEFTINCK

1937 Mortonagrion appendiculatum LIEFTINCK, Treubia, 16 (1): 102. Type male: Buitenzorg Museum; type-locality: Billiton Island. 1953 Mortonagrion appendiculatum - LIEFTINCK, Treubia, 22: 395. 1954 Mortonagrion appendiculatum - LIEFTINCK, Treubia, 22 Suppl.: 70.

Distribution: Billiton Island. arthuri

FRÄSER

1942 Mortonagrion arthuri FRÄSER, Proc. R. ent. Soc. London, (B) 11: 97. Type male: British Museum (Nat. Hist.), London; type-locality: Malaysia. 1954 Mortonagrion arthuri - LIEFTINCK, Treubia, 22 Suppl.: 70 (Malaysia). 1965 Mortonagrion arthuri - ASAHINA, Kontyü, 33 (2): 226.

Distribution: Malaysia. ceylonicum

LIEFTINCK

1984 Mortonagrion ceylonicum LIEFTINCK, 1970 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 52.

Distribution: Sri Lanka. falcatum

LIEFTINCK

1934 Mortonagrion falcatum LIEFTINCK, Stylops, 3: 12. Type male: Buitenzorg Museum; type-locality: Karimondjawa Island. 1934 Mortonagrion falcatum - LIEFTINCK, Treubia, 14 (4): 402. 1954 Mortonagrion falcatum - LIEFTINCK, Treubia, 22 Suppl.: 70 (Malaya, Sumatra, Java, Borneo).

Distribution: Malaysia, Java, Sumatra, Borneo. forficulatum

LIEFTINCK

1953 Mortonagrion forficulatum LIEFTINCK, Treubia, 22: 392. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Borneo. 1954 Mortonagrion forficulatum - LIEFTINCK, Treubia, 22 Suppl.: 71 (Borneo).

Distribution: Borneo.

190 gautama

(FRÄSER)

1922 Indagrion gautama F R Ä S E R , Mem. Dept. Agric. India, 7: 50 (as generotype of Indagrion 1922). Type female: British Museum (Nat. Hist.), London; type-locality: India. 1966 Μortonagrion gautama - K I M M I N S , Bull. Brit. Mus. nat. Hist. (Ent.), 1 8 : 194.

FRASER,

Distribution: India. hirosei

ASAHINA

1972 Mortonagrion hirosei locality: Japan.

ASAHINA,

Kontyü, 40 (1): 13. Type male: Asahina's Collection; type-

Distribution: Japan. martini (Ris) 1900 Argiocnemis martini Ris, Arch. Naturg., 66: 199. Type male: Mus. Naturk., Berlin; typelocality: New Britain. 1932 Mortonagrion martini - LIEFTINCK, Nova Guinea, 15 (5): 587. 1949 Mortonagrion martini - LIEFTINCK, Treubia, 20 (2): 346 (Admirality Island).

Distribution: Bismarck Archipelago. selenion (Ris) 1916 Agriocnemis selenion Ris, Suppl. ent., Berlin, 5: 26. Type male: Naturhist. Riksmus., Stockholm; type-locality: Japan. 1930 Mortonagrion selenion - Ris, Ark. Zool. Uppsala, 21A (31): 14 (Sumatra).

Distribution: Japan, Sumatra. simile Ris 1930 Mortonagrion simile Ris, Ark. Zool., Uppsala, 21A (31): 7. Type male: Naturhist. Riksmus., Stockholm; type-locality: Sumatra. 1935 Mortonagrion simile - LIEFTINCK, Misc. Zool. Sum., 92-93: 12. 1954 Mortonagrion simile - LIEFTINCK, Treubia, 22 Suppl.: 71 (Sumatra, Borneo).

Distribution: Sumatra, Borneo, Malaysia. stygium (Fraser) 1954 Agriocnemis stygia F R A S E R , Rev. Zool.-bot. afr., 50: 274. Type male: Mus. Afr. Centr., Tervuren; type-locality: Belgian Congo. 1957 Mortonagrion stygium - F R A S E R , Revue Zool.-bot. afr., 56: 220 (description of female). 1962 Mortonagrion stygium - Pinhey, Publ. cult. Comp. Diam. Angola, 59: 141.

Distribution: Congo Republic. varralli

FRASER

1920 Mortonagrion varralli F R A S E R , J. Bombay, nat. hist. Soc., 27:148. Type male: British Museum (Nat. Hist.), London; type-locality: India (as generotype of Mortonagrion F R A S E R , 1920). 1924 Mortonagrion varralli - F R A S E R , Ree. Indian Mus., 26: 490 (West India). 1930 Mortonagrion varralli - Ris, Ark. Zool., Uppsala, 21A (31): 13 (Sumatra). 1931 Mortonagrion varralli - F R A S E R , Ree. Indian Mus., 33: 464 (Malabar). Distribution: f r o m I n d i a t o S u m a t r a .

191 Subfamily Argiinae Discoidal field of fore wing short and broad, wing briefly petiolate; Dc short and wide, dilating outwardly; antenodals close and coverging posteriorly; arc at level of distal antenodal, anal vein arises proximal to Ac\ females with no vulvar spine. 1938 Argiinae TILLYARD & FRASER, Austr. Zool., 9: 166 (with genera Hyponeura, Argia, Onychoargia, Paraiargia, Argiallagma, Diargia). - 1957 Argiinae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 50 (with genera Argia, Hyponeura, Diargia, Onychargia, Palaiargia, Papuargia, Archboldargia, Hylaeargia, Argiallagma).

Genus Archboldargia

LIEFTINCK

1949 Archboldargia LIEFTINCK, Nova Guinea (N.S.) 5: 139. Type species: Archboldargia LIEFTINCK, 1949 (original designation).

gloriosa

miriftca

LIEFTINCK

1949 Archboldargia gloriosa LIEFTINCK, Nova Guinea (N.S.) 5: 145. Type male: Rijksmus. Natuurl. Hist.nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. mirifica

LIEFTINCK

1949 Archboldargia mirifica LIEFTINCK, Nova Guinea (N.S.) 5:142 (as generotype of Archboldargia LIEFTINCK, 1949). Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Argia

RAMBUR

1842 Argia RAMBUR, Hist. nat. Ins. Neuropt., Paris: 254. Type species: Agrion fumipennis BURMEISTER, 1839 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 137). - 1854 Hyponeura SELYS, Mon. Cal.: 275. Type-species: Hyponeura funcki SELYS, 1854 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 137). - 1865 Argia - SELYS, Bull. Acad. r. Belg., (2) 20: 382 (revision). - 1903 Argia - NEEDHAM, Bull. 68 N.Y. State Mus.: 236. - 1905 Agria (misprint) - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 812. - 1909 Diargia CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 154. Type species: Diargia bicellulata CALVERT, 1909 (original designation). - 1910 Argia - MUTTKOWSKI, Bull. Publ.Mus. Milwaukee, 1 (1): 44. - 1920 Argia (Cyanargia) KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type species: Agrion lacrimans HÄGEN, 1861 (original designation). - 1920 Argia (Heliargia) KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type species: Argia vivida HÄGEN, 1865 (original designation). - J920 Argia (Micrargia) KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type species: Argia thisma CALVERT, 1909 (original designation). - 1920 Argia (Leptargia) KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type species: Argia mollis SELYS, 1865 (original designation). - 1920 Argia (Chalcargia) KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type-species: Argia aenea SELYS, 1865 (original designation). - 1946 Argia - FRASER, Trans. R. ent. Soc. London, 97: 444. - 1988 Argia MICHALSKI, Univ. West Indies Dep. Zool. Occ. Pap., 6: 39 (key to species).

adamsi CALVERT 1902 Argia adamsi CALVERT, Biol. Centr.-Amer. Neur.: 80. Type male: Calvert's Collection; typelocality: Costa Rica. 1916 Argia adamsi - Ris, Arch. Naturg., (A) 82: 106 (Panama).

Distribution: Costa Rica, Panama.

192 aenea

SELYS

1865 Argia aenea SELYS, Bull. Acad. r. Belg., (2), 20:407. Type male: Selys's Collection; type-locality: Mexico. 1902 Argia aenea - CALVERT, Biol. Centr.-Amer. Neur.: 73 (California, Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Colombia). 1916 Argia aenea - Ris, Arch. Naturg., (A) 82: 114 (Mexico, Guatemala, Panama). 1920 Chalcargia aenea - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Chalcargia KENNEDY, 1920).

Distribution: Central America. agrioides

CALVERT

1895 Argia agrioides CALVERT, Proc. Cal. Acad., (2) 4: 476. Type male: Calif. Acad.; type-locality: California. 1910 Argia agrioides agrioides - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 45 (California, Arizona, Texas, Mexico).

Distribution: South U.S.A., Mexico, alberta

KENNEDY

1918 Argia alberta KENNEDY, Canad. Ent., 50: 257. Type male: Kennedy's Collection; type-locality: U.S.A..

Distribution: U.S.A. albistigma

HÄGEN

1865 Argia albistigma HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 20: 402. Type male: Hagen's Collection; type-locality: Montevideo.

Distribution: Uruguay. ambigua

NAVÄS

1934 Argia ambigua NAVÄS, Rev. Acad. Madrid, 31:179. Type male: Madrid Museum; type-locality: Colombia. 1 9 8 4 Argia ambigua - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 5 2 (Brazil).

Distribution: Colombia, Brazil. apicalis (SAY) Agrion apicalis Say (nec MATSUMURA, 1 9 1 3 ) , J1 Acad. nat. Sei. Philad., 8 : 4 1 0 . Type: unknown; type-locality: U.S.A. 1865 Argia apicalis - SELYS, Bull. Acad. r. Belg., (2) 20: 414. 1890 Argia apicalis - KIRBY, Syn. Cat. Neur.-Odon., London: 139. 1910 Argia apicalis - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 45. Distribution: U . S . A . 1839

augustana

NAVÄS

1934 Argia augustana NAVÄS, Rev. Acad. Madrid, 31: 180. Type male: Madrid Museum; type-locality: Colombia. Distribution: C o l o m b i a .

193 barretti

CALVERT

1902 Argia barretti CALVERT, Biol. Centr.-Amer. Neur.: 87. Type male: Calvert's Collection; typelocality: Mexico.

Distribution: Mexico. bicellulata

(CALVERT)

1909 Diargia bicellulata CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 155. Type male: Mus. Comp. Zool., Cambridge; type-locality: Brazil (as generotype of Diargia CALVERT, 1909). 1984 Argia bicellulata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 52.

Distribution: Brazil. bipunctulata

(HÄGEN)

1861 Agrion bipunctulatum HAGEN, Syn. Neur. N.Amer., Washington: 90. Type male: Mus. Comp. Zool., Cambridge; type-locality: Georgia. 1865 Argia bipunctulata - SELYS, Bull. Acad. r. Belg., (2) 20: 415. 1 8 9 0 Argia bipustulata SELYS, 1 8 6 5 (probably misprint) - loc. cit. K I R B Y , Syn. Cat. Neur.-Odon., London: 139.

Distribution: botacudo

U.S.A.

CALVERT

1909 Argia botacudo CALVERT, Coll. Zool. Selys de Longchamps, 1909: 148. Type male: R. Mus., Brussels; type-locality: Brazil.

Distribution: Brazil. chapadae

CALVERT

1909 Argia chapadae CALVERT, Coll. Zool. Selys de Longchamps, 1909: 146. Type male: R. Mus., Brussels; type-locality: Brazil.

Distribution: Brazil. charalana

NAVÄS

1924 Argia charalana NAVÄS, Mem. R. Acad. Ci. Barcelona, (3) 18 (13): 16. Type male: Madrid Museum; type-locality: Colombia. 1935 Argia charalana - NAVÄS, Bol. Soc. ent. Esp., 17: 33 (description of female).

Distribution: Colombia. chelata

CALVERT

1902 Argia chelata CALVERT, Biol. Centr.-Amer. Neur.: 88. Type male: Calvert's Collection; typelocality: Costa Rica. 1916 Argia chelata - Ris, Arch. Naturg., (A) 82: 114. (Costa Rica, Panama).

Distribution: Costa Rica, Panama. clausseni

SELYS

1865 Argia clausseni SELYS, Bull. Acad. r. Belg., (2) 20: 386. Type male: R. Mus., Brussels; typelocality: Minas Geraes.

Distribution: Brazil.

194

collata

SELYS

1865 Argia collata SELYS, Bull. Acad. r. Belg., (2) 20:395, Type male: R. Mus., Brussels; type-locality: South America.

Distribution: ? South America. columbiana

NAVAS

1924 Argia columbiana NAVAS, Mem. R. Acad. Ci. Barcelona, (3) 18 (13): 15. Type male: Madrid Museum; type-locality: Colombia.

Distribution: Colombia. concinna

(RAMBUR)

1842 Agrion concinnum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 270. Type: sex not given; unknown; ? Cape. 1865 Argia concinna - SELYS, Bull. Acad. r. Belg., (2) 20: 391. 1902 Argia concinna - HÄGEN & CALVERT, Bull. Mus. comp. Zool. Harward Coll., 39: 107 (Dominica).

Distribution: West Indies. croceipennis

SELYS

1865 Argia croceipennis SELYS, Bull. Acad.r. Belg., (2) 20: 393. Type male: Selys's Collection; typelocality: Brazil.

Distribution: Brazil. cupraurea

CALVERT

1902 Argia cupraurea CALVERT, Biol. Centr.-Amer., Neur.: 85. Type male: Calvert's Collection; type-locality: Panama. 1916 Argia cupraurea - RIS, Arch. Naturg., (A) 82: 113 (Panama, Venezuela).

Distribution: Panama, Venezuela. cuprea

(HÄGEN)

1861 Agrion cupreum HAGEN, Syn. Neur. N.Amer., Washington: 96. Type male: Mus. Comp. Zool., Cambridge; type-locality: Mexico. 1865 Argia cuprea - SELYS, Bull. Acad. r. Belg., (2) 20: 407. 1910 Argia cuprea - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46 (Mexico, Guatemala, Honduras). 1916 Argia cuprea - Ris, Arch. Naturg., (A) 82: 110 (Mexico, Guatemale, Honduras, Nicaragua). 1946 Argia cuprea - FRASER, Trans. R. ent. Soc. London, 97: 445 (with form c, male from Peru - Leeds Museum).

Distribution: from Mexico to Peru. cyathigera

NAVAS

1934 Argia cyathigera NAVAS, Rev. Acad. Ci. Madrid, 31: 181. Type male: Madrid Museum; typelocality: Colombia.

Distribution: Colombia. dagnina

FÖRSTER

1914 Argia dagnina FÖRSTER, Arch. Naturg., (A2) 80: 62. Type male: Förster's Collection; typelocality: Colombia.

Distribution: Colombia.

195

deami

CALVERT

1902 Argia deami CALVERT, Biol. Centr.-Amer., Neur.: 71. Type male: A d a m s ' s Collection; typelocality: Mexico. 1910 Argia deami - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46.

Distribution: Mexico. difficilis

SELYS

1865 Argia difficilis SELYS, Bull. Acad. r. Belg., (2) 20: 413. Type female: R. M u s . Brussels; typelocality: Peru. 1902 Argia difficilis - CALVERT, Biol. Centr.-Amer., 2: 108 ( H o n d u r a s , C o s t a Rica, P a n a m a , Colombia, E c u a d o r , Venezuela). 1914 Argia difficilis - FÖRSTER, Arch. N a t u r g . , (A2) 80: 66 (Peru). 1916 Argia difficilis - Ris, Arch. Naturg., (A) 82: 107 ( P a n a m a , Colombia). 1932 Argia difficilis - GEIJSKES, Zool. Meded., Leiden, 14: 240 (Trinidad). 1943 Argia difficilis - SCHMIDT, Beitr. F a u n a Perus, 2: 233 (Peru).

Distribution: Central America. dimissa

SELYS

1865 Argia dimissa SELYS, Bull. Acad. r. Belg., (2) 20: 388. Type male: Selys's Collection; typelocality: Brazil.

Distribution: Brazil. dives

FÖRSTER

1914 Argia cuprea dives FÖRSTER, Arch. Naturg., (A2) 80: 61. Type male: Förster's Collection; type-locality: Peru. 1918 ? Argia cuprea f o r m b Ris, Arch. Naturg., (A) 82: 112. Female f r o m Bolivia. 1943 Argia dives - SCHMIDT, Beitr. F a u n a Perus, 2: 233.

Distribution: Peru, Bolivia. eliptica

SELYS

1865 Argia eliptica SELYS, Bull. Acad. r. Belg., (2) 20: 293. Type male: Selys's Collection; typelocality: Brazil.

Distribution: Brazil. emma

KENNEDY

1915 Argia emma KENNEDY, Proc. U . S . natn. Mus., 49: 271. Type male: U . S . natn. Mus., Washington; type-locality: Washington.

Distribution: U.S.A. espinalensis

NAVAS

1935 Argia espinalensis NAVAS, Bol. Soc. ent. Esp., 17: 36. Type male: M a d r i d M u s e u m ; typelocality: Colombia.

Distribution: Colombia. euphorbia

FRASER

1946 Argia extranea euphorbia FRÄSER, Trans. R . ent. Soc. L o n d o n , 97: 457. Type male: Leeds M u s e u m , type-locality: Peru. 1966 Argia extranea euphorbia - KIMMINS, Bull. Brit. M u s . nat. Hist. (Ent.), 18: 191 [type male in British M u s e u m ( N a t . Hist.), London].

196 1984

Argia euphorbia -

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 5 3 .

Distribution: Peru. extranea

(HÄGEN)

1861 Agrion extraneum HAGEN, Syn. Neur. N.Amer., Washington: 92. Type male: Saussure's Collection; type-locality: Mexico. 1861 Agrion calidum HÄGEN, Syn. Neur. N.Amer., Washington: 93. Type male: Mus. Comp. Zool., Cambridge; type-locality: Mexico. 1865 Argia extranea - SELYS, Bull. Acad. r. Belg., (2) 20: 399. 1865 Argia calida - SELYS, Bull. Acad. r. Belg., (2) 20: 390. 1865 Argia variabilis SELYS, Bull. Acad. r. Belg., (2) 20: 406. Type female: Mus. Comp. Zool., Cambridge; type-locality: Mexico. 1910 Argia extranea - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46 (Mexico to Guyana). 1910 Agrion calidum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46 [proposed synonymy with extranea ( H Ä G E N , 1 8 6 1 ) ] . 1910 Argia variabilis - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46 [proposed synonymy with extranea ( H Ä G E N , 1 8 6 1 ) ] . 1916 Argia extranea - Ris, Arch. Naturg., (A) 82: 116 (Mexico, Honduras, Costa Rica, Panama). 1946 Argia extranea - FRASER, Trans. R. ent. Soc. London, 97: 456. 1 9 8 4 Argia extranea - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 5 3 . 1984 Argia calida - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 53 (as good species). 1984 Argia variabilis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 56 (as good species).

Distribution: from Mexico to Guyana, fissa

SELYS

1865 Argia fissa Bogota. 1910 Argia fissa Colombia, 1916 Argia fissa mala). 1943 Argia fissa

SELYS, Bull. Acad. r. Belg., (2) 20: 401. Type male: Selys's Collection; type-locality: - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46 (Costa Rica, Guatemala, Mexico). - RIS, Arch. Naturg., (A) 82: 114 (Mexico, Costa Rica, Peru, Colombia, Guate- SCHMIDT, Beitr. F a u n a Perus, 2: 233.

Distribution: from Mexico to Peru. forficula

FRASER

1946 Argia extranea forficula FRASER, Trans. R. ent. Soc. L o n d o n , 97: 456. Type male: Leeds M u seum; type-locality: Peru. 1966 Argia extranea forficula - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 193 (lectotype male: British Museum (Nat. Hist.), London). 1984 Argia forficula - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 53.

Distribution: Peru. fosteri

CALVERT

1909 Argia fosteri CALVERT, Ann. Carnegie Mus., Pittsbourgh, 6:128. Type: unknown; type-locality: Paraguay.

Distribution: Paraguay. fraudatricula

FÖRSTER

1914 Argia fraudatricula FÖRSTER, Arch. Naturg., (A2) 80: 64. Type male: Förster's Collection; type-locality: Peru.

197 1943 Argia fraudatricula

- SCHMIDT, Beitr. F a u n a Perus, 2: 236.

Distribution: Peru. frequentula CALVERT 1907 Argia frequentula CALVERT, Biol. Centr.-Amer., 1907:365. Type male: Williamson's Collection; type-locality: Mexico. 1910 Argia frequentula - MUTTKOWSKI, Bull. Publ.Mus. Milwaukee, 1 (1): 47 (Mexico, Vera Cruz to Panama). 1916 Argia frequentula - Ris, Arch. Naturg., (A) 82: 104 (Mexico, Guatemala, Costa Rica).

Distribution: Central America. fulgida NAVÄS 1934 Argia fulgida NAVÄS, Broteria, 30 (3): 137. Type male: not given; type-locality: Colombia.

Distribution: Colombia. fumigata

SELYS

1865 Argia fumigata SELYS, Bull. Acad.r. Belg., (2) 20: 394. Type male: Selys's Collection; typelocality: Guiana.

Distribution: Guyana. fumipennis fumipennis (BURMEISTER) 1839 Agrion fumipenne Burmeister, H a n d b . Ent., 2: 819. Type female: Mus. Comp. Zool., Cambridge; type-locality: Kentucky. 1842 Argia fumipennis - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 254 (as generotype of Argia RAMBUR, 1 8 4 2 ) .

1842 Argia obscura RAMBUR, Hist. nat. Ins. Neuropt., Paris: 256. Type: sex not given: Serville's Collection; type-locality: Amer. septent. 1890 Argia fumipennis - KIRBY, Syn. Cat. Neur.-Odon., London: 138). 1898 Argia fumipennis - CALVERT, Trans. Am. ent. Soc., 25: 38. 1910 Argia fumipennis - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 47. 1910 Argia obscura - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 47 [proposed synonymy w i t h fumipennis

fumipennis

(BURMEISTER, 1839)].

Distribution: U.S.A. fumipennis atra

GLOYD

1984 Argia fumipennis atra GLOYD, 1968 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 52.

Distribution: North America. funcki (SELYS) 1854 Hyponeura funcki SELYS, Monogr. Cal.: 275. Type male: Selys's Collection; type-locality: Colombia (as generotype of Hyponeura SELYS, 1854). 1910 Hyponeura funcki - MUTTKOWSKI, Bull. Publ. Mus. Milwaukeen, 1 (1): 43 (Mexico to Venezuela). 1984 Argiafuncki-DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 53 (Central America).

Distribution: from Mexico to Venezuela.

198 funebris

(HAGEN)

1861 Agrion funebre HAGEN, Syn. Neur. N.Amer., Washington: 92. Type male: Hagen's Collection; type-locality: Mexico. 1865 Argia funebris - SELYS, Bull. Acad. r. Belg., (2) 20: 398.

Distribution: Mexico. gaumeri

CALVERT

1907 Argia gaumeri CALVERT, Biol. Centr.-Amer., Neur.: 363. Type male: Calvert's Collection; type-locality: Central America.

Distribution: Central America, gerhardi gerhardi

CALVERT

1909 Argia gerhardi CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 119. Type male: Ann Arbor Museum; type-locality: Bolivia. 1916 Argia gerhardi - Ris, Arch. Naturg., (A) 82: 107 (Peru, Bolivia). 1943 Argia gerhardi - SCHMIDT, Beitr. Fauna Perus, 2: 232. 1984 Argia gerhardi gerhardi - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 53.

Distribution: Colombia, Bolivia, Peru. gerhardi nigrior CALVERT 1909 Argia gerhardi nigrior CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 132. Type: unknown; type-locality: Peru.

Distribution: Peru. hamulata

FRÄSER

1946 Argia hamulata FRASER, Trans. R. ent. Soc. London, 97: 447. Type male: Leeds Museum; type-locality: Peru. 1966 Argia hamulata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 194 [type in British Museum (Nat. Hist.), London].

Distribution: Peru. harknessi

CALVERT

1899 Argia harknessi CALVERT, Proc. Cal. Acad., (3) 1: 378. Type male: Calif. Acad.; type-locality: Mexico. 1910 Argia harknessi - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 47.

Distribution: Mexico. hasemani

CALVERT

1909 Argia hasemani CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 137. Type: unknown; type-locality: Brazil.

Distribution: Brazil. hebdomatica

NAVÄS

1934 Argia hebdomatica NAVÄS, Broteria, 30 (3): 140. Type male: not given; type-locality: Colombia. Distribution: C o l o m b i a .

199 herberti

CALVERT

1902 Argia herberti CALVERT, Biol. Centr.-Amer., Neur., 1902:70. Type male: G o d m a n ' s Collection; type-locality: Mexico. 1910 Argia herberti - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 47.

Distribution: Mexico. hinei

KENNEDY

1918 Argia hinei KENNEDY, Canad. Ent., 50: 258. Type male: Kennedy's Collection; type-locality: U.S.A. Distribution: U . S . A .

huallaga

FRÄSER

1946 Argia huallaga FRASER, Trans. R. ent. Soc. London, 97: 450. Type male: Leeds Museum; type-locality: Peru. 1966 Argia huallaga - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 196 [lectotype male: British Museum (Nat. Hist.), London].

Distribution: Peru. huanacina

FÖRSTER

1914 Argia huanacina FÖRSTER, Arch. Naturg., (A) 80: 67. Type male: Mus. Naturk., Berlin; typelocality: Peru. 1943 Argia huanacina - SCHMIDT, Beitr. F a u n a Perus, 2: 233.

Distribution: Peru. icterica

NAVÄS

1934 Argia icterica NAVAS, Broteria, 30 (3): 138. Type male: not given; type-locality: Colombia.

Distribution: Colombia. immunda

(HÄGEN)

1861 Agrion immundum HAGEN, Syn. Neur.-N.Amer., Washington: 93. Type male: Mus. Comp. Zool., Cambridge; type-locality: Mexico. 1865 Argia immunda - SELYS, Bull. Acad. r. Belg., (2) 20: 401. 1910 Argia immunda - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 47.

Distribution: Mexico. impura

RAMBUR

1842 Argia impura RAMBUR, Hist. Nat. Ins. Neuropt., Paris: 255. Type: sex not given: Selys's Collection; type-locality: Amazons. 1910 Argia impura - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 48 (tropical America). 1917 Argia impura - SJÖSTEDT, Ark. Zool., 11 Uppsala, (15): 10.

Distribution: Brazil. inculta

HÄGEN

1865 Argia inculta type-locality: 1890 Argia inculta 1943 Argia inculta

HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 20: 400. Type male: Selys's Collection; Lima. - KIRBY, Syn. Cat. Neur.-Odon., London: 138. - SCHMIDT, Beitr. F a u n a Perus, 2: 233.

Distribution: Peru.

200 indicatrix

CALVERT

1902 Argia indicatrix CALVERT, Biol. Centr.-Amer., Neur.: 73. Type male: Calvert's Collection; type-locality: Mexico. 1916 Argia indicatrix - Ris, Arch. Naturg., (A) 82: 104 (Mexico, Colombia, Guatemala, Honduras, Nicaragua, Costa Rica).

Distribution: Central America. indocilis

NAVÄS

1934 Argia indocilis NAVÄS, Rev. Acad. Madrid, 31:183. Type male: Madrid Museum; type-locality: Colombia.

Distribution: Colombia. infrequentula

FRASER

1946 Argia infrequentula FRASER, Trans. R. ent. soc. London, 97: 451. Type male: Leeds Museum; type-locality: Colombia. 1946 Argia infrequentula - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 197 [lectotype male: British Museum (Nat. Hist.), London],

Distribution: Colombia. infumata

SELYS

1865 Argia infumata SELYS, Bull. Acad. r. Belg., (2) 20: 392. Type male: unknown; type-locality: Para.

Distribution: Brazil. insipida

SELYS

1865 Argia insipida SELYS, Bull. Acad. r. Belg., (2) 20: 389. Type male: Selys's Collection; typelocality: Guiana. 1932 Argia ierea GEIJSKES, Zool. Meded., Leiden, 14: 244. Type male: Geijskes's Collection; typelocality: Trinidad.

Distribution: Guyana, Trinidad. iralai

CALVERT

1909 Argia iralai CALVERT, Coll. Zool. Selys de Longchamps, 1909: 153. Type male: Calvert's Collection; type-locality: Paraguay.

Distribution: Paraguay. jocosa

SELYS

1865 Argia jocosa SELYS, Bull. Acad. r. Belg., (2) 20:408. Type male: R. Mus., Brussels; type-locality: Bogota.

Distribution: Colombia. joergenseni Ris 1913 Argia joergenseni Ris, Mem. Soc. ent. Belg., 22: 94. Type male: Mus. Naturk., Berlin; typelocality: Argentina. 1916 Argia joergenseni - Ris, Arch. Naturg., (A) 82: 114. 1948 Argia joergenseni - FRASER, Acta Zool. lilloana, 5: 61 (Argentina).

Distribution: Argentina.

201 johannella

CALVERT

1907 Argia johannella CALVERT, Biol. Centr.-Amer., Neur.: 370. Type male: Calvert's Collection; type-locality: Central America.

Distribution: Central America. jujuya Ris 1913 Argia jujuya Ris, Mem. Soc. ent. Belg., 22: 94. Type male: Mus. Naturk., Berlin; type-locality: Argentina. 1916 Argia jujuya - Ris, Arch. Naturg., (A) 82: 108.

Distribution: Argentina. kiautai

STEINMANN

nom.nov.

1913 Argia apicalis MATSUMURA (nec SAY, 1839), Thons. Ins. Japan, Addit., 1: 51 (in doubtful position). Argia kiautai STEINMANN nom. nov. [new name for Argia apicalis MATSUMURA, 1913, a junior secondary homonymy of Argia apicalis (SAY, 1839)].

Distribution: Japan. kokama

CALVERT

1909 Argia kokama CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 119. Type male: unknown; typelocality: Iquitos. 1943 Argia kokama - SCHMIDT, Beitr. Fauna Perus, 2: 232.

Distribution: Peru. lacrimans

(HÄGEN)

1861 Agrion lacrimans HAGEN, Syn. N.Amer. Neur., Washington: 95. Type female: Mus. Comp. Zool., Cambridge; type-locality: Mexico. 1865 Argia lacrimans - SELYS, Bull. Acad. r. Belg., (2) 20: 386. 1910 Argia lacrymans (misprint) - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 48 (Mexico). 1920 Argia (Cyanargia) lacrimans - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Cyanargia

KENNEDY, 1 9 2 0 ) .

Distribution: Mexico. lilacina

SELYS

1865 Argia lilacina SELYS, Bull. Acad. r. Belg., (2) 20: 405. Type male: Selys's Collection; typelocality: Brazil.

Distribution: Brazil. limitata

NAVÄS

1924 Argia limitata NAVÄS, Mem. R. Acad. Ci. Barcelona, (3) 18 (13): 19. Type male: unknown; type-locality: Peru. 1943 Argia limitata - SCHMIDT, Beitr. F a u n a Perus, 2: 235.

Distribution: Peru. lobata

NAVÄS

1924 Argia lobata NAVÄS, Mem. R. Acad. Ci. Barcelona, (3) 18 (13). 17. Type male: unknown; type-locality: Colombia.

Distribution: Colombia.

202 lugens

(HAGEN)

1861 Agrion lugens HAGEN, Syn. Neur., N.Amer. Washington; 95. Type female: Mus. Naturk., Berlin; type-locality: Mexico. 1865 Hyponeura lugens - SELYS, Bull. Acad. r. Belg., (2) 20: 382. 1910 Hyponeura lugens - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 44 (holotype in Mus. Comp. Zool. Cambridge) (New Mexico, Arizona, Northern Mexico). 1 9 8 4 Argia lugens - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 5 4 .

Distribution: Central America, and South U.S.A. machadina

FÖRSTER

1914 Argia machadina FÖRSTER, Arch. Naturg., 80 (A): 63. Type male: Förster's Collection; typelocality: Matto Grosso.

Distribution: Brazil. makoka

FRASER

1946 Argia makoka FRASER, Trans. R. ent. Soc. London, 97: 455. Type male: Leeds Museum; type-locality: Peru. 1966 Argia makoka KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18:202 (type male: British Museum (Nat. Hist.), London).

Distribution: Peru. mariensis

CALVERT

1961 Argia mariensis CALVERT, Proc. Acad. nat. Sei. Philad., 113: 5. Type male: Acad. nat. Sei. Philad.; type-locality: Peru.

Distribution: Peru. medinensis

NAVÄS

1935 Argia medinensis NAVÄS. Bol. Soc. ent. Esp., 17: 35. Type male: unknown; type-locality: Colombia.

Distribution: Colombia. medullaris

SELYS

1865 Argia medullaris SELYS, Bull. Acad. r. Belg., (2) 20: 412. Type male: R. Mus., Brussels; typelocality: Bogota.

Distribution: Colombia. mishuyaca

FRASER

1946 Argia mishuyaca FRASER, Trans. R. ent. Soc. London, 97: 453. Type male: Leeds Museum; type-locality: Peru. 1966 Argia mishuyaca - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 204 (lectotype male: British Museum (Nat. Hist.), London).

Distribution: Peru. missionensis

FRASER

1948 Argia missionensis FRASER, Acta Zool. lilloana, 5: 58. Type male: Miguel Lille Inst., Tucuman; type-locality: Argentina.

Distribution: Argentina.

203 mista

NAVÄS

1934 Argia mista NAVÄS, Bol. Soc. ent. Esp., 16: 70. Type male: unknown; type-locality: Mexico.

Distribution: Mexico. moesta

(HAGEN)

1861 Agrion moestum HAGEN, Syn. Neur. N.Amer., Washington: 94. Type male: Mus. Comp. Zool., Cambridge; type-locality: Texas. 1861 Agrion putridum HAGEN, Syn. Neur. N.Amer., Washington: 96. Type male: Mus. Comp. Zool. Cambridge; type-locality: Wisconsin. 1865 Argia moesta - SELYS, Bull. Acad. r. Belg., (2) 20: 384. 1865 Argia putrida - SELYS, Bull. Acad. r. Belg., (2) 20: 385. 1910 Argia moesta moesta - MUTTKOWSKI, Bull.Publ. Mus. Milwaukee, 1 (1): 48 (Texas, Arizona, N. Mexico). 1910 Argia moesta putrida - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 48 (Canada, Ν. Dakota, Texas, Florida). 1912 Argia intruda WILLIAMSON, Ent. News, 23: 200. Type male: Williamson's collection; typelocality: Oklahoma. 1912 Argia putrida - WILLIAMSON, Ent. News, 23: 200 [proposed synonymy with moesta (HAGEN, 1861)].

Distribution: North America. modesta

SELY

1865 Argia modesta SELYS, Bull. Acad. r. Belg., (2) 20: 388. Type male: R. Mus., Brussels; typelocality: Brazil.

Distribution: Brazil. mollis

SELYS

1865 Argia mollis SELYS, Bull. Acad. r. Belg., (2) 20: 398. Type male: R. Mus., Brussels; type-locality: Minas Geraes. 1920 Argia (Leptargia) mollis - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Leptargia KENNEDY, 1 9 2 0 ) .

Distribution: Brazil. mollusca

FRÄSER

1946 Argia mollusca FRASER, Trans. R. ent. Soc. London, 197: 454. Type male: Leeds Museum; type-locality: Peru. 1966 Argia mollusca - KIMMINS, Bull. Brit. Mus. Nat. Hist. (Ent.), 18: 204 [lectotype male: British Museum (Nat. Hist.), London],

Distribution: Peru. munda

CALVERT

1901 Argia vivida munda CALVERT, Biol. Centr.-Amer. Neur.: 96. Type male: Mus. comp. Zool. Cambridge; type-locality: Mexico. 1910 Argia vivida munda - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 52 (Arizona, Mexico). 1915 Argia vivida munda „HAGEN" - KENNEDY, Proc. U.S. natn. Mus., 49: 288 (Washington, Colombia). 1919 Argia rita KENNEDY, Canad. Ent., 51: 17. Type male: unknown; type-locality: Arizona. 1958 Argia munda - GLOYD, Occ. Pap. Mus. Zool. Univ. Mich., 593: 17. 1958 Argia rita - GLOYD, Occ. Pap. Mus. Zool. Univ. Mich., 593: 17 (proposed synonymy with munda

CALVERT, 1 9 0 1 ) .

Distribution: South U.S.A., and Mexico.

204 muzana

NAVÄS

1934 Argia muzana NAVÄS, Broteria, 30 (3): 139. Type female: not given; type-locality: Colombia.

Distribution: Colombia. nahuana

CALVERT

1901 Argia nahuana CALVERT, Biol. Centr.-Amer. Neur.: 72. Type male: Calvert's Collection; typelocality: Mexico. 1910 Argia agrioides nahuana - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 45. 1943 Argia saalasi VALLE, Ann. ent. fenn., 8: 164. Type: unknown; type-locality: California. 1984 Argia nahuana - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 55.

Distribution: California, Mexico. oculata

HÄGEN

1865 Argia oculata HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 20: 409. Type male: Mus. comp Zool. Cambridge; type-locality: Mexico. 1910 Argia oculata - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 49 (Mexico, Venezuela Brazil). 1916 Argia oculata - Ris, Arch. Naturg., (A) 82: 105 (Mexico, Guatemala, Honduras, Costa Rica Panama, Venezuela). 1943 Argia oculata - SCHMIDT, Beitr. Fauna Perus, 2: 233 (Venezuela, Peru). 1984 Argia oculata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 55 (Brazil).

Distribution: Mexico to Brazil. oenea

HÄGEN

1865 Argia oenea HAGEN, in SELYS, Bull. Acad.r. Belg., (2) 20: 407. Type male: Mus. comp. Zool Cambridge; type-locality: Mexico. 1890 Argia aenea (misprint) - KIRBY, Syn. Cat. Neur.-Odon., London: 138. 1910 Argia oenea - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 49 (Mexico, Guatemala Colombia, Nicaragua). 1920 Chalcargia oenea - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Chalcargit KENNEDY, 1 9 2 0 ) .

Distribution: Central America. orichalcea

HÄGEN

1865 Argia orichalcea HAGEN, in SELYS, Bull. Acad.r. Belg., (2) 20:408. Type male: Selys's Collection type-locality: Venezuela. 1932 Argia orichalcea - GEIJSKES, Zool. Meded., Leiden, 14: 242 (Trinidad).

Distribution: Central America. pallens

CALVERT

1902 Argia violacea pallens CALVERT, Biol. Centr.-Amer. Neur.: 98. Type male: Godman's Collection; type-locality: Arizona. 1910 Argia violacea pallens - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 49. 1918 Argia solita KENNEDY, Canad. Ent., 50: 256. Type male: unknown; type-locality: U.S.A. Distribution: N o r t h A m e r i c a .

205 percellulata

CALVERT

1902 Argia percellulata CALVERT, Biol. Centr.-Amer. Neur.: 74. Type male: Godman's Collection; type-locality: Vera Cruz. 1916 Argia percellulata - Ris, Arch. Naturg., (A) 82: 102.

Distribution: Mexico. pipila

CALVERT

1907 Argia pipila CALVERT, Biol. Centr.-Amer. Neur.: 373. Type male: Calvert's Collection; typelocality: Central America.

Distribution: Central America. plana

CALVERT

1901 Argia vivida plana CALVERT, Biol. Centr.-Amer. Neur.: 96. Type male: Mus., Comp. Zool. Cambridge; type-locality: Mexico. 1910 Argia vivida plana - M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 (1): 52 (Arizona, Mexico). 1 9 8 4 Argia plana - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 5 5 .

Distribution: Arizona, Mexico. pocomana

CALVERT

1907 Argia pocomana CALVERT, Biol. Centr. Amer., Neur.: 375. Type male: Calvert's Collection; type-locality: Central America.

Distribution: Central America. popoluca

CALVERT

1902 Argia popoluca CALVERT, Biol. Centr.-Amer. Neur.: 82. Type male: Godman's Collection; type-locality: Mexico.

Distribution: Mexico. pulla

HÄGEN

Argia pulla H A G E N , in SELYS, Bull. Acad. r. Belg., ( 2 ) 20: 4 1 0 . Type male: Mus. comp. Zool. Cambridge; type-locality: Venezuela. 1902 Argia pulla - CALVERT, Biol. Centr.-Amer. Neur.: 70 (Mexico, Guatemala, Honduras, Nicaragua, Costa Rica, Panama, Venezuela). 1916 Argia pulla - Ris, Arch. Naturg., (A) 82: 104 (Guatemala, Panama). 1 9 4 8 Argia pulla - FRASER, Acta Zool. lilloana, 5 : 5 8 (Argentina).

1865

Distribution: neotropical fauna region. reclusa

SELYS

1865 Argia reclusa SELYS, Bull. Acad. r. Belg., (2) 20: 395. Type male: Selys's Collection; typelocality: Para.

Distribution: Brazil. rectangula

NAVÄS

1920 Argia rectangula lombia.

NAVAS,

Mem. Acc. Nuovi Lincei, 5: 1. Type: unknown; type-locality: Co-

Distribution: Colombia.

206 rhoadsi CALVERT 1902 Argia rhoadsi CALVERT, Biol. Centr.-Amer. Neur.: 92. Type male: Godman's Collection; typelocality: Mexico. 1910 Argia rhoadsi - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 50.

Distribution: Mexico. rogersi CALVERT 1902 Argia rogersi CALVERT, Biol. Centr.-Amer. Neur.: 83. Type male: Godman's Collection; typelocality: Costa Rica.

Distribution: Costa Rica, sedula (HÄGEN) 1861 Agrion sedulum HAGEN, Syn. Neur. N.Amer. Washington: 94. Type male: Mus. comp. Zool. Cambridge; type-locality: Berkeley Springs. 1865 Argia sedula - SELYS, Bull. Acad. r. Belg., (2) 20: 411 (Virginia, Texas, California). 1899 Argia sedula - KELLICOTT, Odon. Ohio: 78. 1910 Argia sedula - MUTTKOWSKI, Bull. Publ. mus. Milwaukee, 1 (1): 50 (C. & S. States: Arizona; Mexico).

Distribution: California, Texas, Mexico, serva HÄGEN 1865 Argia serva HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 20: 387. Type male: unknown; typelocality: Brazil.

Distribution: Brazil. smithiana CALVERT 1909 Argia smithiana CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 140. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil. sordida SELYS 1865 Argia sordida SELYS, Bull. Acad. r. Belg., (2) 20: 387. Type male: Selys's Collection; typelocality: Brazil.

Distribution: Brazil. spegazzinii NAVÄS 1934 Argia spegazzinii NAVÄS, Bol. Soc. ent. Esp., 16: 69. Type male: unknown; type-locality: Mexico.

Distribution: Mexico, stigmatica NAVÄS 1934 Argia stigmatica NAVÄS, Broteria, 30 (3): 141. Type male: not given; type-locality: Colombia.

Distribution: Colombia. subapicalis CALVERT 1909 Argia subapicalis CALVERT, Coll. Zool. Selys de Longchamps, 1909: 151. Type male: Selys's Collection; type-locality: Brazil. Distribution: B r a z i l .

207 talamanca

CALVERT

1907 Argia talamanca CALVERT, Biol. Centr.-Amer. Neur.: 359. Type male: unknown; type-locality: Costa Rica. 1916 Argia talamanca - Ris, Arch. Naturg., (A) 82: 108 (redescription) (Colombia).

Distribution: Costa Rica, Colombia. tamoyo

CALVERT

1909 Argia tamoyo CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 139. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil. tarascana

CALVERT

1902 Argia tarascana CALVERT, Biol. Centr.-Amer. Neur.: 90. Type male: Godman's Collection; type-locality: Mexico. 1910 Argia tarascana - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 50.

Distribution: Mexico. terira

CALVERT

1907 Argia terira CALVERT, Biol. Centr.-Amer. Neur.: 358. Type male: Calvert's Collection; typelocality: Costa Rica. 1916 Argia terira - Ris, Arch. Naturg., (A) 82: 104.

Distribution: Costa Rica. ternaria

NAVÄS

1934 Argia ternaria NAVÄS, Broteria, 30 (3): 142. Type male: not given; type-locality: Colombia.

Distribution: Colombia. tezpi

CALVERT

1902 Argia tezpi CALVERT, Biol. Centr.-Amer. Neur.: 77. Type male: Godman's Collection; typelocality: Mexico. 1905 Argia cupraea CALVERT, Proc. Calif. Acad., (2) 4: 479. Type male: Calif. Acad.; type-locality: Mexico. 1910 Argia tezpi - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 50 (California, Mexico, Venezuela). 1910 Argia cupraea - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 50 (proposed synonymy w i t h tezpi

CALVERT, 1 9 0 2 ) .

1989 Argia tezpi - MAES, Rev. Nica. Ent., Leon, 8: 6 (U.S.A., Mexico, Guatemala, El Salvador, Honduras, Nicaragua, Costa Rica).

Distribution: Central America. theodori Naväs 1934 Argia theodori NAVÄS, Rev. Acad. Madrid, 31: 182. Type male: unknown; type-locality: Colombia.

Distribution: Colombia. thespis

SELYS

1865 Argia thespis SELYS, Bull. Acad. r. Belg., (2) 20:397. Type male: Selys's Collection; type-locality: Bahia.

Distribution: Brazil.

208 thisma CALVERT 1909 Argia thisma CALVERT, Coll. Zool. Selys des Longchamps, 1909: 141. Type male: Selys's Collection; type-locality: Brazil. 1920 Argia (Micrargia) thisma - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Micrargia

KENNEDY, 1 9 2 0 ) .

Distribution: Brazil. tibialis (RAMBUR) 1842 Platycnemis tibialis RAMBUR, Hist. nat. Ins. Neuropt., Paris: 241. Type female: Serville's Collection; type-locality: Amer. septentr. 1861 Trichocnemis tibialis - HAGEN, Syn. Neur. N.Amer., Washington: 72. 1861 Agrion fontium HÄGEN, Syn. Neur. N.Amer., Washington: 91. Type male: Mus. comp. Zool. Cambridge; type-locality: California. 1862 Agrion binotatum WALSH, Proc. Acad. nat. Sei. Philad., 1862: 387. Type: lost; type-locality: Wisconsin. 1890 Argia tibialis - KIRBY, Syn. Cat. Neur.-Odon. London: 139. 1890 Agrion fontium-KmsY, Syn. Cat. Neur.-Odon. London: 139 [proposed synonymy with tibialis (RAMBUR, 1842)].

1890 Agrion binotatum - KIRBY, Syn. Cat. Neur.-Odon. London: 139 [proposed synonymy with tibialis

(RAMBUR, 1842)].

1910 Argia tibialis - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 50.

Distribution: Central to Southern U.S.A. tinctipennis SELYS 1865 Argia tinctipennis SELYS, Bull. Acad. r. Belg., (2) 20: 396. Type male: Mus. comp. Zool. Cambridge; type-locality: Amazonia.

Distribution: Brazil. tonto CALVERT 1902 Argia tonto CALVERT, Biol. Centr.-Amer. Neur.: 89. Type male: Calvert's Collection; typelocality: Arizona. 1910 Argia tonto - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 51.

Distribution: U.S.A. tractipennis SELYS 1865 Argia tractipennis SELYS, Bull. Acad. r. Belg., (2) 20: 396. Type male: Selys's Collection; typelocality: Amazons.

Distribution: Brazil. translate HÄGEN 1865 Argia translate HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 20: 410. Type male: Mus. comp. Zool. Cambridge; type-locality: Venezuela. 1910 Argia translata - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 51 (C. America, Central & Southern U.S.). 1916 Argia translata - Ris, Arch. Naturg., (A) 82: 104 (U.S.A., Texas, Mexico, Panama). 1989 Argia translata - MAES, Rev. Nica. Ent., Leon, 8: 6 (U.S.A., Mexico, Guatemala, Belize, El Salvador, Honduras, Nicragua, Costa Rica, Panama, Suramerica).

Distribution: Central America, and Southern U.S.A.

209 trifoliata

FRÄSER

1946 Argia trifoliata FRÄSER, Trans. R. ent. Soc. London, 97: 449. Type male: Leeds Museum; type-locality: Colombia. 1966 Argia trifoliata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 216 (lectotype male: British Museum (Nat. Hist.), London).

Distribution: Colombia. trina

NAVÄS

1934 Argia trina NAVÄS, Broteria, 30 (3): 143. Type male: not given; type-locality: Colombia.

Distribution: Colombia. tupi

CALVERT

1909 Argia tupi CALVERT, Coll. Zool. Selys de Longchamps 1909: 150. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil. ulmeca

CALVERT

1902 Argia ulmeca CALVERT, Biol. Centr.-Amer. Neur.: 80. Type male: Godman's Collection; typelocality: Mexico. 1910 Argia ulmeca - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 51 (Honduras, Mexico). 1989 Argia ulmeca - MAES, Rev. Nica. Ent., Leon, 8: 6 (Mexico, Guatemala, Honduras, Nicaragua, Costa Rica).

Distribution: Central America. umbriaca

FRASER

1946 Argia umbriaca FRASER, Trans. R. ent. Soc. London, 97: 451. Type male: Leeds Museum; type-locality: Colombia. 1966 Argia umbriaca - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 217 (lectotype male: British Museum (Nat. Hist.), London).

Distribution: Colombia. underwoodi

CALVERT

1907 Argia underwoodi CALVERT, Biol. Centr.-Amer. Neur.: 370. Type male: Calvert's Collection; type-locality: Central America.

Distribution: Central America. variata 1984

NAVÄS

Argia variata

NAVÄS, 1 9 3 5

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World, Utrecht,

1: 5 6 .

Distribution: Colombia. variegata

FÖRSTER

1914 Argia varigata FÖRSTER, Arch. Naturg., 80 (A): 65. Type male: Förster's Collection; typelocality: Ecuador.

Distribution: Ecuador.

210 violacea

(HAGEN)

1861 Agrion violaceum HAGEN, Syn. Neur. N.Amer., Washington: 90. Type male: Mus. comp. Zool. Cambridge; type-locality: Maryland. 1865 Argia violacea - SELYS, Bull. Acad. r. Belg., (2) 20: 404 (Maryland, Virginia, Washington, Texas, Massachusetts, Connecticut, New York, Illinois, New Jersey, Pennsylvania).

Distribution: U.S.A. vivida

HÄGEN

1865 Argia vivida HÄGEN, Bull. Acad. r. Belg., (2) 20: 406. Type male (only): Mus. comp. Zool. Cambridge; type-locality: Mexico. 1865 Argia kurilis SELYS, Bull. Acad. r. Belg., (2) 20:400. Type male: R. Mus., Brussels; type-locality: Kurile Island. 1901 Argia vivida vivida - CALVERT, Biol. Centr.-Amer. Neur.: 94 (redescription). 1 9 0 5 Agria (misprint) kurilis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 2 . 1910 Argia vivida - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 46 [incorrect synonymy with immunda ( H Ä G E N , 1 8 6 1 ) ] . 1 9 2 0 Argia (Heliargia) VM'RFA - K E N N E D Y , Ohio J . Sei. Columbus, 21: 8 5 (as generotype of Heliargia KENNEDY, 1 9 2 0 ) .

1988 Argia vivida - PRITCHARD, Notul. odonatol., 3 (1): 8 (Canada).

Distribution: from Canada to Mexico. wilsoni

CALVERT

1902 Argia wilsoni CALVERT, Biol. Centr.-Amer. Neur.: 75. Type male: Calvert's Collection; typelocality: Guatemala.

Distribution: Guatemala. Genus Hylaeargia

LIEFTINCK

1949 Hylaeargia LIEFTINCK, Nova Guinea (N.S.) 5: 133. Type species: Hylaeargia simulatrix TINCK, 1949 (original designation).

simulatrix

LIEF-

LIEFTINCK

1949 Hylaeargia simulatrix LIEFTINCK, Nova Guinea (N.S.) 5: 136 (as generotype of Hylaeargia LIEFTINCK, 1949). Type male: Rijksmus. natuurl. Hist. nat. Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Moroagrion 1939

Moroagrion N E E D H A M

NEEDHAM & GYGER, 1 9 3 9

danielli

NEEDHAM & GYGER

Philipp. J . Sei., 70 ( 3 ) : (original designation).

& GYGER,

279.

Type species: Moroagrion danielli

NEEDHAM & GYGER

1939 Moroagrion danielli N E E D H A M & G Y G E R , Philipp. J. Sei. 70 (3): 279. Type male: not given: type-locality: Guara (as generotype of Moroagrion N E E D H A M & G Y G E R , 1939).

Distribution: Philippine Islands.

Genus Onychargia

SELYS

1865 Onychargia SELYS, Bull. Acad. r. Belg., (2) 20: 416. Type species: Onychargia atrocyana 1865 (monotypy). - 1954 Onychargia - LIEFTINCK, Treubia, 22 Suppl.: 53.

SELYS,

211 atrocyana

SELYS

1865 Onychargia atrocyana SELYS, Bull. Acad. r. Belg., (2) 20: 416 (as generotype of Onychargia SELYS, 1 8 6 5 ) . Type male: Selys's Collection; type-locality: Singapoor. 1919 Onychargia atrocyana - LAIDLAW, Ree. Indian Mus., 16: 172 (Assam). 1924 Onychargia atrocyana - FRÄSER, Ree. Indian Mus., 26: 489 (W. India). 1927 Onychargia atrocyana - Ris, Zool. Meded., Leiden, 10 (1): 21 (Sumatra). 1934 Onychargia atrocyana - LIEFTINCK, Treubia, 14 (4): 393 (Java). 1935 Onychargia atrocyana - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 9 (Sumatra). 1954 Onychargia atrocyana - LIEFTINCK, Treubia, 22 Suppl.: 53 (Siam, Malaya, Sumatra, Java, Borneo). 1955 Onychargia atrocyana - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 73 (Ceylon).

Distribution: Oriental faunal region. indica 1984

SAHNI

Onychargia indica SAHNI, 1 9 6 4 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 57.

Distribution: India. vittigera

SELYS

1984 Onychargia vittigera SELYS, 1891 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 57.

Distribution: India. Genus Palaiargia

FÖRSTER

1903 Argia (Palaiargia) FÖRSTER, Annls hist.-nat. Mus. natn. hung., 1903: 549. Type species: Onychargia flavovittata SELYS, 1878 (original designation). - 1932 Palaiargia - LIEFTINCK, Nova Guinea, 15 (5): 73 (revision, key to species). - 1949 Palaiargia - LIEFTINCK, Nova Guinea (N.S.) 5: 113 (key to species). - 1972 Palaiargia - LIEFTINCK, Deutsch, ent. Z., 19 ( 1 - 3 ) : 251 (key to species). - 1984 Palaiargia - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 57.

alcedo

LIEFTINCK

1949 Palaiargia alcedo LIEFTINCK, N o v a Guinea (N.S.), 5: 119. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. arses

LIEFTINCK

1957 Palaiargia arses LIEFTINCK, N o v a Guinea (N.S.), 5: 54. Type male: Rijksmus. Natuurl. Hist, nat.; Leiden; type-locality: New Guinea.

Distribution: New Guinea. carnifex carnifex

LIEFTINCK

1932 Palaiargia carnifex LIEFTINCK, Nova Guinea, 15 (5): 79. Type male: Buitenzorg Museum; type-locality: New Guinea. 1972 Palaiargia carnifex carnifex - LIEFTINCK, Deutsch, ent. Z., 19 ( 1 - 3 ) : 250.

Distribution: New Guinea.

212

carnifex praeclava

LIEFTINCK

1972 Palaiargia carnifex praeclava LIEFTINCK, Deutsch, ent. Z., 19 (1-3): 250. Type male: unknown; type-locality: New Guinea.

Distribution: New Guinea, ceyx ceyx

LIEFTINCK

1949 Palaiargia ceyx ceyx LIEFTINCK, Nova Guinea (N.S.) 5: 125. Type male: Rijksmus. Natuurl. Hist. nat. Leiden; type-locality: New Guinea.

Distribution: New Guinea. ceyx flammula

LIEFTINCK

1949 Palaiargia ceyx flammula LIEFTINCK, Nova Guinea (M.S.) 5: 125. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea, charmosyna charmosyna

LIEFTINCK 1932 Palaiargia charmosyna LIEFTINCK, Nova Guinea, 15 (5): 76. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea. 1949 Palaiargia charmosyna charmosyna - LIEFTINCK, Nova Guinea (N.S.) 5: 118.

Distribution: New Guinea. charmosyna cyclopica

LIEFTINCK

1949 Palaiargia charmosyna cyclopica LIEFTINCK, Nova Guinea (N.S.) 5:118. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea, charmosyna miniata

LIEFTINCK

1949 Palaiargia charmosyna miniata LIEFTINCK, Nova Guinea (N.S.) 5: 118. Type male: Rijksmus. Natuurl. Hist. nat. Leiden; type-locality: New Guinea.

Distribution: New Guinea, eclecta

LIEFTINCK

1949 Palaiargia eclecta LIEFTINCK, New Guinea (N.S.) 5: 127. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. e o s LIEFTINCK 1938 Palaiargia eos LIEFTINCK, Nova Guinea (N.S.) 2: 95. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea, ernstmayri Lieftinck 1972 Palaiargia ernstmayri LIEFTINCK, Deutsch, ent. Z., 19 (1-3): 246. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea.

213 halcyon

LIEFTINCK

1938 Palaiargia halcyon LIEFTINCK, Nova Guinea (N.S.) 2: 97. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. humida

FÖRSTER

1903 Palaiargia flavovittata humida FÖRSTER, Annls hist.-nat. Mus. natn. hung., 1: 550. Type male: Termeszett. Müz., Budapest; type-locality: New Guinea. 1932 Palaiargia humida - LIEFTINCK, Nova Guinea, 15 (5): 81. 1984 Palaiargia humida - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 57 (as generotype of Palaiargia FÖRSTER, 1 9 0 3 ) .

Distribution: New Guinea. melidora

LIEFTINCK

1953 Palaiargia melidora LIEFTINCK, Treubia, 22: 241. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Waigeu Island.

Distribution: Indonesia: Waigeu Island. micropsitta

LIEFTINCK

1957 Palaiargia micropsitta LIEFTINCK, N o v a Guinea (N.S.) 8: 60. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Misool, Island.

Distribution: Misool Island (near New Guinea), myzomela

LIEFTINCK

1957 Palaiargia myzomela LIEFTINCK, Nova Guinea (N.S.) 8: 51. Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea, nasiterna

LIEFTINCK

1938 Palaiargia nasiterna LIEFTINCK, N o v a Guinea (N.S.) 2: 99. Type male: Rijksmus. Natuurl., Hist, nat., Leiden; type-locality: New Guinea.

Distribution: New Guinea. obiensis LIEFTINCK 1957 Palaiargia obiensis LIEFTINCK, Nova Guinea (N.S.) 8: Type male: Rijksmus. Natuurl. Hist, nat., Leiden; type-locality: Moluccas.

Distribution: Moluccas. optata

(SELYS)

1865 Argia optata SELYS, Bull. Acad. r. Belg., (2) 20: 390. Type male: Selys's Collection; type-locality: Moluccas. 1949 Palaiargia optata - LIEFTINCK, Nova Guinea (N.S.) 5: 131. 1957 Palaiargia optata - LIEFTINCK, Nova Guinea (N.S.) 8: 58 (redescription of female).

Distribution: Moluccas.

214

perimecosoma

LIEFTINCK

1957 Palaiargia perimecosoma LIEFTINCK, Nova Guinea (N.S.) 8:44. Type male: Rijksmus. Natuurl. Hist. nat. Leiden; type-locality: Moluccas.

Distribution: Moluccas. rubropunctata

(SELYS)

1878 Onychargia rubropunctata SELYS, Mitt. Zool. Mus. Dresden, 3: 321. Type male: not given; typelocality: New Guinea. 1878 Onychargia ? flavovittata SELYS, Mitt. Zool. Mus. Dresden, 3: 321. Type female: not given; typelocality: New Guinea. 1903 Argia (Palaiargia) flavovittata - FÖRSTER, Annls hist.-nat. Mus. natn. hung., 1903: 550. 1949 Palaiargia flavovittata - LIEFTINCK, Nova Guinea (N.S.) 5: 126. 1957 Palaiargia rubropunctata - LIEFTINCK, Nova Guinea (N.S.) 8: 67. 1957 Onychargia flavovittata - LIEFTINCK, Nova Guinea (N.S.) 8: 67 [proposed synonymy with rubropunctata (SELYS, 1 8 7 8 ) ] .

Distribution: New Guinea. stellata (Ris) 1916 Onychargia stellata Ris, Nova Guinea, 13 (2): 95. Type male: Mus. Naturk., Berlin; typelocality: New Guinea. 1957 Palaiargia stellata - LIEFTINCK, Nova Guinea (N.S.) 8: 60 (redescription).

Distribution: New Guinea. tanysiptera

LIEFTINCK

1953 Palaiargia tanysiptera LIEFTINCK, Treubia, 22: 239. Type male: Manis & Amsari's Collection; type-locality: Halmahera Island.

Distribution: Halmahera Islands. Genus Papuargia

LIEFTINCK

1938 Papuargia LIEFTINCK, Nova Guinea (N.S.) 2:101. Type species: Papuargia stueberi 1938 (original designation).

stueberi

LIEFTINCK,

LIEFTINCK

1938 Papuargia stueberi LIEFTINCK, Nova Guinea (N.S.) 2: 102. Type male: Rijksmus. Natuurl. Hist. nat.. Leiden; type-locality: New Guinea (as generotype of Papuargia LIEFTINCK, 1938).

Distribution: New Guinea.

Subfamily Coenagrioninae Least petiolate subfamily. Anal vein separated from wing border well proximal to Ac: Dc short. No vulvar spine. 1860 Agrioninae SELYS, Bull. Acad. r. Belg., (2) 10:11 (defined as subfamily). - 1890 Coenagrioninae KIRBY, Syn. Cat. Neur.-Odon., London: 1 1 9 . Type-genus: Coenagrion KIRBY, 1 8 9 0 . - 1 9 0 5 Agrioninae - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 2 (with genera Agria (misprint: Argia), Ischnura, Enallagma, Agrion, Nehalennia, Pyrhossoma, Erythromma, Pseudagrion, Ceriagrion). - 1910 Coenagrioninae - MUTTKOWSKI, Bull. Publ. Milwaukee, 1 (1): 42 (with genera Paraphlebia, Hyponeura, Argia, Argiallagma, Hesparagrion, Oxyagrion, Anisagrion, Enallagma,

Telebasis,

Telagrion,

Leptobasis,

Nehalennia,

Coenagrion,

Amphiagrion,

Chrom-

215 agrion, Ischnura, Anomalagrion, Ceratura, Neoneura, Microneura, Protoneura). - 1919 Coenagrioninae - MUNZ, Mem. Amer. ent. Soc., 3: 53 (key to genera Metacnemis, Mesocnemis, Calicnemis, Stenocnemis, Copera, Platycnemis, Idiocnemis, Coeliccia, Allocnemis, Tatocnemis, Nesocnemis, Prionocnemis, Leptocnemis, Paracnemis, Antiagrion, Onychargia, Hyponeura, Palaiargia, Argia, Argiallagma, Diargia, Skiallagma, Tigriagrion, Amphiagrion, Hesperagron, Thermagrion, Xiphiagrion, Oxyagrion, Leptobasis, Oreagrion, Pyrrhosoma, Erythromma, Chromagrion, Aeolagrion, Leptagrion, Telebasis, Metalaptobasis, Leptobasis, Hemiphlebia, Ceriagrion, Nesobasis, Papuagrion, Nesobasis, Argiagrion, Teinobasis, Pericnemis, Amphicnemis, Anomalagrion, Ischnura, Celaeura, Ceratura, Austrocnemis, Argiocnemis, Nehalennia Cercion, Megalagrion, Coenagrion, Enallagma, Zoniagrion, Leptobasis, Acanthagrion, Mombagrion, Aciagrion, Telagrion, Anisagrion, Caliagrion, Pseudagrion, Xanthagrion, Xanthocnemis, Austroagrion, Archibasis). - 1926 Coenagriinae TILLYARD & FRASER, Ins. Austr. and New Zeal.: 79. Type-genus: Coenagrion KIRBY, 1890. - 1938 Coenagriinae - TILL YARD & FRASER, Austr.

Zool., 9: 166 (with genera Ischnura, Argiocnemis, Agriocnemis, Caliagrion, Austroagrion, Xanthagrion, Pseudagrion, Ceriagrion, Aciagrion, Archibasis, Coenagrion, Pyrrhosoma, Erythromma, Nehalennia, Thermagrion, Mombagrion, Argiagrion, Xiphiagrion, Teinobasis, Mortonagrion, Enallagma, Himalagrion, Amphiagrion, Tigriagrion, Hesperagrion, Chromagrion, Anomalagrion, Oxyagrion, Acanthagrion, Anisagrion, Telagrion, Leptagrion, Megalagrion, Hylaeagrion, Aeolagrion, Antiagrion, Telebasis, Metaleptobasis, Ceratura). - 1948 Coenagriinae - ZIMMERMANN, Ins. Hawaii 2: 342 (with genera Megalagrion, Ischnura, Enallagma). - 1957 Coenagriinae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 49 (with genera Coenagrion, Cercion, Austroagrion, Nehalennia, Pyrrhosoma, Erythromma, Himalagrion). - 1962 Coenagriinae PINHEY, Publ. cult. Comp. Diam. Angola, 59: 116 (key to genera Millotagrion, Teinobasis, Argiagrion, Ceriagrion, Ischnuragrion, Pseudagrion, Aciagrion, Mortonagrion, Agriocnemis, Coenagriocnemis, Ischnura, Enallagma).

Genus Austroagrion

TILL YARD

1913 Austroagrion TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 466. Type species: Pseudagrion cyane SELYS, 1876 (original designation). - 1982 Austroagrion - LIEFTINCK, Odonatologica, 11 (4): 289 (key to species).

cyane

(SELYS)

1876 Agrion (Pseudagrion) cyane SELYS (nec Ris, 1910 et TILL YARD, 1913), Synopsis des Agrionines,

Add. 5: 218. Type male: Selys's Collection; type-locality: Australia. 1890 Pseudagrion cyane - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1901 Pseudagrion cyane - MARTIN, Mem. Soc. zool. France, 14: 246 (Victoria). 1908 Pseudagrion coeruleum TILLYARD, Proc. Linn. Soc. N.S. Wales, 32: 739. Type male: Tillyard's Collection; type-locality: Perth. 1913 Austroagrion cyane - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 466 (as generotype of Austroagrion

TILLYARD, 1913).

1913 Austroagrion coeruleum - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 467 (South-Central Australia). 1917 Austroagrion cyane - SJÖSTEDT, Ark. Zool., Uppsala 11 (11) 37 (Queensland). 1982 Austroagrion coeruleum - LIEFTINCK, Odonatologica, 11 (4): 290 [proposed synonymy with cyane (SELYS, 1876)].

Distribution: Australia. exlamationis

CAMPION

1915 Austroagrion exlamationis CAMPION, Ann. Mag. nat. Hist., (6) 16: 106. Type male: not given; type-locality: N.E. Australia.

Distribution: Australia.

216 pindrina

WATSON

1984 Austroagrion pindrina WATSON, 1969 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the W o r l d ,

Utrecht, 1: 58.

Distribution: Australasia. watsoni

LIEFTINCK

1982 Austroagrion watsoni LIEFTINCK, Odonatologica, 11 (4): 287. New name for Austroagrion

cyane

R i s , 1 9 1 0 n e c TILLYARD, 1 9 1 3 .

Distribution: Australia. Genus Austrocoenagrion

KENNEDY

1920 Austrocoenagrion KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type species: Agrion lyelli TILLYARD, 1913 (original designation).

lyelli

(TILLYARD)

1913 Agrion lyelli TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 449. Type male: Tillyard's Collection; type-locality: Victoria. 1920 Austrocoenagrion lyelli - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Austrocoenagrion KENNEDY, 1920).

Distribution: Australia. Genus Cercion 1907

NAVAS

Cercion NAVÄS, Broteria, 6: 5 5 . Type species: Agrion lindeni SELYS, 1 8 4 0 (monotypy). - 1 9 5 0 Cercion - MOREIRA, Inst. Esp. Ent. Madrid: 84. - 1965 Cercion - ASAHINA, Kontyü, 33 (4): 495. - 1984 Cercion - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1 (1): 58.

calamorum (Ris) 1916 Agrion calamorum Ris, Suppl. ent., 5: 32. Type male: Ris's Collection; type-locality: China. 1931 Agrion calamorum - SCHMIDT, Konowia, 10 (3): 183 (Shanghai). 1984 Cercion calamorum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 58 (Japan).

Distribution: China, Japan. hieroglyphicum

(BRAUER)

1865 Agrion hieroglyphicum BRAUER, Verh. Zool.-Bot. Ges., Wien, 15: 510. Type female: not given; type-locality: China (Hong Kong). 1876 Agrion (Agrion) lineolatum SELYS, Synopsis des Agrionines, Add. 5: 172. Type male: Nation. Hist. Mus., Paris; type-locality: Japan. 1890 Coenagrion hieroglyphicum - KIRBY, Syn. Cat. Neur. Odon., London: 150. 1905 Agrion hieroglyphicum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 824 (China). 1916 Agrion hieroglyphidum - Ris, Suppl. ent. Berlin, 5: 37 (China, Japan). 1931 Agrion hieroglyphicum - SCHMIDT, Konowia, 10 (3): 183 (Tche-kiang). 1965 Cercion hieroglyphicum - ASAHINA, Kontyü, 33 (4): 495 (Hong Kong).

Distribution: East Asia. lindeni lindeni

(SELYS)

1840 Agrion lindeni SELYS, Monogr. Libell. europ. Paris: 167. Type male: unknown; type-locality: Belgium. 1876 Agrion (Agrion) lindenii - SELYS, Synopsis des Agrionines, Add. 5: 169 (Belgium, France, Switzerland, Italy, Sicily, Dalmatia, Algeria, Asia Minor).

217 1890 Coenagrion lindenii - KIRBY, Syn. Cat. Neur.-Odon. London: 150 (Asia Minor). 1 9 0 5 Agrion lindeni - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 4 (Europe, Algeria). 1 9 0 7 Cercion lindeni - NAVAS, Broteria, 6 : 5 5 (as generotype of Cercion Naväs, 1 9 0 7 ) . 1923 Cercion lindeni var. nigriceps NAVÄS, Arq. Inst. Cienc., 8 (1920): 10. Male from Spain. 1924 Agrion lindeni - MORTON, Trans, ent. Soc. London, 1924: 33 (Palestine). 1938 Agrion lindeni - SCHMIDT, Sitzber. Akad. Wiss. Wien, (I) 147: 143. 1940 Agrion lindeni - NIELSEN, Mem. Soc. ent. ital., 19: 241 (Sardinia). 1948 Puella lindeni - AKRAMOVSKI, Zool. Zbom., Moskow, 5: 158 (Armenia). 1950 Cercion lindeni - MORERA, Inst. Esp. ent. Madrid: 84 (Spain). 1 9 5 0 Cercion lindeni var. exlis N A V A S , in M O R E R A Inst. Esp. Ent. Madrid: 8 5 . Male from Spain. 1950 Cercion lindeni VAT. nigriceps - MORERA, Inst. Esp. Ent. Madrid: 85 (Spain). 1984 Coenagrion lindeni - STEINMANN, Fauna hung., 160: 26 (Spain, France, Germany, Suisse, Yugoslavia). 1984 Cercion lindeni - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 58.

Distribution: Southern Europe, Asia Minor, and North Africa. lindeni zernyi (SCHMIDT) 1938 Agrion lindeni zernyi SCHMIDT, Sitz. Akad. Wiss. Wien (I) 147:143. Type male: Naturh. Mus., Wien; type-locality: Palestine (Lebanon). 1984 Cercion lindeni zerneyi (misprint) - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 58 (Palestine).

Distribution: Lebanon. luzonicum ASAHINA 1984 Cercion luzonicum Utrecht, 1: 58.

ASAHINA,

1968 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Mindanao. malayanum SELYS 1984 Cercion malayanum Utrecht, 1: 58.

SELYS,

1870 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: S.E. Asia. plagiosum (NEEDHAM) 1930 Coenagrion plagiosum NEEDHAM, Zool. Sinica, (A) 11: 268. Type: unknown; type-locality: China. 1984 Cercion plagiosum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 58.

Distribution: China. quadrigerum (SELYS) 1883 Agrion quadrigerum SELYS, Ann. Soc. Ent. Belg., 27: 136. Type male: R. Mus. Brussels; typelocality: Japan. 1890 Coenagrion quadrigerum - KIRBY, Syn. Cat. Neur.-Odon. London: 150. 1 9 0 5 Agrion quadrigerum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 4 . 1 9 8 4 Cercion quadriferum (misprint) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 5 8 . [proposed synonymy with sieboldi (SELYS, 1 8 7 6 ) ] .

Distribution: Japan.

218 sexlineatum

(SELYS)

1883 Agrion sexlineatum SELYS, Ann. Soc. Ent. Belg., 27: 135. Type male: R. Mus. Brussels; typelocality: Japan. 1890 Coenagrion sexlineatum - KIRBY, Syn. Cat. Neur.-Odon. London: 150. 1916 Agrion sexlineatum - Ris, Suppl. ent. Berlin, 5: 35 (Formosa, China). 1931 Agrion sexlineatum - SCHMIDT, Konowia, 10 (3): 183 (Pu-To Island, Tchekinag, China). 1933 Coenagrion trilineatum NAVÄS, Mus. Heude Notes ent. chinoise, 9: 5. Type: unknown; typelocality: China. 1933 Coenagrion admirationis NAVÄS, Mus. Heude Notes ent. chinoise, 9: 6. Type: unknown; typelocality: China. 1962 Coenagrion trilineatum - CHAO, Acta ent. sinica, Suppl.: 11: 28. 1962 Coenagrion admirationis - CHAO, Acta ent. sinica, Suppl., 11: 28 (proposed synonymy with trilineatum

NAVÄS, 1 9 3 3 ) .

1984 Cercion sexlineatum

-

DAVIES & TOBIN,

The Dragonflies of the World, Utrcht 1: 58.

Distribution: Japan, Taiwan, China. sieboldi

(SELYS)

1876 Agrion (Agrion) sieboldii SELYS, Synopsis des Agrionines, Add. 5: 171. Type female: Selys's Collection; type-locality: Japan. 1890 Coenagrion sieboldi - KIRBY, Syn. Cat. Neur.-Odon. London: 150. 1916 Agrion sieboldi - Ris, Suppl. ent. Berlin, 5: 33. 1916 Agrion sauteri Ris, Suppl. ent. Berlin, 5: 34. Type male: Deutsch. Ent. Inst.; type-locality: Japan. 1984 Cercion sieboldi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 58.

Distribution: Japan. v-nigrum

(NEEDHAM)

1930 Coenagrion v-nigrum NEEDHAM, Zool. Sinica, (A) 11: 269. Type male: unknown; type-locality: Szechwan. 1930 Coenagrion barbatum NEEDHAM, Zool. Sinica, (A) 11: 270. Type male: unknown; type-locality: Szechwan. 1956 Cercion v-nigrum - ASAHINA, Ent. Medd., 27 ( 4 - 5 ) : 219. 1 9 5 6 Coenagrion barbatum - ASAHINA, Ent. Medd., 27 ( 4 - 5 ) : 2 1 9 [proposed synonymy with v-nigrum

(NEEDHAM, 1 9 3 0 ) ] .

Distribution: China. Genus Chromagrion

NEEDHAM

1903 Chromagrion NEEDHAM, Bull. 68 Ν.Υ. State Mus.: 246. Type species: Erythromma conditum H Ä G E N , 1 8 7 6 (monotypy). - 1 9 1 0 Chromagrion - M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 (1): 67.

conditum

(HÄGEN)

1876 Agrion (Erythromma) conditum (HÄGEN MS) in SELYS, Synopsis des Agrionines, Add. 5: 195. Type male: Hagen's Collection; type-locality: Maryland. 1890 Erythromma conditum - KIRBY, Syn. Cat. Neur.-Odon., London: 152. 1903 Chromagrion conditum - NEEDHAM, Bull. 68 N.Y. State Mus.: 246 (as generotype of Chromagrion NEEDHAM, 1903). 1910 Chromagrion conditum - MUTTKOWKI, Bull. Publ. Mus., Milwaukee, 1 (1): 67 (Carolina, Indiana, New Jersey).

Distribution: U.S.A.

219

Genus Coenagrion

KIRBY

1775 Agrion FABRICIUS (partim), Syst. Ent.: 425. Type species: Agrion puella LINNEAUS, 1758 (first emendation as Libellula). - 1815 Agrion - LEACH, Edinb. Encycl., 9: 137. - 1836 Agrion STEPHENS, 111. Brit. Ent., 6: 71. - 1850 Agrion - SELYS & HAGEN, Revue des Odonates Paris: 171 (with subgenera Nehalennia, Erythromma, Ischnura, Agrion s Str.). - 1876 Agrion (Agrion) - SELYS, Synopsis des Agrionines, Add. 5: 136 (revision). - 1890 Coenagrion KIRBY, Syn. Cat. Neur.-Odon. London: 148. Type species: Libellula puella LINNEAUS, 1758 (new name for Agrion FABRICIUS, 1775). - 1905 Agrion - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 818 (key to species). - 1910 Coenagrion - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 65 (on applicability of name, Agrion versus Coenagrion, the former to be preferred). - 1950 Agrion - MORERA, Inst. Esp. Ent., Madrid: 85 (key to Spanish species). - 1954 Coenagrion - LIEFTINCK, Treubia, 22 Suppl.: 69. - 1954 Cercion - LIEFTINCK, Treubia, 22 Suppl.: 69 (incorrect synonymy with Coenagrion KIRBY, 1890). - 1984 Coenagrion - STEINMANN, Fauna hung., 160: 19 (key to Hungarian species). - 1984 Coenagrion - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 58.

adelais

(FOURCROIX)

1785 Libellula adelais FOURCROIX, Ent. Par., 2: 346. Type: unknown; type-locality: France. 1890 Coenagrion adelais - KIRBY, Syn. Cat. Neur.-Odon. London: 151.

Distribution: France. angulatum

WALKER

1912 Coenagrion angulatum WALKER, Canad. Ent., 44: 256. Type male: Walker's Collection; typelocality: Canada. 1913 Enallagma lunifera MUTTKOWSKI, Bull. Wisconsin Soc., 10:164. Type: unknown; type-locality: Iowa. 1927 Enallagma lunifera - BYERS, Trans. Am. ent. Soc., 53: 249 (proposed synonymy with angulatum W a l k e r , 1912).

Distribution: Canada, and Northern U.S.A. armatum

(CHARPENTIER)

1840 Agrion armatum CHARPENTIER, Libell. europ. Lipsiae: 164. Type male: not given; type-locality: Germany: Lüneburg. 1850 Agrion armatum (HEYER) - SELYS & HÄGEN, Revue des Odonata, Paris: 193 (redescription) (Sweden, Germany). 1876 Agrion (Agrion) armatum - SELYS, Synopsis des Agrionines, Add. 5: 160 (Sweden, Germany, Poland). 1890 Coenagrion armatum - KIRBY, Syn. Cat. Neur.-Odon. London: 149. 1905 Agrion armatum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 819 (Germany, Siberia, Russia). 1948 Puella armata - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 156 (Armenia). 1956 Coenagrion armatum - FRÄSER, R. Ent. Soc. London, 1 (10): 29 (England). 1973 Agrion armatum - BELYSHEV, The Dragonflies of Siberia, 1 (2): 542. 1984 Coenagrion armatum - STEINMANN, Fauna Hung., 160: 21 (from East Asia to Central Europe).

Distribution: Mongolia and Siberia, East, Central and West Europe. brisbanense

(TILLYARD)

1917 Agrion brisbanense TILLYARD, Proc. Linn. Soc. N.S. Wales, 42: 477. Type male: Tillyard's Collection: type-locality: Queensland. 1984 Coenagrion brisbanense - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 59.

Distribution: Australia.

220 caerulescens caerulescens

(FONSCOLOMBE)

1838 Agrion caerulescens FONSCOLOMBE, Ann. Soc. Ent. France, 7: 569. Type male: Nation. Hist. Mus., Paris; type-locality: France. 1838 Agrion pulchella FONSCOLOMBE, Ann. Soc. Ent. France, 7: 569. Type male: Nation. Hist. Mus., Paris; type-locality: France. 1842 Agrion aquisextanum RAMBUR, Hist. nat. Ins. Neuropt. Paris: 267. Type: sex and locality not given (Spain ?). 1876 Agrion (Agrion) caerulescens - SELYS, Synopsis des Agrionines, Add. 5: 161 (France, Spain, Sardinia). 1890 Coenagrion caerulescens - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1905 Agrion caerulescens - JACOBSON & BIANKI, Pryam. lozhnos, Ross, imp.: 823 (France, Sardinia, Sicily). 1919 Coenagrion caerulescens var. pygmaea NAVÄS, Mem. Ac. Barcelona, 15 (5): 10. Male from Lerida. 1940 Coenagrion caerulescens - NIELSEN, Mem. Soc. ent. ital., 19: 341 (Sardinia). 1 9 5 0 Agrion coerulescens - M O R E R A - Inst. Esp. Ent. Madrid: 8 6 (Spain). 1959 Agrion caerulescens caerulescens - SCHMIDT, Bull. Inst. Sei. ent. Belg., (2) 35: 12 (key to subspecies).

Distribution: West Mediterranean region, caerulescens caesarum

(SCHMIDT)

1959 Agrion caerulescens caesarum SCHMIDT, Bull. Inst. Sei. ent. Belg., 35 (29): 13. Type male: R. Mus., Brussels; type-locality: Italy.

Distribution: Italy. caerulescens theryi

(SCHMIDT)

1959 Agrion caerulescens theryi SCHMIDT, Bull. Inst. Sei. ent. Belg., 35 (29): 13. Type male: Schmidt's Collection; type-locality: Algeria.

Distribution: Algeria. Morocco, and Tunisia. castellani

ROBERT

1948 Coenagrion castellani ROBERT, Proc. R. ent. Soc. London, (B) 17: 63. Type male: unknown; type-locality: Italy. 1989 Coenagrion castellani - A z z o u z , G U E M M O U H & AGUESSE, Odonatologica, 18 (3): 280 (Morocco).

Distribution: Italy, and Morocco, chusanicum

NAVÄS

1933 Coenagrion chusanicum N A V Ä S , M U S . Heude Notes ent. chinoise, 9 : 4 . Type: unknown, perhaps in Mus. Heude; type-locality: China.

Distribution: China. concinnum

(JOHANSSON)

Agrion concinnum JOHANSSON (nec R A M B U R , 1 8 4 2 ) , Odon. Suec., Nr. 7 : 1 0 6 . Type male: unknown; type-locality: Suede. 1890 Coenagrion concinnum - KIRBY, Syn. Cat. Neur.-Odon. London: 148. 1894 Agrion johanssoni WALLENGREN, Ent. Tidschr., 15: 267 (new name for Agrion concinnum JOHANSSON, 1 8 5 9 , a junior secondary homonymy of Agrion concinnum R A M B U R , 1 8 4 2 ) . 1 9 0 5 Agrion concinnum - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 1 9 . 1905 Agrion johanssoni-JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 819 (proposed synonymy with concinnum JOHANSSON, 1 8 5 9 ) . 1859

221 1954 Coenagrion johanssoni - LONGFIELD, Entomologist's month. Mag., 90: 145. 1984 Coenagrion concinnum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 59.

Distribution: Scandinavia (exept Denmark), Russia, and Siberia to north of the Artie Circle. convalescens

(BARTENEF)

1914 Agrion convalescens BARTENEF, Hör. Soc. ent. ross., 41: 29. Type male: Zool. Mus. Acad. Sei. St. Petersburg; type-locality: Manchuria.

Distribution: Manchuria, dorothea 1785 1890 1924 1984

(FOURCROIX)

Libellula dorothea FOURCROIX, Ent. Par., 2: 344. Type: unknown; type-locality: ? France. Coenagrion dorothea - KIRBY, Syn. Cat. Neur.-Odon. London: 151. Coenagrin dorothea FRASER (as sp. nov.) J. Asiat. Soc. Bengal., 19: 462 (Yunnan). Coenagrion dorothea FRASER, 1924 - loc. cit. DAVIES & TOBIN, the Dragonflies of the World, Utrecht, 1: 59 (China).

Distribution: ? France, ? China. dyeri

(FRASER)

1919 Argiocnemis dyeri FRASER, Ree. Indian Mus., 16: 451. Type male: not given; type-locality: India. 1919 Argiocnemis gravelyi FRASER, Ree. Indian Mus., 16: 451. Type female: not given; type-locality: India. 1924 Argiocnemis dyeri - FRASER, Ree. Indian Mus., 26: 498 (W. India). 1924 Coenagrion violacea FRASER, Mem. Dept. Agr. India, 8: 84. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1953 Coenagrion dyeri - LIEFTINCK, Verh. Naturf. Ges. Basel, 64: 162. 1954 Coenagrion dryeri- LIEFTINCK, Treubia, 22 Suppl.: 69 (India). 1954 Argiocnemis gravelyi - LIEFTINCK, Treubia, 22 Suppl.: 69 [proposed synonymy with dyeri (FRASER, 1919)].

1966 Coenagrion violacea - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 218 (as valid sp.).

Distribution: India. ecornutum

(SELYS)

1872 Agrion ecornutum SELYS, Ann.Soc.Ent. Belg., 15: 22. Type male: Selys's Collection; type-locality: Amur. 1876 Agrion (Agrion) ecornutum - SELYS, Synopsis des Agrionines, Add. 5: 168. 1890 Coenagrion exornatum (misprint) - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1905 Agrion ecornutum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 824 (Siberia). 1958 Agrion ecornutum - ASAHINA, Insecta matsum., 22: 64 (Kurile Island). 1984 Coenagrion ecornutum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 59. 1984 Coenagrion exornatum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 59 (as good sp.).

Distribution: Siberia, and Kuril Islands, glaciale glaciale

(HÄGEN) 1872 Agrion glaciale HAGEN, in SELYS, Ann. Soc. Ent. Belg., 15: 41. Type male: Hagen's Collection; type-locality: Irkutzk. 1876 Agrion (Agrion) glaciale - SELYS, Synopsis des Agrionines, Add. 5: 158 (Irkutzk). 1890 Coenagrion glaciale - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1905 Agrion glaciale ~ JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 820 (Siberia).

222 1973 Agrion glaciale - BELYSHEV, The Dragonflies of Siberia, 1 (2): 246.

Distribution: Siberia, and Mongolia. glaciale Orientale

BELYSHEV

1973 Coenagrion glaciale Orientale BELYSHEV, The Dragonflies of Siberia, 1 (2): 101. Type male: unknown; type-locality: Russia.

Distribution: Russia. hastulatum 1825

1876 1890 1905

1940 1956 1961 1984

(CHARPENTIER)

Agrion hastulatum CHARPENTIER (nec BURMEISTER, 1 8 3 9 ) Hor. Ent., Paris: 2 0 . Type male: unknown; type-locality: Lapland. Agrion (Agrion) hastulatum - SELYS, Synopsis des Agrionines, Add. 5: 150 (Finland, Sweden, Russia, Germany, Belgium, Switzerland, Anatolia). Coenagrion hastulatum - KIRBY, Syn. Cat. Neur.-Odon. London: 149. Agrion hastulatum- JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 0 (Siberia, to 6 0 - 6 2 ° ) . Agrion hastulatum - NIELSEN, Mem. Soc. ent. ital., 19: 241 (Sardinia). Coenagrion hastulatum - FRÄSER, R. Ent. Soc. Lond. 1 (10): 29 (England). Agrion hastulatum - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 40 (Kazakhstan). Coenagrion hastulatum - STEINMANN, Fauna Hung., 160: 22 (N. & C. Europe, Hungary, France, Spain).

Distribution: from Siberia to South Europe. holdereri

FÖRSTER

1900 Agrion (Coenagrion) holdereri FÖRSTER, Wiener ent. Ztg., 19: 264. Type male: Förster's Collection; type-locality: Central Asia. 1 9 0 5 Agrion holdereri - JACOBSON & BIANKI, Pryam. lozhnos Ross, imp.: 8 2 0 . 1984 Coenagrion holderevei (misprint) - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 59.

Distribution: Central Asia, hylas

(TRYBOM)

1889 Agrion hylas TRYBOM, Bih. Svensk. Vet. Akad. Handl., 15(4): 12. Type male: Naturh. Riksmus., Stockholm; type-locality: Jenisei. 1 9 0 5 Agrion hylas - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 9 (Siberia). 1955 Agrion ? freyi BILEK, Nachr. Bay. Ent., 3 (10): 98. Type female: Zool. Samml. Bayer. Staat. München; type-locality: Zwingsee (Germany). 1956 Agrion ( = Coenagrion) freyi - BILEK, Nachr. Bay. Ent., 4 (9): 1 (description of male). 1 9 5 6 Coenagrion hylas ussurense BELYSHEV, The Dragonflies of Siberia, 3 : 1 9 6 . Type male: unknown; type-locality: Russia. 1964 Coenagrion hylas - LIEFTINCK, Tijdschr. Ent., 107: 159. 1 9 6 4 Agrion freyi - LIEFTINCK, Tijdschr. Ent., 1 0 7 : 1 5 9 [proposed synonymy with hylas (TRYBOM, 1889)].

1984 Coenagrion hylas - STEINMANN, Fauna Hung., 160: 25 (West Asia, and Germany).

Distribution: Western Asia, and ? Germany (Alps). impar

NEEDHAM

1930 Coenagrion impar NEEDHAM, Zool. Sinica, (A) 11: 271. Type male: Peking Museum; typelocality: China.

Distribution: China.

223 interrogatum

(HAGEN)

1876 Agrion (Agrion) interrogatum HAGEN, in SELYS, Synopsis des Agrionines, Add. 5: 144. Type female: Mus. Comp. Zool., Cambridge; type-locality: Saskatchewan. 1890 Coenagrion interrogatum - KIRBY, Syn. Cat. Neur.-Odon., London: 148.

Distribution: North America. kagiensis

(MATSUMURA)

1911 Agrion (Ischnura) kagiensis MATSUMURA, Mem. Soc. ent. Belg., 18:147. Type male: unknown; type-locality: Formosa.

Distribution: Taiwan. kashmirum 1984

CHOWDHARY

Coenagrion kashmirum C H O W D H A R Y , the World, Utrecht, 1: 60.

1975

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of

Distribution: Kashmir. lanceolatum

(SELYS)

1872 Agrion (Agrion) lanceolatum SELYS, Ann. Soc. Ent. Belg., 15:43. Type male: Selys's Collection; type-locality: Amur. 1890 Coenagrion lanceolatum - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1 9 0 5 Agrion lanceolatum - JACOBSON & B I A N K I , Pryam. lozhnos, Ross, imp.: 8 2 0 (Amur). 1 9 8 4 Coenagrion lanceolatum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 0 ) .

Distribution: Central Asia. lehmanni 1984

KOLENATI

Coenagrion lehmanni Utrecht, 1: 60.

KOLENATI, 1 8 5 6

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Russia. malayanum

(SELYS)

1876 Agrion (Enallagma) malayanum SELYS, Synopsis des Agrionines, Add. 5:119. Type male: Selys's Collection; type-locality: Java. 1890 Enallagma malayanum - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1919 Enallagma malayanum - LAIDLAW, Ree. Indian Mus., 16: 183 (India). 1933 Enallagma malayanum - FRASER, The Fauna of brit. India, Odon., 1: 372. 1934 Agrion malayanum - SCHMIDT, Arch. Hydrobiol. Suppl.: 13: 350. 1953 Coenagrion malayanum - LIEFTINCK, Treubia, 22 Suppl.: 69 (Java). 1955 Coenagrion malayanum - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 75 (Ceylon).

Distribution: India, Sri Lanka, Java. melanoproctum

(SELYS)

1876 Agrion (Agrion) melanoproctum SELYS, Synopsis des Agrionines, Add. 5: 177. Type female: Selys's Collection; type-locality: Melanesia. 1890 Coenagrion melanoproctum - KIRBY, Syn. Cat. Neur. Odon., London: 150. 1 9 8 4 Coenagrion melanoproctum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 0 (Polynesia).

Distribution: Oceania.

224 mercuriale mercuriale

(CHARPENTIER)

1840 Agrion mercuriale CHARPENTIER, Libell. europ. Lipsae: 159. Type male: Nation. Hist. Nat. Mus. Paris; type-locality: Germany. 1842 Agrion fonscolombii RAMBUR, Hist. nat. Ins. Neuropt., Paris: 266. Type male: unknown; typelocality: Germany. 1876 Agrion (Agrion) mercuriale - SELYS, Synopsis des Agrionines, Add. 5:165 (Germany, Belgium, England, France, Suisse, Spain, Portugal). 1876 Agrion fonscolombiiSELYS, Synopsis des Agrionines, Add. 5: 165 [proposed synonymy with mercuriale (CHARPENTIER, 1 8 4 0 ) ] . 1890 Coenagrion mercuriale - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1905 Agrion mercuriale mercuriale - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 823 (Europe). 1948 Puella mercuriale - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 158 (Armenia). 1950 Agrion mercuriale - MORERA, Inst. Esp. Ent., Madrid, 89 (Spain). 1956 Coenagrion mercuriale - FRÄSER, R. Ent. Soc. London, 1 (10): 28 (England). 1984 Coenagrion mercuriale - SEINMANN, F a u n a Hung., 160: 27 (Italy, Austria, Germany, Poland, England).

Distribution: from Armenia to Spain, and North Africa. mercuriale hermeticum

(SELYS)

1872 Agrion hermeticum SELYS, Ann. Soc. Ent. Belg., 15: 45. Type male: R. Mus. Brussels; typelocality: Algeria. 1876 Agrion (Agrion) mercuriale race hermeticum - SELYS, Synopsis des Agrionines, Add. 5: 167. 1890 Coenagrion hermeticum - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1 9 0 5 Agrion mercuriale hermeticum - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 2 3 (Algeria). 1884 Coenagrion hermeticum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 59 (as good species). 1 9 8 9 Coenagrion mercuriale hermeticum - A z z o u z , G U E M M O U H & AGUESSE, Odonatologica, 1 8 ( 3 ) : 282 (Morocco).

Distribution: Algeria, Morocco, and Tunisia. minutissimum

(SELYS)

1876 Agrion (Agrion) minutissimum (BATES MS) SELYS, Synopsis des Agrionines, Add. 5: 140. Type female: Bates's Collection; type-locality: Amazons. 1890 Coenagrion minutissium - KIRBY, Syn. Cat. Neur.-Odon., London: 148.

Distribution: Brazil. ornatum

(SELYS & H Ä G E N )

1850 Agrion ornatum ( H E Y E R M S ) SELYS & H Ä G E N , Revue Odon. Libell. europ. Paris: 203. Type male: Selys's Collection; type-locality: Hannover. 1876 Agrion (Agrion) ornatum - SELYS, Synopsis des Agrionines, Add. 5: 155 (Germany, Holland, Hungary). 1890 Coenagrion ornatum - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1905 Agrion ornatum - JACOBSON & B I A N K I , Pram, lozhnos. Ross, imp.: 822 (Hungary, Germany, Balkan, Russia, Mesopotamia). 1948 Puella ornata - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 156 (Armenia). 1956 Agrion ornatum - ADAMOVIC, Bull. Mus. Hist. nat. Serbe, (Β) 8: 104 (Serbia). 1984 Coenagrion ornatum - STEINMANN, F a u n a Hung., 160: 23 (Hungary). Distribution: C e n t r a l a n d E a s t E u r o p e t o A r m e n i a .

225 pendulum 1939

NEEDHAM & GYGER

Coenagrion pendulum type-locality: Luzon.

NEEDHAM & GYGER,

Philipp.

J.

Sei., 70

(3): 2 8 7 .

Type male: not given;

Distribution: Philippine Islands. ponticum

(BARTENEF)

comb. nov.

1929 Agrion ponticum BARTENEF, Zool. Anz., 85: 64. Type male: not given; type-locality: Caucasus. 1930 Agrion ponticum - BARTENEF, Arb. Nord.-kauk. Assoc. wiss. Inst., 72: 65 (as sp. nov. from Zoological Record).

Distribution: Caucasus. puella puella 1758 1823 1823 1825 1832 1836 1876 1890

(LINNEAUS)

Libellula puella LINNAEUS, Syst. Nat., 1: 546. Type: unknown; type-locality: Sweden. Agrion pupa HANSEMANN, Wiedem. Zool. Mag., 2 (1): 155. Type: unknwon. Agrion puella - VAN DER LINDEN, Opusc. Sei., 4: 104. Agrion furcatum CHARPENTIER, Hor. Ent., Paris, 2:18. Type male: Halle Museum; type-locality: Germany. Puella irina BRÜLLE, Expl. Moree, 2: 105. Type: unknown; type-locality: Greece. Agrion annulare STEPHENS, 111. Brit. Ent. Mand., 6: 74. Type male: British Museum (Nat. Hist.), London; type-locality: unknown. Agrion (Agrion) puella - SELYS, Synopsis des Agrionines, Add. 5: 147 (Europe, West Asia, North Africa). Coenagrion puella- KIRBY, Syn. Cat. Neur.-Odon., London: 148 (as generotype of Coenagrion KIRBY, 1890).

1890 Coenagrion irina - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1905 Agrion puella - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 822 (Europe, Corse, Sicilia, Sardinia). 1928 Agrion puella - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 25 (Egypt). 1948 Puella puella - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 157 (Armenia). 1950 Agrion puella - MORERA, Inst.Esp.Ent. Madrid: 88 (Spain). 1956 Agrion puella - ADAMOVIC, Bull.Mus. Hist. nat. Serbe, (Β) 8: 116 (Serbia). 1 9 5 6 Coenagrion puella - FRÄSER, R . Ent. Soc. Lond., 1 ( 1 0 ) : 2 9 (England). 1961 Agrion puella puella - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 39 (Kazakhstan). 1984 Coenagrion puella puella - STEINMANN, Fauna Hung., 160: 24 (Hungary).

Distribution: Europe (except Iceland), West Asia, North Africa. puella kocheri (SCHMIDT) comb. nov. I960 Agrion puella kocheri SCHMIDT, C.R. SOC. Sei. nat. Moroc., 26: 124. Type: unknown; typelocality: Morocco.

Distribution: Morocco, puella syriacum (MORTON) comb. nov. 1924 Agrion puella syriaca MORTON, Trans, ent. Soc. London, 1924: 32. Type male: Morton's Collection; type-locality: Palestine. 1960 Agrion puella syriacum - SCHMIDT, C.R. Soc. Sei. nat. Moroc, 26: 123 (Morocco).

Distribution: Morocco, Israel, Turkey to Caucasus. pulchellum (VAN

DER LINDEN)

1823 Agrion pulchella VAN DER LINDEN, Opusc. Sei., 4: 103. Type male: Mus. Naturk., Berlin; typelocality: Germany.

226 1825 Agrion interruptum CHARPENTIER, Hor. Ent. 2: 16. Type female: Halle Museum; type-locality: Germany. 1840 Agrion elegantulum ZETTERSTEDT (nom.nud.) Ins. lapp. descr.: 1037. 1876 Agrion (Agrion) pulchellum - SELYS, Synopsis des Agrionines, Add. 5: 145 (Europe, Asia Minor, Turkestan, Algeria). 1890 Coenagrion pulchellum - KIRBY, Syn. Cat. Neur.-Odon., London: 148. 1905 Agrion pulchellum - JACOBSON & BIANKI, Pryam. lozhnos. Ross imp.: 821 (Corse, Anatolia, Pyrennes). 1913 Agrion pulchellum ab. bilineatum SCHIRMER, Arch. Naturg., 79: 20. Male from Germany. 1948 Puella pulchella - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 157 (Armenia). 1950 Agrion pulchellum - MORERA, Inst. Esp. Ent., Madrid, 87 (Spain). 1956 Coenagrion pulchellum - FRÄSER, R. ent. Soc. Lond., 1 (10): 29 (England). 1984 Coenagrion pulchellum

interruptum

- STEINMANN, F a u n a Hung., 160 24 (Hungary).

Distribution: Europe, Asia Minor, West Asia, Algeria. resolutum

(HÄGEN)

1876 Agrion (Agrion) resolutum HAGEN, in SELYS, Synopsis des Agrionines, Add. 5:155. Type male: Mus. Comp. Zool., Cambridge; type-locality: H u d s o n ' s Bay. 1890 Coenagrion resolutum - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1911 Enallagma walkeri MUTTKOWSKI, Bull. Wise. nat. Hist. Soc., 9: 33. Type male: unknown; type-locality: Wisconsin. 1912 Enallagma walkeri - MUTTKOWSKI, Bull. Wise. nat. Hist. Soc., 9: 166 [proposed synonymy w i t h resolutum

(HÄGEN, 1 8 7 6 ) ] .

1943 Coenagrion resolutum - WALKER, Canad. Ent., 75: 81.

Distribution: North America. scitulum

(RAMBUR)

1842 Agrion scitulum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 266. Type male: Mus. nat. Hist., Paris; type-locality: France. 1842 Agrion distinctum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 269. Type female: Mus. nat. Hist., Paris; type-locality: Sardinia. 1876 Agrion (Agrion) scitulum - SELYS, Synopsis des Agrionines Add. 5: 163 (Corsica, Sardinia, Algeria, France, Spain, Belgium). 1887 Agrion scitulum - SELYS, Ann. Soc. ent. Belg., 31:47 (France, Italy, Creta, Algeria, Asia Minor). 1890 Coenagrion scitulum - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1905 Agrion scitulum - JACOBSON & BIANKI, Pryam. lozhnos. Ross.imp.: 822 (Belgium, Italy, Algeria, Syria). 1948 Puella scitula - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 158 (Armenia). 1950 Agrion scitulum - MORERA, Inst. Esp. Ent., Madrid: 88 (Spain). 1956 Coenagrion scitulum - FRÄSER, R. Ent. Soc. London, 1 (10): 28 (England). 1984 Coenagrion scitulum - STEINMANN, F a u n a Hung., 160: 27 (Hungary).

Distribution: Mediterranean region of Europe, Asia Minor, Algeria. simillimum

(BARTENEF)

1912 Agrion simillimum BARTENEF, Ann. Mus. Zool. Acad. Imp. Sei. St.-Petersburgh, 16: 444. Type male: Zool. Mus. Acad. Sei. St Petersburg; type-locality: Hakodate. 1984 Coenagrion simillimum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 60.

Distribution: Japan. sophia

(FOURCROIX)

1 7 8 5 L i b e l l u l a s o p h i a FOURCROIX, E n t . P a r . , 2 : 3 4 5 . T y p e m a l e : u n k n o w n ; t y p e - l o c a l i t y : F r a n c e .

227 1890 Coenagrion sophia-

KIRBY, Syn. Cat. Neur.-Odon., London: 151.

Distribution: France. vernale

(HÄGEN)

1839 Agrion vernale HAGEN, Preuss. Provinzbl., 21: 56. Type: unknown; type-locality: Silesia. 1840 Agrion lunulatum CHARPENTIER, Libell. eur. Lipsiae: 162. Type female: Mus. Hist., nat., Paris; type-locality: Silesia. 1876 Agrion (Agrion) vernale - SELYS, Synopsis des Agrionines, Add. 5: 163 1890 Coenagrion vernale - KIRBY, Syn. Cat. Neur.-Odon., London: 149. 1 9 0 5 Agrion vernale - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 1 (Russia. Siberia). 1948 Puella lunulata - AKRAMOVSKI, Zool. Zbom., Moscow 5: 156 (Armenia). 1954 Coenagrion vernale - LONGFIELD, Entomologist's month. Mag., 90: 146. 1961 Agrion vernale - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 40 (Kazakhstan). 1974 Agrion vernale - MIELEWCZYK, Odonatologica, 3 (4): 267 (nom. nud.; replaced name by Agrion lunulatum

CHARPENTIER,

1840).

1984 Coenagrion vernale - STEINMANN, Fauna Hung., 160: 21 (Hungary).

Distribution: from Central Asia to Central Europe. Genus Erythromma

CHARPENTIER

1840 Agrion (Erythromma) CHARPENTIER, Libell. europ. Lipsiae: 20. Type species: not given, first emendation: Agrion chloridion CHARPENTIER, 1 8 2 5 . - 1 8 5 0 Agrion (Erythromma) - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 7 4 (type species: Agrion najas HANSEMANN, 1 8 2 3 ) . 1876 Agrion (Erythromma) - SELYS, Synopsis des Agrionines, Add. 5 : 1 9 0 . - 1890 Erythromma - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 5 2 (designated type: Agrion najas HANSEMANN, 1 8 2 3 ) . - 1 9 0 5 Erythromma - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 9 (key to species). - 1920 Pseuderythromma KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type species: Agrion viridulum CHARPENTIER, 1 8 4 0 (original designation). - 1 9 6 2 Erythromma - PINHEY, Publ. cult. Diam. Angola, 59: 131. - 1984 Erythromma - STEINMANN, Fauna Hung., 160: 30.

humerale

SELYS

1887 Erythromma humerale SELYS, Ann. Soc. Ent. Belg., 31: 61. Type male: R. Mus. Brussels; typelocality: Irkutzk. 1 9 0 5 Erythromma humerale- JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 9 (Irkutzk, Amur).

Distribution: Siberia. najas

(HANSEMANN)

1823 Agrion najas HANSEMANN, Widern. Zool. Mag., 2 (1): 158. Type male: unknown; type-locality: Germany. 1825 Agrion analis VAN DER LINDEN, Mon. Libell. Europ.: 40. Type: unknown. 1825 Agrion chloridion CHARPENTIER, Hor. Ent. Paris: 14. Type male: Mus. nat. Hist., Paris; typelocality: Europe. 1836 Agrion lincolniense STEPHENS, 111. Brit. Ent. Mand., 6: 74, emendation. 1840 Eryhromma najas - CHARPENTIER, Libell. europ. Lipsiae: 20 (as generotype of Erythromma CHARPENTIER, 1840).

Agrion (Erythromma) najas - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 7 7 (Sweden, England, Ireland, Belgium, France, Italy, Portugal, Poland). 1876 Agrion (Erythromma) najas - SELYS, Synopsis des Agrionines, Add. 5:192 (North and Central Europe). 1890 Erythromma najas - KIRBY, Syn. Cat. Neur.-Odon., London: 152 (as generotype of Erythromma CHARPENTIER, 1840). 1890 Agrion analis - KIRBY, Syn. Cat. Neur.-Odon., London: 152 [proposed synonymy with najas 1850

(HANSEMANN, 1823)].

228 1890 Agrion chloridion - KIRBY, Syn. Cat. Neur.-Odon., London: 152 [proposed synonymy with najas

(HANSEMANN, 1 8 2 3 ) ] .

1890 Agrion lincolniense - KIRBY, Syn. Cat. Neur.-Odon., London: 152 [proposed synonymy with najas

(HANSEMANN, 1 8 2 3 ) ] .

Erythromma najas - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 2 9 (Europe, Siberia). 1956 Erythromma najas - FRÄSER, R. Ent. Soc. London, 1 (10): 32 (England). 1984 Erythromma najas - STEINMANN, F a u n a Hung., 160: 31 (Hungary). 1 9 8 4 Erythromma najas f. baicalensis BELYSHEV, 1 9 6 4 - loc. cit. D A VIES & TOBIN The Dragonflies of the World, Utrecht, 1: 61. (Japan). 1 9 8 4 Erythromma najas f. bipunctatum SCHIRMER, 1 9 1 3 - loc cit. D A VIES & TOBIN The Dragonflies of the World, Utrecht, 1: 61. 1 9 8 4 Erythromma najas f. immaculatum SCHIRMER, 1 9 1 3 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 61. 1 9 8 4 Erythromma najas f. puellare SCHIRMER, 1 9 1 3 - loc. cit. D A VIES & TOBIN, The Dragonflies of 1905

the World, Utrecht, 1: 61.

Distribution: Europe, but almost absent from the Mediterannean region, and Palaerctical West Asia. viridulum

(CHARPENTIER)

1840 Agrion viridulum CHARPENTIER, Libell. europ. Lipsiae: 149. Type male: Mus. Nat. Hist., Paris; type-locality: Silesia. 1842 Agrion bremii RAMBUR, Hist. nat. Ins. Neuropt. Paris: 265. Type male: not given; type-locality: Sicily. 1876 Agrion (Erythromma) viridulum - SELYS, Synopsis des Agrionines, Add. 5: 194 (Silesia, Belgium, Sicily, Asia Minor). 1890 Erytromma viridulum - KIRBY, Syn. Cat. Neur.-Odon., London: 152. 1 9 0 5 Erythromma viridulum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 0 (Asia Minor, Europe, Russia, Turkestan). 1920 Pseuderythromma viridulum - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Pseuderythromma K E N N E D Y , 1 9 2 0 ) . 1948 Erythromma viridulum - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 158 (Armenia). 1950 Erythromma viridulum - MORERA, Inst. Esp. Ins. Ent. Madrid: 83 (Spain). 1984 Erythromma viridulum - STEINMANN, F a u n a Hung., 160: 32 (Hungary). 1 9 8 4 Erythromma viridulum orientate SCHMIDT, 1 9 6 0 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 61 (Middle East). 1 9 8 4 Erythromma viridulum f. latistrigii L E R O I , 1 9 1 4 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 61 (Slovenia). 1 9 8 8 Erythromma viridulum Orientale - D U M O N T , DEMIRSOY & M E R T E N S , Notul. odonatol., 3 ( 2 ) : 24 (Turkey).

Distribution: Morocco, Egypt, Anatolia, Turkestan, Armenia, Europe (except North-west territories).

Genus Himalagrion

FRASER

1919 Himalagrion FRÄSER, Ree. Indian Mus., 16: 452. Type-species: Himalagrion exlamationis SER, 1919 (original designation).

exclamationis

FRÄ-

FRASER

1919 Himalagrion exclamationis FRASER, Ree. Indian Mus., 16: 453. Type male: Indian Mus.; typelocality: Darjeeling (as generotype of Himalagrion FRASER, 1919). Distribution: I n d i a .

229 pithoragarhicum 1984

SAHNI

Himalagrion pithoragarhicum World, Utrecht, 1: 61.

SAHNI, 1 9 6 4

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: India. Genus Nehalennia

SELYS

1850 Agrion (Nehalennia) SELYS, Revue des Odonates, Paris: 172. Type-species: Agrion speciosum CHARPENTIER, 1840 (original designation). - 1876 Agrion (Nehalennia)-SELYS, Synopsis des Agrionines, Add. 5: 125. - 1876 Argiallagma SELYS, Bull. Acad. r. Belg., (2) 41: 498. Typespecies: Trichocnemis minutum SELYS, 1 8 5 7 (original designation). - 1 8 9 0 Nehalennia-KIRBY, Syn. Cat. Neur.-Odon., London: 1 4 7 . - 1 9 0 5 Nehalennia- JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 827 (key to species). - 1919 Agrion (Nehalennia)-CALVERT, Trans. Am. ent. Soc., 45: 350. - 1957 Argiallagma - LIEFTINCK, N o v a Guinea, (N.S.) 8: 41 (from Argiinae to Coenagrioninae). - 1 9 8 4 Nehalennia „ C A L V E R T , 1 9 0 7 " (misprint) - D A VIES & TOBIN The Dragonflies of the World, Utrecht, 1: 61.

arawakai

MATSUMURA

1931 Nehalennia arawakai MATSUMURA, 6000 111. Ins. Japan: 1461. Type male: M a t s u m u r a ' s Collection; type-locality: Japan.

Distribution: Japan. atrinuchalis

SELYS

1876 Agrion (Nehalennia) atrinuchalis SELYS, Synopsis des Agrionines, Add. 5: 127. Type male: Mus. Nat. Hist., Paris; type-locality: Shanghai. 1890 Nehalennia atrinuchalis - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1905 Nehalennia atrinuchalis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 827.

Distribution: China. gracilis

MORSE

1895 Nehalennia gracilis MORSE, Psyche, 7: 274. Type male: Mus. Comp. Zool. Cambridge; typelocality: Carolina.

Distribution: integricollis

U.S.A.

CALVERT

1913 Nehalennia intergicollis CALVERT, Ent. News, 24: 321. Type male: Calvert's Collection; typelocality: New Jersey.

Distribution: irene

U.S.A.

HÄGEN

1861 Agrion (Nehalennia) irene HÄGEN, Syn. Neur. N. Amer., Washington: 74. Type male: Hagen's Collection; type-locality: Chicago. 1876 Agrion (Nehalennia) irene - SELYS, Synopsis des Agrionines, Add. 5: 130 (Florida, Visconsin, Illinois, New Jersey, Maine, Massachussett). 1890 Nehalennia irene, - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1914 Nehalennia carlotta BUTLER, Canad. Ent., 46: 347. Type male: Butler's Collection; type-locality: New York. 1 9 2 8 Nehalennia carlotta - BYERS, Ent. News, 39: 2 3 0 (proposed synonymy with irene H Ä G E N , 1 8 6 1 ) .

Distribution:

U.S.A.

230 minuta (SELYS) 1910 Trichocnemis minutum SELYS, 1857 - loc. cit. MUTTKOWSKI, Publ. Mus. Milwaukee, 1 (1): 54. Type male: Selys's Collection; type-locality: C. America. 1861 Agrion aduncum HAGEN, Syn. Neur. N. Amer., Washington: 79. Type male: Hagen's Collection; type-locality: N. America. 1876 Argiallagma minutum - SELYS, Bull. Acad. r. Belg., (2) 41: 498 (as generotype of Argiallagma SELYS, 1 8 7 6 ) .

Distribution: North America. pallidula

CALVERT

1913 Nehalennia pallidula CALVERT, Ent. News, 24: 373. Type male: Calvert's Collection; type-locality: Florida.

Distribution: U.S.A. selysi

KIRBY

1876 Agrion (Nehalennia) sophia SELYS (nec SELYS, 1840), Synopsis des Agrionines, Add. 5: 131. Type male: Selys's Collection: type-locality: Minas Geraes. 1890 Nehalennia selysi KIRBY, Syn. Cat. Neur.-Odon., London: 147 (new name for sophia SELYS, 1876, a junior secondary homonymy of sophia SELYS, 1840). 1984 Nehalennia minuta selysi - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 61.

Distribution: Brazil. speciosa (CHARPENTIER) 1840 Agrion (Ischnura) speciosum CHARPENTIER, Libell. europ., Lipsiae: 151. Type: not given; typelocality: Germany. 1840 Agrion sophia SELYS (nec SELYS, 1876), Monogr. Libell. europ. Paris: 213. Type: unknown. 1850 Agrion (Nehalennia) speciosum - SELYS & HÄGEN, Revue des Odonates, Paris: 173 (as generotype of Nehalennia SELYS, 1850) (Belgium, Switzerland, Germany, Sweden) (redescription). 1876 Agrion (Nehalennia) speciosa - SELYS, Synopsis des Agrionines, Add. 5: 128 (Belgium, Germany, Sweden). 1890 Nehalennia speciosa - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1905 Nehalennia speciosa - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 827 (Europe). 1960 Nehalennia speciosa - HOFFMANN, Archives, 27: 223 (Luxemburg).

Distribution: Central and East Europe to Japan.

Genus Neoerythromma

KENNEDY

1920 Neoerythromma KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type-species: Agrion cultellatum SELYS, 1876 (original designation).

(Enallagma)

cultellatum (SELYS) 1876 Agrion (Enallagma) cultellatum (HÄGEN MS) SELYS, Synopsis des Agrionines, Add. 5: 107. Type male: Hagen's Collection; type-locality: Cuba. 1890 Enallagma cultellatum - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1919 Agrion (Enallagma) - CALVERT, Trans. Am. ent. Soc., 45: 353 (Guatemala). 1920 Neoerythromma cultellatum - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Neoerythromma KENNEDY, 1920).

Distribution: Cuba, Jamaica, Haiti, Guatemala.

231 gladiolatum WILLIAMSON 1930 Neoerythromma gladiolatum WILLIAMSON, OCC. P a p . M u s . Zool. Univ. Michigan, 216: 7. Type male: Williamson's Collection; type-locality: Mexico.

Distribution: Mexico. Genus Pyrrhosoma

CHARPENTIER

1840 Agrion(Pyrrhosoma) CHARPENTIER, Libell. europ. Lipsiae: 19. Type-species: Libellula H A R R I S , 1 8 7 2 ( m o n o t y p y ) . - 1 8 5 0 Agrion

(Pyrrhosoma)

minimum

- SELYS & H Ä G E N , R e v u e d e s O d o -

natates, Paris: 178. - 1876 Agrion (Pyrrhosoma) - SELYS, Synopsis des Agrionines, A d d . 5: 183. - 1890 Pyrrhosoma - KIRBY, Syn. C a t . N e u r . - O d o n . , L o n d o n : 151 (type: Libellula nymphula

S U L Z E R , 1 7 7 6 ) . - 1 9 0 5 Pyrrhosoma

- JACOBSON & BIANKI, P r y a m . l o z h n o s . R o s s , i m p . :

827 (key to species). - 1919 Agrion (Pyrrhosoma) - CALVERT, Trans. A m . ent. Soc., 45: 354. - 1950 Pyrrhosoma - MORERA, Inst. Esp. Ent. M a d r i d : 82 (key to Spanish species).

nymphula nymphula (SULZER) 1776 Libellula nymphula SULZER, Gesch. Ins.: 24. Type: u n k n o w n ; type-locality: Switzerland. 1782 Libellula minimum HARRIS, Exp. Eng. Ins.: 29. Type male: Halle M u s e u m ; type-locality: Germany. 1823 Agrion amazon HANSEMANN, Wiedem. Zool. M a g . , (2) 1: 152. Type: u n k n o w n . 1825 Agrion minimum - CHARPENTIER. H o r . Ent.: 13. 1825 Agrion sanguineum VAN DER LINDEN, M o n . Libell. Eur.: 41. Type male: M u s . N a t u r k . , Berlin, type-locality: G e r m a n y . 1836 Agrion fulvipes STEPHENS, 111. Brit. E n t . M a n d . , 6: 75. T y p e j u v . : British M u s e u m ( N a t . Hist.), L o n d o n , type-locality: u n k n o w n . 1845 Erythromma chloridum EVANS, Brit. Libell.: 6. Type: u n k n o w n . 1850 Agrion (Pyrrhosoma) minimum - SELYS & HÄGEN, Revue des O d o n a t e s , Paris: 178 (as generotype of Pyrrhosoma CHARPENTIER, 1840). (Sweden, England, H o l l a n d , Belgium, Italy, G e r m a n y , Spain, France). 1876 Agrion (Pyrrhosoma) minimum - SELYS, Synopsis des Agrionines, A d d . 5: 186 (Sweden, England, H o l l a n d , Belgium, France, G e r m a n y , Italy, Serbia, C o r f u , Spain). 1890 Pyrrhosoma nymphula - KIRBY, Syn. C a t . N e u r . - O d o n . , L o n d o n : 151 (as generotype of Pyrrhosoma

1905 1950 1954 1956

CHARPENTIER, 1 8 4 0 ) .

Pyrrhosoma Pyrrhosoma Pyrrhosoma Pyrrhosoma

nymphula nymphula nymphula nymphula

-

JACOBSON & BIANKI, P r y a m . lozhnos, Ross, imp.: 838 (Russia). MOREIRA, Inst. Esp. Ent. M a d r i d : 82 (Spain). LONGFIELD, Entomologist's m o n t h . M a g . , 90: 145. FRÄSER, R. Ent. Soc. L o n d o n , 1 (10): 31 (England).

1984 Pyrrhosoma

nymphula

- STEINMANN, F a u n a Hung., 160: 19.

Distribution: Europe to Siberia. nymphula elisabethae

SCHMIDT

1948 Pyrrhosoma elisabethae SCHMIDT, Opusc. ent. L u n d , 13. 72. Type male: L u n d M u s e u m ; typelocality: Greece. 1954 Pyrrhosomanymphulaxa.eeelisabethae-BUCHHOLTZ,

B o n n . z o o l . Beitr. S o n d e r b a n d , 1954: 58.

Distribution: Greece. nymphula interposita

VARGA

1968 Pyrrhosoma nymphula interposita VARGA, A c t a Biol., Debrecina, 6: 187. Type male: Varga's Collection; type-locality: H u n g a r y .

Distribution: Hungary.

232 tinctipennis (MCLACHLAN) 1894 Erythromma tinctipennis MCLACHLAN, Ann. Mag. nat. Hist., (6) 13: 436. Type female: British Museum (Nat. Hist.), London; type-locality: West China. 1896 Erythromma tinctipennis-MCLACHLAN, Ann. Mag. nat. Hist., (6) 17:373 (description of male). 1905 Pyrrhosoma tinctipennis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 829.

Distribution: China. Subfamily Ischnurinae Wings petiolate to level of Ac. Anal vein separate from wing border proximal to Ac; arc at level of distal antenodal. Stigma differs in fore and hind wings of male, latter being like in female. Male seg. 10 with raised bifid tubercle; female usually with vulvar spine. 1957 Ischnurinae FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 49. Type-genus: Ischnura CHARPENTIER 1840 (with genera Ischnura, Pacificagrion, Amorphostigma, Enallagma, Rhodischnura, Aciagrion, Oreagrion, Acanthagrion, Anomalagrion, Amphiagrion, Oxyagrion, Skiallagma, Tigriagrion, Hesperagrion, Argiallagma, Mombagrion, Ischnuragrion, Ceratura).

Genus Acanthagrion SELYS 1876 Agrion (Acanthagrion) SELYS, Synopsis des Agrionines, Add. 5: 60. Type-species: Acanthagrion apicale SELYS, 1876 (designated by KIRBY, Syn. Cat. Neur.-Odon., London: 144). - 1890 Acanthagrion - KIRBY, Syn. Cat. Neur.-Odon., London: 144. - 1916 Acanthagrion - Ris, Arch. Naturg., (A) 82: 120. - 1946 Acanthagrion - FRASER, Trans. R. ent. Soc. London, 96: 38. - 1914 Myagrion FÖRSTER, Arch. Naturg., (A2) 80: 68. Type-species: Myagrion obsoletum FÖRSTER, 1914 (original designation). - 1977 Acanthagrion - LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 152: 18 (revision, key to species). - 1977 Myagrion - LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 56 (proposed synonymy with Acanthagrion SELYS, 1876).

ablutum CALVERT 1909 Acanthagrion gracile ablutum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 161. Type male: Calvert's Collection; type-locality: Bolivia. 1916 Acanthagrion ablutum - KENNEDY, Ent. News, 27: 325. 1916 Acanthagrion ablutum - Ris, Arch. Naturg., (A), 82: 123. (Bolivia). 1948 Acanthagrion ablutum - FRASER, Acta. Zool. lilloana, 5: 50 (Argentina).

Distribution: Bolivia, Argentina. abunae LEONARD 1977 Acanthagrion abunae LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 37. Type male: Leonard's Collection; type-locality: Brazil.

Distribution: Brazil. adustum WILLIAMSON 1916 Acanthagrion adustum WILLIAMSON, Ent. News, 27: 320. Type male: Williamson's Collection; type-locality: Amer. merid. Distribution: G u y a n a .

233 amazonicum

SJÖSTEDT

1917 Acanthagrion amazonicum SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 16. Type male: Riksmus. Stockholm; type-locality: Amazonas.

Distribution: Brazil, apicale

SELYS

1876 Agrion (Acanthagrion) apicale (BATES MS) SELYS, Synopsis des Agrionines, Add. 5: 62. Type male: British Museum (Nat. Hist.), London, type-locality: Para. 1890 Acanthagrion apicale - KIRBY, Syn. Cat. Neur.-Odon. London: 144. 1943 Acanthagrion apicale - SCHMIDT, Beitr. F a u n a Perus, 2: 236 (Peru). 1946 Acanthagrion apicale apicale - FRÄSER, Trans. R. ent. Soc. London, 96: 38. 1946 Acanthagrion apicale descendens FRÄSER, Trans. R. ent. Soc. London, 96, 38. Type male: Leeds Mus., type-locality: Peru. 1966 Acanthagrion apicale descendens - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 189.

Distribution: Peru, Brazil. ascendens

CALVERT

1909 Acanthagrion gracile ascendens CALVERT, Ann. Carnegie Mus. Pittsburgh, 6: 161. Type male: US natn. Mus. Washington, type-locality: M a t o Grosso. 1916 Acanthagrion ascendens - WILLIAMSON, Ent. News, 27: 351. 1916 Acanthagrion ascendens - Ris, Arch. Naturg., (A) 82: 125 (Trinidad). 1932 Acanthagrion ascendens - GEIJSKES, Zool. Meded. Leiden, 14: 255.

Distribution: Brazil, Trinidad. chacoense

CALVERT

1909 Acanthagrion chacoense CALVERT, Ann. Carnegie Mus., Pittsburg, 6:172. Type male: Calvert's Collection; type-locality: Brazil. 1961 Acanthagrion chacoense - CALVERT, Proc. Acad. nat. Sei. Philad., 113: 7 (Peru).

Distribution: Brazil, Peru. chararum

CALVERT

1909 Acanthagrion chararum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6:170. Type male: Calvert's Collection; type-locality: Brazil.

Distribution: Brazil, egleri

SANTOS

1961 Acanthagrion egleri SANTOS, Bol. Mus. Goeldi (Zool.) N.S., 38: 2. Type male: Santos's Collection; type-locality: Brazil.

Distribution: Brazil. gracile gracile

(RAMBUR)

1842 Agrion gracile RAMBUR, Hist. nat. Ins. Neuropt. Paris: 260. Type male: Serville's Collection; type-locality: Brazil. 1876 Agrion (Acanthagrion) gracile - SELYS, Synopsis des Agrionines, Add. 5: 63 (as generotype o f Acanthagrion

1890 Acanthagrion

SELYS, 1 8 7 6 ) .

gracile - KIRBY, Syn. Cat. Neur.-Odon. London: 144.

234 1916 1917 1948 1961 1984

Acanthagrion gracile - Ris, Arch. Naturg., (A) 82: 126 (Guatemala, Costa Rica). Acanthagrion gracile - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 18 (Amazonas). Acanthagrion gracile - FRASER, Acta Zool. lilloana, 5: 51 (Argentina). Acanthagrion gracile - CALVERT, Proc. Acad.nat. Sei. Philad., 113: 7 (Peru). Acanthagrion maculae SJÖSTEDT, 1 9 1 8 - loc. cit. D A VIES & TOBIN The Dragonflies of the World, Utrecht, 1: 63.

Distribution: South America. gracile floridense

FRASER

1946 Acanthagrion gracile floridense FRASER, Trans. R. ent. Soc. London, 96: 37. Type male: Leeds Museum; type-locality: Florida. 1966 Acanthagrion gracile floridense KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 192 [type in: British Museum (Nat.Hist.), London]. 1984 Acanthagrion floridense - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 62.

Distribution: Florida. gracile minarum

SELYS

1876 Agrion (Acanthagrion) gracile race minarum SELYS, Synopsis des Agrionines, Add. 5: 65. Male from Brazil. 1890 Acanthagrion minarum - KIRBY, Syn. Cat. Neur.-Odon., London: 144.

Distribution: Brazil. hermosae

LEONARD

1977 Acanthagrion hermosae LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 51. Type male: Zool. Univ. Mich.; type-locality: Peru.

Distribution: Peru. hildegarda 1984

GLOGER

Acanthagrion hildegarda World, Utrecht, 1: 63.

GLOGER, 1 9 6 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Argentina. indefensum

WILLIAMSON

1916 Acanthagrion indefensum WILLIAMSON, Ent. News, 27: 319. Type male: Williamson's Collection, type-locality: Guyana.

Distribution: Guyana. inexpectum

LEONARD

1977 Acanthagrion inexpectum LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 41. Type male: Zool. Univ. Mich., type-locality: Panama.

Distribution: Panama. jessei

LEONARD

1977 Acanthagrion jessei LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 32. Type male: Zool. Univ. Mich., type-locality: Brazil.

Distribution: Brazil.

235 kennedyi

WILLIAMSON

1916 Acanthagrion kennedii WILLIAMSON, Ent. News, 27: 314. Type male: Williamson's Collection; type-locality: Trinidad. 1932 Acanthagrion kennedii - GEIJSKES, Zool. Meded., Leiden, 14: 254 (Trinidad).

Distribution: Trinidad. lanceum

SELYS

1876 Agrion (Acanthagrion) gracile lancea SELYS, Synopsis des Agrionines, Add. 5: 66. Type male: Selys's Collection; type-locality: Argentina. 1890 Acanthagrion lancea - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1909 Acanthagrion cuyabae CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 166. Type: unknown; type-locality: Brazil. 1909 Acanthagrion flmense CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 168. Type: unknown; type-locality: Brazil. 1909 Acanthagrion freiense CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 168. Type: unknown; type-locality: Brazil. 1977 Acanthagrion cuyabae - LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 102 (proposed synonymy with lanceum SELYS, 1876).

Distribution: Brazil, Argentina. latapistylum

CALVERT

1902 Acanthagrion latapistylum CALVERT, Ann. Mus. Buenos Aires, 7: 26. Type male: Calvert's Collection; type-locality: Paraguay.

Distribution: Paraguay. leonardi

JURZITZA

1 9 8 4 A c a n t h a g r i o n leonardi JURZITZA, 1 9 8 0 -

l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f

the

World, Utrecht, 1: 63.

Distribution: Argentina. longispinosum

LEONARD

1977 Acanthagrion longispinosum LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 147. Type male: Zool. Univ. Mich., type-locality: Brazil.

Distribution: Brazil. luteum

RACENIS

1958 Acanthagrion luteum RACENIS, Acta biol. venezuel., 2: 184. Type: unknown, type-locality: Venezuela.

Distribution: Venezuela. minutum

LEONARD

1977 Acanthagrion minutum LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 119. Type male: Zool. Univ. Mich., type-locality: Venezuela.

Distribution: Venezuela. obsoletum

(FÖRSTER)

1914 Myagrion obsoletum FÖRSTER, Arch. Naturg. (A) 80: 69 (as generotype of Myagrion FÖRSTER, 1914). Type male: Förster's Collection; type-locality: M a d r e de Dios.

236 1916 Acanthagrion luna Ris, Arch. Naturg. (A) 82: 122. Type male: not given; type-locality: Colombia. 1943 Acanthagrion obsoletum - SCHMIDT, Beitr. F a u n a Perus, 2: 240 (Peru). 1946 Acanthagrion apicale luna - FRASER, Trans. R. ent. Soc. London, 96: 38. 1977 Acanthagrion luna - LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 56 [proposed synonymy with obsoletum (FÖRSTER, 1914)]. 1977 Acanthagrion leonora - LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 56 [proposed synonymy with obsoletum (FÖRSTER, 1914)]. 1 9 8 4 Acanthagrion leonora G L O G E R , 1 9 6 7 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 63.

Distribution: Colombia, Peru. peruanum

SCHMIDT

1943 Acanthagrion peruanum SCHMIDT, Beitr. Fauna Perus, 2: 236. Type male: Mus. Naturk. Berlin; type-locality: Peru. 1977 Acanthagrion deceptum LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 110. Type male: Zool. Univ. Mich., type-locality: Peru.

Distribution: Peru. peruvianum

LEONARD

1977 Acanthagrion peruvianum LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 114. Type: unknown; type-locality: Peru.

Distribution: Peru. phallicornis

LEONARD

1977 Acanthagrion phallicornis LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153:114. Type male: Zool. Univ. Mich., type-locality: Brazil.

Distribution: Brazil. quadratum

SELYS

1876 Agrion (Acanthagrion) gracile quadratum SELYS, Synopsis des Agrionines, Add. 5: 65. Type: not designated; type-locality: Buenos Aires. 1984 Acanthagrion quadratum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 63.

Distribution: Argentina. rubrifrons

LEONARD

1977 Acanthagrion rubrifrons LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 25. Type male: Zool. Univ. Mich., type-locality: Brazil.

Distribution: Brazil. taxaense 1984

SANTOS

Acanthagrion taxaense Utrecht, 1: 63.

SANTOS, 1 9 6 5

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Argentina. temporale

SELYS

1876 Agrion (Acanthagrion) temporale SELYS, Synopsis des Agrionines, Add. 5: 68. Type male: Selys's Collection; type-locality: Minas. 1890 Acanthagrion temporale - KIRBY, Syn. Cat. Neur.-Odon., London: 145.

Distribution: Brazil.

237 trilobatum

LEONARD

1977 Acanthagrion trilobatum LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 85. Type male: Zool. Univ. Mich., type-locality: Colombia.

Distribution: Colombia. truncatum

SELYS

1876 Agrion (Acanthagrion) truncatum SELYS, Synopsis des Agrionines, Add. 5: 67. Type male: Selys's Collection; type-locality: Minas. 1890 Acanthagrion truncatum - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1919 Agrion (Enallagma) truncatum - CALVERT, Trans. Am. Ent. Soc., 45: 376 (type in Instituto Havanna) (Cuba).

Distribution: Brazil, Cuba. viduum

SELYS

1876 Agrion (Acanthagrion) gracile vidua SELYS, Synopsis des Agrionines, Add. 5: 66. Type male: Selys's Collection; type-locality: Venezuela. 1890 Acanthagrion vidua - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1977 Acanthagrion risi LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 69. Type male: Zool. Univ. Mich., type-locality: Venezuela. 1 9 8 4 Acanthagrion risi - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 3 (proposed synonymy with viduum SELYS, 1 8 7 6 ) .

Distribution: Venezuela. viridescens

LEONARD

1977 Acanthagrion viridescens LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 98. Type male: Univ. Mich., type-locality: Brazil.

Distribution: Brazil. williamsoni

LEONARD

1977 Acanthagrion williamsoni LEONARD, Misc. Publ. Mus. Zool. Univ. Mich., 153: 66. Type male: Zool. Univ. Mich., type-locality: Colombia.

Distribution: Colombia. yungarum Ris 1916 Acanthagrion yungarum Ris, Arch. Naturg., (A) 82: 124. Type male: not given, type-locality: Peru. 1943 Acanthagrion yungarum - SCHMIDT, Beitr. F a u n a Perus, 2: 236.

Distribution: Peru.

Genus Acanthallagma

WILLIAMSON & WILLIAMSON

1924 Acanthallagma WILLIAMSON & WILLIAMSON, Occ. Pprs. Mus. Zool. Univ. Mich., 154:1. Typespecies: Acanthallagma strohmi, WILLIAMSON & WILLIAMSON, 1924 (original designation).

caeruleum

WILLIAMSON & WILLIAMSON

1924 Acanthallagma caeruleum WILLIAMSON & WILLIAMSON, Occ. Pprs. Mus. Zool. Univ. Mich., 154: 13. Type male: Zool. Univ. Mich., type-locality: Brazil.

Distribution: Brazil.

238 luteum 1924

WILLIAMSON &

WILLIAMSON

Acanthallagma luteum WILLIAMSON & WILLIAMSON, OCC. Pprs. Mus. Zool. Univ. Mich., 154: 16. Type male: Zool. Univ. Mich.; type-locality: Brazil.

Distribution: Brazil, strohmi 1924

WILLIAMSON &

WILLIAMSON

Acanthallagma strohmi WILLIAMSON & WILLIAMSON, OCC. Pprs. Mus. Zool. Univ. Mich., 154: 9 (as generotype of Acanthallagma WILLIAMSON & WILLIAMSON, 1 9 2 4 ) . Type male: Zool. Univ. Mich.; type-locality: Brazil.

Distribution: Brazil. Genus Aciagrion 1891

Agrion (Aciagrion) SELYS, Ann. Mus. civ. Stor. nat. Genova, 3 0 : 5 0 9 . Type-species: Agrion (Pseudagrion) hisopa SELYS, 1 8 7 6 (monotypy). - 1 8 9 2 Aciagrion - SELYS, Odonates de Birmanie: 7 7 . - 1 9 0 9 Mombagrion SJÖSTEDT, Kiliman.-Meru Exp., 2 ( 1 4 ) : 4 4 . Type-species: Mombagrion gracile SJÖSTEDT, 1 9 0 9 (monotypy). - 1 9 1 3 Aciagrion - TILLYARD, Proc. Linn. Soc. N . S . Wales, 3 7 : 4 7 2 . - 1 9 1 7 Mombagrion - SJÖSTEDT, Ark. Zool. Uppsala, 1 1 ( 1 4 ) : 1 4 (redescription). - 1 9 2 4 Aciagrion - L A I D L A W , Proc. U.S. natn. Mus., 6 6 : 1 (key to species). - 1 9 3 6 Aciagrion - LONGFIELD, Trans. R . ent. Soc. London, 85: 4 7 2 . - 1 9 3 7 Aciagrion LIEFTINCK, Nova Guinea (N.S.) 1: 3 2 9 (key to New Guinean species). - 1 9 4 5 Ischnuragrion LONGFIELD, Arq. Mus. Bocage, 16: 6 . Type-species: Ischnuragrion varum LONGFIELD, 1 9 4 7 (original designation). - 1 9 5 4 Aciagrion - LIEFTINCK, Treubia, 22 Suppl.: 7 5 . - 1 9 6 2 Aciagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 3 2 (revision). 1 9 6 2 Ischnuragrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 3 2 (proposed synonymy with Aciagrion SELYS, 1 8 9 1 ) . - 1 9 7 2 Aciagrion - PINHEY, Occ. Pap. natn. Mus. Sth. Rhod., ( 8 ) , 5 ( 1 ) : 1 6 (key to species). - 1 9 7 6 Aciagrion - PINHEY, OCC. Pap. natn. Mus. Monum. Rhod. (Nat.Sei.) 5 ( 1 0 ) : 5 4 5 (key to species). - 1 9 8 2 Aciagrion - LEGRAND, Revue fr. Ent., (N.S.) 4 ( 4 ) : 1 6 1 .

aciculare 1929

1934 1934 1935 1954

SELYS

LIEFTINCK

Aciagrion aciculare LIEFTINCK, Tijdschr. Ent., 72: 1 1 7 . Type male: Buitenzorg Mus.; typelocality: Java. Aciagrion aciculare - SCHMIDT, Arch. Hydrob. Suppl. 13: 344 (Java). Aciagrion aciculare - LIEFTINCK, Treubia, 14 (14): 402 (Java). Aciagrion aciculare - LIEFTINCK, Misc. Zool. Sum., 92-93: 12 (Sumatra). Aciagrion aciculare - LIEFTINCK, Treubia, 22 Suppl.: 75 (Sumatra, Java).

Distribution: Sumatra, Java. africanum

MARTIN

1908 Aciagrion africanum M A R T I N , Ann. Mus. civ. Stor. nat. Genova, 43: 689. Type male: Mus. civ. Stor. nat. Genova; type-locality: Guinea. 1936 Pseudagrion pseuderythromma SCHMIDT, Abh. Senckenb. naturf. Ges.; 433: 67. Type male: Riksmus. Stockholm; type-locality: Congo. 1 9 4 6 Pseudagrion pseuderythromma - FRASER, Entomologist's month. Mag., 8 2 : 2 0 1 [proposed synonymy with congoense (SJÖSTEDT, 1 9 1 7 ) ] . 1 9 6 2 Aciagrion africanum PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 3 2 (Belg. Congo, French Congo, Portuguese Guinea, Cameroon, Uganda). 1962 Pseudagrion pseuderythromma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 132 (proposed synonymy with africanum M A R T I N , 1908). 1972 Aciagrion africanum - PINHEY, Occ. Pap. natn. Mus. South Rhod., (8) 5 (1): 19. Distribution: C e n t r a l A f r i c a .

239 approximans

(SELYS)

1876 Agrion (Pseudagrion) approximans SELYS, Synopsis des Agrionines, Add. 5: 217. Type male: Dresden Mus., type-locality: Malaysia. 1890 Pseudagrion approximans - K I R B Y , Syn. Cat. Neur.-Odon., London: 153 (E. Indies). 1984 Aciagrion approximans - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 64.

Distribution: Malaysia. azureum

FRASER

1922 Aciagrion azureum FRASER, Mem. Dept. Agr. India, 7: 51. Type male: British Museum (Nat.Hist.), London: (ex Pusa Collection); type-locality: Assam. 1964 Aciagrion azureum - SCHMIDT, Ent. Tidschr., 85: 150 (Burma).

Distribution: India, Burma, balachowskyi

LEGRAND

1982 Aciagrion balachowskyi L E G R A N D , Revue fr. Ent. (N.S.), 4 (4): 161. Type male: Nat. Hist., natn. Mus., Paris; type-locality: Gabon.

Distribution: Gabon. borneense Ris 1911 Aciagrion borneense Ris, Ann. Soc. ent. Belg., 55: 234. Type male: Mus. Naturk., Berlin; type-locality: Borneo. 1924 Aciagrion borneense - L A I D L A W , Proc. U.S. natn. Mus., 66 7 (Malaysia, Siam). 1927 Aciagrion borneense - Ris, Zool. Meded., Leiden, 10 (1): 22 (Sumatra). 1935 Aciagrion borneense - LIEFTINCK, Misc. Zool. Sum., 92-93: 1 2 (Sumatra). 1954 Aciagrion borneense - LIEFTINCK, Treubia, 22 Suppl.: 76 (Siam, Malaya, Sumatra, Borneo).

Distribution: Borneo, Sumatra, Malaysia, Thailand. brosetti 1982

LEGRAND

Aciagrion brosetti L E G R A N D , Revue fr. Ent. (N.S.) 4 Mus. Paris; type-locality: Gabon.

(4): 163.

Type male: Nat. Hist. natn.

Distribution: Gabon. congoense

(SJÖSTEDT)

1917 Mombagrion congoense SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 15. Type male: Riksmus. Stockholm; type-locality: Congo. 1936 Mombagrion congoense - LONGFIELD, Trans. R. ent. Soc. London, 8 5 : 472 (Uganda). 1972 Aciagrion congoense P I N H E Y , Occ. Pap. natn. Mus. Sth. Rhod., ( B ) 5 (1): 25. 1 9 8 4 Mombagrion congoense - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 4 .

Distribution: Congo Republic, Uganda. fasciculare

LIEFTINCK

1934 Aciagrion fasciculare LIEFTINCK, Stylops, 3: 15. Type male: Buitenzorg Mus.; type-locality: Java. 1934 Aciagrion fasciculare - LIEFTINCK, Treubia, 1 4 (4): 403 (Java). 1954 Aciagrion fasciculare - LIEFTINCK, Treubia, 2 2 Suppl: 7 6 (Java).

Distribution: Java.

240 feuerborni

SCHMIDT

1934 Aciagrion feuerborni SCHMIDT, Arch. Hydrob. Suppl., 13: 344. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1935 Aciagrion feuerborni - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 1 2 . 1954 Aciagrion feuerborni

- LIEFTINCK, Treubia, 22 Suppl.: 76 (Java, Sumatra).

Distribution: Java, Sumatra. fragile

(TILLYARD)

1906 Ischnura fragilis TILLYARD, Proc. Linn. Soc. N.S. Wales, 31 (2): 186. Type male: Tillyard's Collection; type-locality: N. Queensland. 1913 Aciagrion fragilis - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 472. 1917 Aciagrion fragilis - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 37. 1924 Ischnura fragilis - LAIDLAW, Proc. U.S. natn. Mus., 66: 5. 1937 Aciagrion fragilis - LIEFTINCK, N o v a Guinea (N.S.), 1: 330 (N. Australia, New Guinea). 1953 Aciagrion fragilis - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 166 (Sumba, New Guinea, N o r t h Australia).

Distribution: North Australia, New Guinea, Sumba. gracile gracile

(SJÖSTEDT)

1909 Mombagrion gracile SJÖSTEDT, Kiliman.-Meru Exp., 2 (14): 44 (as generotype of Mombarion SJÖSTEDT, 1909). Type male: Riksmus. Stockholm; type-locality: U s a m b a r a . 1962 Aciagrion gracile - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 132 (Tanganyika, Zanzibar, Kenya). 1972 Aciagrion (Mombagrion) gracile gracile - PINHEY, OCC. Pap. natn. Mus. Sth. Rhod., ( B ) , 5 (1): 33 (redescription).

Distribution: Central Africa. gracile attenuatum

FRASER

1928 Aciagrion attenuatum FRASER, Trans, ent. Soc. London, 76: 126. Type male: British Museum (Nat. Hist.), London, type-locality: Nyasaland. 1962 Aciagrion attenuatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 132 (type in Fraser's Collection) (Rhodesia, Nyasaland). 1966 Aciagrion attenuatum - KIMMINS, Bull. Brit. Mus., nat. Hist. (Ent.), 18: 180. 1972 Aciagrion (Mombagrion) gracile attenuatum - PINHEY, OCC. Pap. natn. Mus. Sth. Rhod., ( B ) 5 (1): 33.

Distribution: Nyasaland, Rhodesia, hamoni

FRASER 1955 Aciagrion hamoni FRASER, Revue fr. Ent., 22 (3): 240. Type male: Nat. Hist. natn. Mus. Paris; type-locality: French Congo. 1962 Aciagrion hamoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 132. 1 9 7 2 Aciagrion hamoni - PINHEY, OCC. Pap. natn. Mus. South. Rhod., ( B ) 5 ( 1 ) : 4 3 .

Distribution: French Congo. hisopa

(SELYS)

1876 Agrion (Pseudagrion) hisopa SELYS, Synopsis des Agrionines, Add. 5:219. Type male: Bouvier's Collection; type-locality: Malaysia. 1890 Pseudagrion hisopa - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1891 Aciagrion hisopa -SELYS, Ann. Mus. civ. Stor. nat. Genova, 30:159 (as generotype of Aciagrion SELYS, 1 8 9 1 ) .

1916 Aciagrion hisopa — Ris, Suppl. ent. Berlin, 5: 39 (Formosa, Japan).

241 1924 1931 1931 1942 1948 1954 1966

Aciagrion Aciagrion Aciagrion Aciagrion Aciagrion Aciagrion Aciagrion

hisopa hisopa hisopa hisopa hisopa hisopa krishna

- LAIDLAW, Proc. U.S. natn. Mus., 66: 5 (Burma, Malaysia). - SCHMIDT, Konowia, 10 (3): 185 (China). - FRÄSER, Ree. Indian Mus., 33: 464. - FRASER, Proc. R. ent. Soc. Lond., (B) 11: 97. - LIEFTINCK, Ark. Zool. Uppsala, 41A (10): 16 (India, China, Japan). hisopa - LIEFTINCK, Treubia, 22 Suppl.: 76 (Malaya). - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 199.

Distribution: from Japan to Malaysia. heterostictum heterostictum

FRASER

1955 Aciagrion heterosticta FRASER, Revue Zool. Bot. Afr., 52: 19. Type male: Tervuren Mus.; type-locality: Belgian Congo. 1962 Aciagrion heterostictum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 132. 1 9 7 2 Aciagrion heterostictum heterostictum - PINHEY, OCC. Pap. natn. Mus. South. Rhod., ( B ) 5 (1): 4 1 .

Distribution: Zaire. heterostictum karamoja 1972

PINHEY

Aciagrion heterostictum karamoja PINHEY, OCC. Pap. natn. Mus. South. Rhod. ( B ) 5 ( 1 ) : 4 2 . Type male: unknown; type-locality: Uganda.

Distribution: Uganda. macrootithenae

PINHEY

1972 Aciagrion macrootithenae PINHEY, OCC. Pap. natn. Mus. South. Rhod., (B) 5 (1): 37. Type male: unknown; type-locality: Zambia.

Distribution: Zambia. migratum

(SELYS)

1876 Agrion (Pseudagrion) migratum SELYS, Synopsis des Agrionines, Add. 5: 217. Type male: Selys's Collection; type-locality: Japan. 1890 Pseudagrion migratum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1 9 0 5 Pseudagrion migratum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 0 . 1 9 8 4 Aciagrion migratum - DAVIES & TOBIES, The Dragonflies of the World, Utrecht, 1: 6 4 .

Distribution: Japan. nodosum

(PINHEY)

1964 Ischnuragrion nodosum PINHEY, Publ. cult. Comp. Diam. Angola, 63:105. Type male: National Mus., Bulawayo; type-locality: Angola. 1972 Aciagrion nodosum - PINHEY, Occ. Pap. natn. Mus. South. Rhod. (B) 5 (1): 46.

Distribution: Angola. occidentale

LAIDLAW

1919 Aciagrion hisopa occidentale LAIDLAW, Ree. Indian Mus., 16: 187. Type male: Zool. Surv. India, Calcutta; type-locality: Ceylon. 1922 Aciagrion paludensis FRASER, J. Bombay nat. Hist. Soc. 28: 698. Type male: British Museum (Nat. Hist.), London: type-locality: Nilgiris. 1924 Aciagrion occidentalis - FRASER, Ree. Indian Mus., 26: 491 (West India). 1955 Aciagrion occidentale - LIEFTINCK, Zool. Meded., Leiden, 45 (5): 76 (Ceylon, India).

242 1966 Aciagrion paludensis- KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 208 (proposed synonymy w i t h occidentale

LAIDLAW, 1919).

Distribution: India, Sri Lanka. olympicum

LAIDLAW

1919 Aciagrion olympicum LAIDLAW, Ree. Indian Mus., 16: 184. Type male: Zool. Surv. India, Calcutta; type-locality: India. 1924 Aciagrion olympicum - LAIDLAW, Proc. U.S. natn. Mus., 66: 8.

Distribution: Himalayas. pallidum

SELYS

1892 Agrion (Aciagrion) pallidum SELYS, Ann. Mus. civ. Stor. nat. Genova, (2) 10: 512. Type female: Mus. civ. Stor. nat. Genova; type-locality: Burma. 1919 Aciagrion pallidum - LAIDLAW, Ree. Indian Mus., 16: 186 (India). 1924 Aciagrion pallidum - FRÄSER, Ree. Indian Mus., 26: 491.

Distribution: Burma, India. rarum

(LONGFIELD)

1945 Ischnuragrion rarum LONGFIELD, Arq. Mus. Bocage, 16: 6 (as generotype of Ischnuragrion LONGFIELD, 1945). Type male: British Museum (Nat.Hist.), London; type-locality: Angola. 1962 Ischnuragrion rarum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 138. 1972 Aciagrion rarum - PINHEY, OCC. Pap. natn. Mus. Sth. Rhod., (B) 5 (1): 49. 1984 Aciagrion rarum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 65 (S.W. Africa).

Distribution: Angola. steeleae steeleae

KIMMINS

1955 Aciagrion steeleae KIMMINS, Entomologist, 88: 109. Type male: British Museum (Nat.Hist.), London; type-locality: North Rhodesia. 1962 Aciagrion steeleae steeleae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 133. 1972 Aciagrion steeleae steeleae - PINHEY, Occ. Pap. natn. South. Rhod., (B) 5 (1): 29.

Distribution: Rhodesia, steeleae abercornensis

PINHEY 1958 Aciagrion steeleae abercornensis PINHEY, OCC. Pap. nat. Mus. S. Rhodesia, 22B: 103. Type male: British Museum (Nat.Hist.), London; type-locality: Rhodesia. 1962 Aciagrion steeleae abercornensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 133. 1984 Aciagrion steeleae abercornensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 64 (proposed synomymy with steeleae KIMMINS, 1955).

Distribution: Rhodesia, tillyardi

LAIDLAW 1919 Aciagrion tillyardi LAIDLAW, Ree. Indian Mus., 16:187. Type male: Zool. Surv. India, Calcutta; type-locality: Assam. 1919 Enallagma assamica FRÄSER, J. Bombay nat. Hist. Soc. 26: 877. Type series originally in British Museum (Nat.Hist.), London, now unknown; type-locality: Assam. 1935 Aciagrion tillyardi - LIEFTINCK, Misc. Zool. Sum., 92-93: 12 (Sumatra). 1954 Aciagrion tillyardi - LIEFTINCK, Treubia, 22 Suppl.: 76 (Sumatra). 1966 Enallagma assamica - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18:180 (proposed synonymy w i t h tillyardi

LAIDLAW, 1919).

Distribution: A s s a m , S u m a t r a .

243 tonsillare

LIEFTINCK -

>

1937 Aciagrion tonsillare LIEFTINCK, Nova Guinea (N.S.) 1: 331. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. zambiense

PINHEY

1972 Aciagrion zambiense PINHEY, OCC. Pap. natn. Mus. South. Rhod. (B) 5 (1): 23. Type male: unknown; type-locality: Zambia.

Distribution: Zambia. Genus Amorphostigma

FRASER

1924 Amorphostigma FRASER, Trans, ent. Soc. London, 1924: 432. Type-species: armstrongi FRASER, 1924 (original designation).

armstrongi

Amorphostigma

FRASER

1924 Amorphostigma armstrongi FRASER, Trans, ent. Soc. London, 1924: 433 (as generotype of Amorphostigma FRASER, 1924). Type male: British Museum (Nat.Hist.) London; type-locality: Samoa.

Distribution: Samoa. auricolor

FRASER

1927 Amorphostigma auricolor FRASER, Insects of Samoa, (7) 1: 30. Type male: British Museum (Nat.Hist.), London; type-locality: West Samoa.

Distribution: West Samoa. Genus Amphiagrion

SELYS

1876 Agrion (Amphiagrion) SELYS, Synopsis des Agrionines, Add. 5:40. Type-species: Agrion discolor BURMEISTER, 1 8 3 9 (designated by K I R B Y , Syn. Cat. Neur.-Odon, London: 1 4 3 , 1 8 9 0 ) . 1890 Amphiagrion - KIRBY, Syn. Cat. Neur.-Odon, London: 143. - 1910 Amphiagrion M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 ( 1 ) : 6 6 . [type-species: Agrion saucium (BURMEISTER, 1 8 3 9 ) ] .

amphion

SELYS

1876 Agrion (Amphiagrion) amphion SELYS, Synopsis des Agrionines, Add. 5:43. Type female: Selys's Collection; type-locality: North America. 1890 Amphiagrion amphion - KIRBY, Syn. Cat. Neur.-Odon, London: 143.

Distribution: North America. saucium

(BURMEISTER)

1839 Agrion saucium BURMEISTER, Handb. Ent., 2: 819. Type male: Mus. Comp. Zool. Cambridge; type-locality: South Caroline. 1839 Agrion discolor BURMEISTER, Handb. Ent., 2: 819. Type female: Halle Museum; type-locality: South Caroline. 1876 Agrion (Amphiagrion) saucium - SELYS, Synopsis des Agrionines, Add. 5: 41 [proposed synonymy with discolor (BURMEISTER, 1 8 3 9 ) (as generotype of Amphiagrion SELYS, 1 8 7 6 ) ] . 1876 Agrion (Pyrrhosoma) abbreviatum SELYS, Synopsis des Agrionines, Add. 5: 189. Type male: Selys's Collection; type-locality: California. 1890 Amphiagrion saucium - KIRBY, Syn. Cat. Neur.-Odon., London: 143 [proposed synonymy with discolor (BURMEISTER, 1 8 3 9 ) ] .

244 1890 Agrion (Pyrrhosoma) abbreviatum - KIRBY, Syn. Cat. Neur.-Odon., London: 143 [proposed synonymy with discolor (BURMEISTER, 1 8 3 9 ) ] . 1898 Agrion discolor - CALVERT, Trans. Amer. ent. Soc., 25: 38. 1910 Amphiagrion saucium - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 66. 1910 Agrion discolor - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 66 [proposed synonymy with saucium (BURMEISTER, 1 8 3 9 ) ] . 1910 Agrion (Pyrrhosoma) abbreviatum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 66 [proposed synonymy with saucium (BURMEISTER, 1839)]. 1919 Agrion (Erythagrion) discolor - CALVERT, Trans. Am. ent. Soc., 45: 345. 1919 Agrion saucium - CALVERT, Trans. Am. ent. Soc., 45: 345 [proposed synonymy with discolor (BURMEISTER, 1 8 3 9 ) ] .

1984 Pyrrhosoma abbreviatum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 65 (as good species). Distribution: U . S . A .

Genus Amphiallagma

KENNEDY

1920 Amphiallagma KENNEDY, Ohio J. Sei. Columbus, 21: 88. Type-species: Agrion parvum SELYS, 1876 (original designation).

parvum

(Enallagma)

(SELYS)

1876 Agrion (Enallagma) parvum SELYS, Synopsis des Agrionines, Add. 5: 120. Type male: Selys's Collection; type-locality: India. 1890 Enallagma parvum - KIRBY, Syn. Cat. Neur.-Odon, London: 147. 1 9 0 5 Enallagma parvum - JACOBSON & BIANKI, Pryam. lozhnos, Ross, imp.: 8 1 7 (India). 1913 Ischnura immsi LAIDLAW, Entomologist, 46: 236. Type male: British Museum (Nat.Hist.), London; type-locality: India. 1919 Enallagma parvum - LAIDLAW, Ree. Indian Mus., 16: 184. 1919 Ischnura immsi - LAIDLAW, Ree. Indian Mus., 16: 184 [proposed synonymy with parvum (SELYS, 1 8 7 6 ) ] .

1920 Amphiallagma parvum - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Amphiallagma KENNEDY, 1920). 1924 Enallagma parvum - FRÄSER, Ree. Indian Mus., 26: 491 (West India). 1955 Enallagma parvum - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 76 (Ceylon). 1984 Amphiallagma parvum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 65.

Distribution: India, Sri Lanka.

Genus Andinagrion

BULLA

1973 Andinagrion BULLA, Physis B. Aires (Sect. C) 32 (85): 512. Type-species: Oxyagrion peterseni Ris, 1908 (original designation).

peterseni (Ris) 1908 Oxyagrion peterseni Ris, Deutsch, ent. Z., 1908: 519. Type male: Mus. Naturk., Berlin; typelocality: Argentina. 1973 Andinagrion peterseni - BULLA, Physis B. Aires (Sect. C) 32 (85): 512 (as generotype of Andinagrion BULLA, 1973).

Distribution: Argentina. salicoti (Ris) 1904 Oxyagrion salicoti Ris, Hamb. Magalhaen. Reise, 7: 9. Type male: Mus. Naturk., Berlin; type-locality: Buenos Aires.

245 1914 Tigriagrion salicoti - FÖRSTER, Arch. Naturg., 80 (A2): 70 (Brazil, Argentina). 1973 Andinagrion salicoti - BULLA, Physis B. Aires (Sect. C) 32 (85): 515. 1984 Andinagrion salicoti - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 65.

Distribution: Brazil, Argentina. Genus Anomalagrion

SELYS

1857 Agrion (Anomalagrion) SELYS, Sagra, Hist. Cuba Ins.: 469. Type-species: Agrion hastatum SAY, 1839 (monotypy). - 1876 Agrion (Anomalagrion) - SELYS, Bull. Acad. r. Belg. (2) 41 (2-3): 10. - 1919 Agrion (Anomalagrion) - CALVERT, Trans. Am Ent. Soc. 45: 356. 1984 Anomalagrion - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 65.

hastatum (SAY) 1839 Agrion hastata SAY, J1 Acad. nat. Sei. Philad., 8: 38. Type male: Boston Mus.; type-locality: Cuba. 1842 Agrion anomalum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 281. Type male: Serville's Colletion; type-locality: Cuba. 1844 Agrion venerinotata HALDEMANN, Proc. Acad. nat. Sei. Philad., 2: 55. Type: unknown; typelocality: unknown. 1857 Agrion (Anomalagrion) hastatum - SELYS, Sagra, Hist. Cuba Ins.: 469 (as generotype of Anomalarion SELYS, 1857). 1861 Agrion hastatum - HAGEN, Syn. N. Amer. Neur., Washington: 77 (Indiana, Maine, Maryland, New Jersey, Savannah, Georgia, Lousiana, Florida, Pennsylvania, Cuba, Venezuela). 1876 Agrion (Anomalagrion) hastatum - SELYS, Synopsis des Agrionines, Add. 5:11 (redescription). 1890 Anomalagrion hastatum - KIRBY, Syn. Cat. Neur.-Odon., London: 140. 1890 Agrion anomalum - KIRBY, Syn. Cat. Neur.-Odon., London [proposed synonymy with hastatum

(SAY, 1839)].

1890 Agrion venerinotatum - KIRBY, Syn. Cat. Neur.-Odon., London: 140 [proposed synonymy with hastatum (SAY, 1 8 3 9 ) ] . 1919 Agrion (Anomalagrion) hastatum - CALVERT, Trans. Am. ent. Soc., 45: 356 (Cuba). 1984 Anomalagrion hastatum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 65.

Distribution: U.S.A., and Cuba, Venezuela. Genus Archaeallagma

KENNEDY

1920 Archaeallagma KENNEDY, Ohio J. Sei. Columbus, 21: 87. Type-species: Enallagma CALVERT, 1909 (original designation).

ovigerum

ovigerum

(CALVERT)

1909 Enallagma ovigerum CALVERT, Ann. Carnegie Mus. Univ. Mich., 6: 159. Type male: Ris's Collection; type-locality: Bogota. 1916 Enallagma ovigerum - Ris, Arch. Naturg., (A) 82: 117 (Colombia) (redescription). 1920 Archaeallagma ovigerum - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Archaeallagma

KENNEDY, 1 9 2 0 ) .

1989 Enallagma ovigerum - DE MARMELS, Odonatologica, 18 (3): 250.

Distribution: Colombia. Genus Argentagrion

FRÄSER

1948 Argentagrion FRASER, Acta Zool. lilloana, 5: 48. Type-species: Acanthagrion ambiguum Ris, 1904 (original designation). - 1920 Homeoura KENNEDY, Ohio J. Sei. Columbus, 21: 88. Typespecies: Agrion (Agrion) nepos SELYS, 1876 (original designation). - 1948 Homeoura - FRASER, Acta Zool. lilloana, 5: 48 (proposed synonymy with Argentagrion FRASER, 1948. - 1971 Ar-

246 gentagrion - B U L L A , Revta soc. ent. argent., 3 3 : 4 9 (revision) & TOBIN, The Dragonflies of the World, Utrecht, 1: 65..

1984

Argentagrion

-

DAVIES

ambiguum (Ris) 1904 Acanthagrion ambiguum Ris, Hamb. Magalhaen. Reise. 7:13. Type male: Mus. Naturk. Berlin; type-locality: Argentina. 1916 Acanthagrion ambiguum - Ris, Arch. Naturg., (A) 82: 120. 1948 Argentagrion ambiguum - FRASER, Acta Zool. lilloana, 5: 48 (as generotype of Argentagrion FRASER, 1 9 4 8 ) .

1971 Acanthagrion ambiguum - BULLA, Revta soc. ent. argent., 33: 49.

Distribution: Argentina. cheliferum (SELYS) 1876 Agrion (Acanthagrion) ? cheliferum SELYS, Synopsis des Agrionines, Add. 5: 75. Type male: Selys's Collection; type-locality: Brazil. 1890 Acanthagrion cheliferum - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1956 Enallagma cheliferum - SANTOS, An. Acad. bras. Cienc., 28: 571. 1958 Argentagrion cheliferum - RACENIS, Acta biol. venezuel., 2: 192.

Distribution: Brazil. lindneri (Ris) 1928 Acanthagrion lindneri Ris, Konowia, 7: 41. Type male: Mus. Naturk. Berlin; type-locality: Argentina. 1958 Argentagrion lindneri - RACENIS, Acta biol. venezuel., 2: 193.

Distribution: Argentina. nepos(SELYS) 1876 Agrion (Agrion) nepos (in BATES M S ) SELYS, Synopsis des Agrionines, Add. 5: 138. Type male: Selys's Collection; type-locality: Amazonas. 1890 Coenagrion nepos - KIRBY, Syn. Cat. Neur.-Odon., London: 148. 1920 Homeoura nepos - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Homeoura KENNEDY, 1 9 2 0 ) .

1948 Argentagrion nepos - FRASER, Acta Zool. lilloana, 5: 48.

Distribution: Brazil. silviae BULLA 1971 Argentagrion silviae BULLA, Revta soc. ent. argent., 33: 51. Type male: Bulla's Collection; type-locality: Paraguay.

Distribution: Paraguay. Genus Austroallagma LIEFTINCK 1953 Austroallagma LIEFTINCK, Verh. naturf. Ges. Basel, 64: 157. Type-species: Aciagrion sagittiferum LIEFTINCK, 1949 (original designation).

sagittiferum (LIEFTINCK) 1949 Aciagrion sagittiferum LIEFTINCK, Nova Guinea (N.S.) 5: 200. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Tenimber Island. 1953 Austroallagma sagittiferum - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 163 (as generotype of Austroallagma LIEFTINCK, 1953) (redescription) (Sumba, Tanimber Island).

Distribution: Sumba, Tanimbar Island.

247 Genus Calvertagrion

ST. QUENTIN

I960 Calvertagrion ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 54. Type-species: Calvertagrion cellulare ST. QUENTIN, 1960 (original designation).

dicellulare I960

di-

ST. QUENTIN

Calvertagrion dicellulare S T . Q U E N T I N , Beitr. neotrop. Fauna, Mus., Wien; type-locality: Brazil.

2 (1): 54.

Type male: Naturhist.

Distribution: Brazil. Genus Cyanallagma

KENNEDY

1920 Cyanallagma KENNEDY, Ohio J. Sei. Columbus, 21: 87. Type-species: Agrion interrupta SELYS, 1 8 7 6 (original designation).

(Acanthagrion)

acuta (Ris) 1916 Acanthagrion acutum Ris, Arch. Naturg., (A) 82: 120. Type male: not given; type-locality: Bolivia. 1984 Cyanallagma acutum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 66. 1989 Acanthagrion acutum - DE MARMELS, Odonatologica, 18 (3): 246.

Distribution: Bolivia. bonariensis (Ris) 1913 Acanthagrion interruptum bonariense Ris, Mem. Soc. ent. Belg., 22: 67. Type male: Mus. Naturk. Berlin, type-locality: Argentina. 1 9 8 4 Cyanallagma bonariensis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 6 .

Distribution: Argentina. interruptum

(SELYS)

1876 Agrion (Acanthagrion) interruptum SELYS, Synopsis des Agrionines, Add. 5: 70. Type male: McLachlan's Collection; type-locality: Chile. 1890 Acanthagrion interruptum - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1920 Cyanallagma interrupta - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Cyanallagma

KENNEDY, 1920).

1948 Acanthagrion interruptum - FRASER, Acta Zool. lilloana, 5: 51 (Argentina). 1 9 8 4 Cyanallagma interruptum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, (as interruptum SELYS, 1 8 9 6 - misprint).

1: 6 6

Distribution: Chile, Argentina. laterale

(SELYS)

1876 Agrion (Acanthagrion) laterale SELYS, Synopsis des Agrionines, Add. 5: 72. Type male: McLachlan's Collection; type-locality: New Granada. 1890 Acanthagrion laterale - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1916 Acanthagrion laterale - Ris, Arch. Naturg. (A) 82: 122 (Colombia). 1 9 8 4 Cyanallagma laterale - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 6 .

Distribution: New Granada, Colombia. nigrinuchale

(SELYS)

1876 Agrion (Acanthagrion) nigrinuchale SELYS, Synopsis des Agrionines, Add. 5: 72. Type male: Selys's Collection; type-locality: Brazil. 1890 Acanthagrion nigrinuchale - KIRBY, Syn. Cat. Neur.-Odon., London: 145.

248 1984 Cyanallagma nigrinuchale - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 66.

Distribution: Brazil. trimaculatum (SELYS) 1876 Agrion (Acanthagrion) trimaculatum SELYS, Synopsis des Agrionines, Add. 5: 69. Type male: Selys's Collection; type-locality: Brazil. 1890 Acanthagrion trimaculatum - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1 9 8 4 Cyanallagma trimaculatum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 6 6 .

Distribution: Brazil. Genus Enacantha DONNELLY & ALAYO 1984 Enacantha DONNELLY & A L A Y O , 1966 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 66. Type-species: Enacantha caribbea DONNELLY & ALAYO, 1966 (original designation).

caribbea DONNELLY & ALAYO 1984

Enacantha caribbea DONNELLY of the World, Utrecht, 1: 66.

& ALAYO, 1 9 6 6

- loc. cit.

DAVIES & TOBIN,

The Dragonflies

Distribution: Guatemala.

Genus Enallagma CHARPENTIER 1840 Enallagma CHARPENTIER, Libell. europ. Lipsiae: 21. Type-species: Agrion cyathigerum CHARPENTIER, 1840 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 145). - 1876 Agrion (Enallagma) - SELYS, Bull. Acad. r. Belg. (2) 4 1 (2-3): 79. - 1890 Enallagma- K I R B Y , Syn. Cat. Neur.-Odon., London: 145. - 1 9 0 6 Thermagrion FÖRSTER, Wiesbaden Jb Ver. Natk., 59: 336. Type-species: Thermagrion webbianum FÖRSTER, 1906 (monotypy). - 1920 Africallagma KENNEDY, Ohio J. Sei. Columbus, 2 1 : 87. Type-species: Agrion glaucum BURMEISTER, 1839 (original designation). - 1920Ischnallagma KENNEDY, Ohio J. Sei. Columbus, 21: 87. Type-species: Ischnura elongata MARTIN, 1906 (original designation). - 1 9 2 0 Proischnura KENNEDY, Ohio J. Sei. Columbus, 2 1 : 87. Type-species: Agrion (Enallagma) subfurcatum SELYS, 1876 (original designation). - 1920 Teleallagma KENNEDY, Ohio J. Sei. Columbus, 21: 87. Type-species: Telagrion daeckii CALVERT, 1903 (original designation). - 1921 Enallagma Ris, Annals S. Afr. Mus., 18: 317 (key to South African species). - 1922 Enallagma - WILLIAMSON, Ent. News, 33: 138 (key to pollotum group). - 1927 Enallagma - BYERS, Trans. Am. ent. Soc., 53: 251 (key to species).- 1928 Libyagrion FRASER, Trans, ent. Soc. London, 76: 126. Type-species: Libyagrion decoloratum FRASER, 1928 (monotypy). - 1936 Enallagma - LONGFIELD, Trans. R . ent. Soc. London, 8 5 : 4 7 4 . - 1962 Enallagma (Africallagma) - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134. - 1962 Enallagma (Ischnallagma) - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135. - 1962 Enallagma (Proischnura) - PINHEY, Publ. cult. Comp. Diam. Angola, 59:133. - 1964 Enallagma (Teleallagma) - DONNELLY, Proc. ent. Soc. London, 66: 108. - 1975 Enallagma - HARITONOV, Norge maloizv. Vidy Faun., 9: 19 (key to Palaearctic species). - 1976 Enallagma - PINHEY, Occ. Pap. natn. Mus. Rhod., (Nat.Sei.), 5 (10): 546 (key to species).

ambiguum NAVAS 1936 Enallagma ambiguum NAVÄS, Mus. Heude Notes ent. chinoise, 3 (4): 47. Type male: not given (perhaps in Mus. Heude); type-locality: China.

Distribution: China.

249 anna

WILLIAMSON

1900 Enallagma anna WILLIAMSON, Ent. News, 11: 344. Type male: Williamson's Collection; typelocality: Arizona. 1927 Enallagma culicinovum BYERS, Trans. Am. ent. Soc., 53: 249. Type male: Cornell Univ.; typelocality: Utah.

Distribution:

U.S.A.

antennatum (SAY) 1839 Agrion antennatum SAY, in Acad. nat. Sei. Philad., 8: 39. Type: unknown. 1862 Protoneura antennata - HAGEN, Syn. Neur. N.Amer., Washington: 73 (Indiana). 1895 Enallagma fischeri KELLICOTT, Jn Cincin. Soc. nat. Hist., 17: 206. Type male: Mus. Ohio State Univ., type-locality: Columbus. 1900 Enallagma antennatum - WILLIAMSON, Drag. Ind.: 274. 1910 Enallagma antennatum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 55 (Iowa to New York). 1920 Enallagma fischeri- MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 55 [proposed synonymy with antennatum (SAY, 1 8 3 9 ) ] .

Distribution: aspersum

U.S.A.

(HÄGEN)

1861 Agrion aspersum HÄGEN, Syn. Neur. N.Amer., Washington: 97. Type male: Hagen's Collection; type-locality: New York. 1876 Agrion (Enallagma) aspersum - SELYS, Synopsis des Agrionines, Add. 5: 101 (New Jersey, Illinois, New York, Chicago. 1890 Enallagma aspersum - KIRBY, Syn. Cat. Neur.-Odon., London: 146.

Distribution: basidens

U.S.A.

CALVERT

1902 Enallagma basidens CALVERT, Biol. Centr. Amer., London: 114. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Texas.

Distribution: belyshevi

U.S.A.

HARITONOV

1975 Enallagma belyshevi HARITONOV, Norge maloizv. Vidy Faun., 9: 17. Type: unknown; typelocality: Russia.

Distribution: Russia. boreale

SELYS

1875 Enallagma boreale SELYS, Entomologist's month. Mag. 11: 242. Type male: McLachlan's Collection; type-locality: Newfoundland. 1876 Agrion (Enallagma) cyathigerum race boreale - SELYS, Synopsis des Agrionines, Add. 5: 90. 1890 Enallagma boreale - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1895 Enallagma calverti MORSE, Psyche, 7: 208. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: N. America. 1919 Enallagma calverti - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 55 (Alaska, Washington, Nevada to Maine). 1 9 2 4 Enallagma calverti - WALKER, Can. Ent., 5 6 : 1 7 3 (proposed synonymy with boreale SELYS, 1875). 1984

Enallagma calverti - DAVIES & synonymy with derserti deserti

Distribution: Canada,

TOBIN,

The Dragonflies of the World, Utrecht,

(SELYS, 1 8 7 1 ) ] .

U.S.A.

1: 6 7

[incorrect

250 buchholzi

PINHEY

1971 Enallagma buchholzi Fernando Po.

PINHEY,

J. ent. Soc. sth. Afr., 34: 218. Type male: unknown; type-locality:

Distribution: Equatorial Africa, brevispina 1876 1890 1904

SELYS

Agrion (Enallagma) brevispina SELYS, Synopsis des Agrionines, Add. 5 : 9 4 . Type female: Selys's Collection; type-locality: Japan. Enallagma brevispina - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 4 6 . Enallagma brevispina - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 1 8 (Japan).

Distribution: Japan. camerunense

KARSCH

1899 Enallagma ? camerunense K A R S C H , Ent. Nachr., 25 (11): 8. Type male: lost; type-locality: Cameroon. 1962 Enallagma ? camerunense - PINHEY; Publ. cult. Comp. Diam. Angola, 59: 135.

Distribution: Cameroon. cardenium 1876 1890 1919 1922

SELYS

Agrion (Enallagma) caecum race cardenium ( H Ä G E N MS) SELYS, Synopsis des Agrionines, Add. 5: 113. Male from Cuba. Enallagma cardenium - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 4 7 . Enallagma caecum cardenium - CALVERT, Trans. Am. ent. Soc., 45: 3 5 3 . Enallagma cardenium - WILLIAMSON, Ent. News, 3 3 : 1 4 3 (Florida).

Distribution: U.S.A., Cuba, carunculatum

MORSE 1895 Enallagma carunculatum M O R S E , Psyche, 7: 208. Type male: Agassiz Museum, Cambridge, (Mass.), type-locality: North America. 1910 Enallagma carunculatum - M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 (1): 56.

Distribution: North America. circulatum

SELYS

1883 Enallagma circulatum SELYS, Ann. Soc. ent. Belg., 27: 133. Type male: R. Mus. Brussels; type-locality: Japan. 1 9 0 5 Enallagma circulatum - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 1 7 . 1984 Enallagma circulatum - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 67 (incorrect synonymy with deserti yezoense A S A H I N A , 1919).

Distribution: Japan. civile

(HÄGEN)

Agrion civile H Ä G E N , Syn. Neur. N. Amer., Washington: 8 8 . Type male: Agassiz Museum, Cambridge (Mass.); type-locality: New York. 1 8 7 6 Agrion (Enallagma) cf'vi'/e - SELYS, Synopsis des Agrionines, Add. 5 : 9 6 (New York, Maryland, Pennsylvania, Virginia, Washington, Texas, California, Mexico, Cuba). 1876 Agrion (Enallagma) civile race simile SELYS, Synopsis des Agrionines, Add. 5: 98. Male from Colombia. 1876 Agrion canadense P R O V A N C H E V , Nat. Canad., 8 : 325. Type: unknown. 1890 Enallagma civile - K I R B Y , Syn. Cat. Neur.-Odon., London: 146. 1861

1890

Enallagma

simile

- KIRBY, S y n . C a t . N e u r . - O d o n . , L o n d o n :

146.

251 1890 Agrion canadense - KIRBY, Syn. Cat. Neur.-Odon., London: 146 [proposed synonymy with civile

(HÄGEN, 1861)].

1910 Enallagma civile - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 56 (North America und West Indies). 1919 Agrion (Enallagma) civile - CALVERT, Trans. Am. ent. Soc., 45: 354 (Cuba). 1948 Enallagma civile - ZIMMERMANN, Ins. Hawaii, 2: 381 (Hawaii).

Distribution: North and Central America, Hawaii. clausuni

MORSE

1895 Enallagma clausuni MORSE, Psyche, 7: 209. Type male: Agassiz Museum, Cambridge (Mass.), type-locality: Nevada.

Distribution: U.S.A. coecum coecum

(HÄGEN)

1861 Agrion coecum HÄGEN, Syn. Neur. N . Amer., Washington: 84. Type male: Hagen's Collection; type-locality: St. Thomas. 1876 Agrion (Enallagma) caecum - SELYS, Synopsis des Agrionines, Add. 5: 111 (Mexico, Cuba, Puerto Rico). 1888 Enallagma krugi KOLBE, Arch. f. Nat., 104: 171. Type male: Mus. Naturk., Berlin; typelocality: Puerto Rico. 1890 Enallagma krugi - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1902 Enallagma coecum - CALVERT, Biol. Centr. Amer., London: 112 (Dominica). 1919 Agrion (Enallagma) coecum - CALVERT, Trans. Am. ent. Soc. 45: 350 (redescription) (Cuba).

Distribution: Central America. coecum novaehispaniae

CALVERT

1895 Enallagma coecum CALVERT, Proc. Acad. Cal., (2) 4: 485 - misidentification. 1907 Enallagma coecum novaehispaniae CALVERT, Biol. Centr. Amer., London: 381 (new name for Enallagma coecum CALVERT, 1895). Type: undesignated. 1932 Enallagma coecum novaehispaniae - GEIJSKES, Zool. Meded., Leiden, 14: 252 (redescription). 1942 Enallagma ypsilon NEEDHAM, Bol. Ent. venezuel., 1: 62. Type male: Needham's Collection; type-locality: Venezuela. 1982 Enallagma ypsilon - DE MARMELS, Notulae odonat., 1 (10): 162 (proposed synonymy with coecum novaehispaniae CALVERT, 1 9 0 7 ) .

Distribution: Venezuela, Trinidad. consisum

WILLIAMSON

1922 Enallagma consisum WILLIAMSON, Ent. News, 33: 117. Type male: Williamson's Collection; type-locality: Florida.

Distribution: U.S.A. cuneistigma 1984

PINHEY

Enallagma cuneistigma Utrecht, 1: 67.

PINHEY, 1 9 6 9

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Rhodesia. cyathigerum cyathigerum

(CHARPENTIER)

1836 Agrion hastulatum STEPHENS (nec BURMEISTER, 1839). 111. Brit. Ent., Mand., 6 : 74. Type: undesignated. 1840 Agrion cyathigerum CHARPENTIER, Libell. europ. Lipsiae: 163. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Silesia.

252 1840 Agrion charpenticri SELYS, Monogr. Libell. eur. Paris: 214. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: France. 1840 Agrion pulchrum HAGEN, Syn. Libell. Eur.: 80. Type female: unknown. 1861 Agrion annexum HÄGEN, Syn. Neur. N. Amer., Washington: 87. Type male: Mus. Naturk., Berlin; type-locality: Sitka. 1876 Agrion (Enallagma) cyathigerum - SELYS, Synopsis des Agrionines, Add. 5: 86. 1876 Agrion charpentieri - SELYS, Synopsis des Agrionines, Add. 5: 86 [proposed synonymy with cyathigerum

cyathigerum

1876 Agrion pulchrum cyathigerum

cyathigerum

1876 Agrion annexum cyathigerum

(CHARPENTIER, 1840)].

- SELYS, Synopsis des Agrionines, Add. 5: 86 [proposed synonymy with (CHARPENTIER, 1840)].

- SELYS, Synopsis des Agrionines, Add. 5: 86 [proposed synonymy with

cyathigerum

(CHARPENTIER, 1840)].

1890 Enallagma cyathigerum - KIRBY, Syn. Cat. Neur.-Odon., London: 145 (as generotype of Enallagma

CHARPENTIER, 1 8 4 0 ) .

1890 Enallagma annexum - KIRBY, Syn. Cat. Neur.-Odon., London: 146 (Irkutsk, N.America). 1 9 0 5 Enallagma cyathigerum cyathigerum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 7 . (Caucasus, Turkestan, Spain, Sardinia). 1910 Enallagma cyathigerum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 57. 1910 Agrion annexum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 57 [proposed synonymy with cyathigerum cyathigerum (CHARPENTIER, 1 8 4 0 ) ] . 1919 Enallagma cyathigerum - LAIDLAW, Ree. Indian Mus., 16: 183 (Kashmir). 1929 Enallagma cyathigerum var. rotundatum BARTENEF, Zool. Anz., 85: 63. Male from Caucasus. 1940 Enallagma cyathigerum - NIELSEN, Mem. Soc. ent. ital., 19: 240 (Sardinia). 1948 Enallagma cyathigerum - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 153 (Armenia). 1950 Enallagma cyathigerum - MORERA, Inst. Esp. Ent., Madrid: 90 (Spain). 1955 Enallagma cyathigerum - SANTOCK & all. Agra Univ. J. Res., 4 , Suppl.: 748 (Himalayas). 1956 Enallagma cyathigerum - FRÄSER, R. Ent. Soc. London, 1 (10): 32 (England). 1984 Enallagma cyathigerum f. nigriprothorax LE ROI, 1914 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 68 (from Germany). 1984 Enallagma cyathigerum - STEINMANN, F a u n a Hung., 160: 35 (Hungary).

Distribution: Palaearctic region, and North America. cyathigerum mongolicum

BENEDEK

1968 Enallagma cyathigerum mongolicum BENEDEK, Reichenbachia, 11 (17): 183. Type male: Termeszett. Μύζ., Budapest; type-locality: Mongolia.

Distribution: Mongolia. cyathigerum possompsei

HEYMER

1984 Enallagma cyathigerum possompsei HEYMER, 1968 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s

of the World, Utrecht, 1: 69.

Distribution: Pyrenees. daeckii

(CALVERT)

1903 Telagrion daeckii CALVERT, Ent. News, 14: 36. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: U.S.A. Atlantic Coast. 1915 Telagrion daeckii - WILLIAMSON, Proc. U S natn. Mus., 48: 616 (New Jersey). 1920 Teleallagma daeckii - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Teleallagma KENNEDY, 1920).

1964 Enallagma

(Teleallagma)

Distribution: U.S.A.

daeckii - DONNELLY, Proc. ent. Soc. Wash., 66: 108.

253 davisi

WESTFALL

1943 Enallagma davisi WESTFALL, Ent. News, 54:103. Type male: unknown; type-locality: Florida.

Distribution: U.S.A. deserti deserti

(SELYS)

1871 Agrion deserti SELYS, Ann. Soc. Ent. Belg., 14:19. Type female: Selys's Collection; type-locality: Algeria. 1876 Agrion (Enallagma) deserti - SELYS, Synopsis des Agrionines, Add. 5: 93. 1890 Enallagma deserti - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1928 Enallagma deserti - Ris, Ent. Mitt., 17: 277 (description of male). 1949 Enallagma deserti deserti - ASAHINA, Insecta matsum., 17: 34.

Distribution: Algeria, Middle East, Southern Russia to Japan. deserti continental

BELYSHEV

1984 Enallagma deserti continentale BELYSHEV, 1956 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 67.

Distribution: East Asia. deserti yezoense

ASAHINA

1949 Enallagma deserti yezoense ASAHINA, Insecta matsum., 17:34. Type male: Asahina's Collection; type-locality: Japan. 1958 Enallagma deserti yezoense - ASAHINA, Insecta matsum., 22: 64 (Kurile Island).

Distribution: Japan. divagans

SELYS

1876 Agrion (Enallagma) divagans SELYS (nec KELLICOTT. 1895), Synopsis des Agrionines, Add. 5: 104. Type male: Selys's Collection; type-locality: Massechusetts. 1890 Enallagma divagans - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1922 Enallagma divagans - WILLIAMSON, Ent. News, 33: 142 (S. Carolina).

Distribution: U.S.A. doubledayi

(SELYS & HÄGEN)

1850 Agrion doubledayi SELYS & HÄGEN, Revue des Odonates Libell. europ. Paris: 209. Type male: Selys's Collection; type-locality: Florida. 1861 Agrion doubledayi - HÄGEN, Syn. Neur. N. Amer. Washington: 89 (redescription). 1876 Agrion (Enallagma) doubledayi - SELYS, Synopsis des Agrionines, Add. 5: 85. 1890 Enallagma doubledayi - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1919 Agrion (?) doubledayi - CALVERT, Trans. Am. ent. Soc., 45: 355 (Cuba). 1922 Enallagma doubledayi - WILLIAMSON, Ent. News, 33: 142 (Florida, South Carolina).

Distribution: Southern U.S.A., Cuba. dubium

ROOT

1924 Enallagma dubium ROOT, Ent. News, 35: 321. Type: unknown; type-locality: Georgia.

Distribution: U.S.A. durum

(HÄGEN)

1861 Agrion durum HÄGEN, Syn. Neur. N. Amer., Washington: 87. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Maryland.

254 1876 Agrion (Enallagma) durum - SELYS, Synopsis des Agrionines, A d d . 5: 85 (Lousiana, Florida, Maryland, Baltimore). 1890 Enallagma durum - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1922 Enallagma durum - WILLIAMSON, Ent. News, 33: 142 (Florida).

Distribution: U.S.A. ebrium

(HÄGEN)

1861 Agrion ebrium HÄGEN, Syn. Neur. N . Amer., Washington: 89. Type male: Hagen's Collection; type-locality: Chicago. 1876 Agrion (Enallagma) ebrium - SELYS, Synopsis des Agrionines, Add. 5: 96 (Illinois). 1890 Enallagma ebrium - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1 9 8 2 Enallagma

ebrium

- MASSEAN & P I L O N , O d o n a t o l o g i c a , 1 1 ( 3 ) : 1 9 2 .

Distribution: U.S.A. eiseni

CALVERT

1895 Enallagma eiseni CALVERT, Cal. Acad. Proc. (2) 4: 486. Type male: Calif. Acad., type-locality: California.

Distribution: U.S.A. elongatum

(MARTIN)

1906 Ischnura elongata MARTIN, Bull. Mus. Paris, 1906: 513. Type female: Naturhist. Riksmus., Stockholm; type-locality: East Africa. 1920 Ischnurallagma elongata - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Ischnurallagma KENNEDY, 1920). 1921 Enallagma elongatum - Ris, Annls S. Afr. Mus., 18 324 (redescription) (Kilimandjaro). 1921 Enallagma fractum Ris, Annls S. Afr. Mus., 18: 329. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Katanga. 1936 Enallagma elongatum - LONGFIELD, Trans. R. ent. Soc. London, 85: 474 (Nairobi). 1936 Enallagma pseudoelongatum LONGFIELD, Trans. R. ent. Soc. London, 85: 475. Type male: British Museum (Nat.Hist.), London; type-locality: Uganda. 1947 Enallagma elongatum - FRASER, Proc. R. ent. Soc. London, (B) 16: 146. 1947 Enallagma fractum - FRASER, Proc. R. ent. Soc. London, (B) 16: 146 [proposed synonymy w i t h elongatum

(MARTIN, 1906)].

1962 Enallagma (Ischnurallagma) elongatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135 (Angola). 1962 Enallagma (Africallagma) pseudoelongatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134 (Tanganyika, Kenya, Uganda).

Distribution: Equatorial Africa. exulans

(HÄGEN)

1861 Agrion exulans HÄGEN, Syn. Neur. N. Amer., Washington: 82. Type male: Agassiz Museum, Cambridge (Mass.), type-locality: Philadelphia. 1876 Agrion (Enallagma) exulans - SELYS, Synopsis des Agrionines, Add. 5: 96 (Philadelphia, Washington, Virginia, Maryland, Texas, Illinois). 1890 Enallagma exulans - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1922 Enallagma exulans - WILLIAMSON, Ent. News, 33: 142 (Florida, South Carolina).

Distribution: U.S.A. geminatum

KELLICOTT

1895 Enallagma geminatum KELLICOTT, Ent. News, 6: 239. Type male: Ohio Univ.; type-locality: Carolina.

255 1895 Enallagma divagans KELLICOTT (nec SELYS, 1876), Jn. Cincin. Soc., 17: 205. Type female: unknown; type-locality: Ohio. 1902 Enallagma piscinarium WILLIAMSON, Rep. Dep. Geol. Indiana, 24: 273. Type male: Williamson's Collection; type-locality: Indiana. 1922 Enallagma geminatum - WILLIAMSON, Ent. News. 33: 142 (South Carolina).

Distribution: U.S.A. glaucum

(BURMEISTER)

1839 Agrion glaucum BURMEISTER, Handb. Ent., 2: 821. Type male: Halle Museum; type locality: Cape of Good Hope. 1876 Agrion (Enallagma) gabonense SELYS, Synopsis des Agrionines, Add. 5; 116. Type male: R. Mus., Brussels; type-locality: Gabon. 1876 Agrion (Enallagma) obliteratum SELYS, Synopsis des Agrionines, Add. 5: 116. Type male: R. Mus. Brussels; type-locality: Cape of Good Hope. 1890 Disparoneura glauca - KIRBY, Syn. Cat. Neur.-Odon, London: 133. 1890 Enallagma gabonense - KIRBY, Syn. Cat. Neur.-Odon, London: 147. 1890 Enallagma obliteratum - KIRBY, Syn. Cat. Neur.-Odon, London: 147. 1906 Pseudagrion sikorae FÖRSTER, Mannheim Jb Ver.-Natk. 71-72: 59. Type male: Ann Arbor Musuem; type-locality: Reunion. 1908 Enallagma schultzei Ris, Jena. Denkschr. med. Ges., 13: 314. Type: unknown; type-locality: Kalahari. 1920 Africallagma glaucum - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Africallagma

KENNEDY, 1 9 2 0 ) :

1921 Enallagma glaucum - Ris, Ann. S. Afr. Mus., 18: 326 (South Africa). 1962 Enallagma glaucum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134 (continental African Region, and Reunion). 1962 Agrion (Enallagma) gabonense — PINHEY, Publ. cult. Comp. Diam. Angola, 59:134 [proposed synonymy with glaucum (BURMEISTER, 1839)]. 1962 Agrion (Enallagma) obliteratum - PINHEY, Publ. cult. Comp. Angola, 59: 134 [proposed synonymy with glaucum (BURMEISTER, 1839)]. 1962 Pseudagrion sikorae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134 [proposed synonymy w i t h glaucum

(BURMEISTER, 1839)].

1962 Enallagma schutzei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134 [proposed synonymy w i t h glaucum

(BURMEISTER, 1839)].

1984 Enallagma glaucum f. fugax PINHEY, 1962 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: Male from Rhodesia.

Distribution: continental Africa, and Reunion. granti

(MCLACHLAN)

1903 Ischnura ? granti MCLACHLAN, Nat. Hist. Sokotra, 24: 402. Type female: British Museum (Nat.Hist).), London; type-locality: Sokotra. 1962 Enallagma (nearer than Ischnura) granti - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135.

Distribution: Sokotra. hageni

(WALSH)

1863 Agrion hageni WALSH, Proc. ent. Soc. Philad., 2: 234. Type male: Hagen's Collection; typelocality: Illinois. 1876 Agrion (Enallagma) hageni - SELYS, Synopsis des Agrionines, Add. 5: 95. 1890 Enallagma hageni - KIRBY, Syn. Cat. Neur.-Odon, London: 146. 1982 Enallagma hageni - MASSEAU & PILON, Odonatologica, 11 (3): 192.

Distribution: U.S.A.

256 insula

FRÄSER

1920 Enallagma insula FRÄSER, Ree. Indian. Mus., 19: 32. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: South India.

Distribution: India. ifanicum

FRASER

1962 Enallagma ifanicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 136 (as undescribed species). Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Senegal.

Distribution: Senegal. kagiensis

MATSUMURA

1931 Enallagma kagiensis MATSUMURA, 6000 111. Ins. Japan: 1461. Type: unknown; type-locality: Japan.

Distribution: Japan. laterale

MORSE

1895 Enallagma laterale MORSE, Psyche, 7: 207. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Indiana.

Distribution: maldiviense

U.S.A.

LAIDLAW

1902 Enallagma ? maldiviensis LAIDLAW, Gardiner F a u n a Maldive, 1: 221. Type male: unknown; type-locality: Maldive Island. 1919 Enallagma maldiviense - LAIDLAW, Ree. Indian Mus., 16: 183.

Distribution: Maldive Island. melanotum

SELYS

1876 Agrion (Enallagma) melanotum SELYS, Synopsis des Agrionines, Add. 5: 121. Type male: Selys's Collection; type-locality: China. 1890 Enallagma melanotum - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1 9 0 5 Enallagma melanotum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 7 (China).

Distribution: China. minusculum

MORSE

1895 Enallagma minusculum MORSE, Psyche, 7: 207. Type male: Agassiz Museum, Cambridge, (Mass.); type-locality: Massachusetts.

Distribution: minutum

U.S.A.

(SELYS)

1857 Trichocnemis minuta SELYS, (nec Ris, 1930), Hist. Cuba Ins.: 464. Type male: Selys & Hagen's Collection; type-locality: Cuba. 1861 Agrion aduncum HÄGEN, Syn. Neur. N. Amer. Washington: 79. Type male: Hagen's Collection; type-locality: Cuba. 1876 Agrion (Enallagma) aduncum - SELYS, Synopsis des Agrionines, Add. 5: 84. 1890 Enallagma minutum - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1907 Argiallagma minutum - CALVERT, Biol. Centr. Amer., London: 376. 1910 Argiallagma minutum - MUTTKOWSKI, Publ. Bull. Mus. Milwaukee, 1 (1): 52. 1910 Agrion aduncum - Muttkowski, Bull. Publ. Mus. Milwaukee, 1 (1): 52 [incorrect synonymy w i t h minutum

F SELYS, 1 8 5 7 ) ] .

257 1919 Agrion (Enallagma) aduncum - CALVERT, Trans. Am. ent. Soc., 45: 354. 1919 Trichocnemis minuta - CALVERT, Trans. Am. ent. Soc., 45: 354 [incorrect synonymy with aduncum (HÄGEN), 1861)].

Distribution: Cuba. moremi 1984 1984

BALINSKY

Enallagma moremi BALINSKY, Utrecht, 1: 68. Enallagma angolicum PINHEY, Utrecht, 1: 68.

1967

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

1966

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: West Africa. nigridorsum nigridorsum

SELYS

1876 Agrion (Enallagma) nigridorsum SELYS, Synopsis des Agrionines, Add. 5: 114. Type male: R. Mus. Brussels, type-locality: Zanzibar. 1890 Enallagma nigridorsum - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1921 Enallagma nigridorsum - Ris, Annls. S. Afr. Mus., 18: 322 (South Africa). 1962 Enallagma nigridorsum nigridorsum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134.

Distribution: subtropical and tropical continental Ethiopian region, and Sokotra. nigridorsum kauderni

(SJÖSTEDT)

1917 Ischnura kauderni SJÖSTEDT, Ark. Zool., Uppsala, 11 (13): 4. Type female: Naturhist. Riksmus., Stockholm; type-locality: Madagascar. 1949 Enallagma kauderni - FRASER, Mem. Inst. sei. Madagascar, A3: 2. 1951 Enallagma nigridorsum kauderni - SCHMIDT, Mem. Inst. sei. Madagascar, A6: 207. 1962 Enallagma (Afriallagma) nigridorsum kauderni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 3 4 .

Distribution: Madagascar, nigrolineatum

HARITONOV

1975 Enallagma nigrolineatum HARITONOV, Norge maloizv. Vidy Faun., 9: 16. Type: unknown; type-locality: Kamchatka peninsula.

Distribution: Russia: Far East. pallidum

ROOT

1923 Enallagma pallidum ROOT, Ent. News, 34: 202. Type: unknown; type-locality: Maryland.

Distribution: U.S.A. pictum

MORSE

1895 Enallagma pictum MORSE, Psyche, 7: 207. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Massachusetts. 1919 Enallagma pictum - CALVERT, Trans. Am. ent. Soc., 45: 385 (redescription).

Distribution: U.S.A. pollutum

(HÄGEN) 1861 Agrion pollutum HÄGEN, Syn. Neur. N. Amer., Washington: 83. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Florida. 1876 Agrion (Enallagma) pollutum - SELYS, Synopsis des Agrionines, Add. 5: 110. 1890 Enallagma pollutum - KIRBY, Syn. Cat. Neur.-Odon., London: 147.

258 1919 Enallagma pollutum - CALVERT, Trans. Am. ent. Soc., 45: 378 (redescription). 1922 Enallagma pollutum - WILLIAMSON, Ent. News, 33: 140. Distribution: U . S . A .

polychromaticum

BARHARD

1937 Enallagma polychromaticum BARHARD, Annl. S. Afr. Mus., 32: 220. Type male: South Afr. Mus.; type-locality: Cape. 1958 Agriocnemis polychromaticum - NIELSEN, Riv. biol. colon., 16: 40. 1962 Enallagma polychromaticum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135.

Distribution: South Africa, praevarum

(HÄGEN)

1861 Agrion praevarum HAGEN, Syn. Neur. N. Amer. Washington: 88. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Mexico. 1876 Agrion (Enallagma) praevarum - SELYS, Synopsis des Agrionines, Add. 5: 99. 1890 Enallagma praevarum - KIRBY, Syn. Cat. Neur.-Odon., London: 146.

Distribution: Mexico. recurvatum

DAVIS

1913 Enallagma recurvatum DAVIS, J. New York Ent. Soc., 21: 15. Type: unknown; type-locality: New York.

Distribution: risi 1961

U.S.A.

SCHMIDT Enallagma risi S C H M I D T (nec PINHEY, 1 9 6 2 ) , Beitr. naturk. Forsch. SW-Dtschl., 19: male: Schmidt's Collection; type-locality: Afghanistan.

409.

Type

Distribution: Afghanistan, rotundipenne Ris 1921 Enallagma rotundipenne Ris, Annals S. Afr. Mus., 18: 321. Type male: Naturhist. Riksmus., Stockholm; type-locality: CafTraria. 1962 Enallagma (Proischnura) rotundipenne - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 133 (type lost). South Africa. Distribution:

robustum

SELYS

1875 Enallagma robustum SELYS, Entomologist's month. Mag. 11: 243. Type female: McLachlan's Collection; type-locality: California. 1876 Agrion (Enallagma) cyathigerum race robustum - SELYS, Synopsis des Agrionines, Add. 5: 92. 1890 Enallagma robustum - KIRBY, Syn. Cat. Neur.-Odon., London: 146. 1984 Enallagma robustum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 69 [incorrect synonymy with cyathigerum (CHARPENTIER, 1 8 4 0 ) ] . Distribution: U . S . A . r u a DONNELLY 1984 Enallagma rua DONNELLY, 1968 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the W o r l d ,

Utrecht, 1: 69. Distribution: C e n t r a l A m e r i c a .

259 rubristigma

SCHMIDT

1951 Enallagma rubristigma SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 209. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Enallagma (Proischnura) rubristigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134.

Distribution: Madagascar. sapphirina

PINHEY

1950 Enallagma sapphirina PINHEY, Ann. Transvaal Mus., 21: 267. Type male: Transvaal Museum; type-locality: Natal. 1962 Enallagma (Africallagma) sapphirinum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135 (Transvaal, Natal).

Distribution: South Africa. schmidti

STEINMANN

nom.nov.

1930 Enallagma minutum Ris (nec SELYS, 1857), Revue suisse Zool., 38: 102. Type male: Mus. Hist, nat. Chaux-de-Fonds; type-locality: Angola. 1962 Enallagma risi PINHEY (nec SCHMIDT, 1961), Publ. cult. Comp. Diam. Angola, 59: 135 (new name for Enallagma minutum Ris, 1930, a secondary homonymy of Trichocnemis minuta SELYS, 1857). Enallagma schmidti STEINMANN nom.nov. (new name for Enallagma risi PINHEY, 1962, a secondary homonymy of Enallagma risi SCHMIDT, 1961).

Distribution: Angola. semicirculare

SELYS

1876 Agrion (Enallagma) semicirculare SELYS, Synopsis des Agrionines, Add. 5: 100. Type male: Selys's Collection; type-locality: Mexico. 1890 Enallagma semicirculare - KIRBY, Syn. Cat. Neur.-Odon., London: 146.

Distribution: Mexico. signatum

(HÄGEN)

1861 Agrion signatum HAGEN, Syn. Neur. N.Amer., Washington: 84. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Georgia. 1863 Agrion dentiferum WALSH, Proc. Ent. Soc. Philad., 2: 236. Type: destroyed; type-locality: unknown. 1876 Agrion (Enallagma) signatum - SELYS, Synopsis des Agrionines, Add. 5: 108 (Georgia, Lousiana, Maryland, Illinois). 1919 Enallagma signatum - CALVERT, Trans. Am. ent. Soc. 45: 377 (redescription) (Maine, New Jersey, Pennsylvania, Indiana). 1922 Enallagma signatum - WILLIAMSON, Ent. News, 33: 140 (Florida).

Distribution: U.S.A. sinuatum Ris 1921 Enallagma sinuatum Ris, Annls S. Afr. Mus., 18: 330. Type male: Transvaal Museum; typelocality: Mashonaland. 1962 Enallagma (Ischnallagma) sinuatum - PINHEY, Publ. cult. Comp. Diam. Angola 59: 135 (type in: Mus. r. Afr. centr., Tervuren) (Belg. Congo, Natal, Rhodesia, South Tanganyika). 1962 Enallagma sinuatum f. fugax PINHEY, OCC. Pap. nat. Mus. S. Rhodesia (B) 3: 899. Male from N o r t h Rhodesia.

Distribution: Central and South Africa.

260 somalicum

LONGFIELD

1931 Enallagma somalicum LONGFIELD, Entomologist, 64: 277. Type male: British Museum (Nat.Hist.), London; type-locality: Somaliland. 1958 Enallagma somalicum - KIMMINS, Bull. Brit. Mus. (Nat.Hist.) Ent., 7: 352 (Ethiopia). 1962 Enallagma (Ischnallagma) somalicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135 (British Somaliland Protect).

Distribution: Ethiopia, Somalia. strouhali

ST. QUENTIN

1962 Enallagma strouhali ST. QUENTIN, Ann. naturh. Mus. Wien, 65: 241. Type male: Naturhist. Mus., Wien; type-locality: Manchuria.

Distribution: Manchuria.

subfurcatum

SELYS

1876 Agrion (Enallagma) subfurcatum SELYS, Synopsis des Agrionines, Add. 5: 117. Type male: Mus. Nation. Hist. Nat. Paris; type-locality: Abyssinia. 1890 Enallagma subfurcatum - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1902 Micronympha bilobata GRUNBERG, S. B. Ges. Nat., Berlin, 9:231. Type male: lost; type-locality: Nyassaland. 1 9 0 6 Ischnura schoana FÖRSTER , Wiesbaden Jb Ver. Natk., 59: 3 3 2 . Type male. Ann Arbor Museum; type-locality: Abyssinia. 1906 Ischnura urnifera MARTIN, Bull. Mus. nat. Hist. Paris, 1906: 513. Type male: Mus. Nation. Hist. Nat., Paris, type-locality: Kenya. 1917 Enallagma abessinicum SJÖSTEDT, Ark. Zool., Uppsala, 11 (14): 17. Type male: Naturhist. Riksmus., Stockholm; type-locality: Erithrea. 1920 Proischnura subfurcatum - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Proischnura

KENNEDY, 1 9 2 0 ) .

1921 Enallagma subfurcatum - Ris, Annls. S. Afr. Mus., 18: 319 (Mashonaland, Erithrea, Abyssinia). 1946 Micronympha bilobata - FRASER, Entomologist's month. Mag., 82: 201 (proposed synonymy with subfurcatum SELYS, 1876). 1962 Enallagma (Proischnura) subfurcatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134. 1962 Ischnura schoana - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134 (proposed synonymy with subfurcatum SELYS, 1876). 1962 Ischnura urnifera - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 134 (proposed synonymy with subfurcatum SELYS, 1 8 7 6 ) .

Distribution: East and South Africa. subtile Ris 1921 Enallagma subtile Ris, Annls. S. Afr. Mus., 18: 332. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Katanga. 1928 Libyagrion decoloratum FRASER, Trans, ent. Soc. London, 76:126. Type male: British Museum (Nat. Hist.) London; type-locality: Nyasaland (as generotype of Libyagrion FRASER, 1928). 1962 Enallagma (Africallagma) subtile- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135 (South Africa, Belg. Congo, Tanganyika, Kenya, S. Rhodesia, N. Nigeria). 1966 Libyarion decoloratum - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 188 (proposed synonymy with subtile Ris, 1921).

Distribution: Central and South Africa.

261 sulcatum

WILLIAMSON

1922 Enallagma sulcatum WILLIAMSON, Ent. News, 33: 114. Type male: Williamson's Collection; type-locality: Florida.

Distribution: U.S.A. traviatum traviatum

SELYS

1876 Agrion (Enallagma) traviatum SELYS, Synopsis des Agrionines, Add. 5: 102. Type male: Morrison's Collection; type-locality: Massachusetts. 1890 Enallagma traviatum - KIRBY, Syn. Cat. Neur.-Odon. London: 146.

Distribution: U.S.A. traviatum westfalli

DONNELLY

1964 Enallagma (Teleallagma) westfalli DONNELLY, Proc. ent. Soc. Wash., 66:103. Type male: Univ. Florida; type-locality: Texas. 1984 Enallagma traviatum westfalli-DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 70.

Distribution: U.S.A. truncatum 1984

GUNDLACH

Enallagma truncatum G U N D L A C H , World, Utrecht, 1: 70.

1888

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Cuba. vaginale vaginale

SJÖSTEDT

1917 Enallagma vaginale SJÖSTEDT, Ark. Zool. Uppsala, 11 (14): 20. Type male: Naturhist. Riksmus., Stockholm; type-locality: Congo. 1963 Enallagma vaginale vaginale - PINHEY, J. ent. Soc. S. Afr., 26: 149.

Distribution: Congo Republic. vaginale longfieldi

FRASER

1947 Enallagma longfieldi FRASER, Proc. R. ent. Soc. London, (B) 16: 146. Type male: Fraser Collection; type-locality: Uganda. 1962 Enallagma (Ischnallagma) longfieldae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 135 (Kenya, Uganda). 1 9 6 3 Enallagma vaginale longfieldi - PINHEY, J . ent. Soc. S. Afr., 2 6 : 1 4 9 .

Distribution: Uganda, Kenya. vansomereni

PINHEY

1956 Enallagma vansomereni PINHEY, Occ. Pap. Coryndon Mus., 4: 26. Type male: Coryndon Mus.; type-locality. Uganda. 1 9 5 1 Enallagma sudanicum FRASER (undescribed species) - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, 1984, 1: 70. 1962 Enallagma (Africallagma) vansomereni - PINHEY, Publ. cult. Comp. Diam. Angola 59: 135. 1962 Enallagma sudanicum - PINHEY, Publ. comp. Cult. Diam. Angola, 59: 136 (as undescribed species - type in: Mus. Nation. Hist, nat., Paris ?), (Sudan). 1 9 8 4 Enallagma sudanicum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 70 (as syn. with vansomereni PINHEY, 1 9 5 6 ) .

Distribution: Uganda, Sudan.

262 vernale 1943

1982

GLOYD

Enallagma vernale G L O Y D , OCC. Pap. Mus. Zool. Mich., 479: 1. Type male: Univ. Zool. Michigan; type-locality: Michigan. Enallagma vernale - MASSEAU & P I L O N , Odonatologica, 1 1 ( 3 ) : 1 9 2 .

Distribution: U.S.A., and Canada. vesperum

CALVERT

1919 Enallagma vesperum CALVERT, Trans. Am. ent. Soc., 45: 379. Type male: Acad. Nat. Sei. Philad., type-locality: South Caroline. 1919 Enallagma laurenti CALVERT, Trans. Am. ent. Soc., 45: 380. Type male: Amer. Ent. Soc.; type-locality: Florida. 1922 Enallagma vesperum - WILLIAMSON, Ent. News, 33: 141 (Florida, S. Caroline).

Distribution: U.S.A. webbianum

(FÖRSTER) 1906 Thermagrion webbianum FÖRSTER, Wiesbaden Jb Ver. Natk., 59: 336. Type female: unknown (as generotype of Thermagrion FÖRSTER, 1 9 0 6 ) ; type-locality: N . O . Afrika. 1962 Enallagma webbianum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 136 (Abyssinia).

Distribution: East Africa. weewa

BYERS

1927 Enallagma weewa BYERS, Ann. Ent. Soc. Amer., 20: 385. Type: unknown; type-locality: Florida.

Distribution: U.S.A.

Genus Helveciagrion

MACHADO

1980 Helveciagrion M A C H A D O , Lundiana, 1: 60. Type species: Helveciagrion vulcanoae 1980 (original designation).

chirihuanum

MACHADO,

(CALVERT)

1909 Acanthagrion chirihuanum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 173. Type male: Carnegie Mus., type-locality: Brazil. 1980 Helveciagrion chirihuanum - MACHADO, Lundiana, 1: 66 (redescription).

Distribution: Brazil. simulacrum

(CALVERT)

1909 Skiallagma simulacrum CALVERT, Ann. Carnegie Mus. Pittsburgh, 6:176. Type male: Carnegie Mus., type-locality: Brazil. 1980 Helveciagrion simulacrum - MACHADO, Lundiana, 1: 71 (redescription).

Distribution: Brazil. vulcanoae

MACHADO

1980 Helveciagrion vulcanoae M A C H A D O , Lundiana, 1: 66 (as generotype of Helveciagrion DO, 1980). Type male: Machado's Collection; type-locality: Brazil.

Distribution: Brazil.

MACHA-

263 Genus Hesperagrion

CALVERT

1902 Hesperagrion CALVERT, Biol. Centr. Amer., London: 103. Type species: Agrion heterodoxum SELYS, 1868 (original designation). - 1984 Hesperagrion - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 70 (as „CALVERT, 1920" - misprint).

heterodoxum

(SELYS)

1868 Agrion heterodoxum SELYS, C.R. SOC. ent. Belg. 11: 119. Type male: Sely's Collection; typelocality: Mexico. 1868 Agrion heterodoxum var. flavescens SELYS, C.R. Soc. ent. Belg., 11: 120. Male from Mexico. 1876 Agrion (Amphiagrion) heterodoxum - SELYS, Synopsis des Agrionines, Add. 5: 44. 1876 Agrion (Amphiagrion) flavenscens - SELYS, Synopsis des Agrionines, Add. 5: 45. 1890 Amphiagrion heterodoxum - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1890 Amphiagrion flavescens - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1902 Hesperagrion heterodoxum - CALVERT, Biol. Centr. Amer., London: 103 (as generotype of Hesperagrion

1910 Hesperagrion

CALVERT, 1 9 0 2 ) .

heterodoxum

- MUTTKOWSKI, Bull. Publ. Mus., Milwaukee 1 (1): 53.

Distribution: Mexico.

Genus Ischnura

CHARPENTIER

1840 Agrion (Ischnura) CHARPENTIER, Libell. Europ. Lipsiae: 20. Type species: not given, first emendation: Agrion (Ischnura) speciosum CHARPENTIER, 1840. - 1850 Agrion (Ischnura) - SELYS & HAGEN, Revue des Odonates, Paris: 182 (Type: Agrion elegans VAN DER LINDEN, 1823). 1876 Agrion (Ischnura) - SELYS, Bull. Acad. r. Belg., (2) 41: ( 2 - 3 ) : 14. - 1876 Agrion (Ceratura) SELYS, Synopsis des Agrionines, Add. 5: 7. Type species: Agrion capraeola HAGEN, 1861 (monotypy). - 1890 Micronympha KIRBY, Syn. Cat. Neur.-Odon., London: 140 (new name for Ischnura CHARPENTIER, 1840, a junior primary homonymy of Ischnura KOCH, 1837). - 1890 Ceratura - KIRBY, Syn. Cat. Neur.-Odon., London: 139. - 1894 Ischnosoma WALLENGREN, Ent. Tidschr., 15: 270. Type species: not given. - 1905 Ischnura - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 813 (key to species). - 1910 Ischnura - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee 1 (1): 67 (type-species: Agrion pumilio CHARPENTIER, 1825). 1910 Ischnosoma - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee 1 (1): 67 (proposed synomymy with Ischnura CHARPENTIER, 1840). - 1916 Ischnura (Ceratura) - Ris, Arch. Naturg. (A) 82: 124. - 1917 Celaeneura KENNEDY, Proc. U.S. natn. Mus., 52: 495. Type species: Agrion denticolle BURMEISTER, 1839 (original designation). - 1919 Agrion (Ischnura) - CALVERT, Trans. Am. ent. Soc., 45: 350. - 1919 Ischnura- LAIDLAW, Ree. Indian Mus., 16:172 (key to species). - 1920 Nanosura KENNEDY, Ohio J. Sei. Columbus, 21: 88. Type species: Agrion (Ischnura) aurora BRAUER, 1865 (original designation). - 1920 Anomalura KENNEDY, Ohio J. Sei. Columbus, 21: 88. Type-species: Agrion prognathum HÄGEN, 1861 (original designation). 1921 Ischnura -Ris, Annls. S. Afr. Mus., 18: 333. - 1932 Ischnura - LIEFTINCK, Nova Guinea, 15: (5): 110. - 1942 Bedfordia MUMFORD, Ann. Mag. nat. Hist., (11) 9: 828. Type-species: Bedfordiahalecarpenteri MUMFORD, 1942 (original designation). - 1 9 5 2 Boninagrion ASAHINA, Mushi, 23 (6): 51. Type-species: Boninagrion azoin ASAHINA, 1952 (original designation). 1962 Ischnura - LIEFTINCK, Insects of Micronesia, 5 (1): 38. - 1956 Boninagrion - LIEFTINCK, Insects of Micronesia, 5 (1): 38 (proposed synonymy with Ischnura CHARPENTIER, 1840). 1962 Nanosura - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 137 (proposed synonymy with Ischnura

CHARPENTIER, 1840). - 1984 Ischnura

- DAVIES & TOBIN, T h e D r a g o n f l i e s of

the World, Utrecht, 1: 70. - 1984 Ischnura (Ischnuridia) DONNELLY, 1965 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 70. Type-species: Agrion posita HÄGEN, 1861 (original designation). - 1988 Ischnura - MICHALSKI, Univ. West Indies Dep. Zool. Occ. Pap., 6: 43 (key to species).

264 abyssinica

MARTIN

1906 Ischnura abyssinica MARTIN, Bull. Mus. Paris, 1906: 512. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Abyssinia. 1962 Ischnura abyssinica - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 137.

Distribution: Abyssinia. acuticauda

LIEFTINCK

1959 Ischnura acuticauda LIEFTINCK, N o v a Guinea, (N.S.) 10: Type: unknown; type-locality: New Guinea.

Distribution: New Guinea. albistigma

FRASER

1927 Ischnura albistigma FRASER, Ins. Samoa, 7: 27. Type male: British Museum (Nat.Hist.), London; type-locality: Samoa. 1953 Ischnura albistigma - FRASER, Proc. R. ent. Soc. London, (B) 22: 122.

Distribution: Samoa. aralensis 1984

CHARITONOV

Ischnura aralensis Utrecht, 1: 71.

CHARITONOV, 1 9 7 9

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Asia. ariel

LIEFTINCK

1949 Ischnura ariel LIEFTINCK, N o v a Guinea (N.S.) 5: 223. Type male: Lieftinck's Collection; typelocality: New Guinea.

Distribution: New Guinea. asiatica

(BRAUER)

1865 Agrion (Ischnura) asiatica BRAUER, Verh. Zool.-Bot. Ges. Wien., 15: 509. Type male: not given, perhaps in Naturhist. Mus. Wien; type-locality: China. 1876 Agrion (Ischnura) orientalis SELYS, Synopsis des Agrionines, Add. 5: 36. Type male: Selys's Collection; type-locality: Japan. 1890 Micronympha asiatica - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1890 Micronympha orientalis - KIRBY, Syn. Cat. Neur.-Odon., London: 143 (China, Japan). 1 9 0 5 Ischnura asiatica - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 1 3 (China). 1 9 0 5 Ischnura orientalis - JACOBSON & BIANKI. Pryam. lozhnos. Ross, imp.: 8 1 3 (Japan, China). 1916 Ischnura asiatica - Ris. Suppl. ent. Berlin, 5: 19 (Japan, China). 1930 Ischnura lobata NEEDHAM, Zool. Sinica, (A) 11: 280. Type male: British Museum (Nat. Hist.), London; type-locality: China. 1931 Ischnura formosana CHUJO, Trans, nat. Hist. Soc. Formosa, 21 (112): 47. Type male: Chujo's Collection; type-locality: Formosa. 1931 Ischnura asiatica - SCHMIDT, Konowia, 10 (3): 184 (Shanghai). 1 9 3 1 Ischnura lobata - SCHMIDT, Konowia, 10 ( 3 ) : 1 8 4 [proposed synonymy with asiatica ( B R A U E R , 1865)].

1931 Ischnura formosana

- SCHMIDT, Konowia, 10 (3): 184 [proposed synonymy with

asiatica

(BRAUER), 1865)].

1933 Ischnura asiatica - SJÖSTEDT, Ark. Zool. Uppsala, 25 (A) (5): 20 (Szechwan, Gansu).

Distribution: China, Taiwan, Japan.

265 aurora aurora

(BRAUER)

1858 Agrion delicatum HAGEN, Verh. Zool.-Bot. Ges. Wien., Abh. 8: 479. Type: undesignated. 1865 Agrion (Ischnura) aurora BRAUER, Verh. Zool.-Bot. Ges. Wien., 15. 510. Type male: not given, perhaps in Naturhist. Mus., Wien; type-locality: Tahiti. 1866 Agrion (Ischnura) aurora - BRAUER, N o v a r a Exp. Zool. 1. Neuropt.: 56. 1890 Micronympha aurora - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1916 Ischnura aurora - LAIDLAW, Ree. Indian Mus., 12: 130 (India, Sri Lanka). 1917 Ischnura aurora - SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 33 (Queensland, N.W. Australia). 1919 Ischnura aurora - LAIDLAW, Ree. Indian Mus., 16: 175. 1920 Ischnura (Nanosura) aurora - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Nanosura

KENNEDY, 1 9 2 0 ) .

1924 Ischnura aurora - FRÄSER, Ree. Indian Mus., 26, 489 (W. India). 1930 Agriocnemis amelia NEEDHAM, Zool. Sinica, (A), 11: 256. Type male: Peking Museum; typelocality: Hainan. 1934 Ischnura aurora - LIEFTINCK, Treubia, 14 (4): 400 (Java). 1938 Ischnura aurora - SCHMIDT, Ent. Soc. Amer. Ann., 31: 324. 1942 Ischnura delicata - SWEZEY & WILLIAMS, Bishop Mus. Bull., 172: 3 (Guam). 1944 Agriocnemis amelia - NEEDHAM, Bull. Mus. comp. Zool. Harv., 94: 162 [proposed synonymy with delicatum ( H Ä G E N , 1 8 5 8 ) ] . 1949 Ischnura aurora aurora - LIEFTINCK, Nova Guinea, 5: 220. 1954 Ischnura aurora - LIEFTINCK, Treubia, 22 Suppl.: 74 (Java). 1955 Ischnura aurora aurora - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 76 (Ceylon). 1962 Ischnura aurora aurora - LIEFTINCK, Insecta of Micronesia, 5 (1): 39 (from India to Polynesia). 1954 Ischnura bhintalensis SAHNI'S, Indian J. Ent., 27: 207. Type male: Sahni's Collection; typelocality: N o r t h India. 1989 Ischnura bhintalensis - HÄMÄLÄINEN, Odonatologica, 18 (1): 17 [proposed synonymy with aurora aurora ( B R A U E R , 1 8 5 8 ) ] .

Distribution: from India to Polynesia and New Zealand. aurora viduata

LIEFTINCK

1949 Ischnura aurora viduata LIEFTINCK, N o v a Guinea (N.S.) 5: 222. Type: unknown; type-locality: New Guinea.

Distribution: New Guinea. barberi

CURRIE

1903 Ischnura barberi CURRIE, Proc. Wash. Ent. Soc., 5: 302. Type male: U.S. natn. Mus., Washington; type-locality: Colorado. 1 9 1 0 Ischnura utahensis M U T T K O W S I , Bull. Wis. nat. Hist. Soc., 8 : 1 7 2 . Type: unknown; type-locality: Utah.

Distribution: bizonata

U.S.A.

SELYS

1876 Agrion (Ischnura) bizonata SELYS, Synopsis des Agrionines, Add. 5: 26. Type male: Selys's Collection; type-locality: Rio. 1890 Micronympha bizonata - KIRBY, Syn. Cat. Neur.-Odon., London: 141.

Distribution: Brazil. blumi L O H M A N N 1984

Ischnura blumi Utrecht, 1: 71.

LOHMANN, 1 9 7 9

Distribution: New Guinea.

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

266 buxtoni

FRÄSER

Ischnuri buxtoni FRÄSER, Ins. Samoa, 7: 2 3 . Type male: British Museum (Nat.Hist.), London; type-locality: Samoa. 1953 Ischnura buxtoni - FRÄSER, Proc. R. ent. Soc. London, (B), 22: 124 (redescription). 1927

Distribution: Samoa. capreola

HÄGEN

1861 Agrion (Ischnura) capreolus H A G E N , Syn. Neur. N. Amer., Washington: 78. Type male: Agassiz Mus., Cambridge (Mass.); type-locality: Puerto Rico. 1876 Agrion (Ceratura) capreola - SELYS, Synopsis des Agrionines, Add. 5: 8 (redescription) (Cuba, Puerto Rico, Brazil). 1890 Ceratura capreola - K I R B Y , Syn. Cat. Neur.-Odon., London: 139. 1916 Ischnura capreola - Ris, Arch. Naturg., (A) 82: 135 (Guatemala, Honduras, Ecuador, Peru, Antillen). 1917 Ischnura capreola - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 24 (Manaos). 1919 Agrion (Ceratura) capreolus - CALVERT, Trans. Am. ent. Soc., 45: 356 (Puerto Rico, Mexico, Brazil, Guatemala, Cuba). 1934 Ischnura capreola- SCHMIDT, Beitr. F a u n a Perus, 2:240 (Puerto Rico, Brazil, Argentina, Peru).

Distribution: Central and South America. cardinalis

KIMMINS

1929 Ischnura cardinalis KIMMINS, Entomologist, 62: 224. Type male: British Museum (Nat.Hist.), London; type-locality: Sociaty Island.

Distribution: Society Island. cervula

SELYS

1876 Agrion (Ischnura) cervula SELYS, Synopsis des Agrionines, Add. 5:18. Type male: McLachlan's Collection; type-locality: California. 1 8 9 0 Micronympha cervula - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 4 0 . 1910 Ischnura cervula - M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 (1): 68 (California, Arizona, Pacific Coast). 1 9 1 5 Ischnura cervula - K E N N E D Y , Proc. U . S . natn. Mus., 49: 2 9 6 (life history). 1946 Ischnura cervula - L A RIVERS, Ent. News, 5 7 : 217 (Nevada).

Distribution: U.S.A. chromostigma 1927

FRASER

Ischnura chromostigma

FRASER,

Ins. Samoa,

7: 2 8 .

Type: unknown; type-locality: Samoa.

Distribution: Samoa. cruzi

DE M A R M E L S

1987 Ischnura (Anomalagrion) cruzi DE MARMELS, Mitt. ent. Ges. Basel, 37 (1): 1. Type male: N a turhist. Mus., Basel; type-locality: Colombia.

Distribution: Colombia. damula

CALVERT

1902 Ischnura damula CALVERT, Biol. Centr.-Amer. London, 126. Type male: Agassiz Mus. Cambridge (Mass.); type-locality: New Mexico.

Distribution: U.S.A.

267 defixa

(HAGEN)

1861 Agrion defixum HAGEN, Syn. Neur. N . Amer., Washington: 80. Type: sex not given: u n k n o w n ; in MUTTKOWSKI: Type male: Agassiz Museum, Cambridge (Mass.); type-locality: N o r t h California. 1876 Agrion (Ischnura) defixa - SELYS, Synopsis des Agrionines, Add. 5: 17 (description of male). 1890 Micronympha defixa - KIRBY, Syn. Cat. Neur.-Odon., London: 140. 1910 Agrion (Ischnura) defixa - MUTTKOWSKI, Publ. Bull. Mus. Milwaukee, 1 (1): 69 (as syn. with Ischnura perparva SELYS, 1876) - 1 (1): 70 [as syn. with Ischnura ramburii credula (HÄGEN, 1861)]. 1984 Ischnura defixa - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 71 (as syn. with perparva SELYS, 1876).

Distribution: delicata

U.S.A.

SELYS

1876 Ischnura locality: 1890 Ischnura Ischnura

delicata SELYS, Bull. Acad. r. Belg., (2) 42: 281. Type male: Selys's Collection; typeAustralia. delicata - KIRBY, Syn. Cat. Neur.-Odon., London: 143 [proposed synonymy with aurora aurora (BRAUER, 1865)].

1901 Ischnura delicata - MARTIN, Mem. Soc. Zool. France, 14: 246 (Victoria).

Distribution: Australia. demorsa

(HÄGEN)

1861 Agrion demorsum HÄGEN, Syn. Neur. N . Amer., Washington: 81. Type male: Saussure's Collection; type-locality: Mexico. 1876 Agrion (Ischnura) demorsa - SELYS, Synopsis des Agrionines, A d d . 5: 17. 1890 Micronympha demorsa - KIRBY, Syn. Cat. Neur.-Odon., London: 140. 1910 Ischnura demorsa - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 68.

Distribution: Mexico. denticollis

(BURMEISTER)

1839 Agrion denticolle BURMEISTER, H a n d b . Ent., 2: 819. Type female: Halle Museum; type-locality: Mexico. 1876 Agrion (Nehalennia) denticollis - SELYS, Synopsis des Agrionines, A d d . 5: 134, and 282. 1890 Nehalennia denticollis - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1895 Ischnura exstriata CALVERT, Proc. Cal. Acad., (2) 4:493. Type male: Calif. Acad., type-locality: California. 1898 Ischnura denticollis - CALVERT, Trans. A m . ent. Soc., 25: 38. 1898 Ischnura exstriata-CALVERT, Trans. A m . ent. Soc., 25: 38 [proposed synonymy with denticollis (BURMEISTER, 1839)]. 1910 Ischnura denticollis - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 68. 1917 Celaenura denticollis - KENNEDY, Proc. U.S. natn. Mus., 52: 500 (as generotype of Celaenura KENNEDY, 1917). 1946 Ischnura denticollis - LA RIVERS, Ent. News, 57: 217 (Nevada).

Distribution: dorothea

U.S.A.

FRASER

1924 Ischnura dorothea FRÄSER, Ree. Indian Mus., 26: 428. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: India.

Distribution: India.

268 elegans elegans (VAN

DER LINDEN)

1823 Agrion elegans VAN DER LINDEN, Opusc. Sei., 4: 104. Type male: Mus. Naturk., Berlin ?; type-locality: Italy: Bologna. 1823 Agrion pupilla HANSEMANN, Wiedem. Zool. Mag., 2 (1), 156. Type: undesignated. 1 8 2 5 Agrion tuberculatum CHARPENTIER, (nec SELYS, 1 8 5 7 ) , Hor. Ent.: 2 1 . Type female: Halle Museum; type-locality: Germany. 1836 Agrion ezonatum STEPHENS, 111. Brit. Ent. Mand., 5: 72. Type male: British Museum (Nat.Hist.), London; type-locality: England. 1836 Agrion rufescens STEPHENS, 111. Brit. Ent. Mand., 5: 72. Type: undesignated. 1838 Agrion aglae FONSCOLOMBE, Ann. Soc. ent. France, 1838: 567. Type male: Mus. Natn. Hist. Nat., Paris; type-locality: France. 1839 Agrion hastulatum BURMEISTER, Handbuch. Ent., 2: 820. Type: undesignated. 1845 Agrion rubens EVANS, Brit. Lib.: 14. Type: undesignated. 1876 Agrion (Ischnura) elegans - SELYS, Synopsis des Agrionines, Add. 5: 33 (redescription) (Europe). 1887 Ischnura elegans - SELYS, Ann. Soc. ent. Belg., 31: 65 (Algeria, Tunis, Morocco). 1890 Micronympha elegans - KIRBY, Syn. Cat. Neur.-Odon., London: 142. 1924 Ischnura elegans var. infuscans KILLINGTON, Entomologist, 57: 278 (emandation). 1924 Ischnura elegans var. obsoleta KILLINGTON, Entomologist, 57: 278 (emandation). 1928 Ischnura elegans - ANDERS, Mem. Soc. R. ent. Egypt., 3 (1): 25 (Egypt). 1939 Ischnura elegans - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 289 (Luzon ?). 1940 Ischnura elegans - NIELSEN, Mem. Soc. ent. ital., 19: 239 (Sardinia). 1950 Ischnura elegans - MORERA, Inst. Esp. Ent. Madrid, 92 (Spain). 1956 Ischnura elegans - FRÄSER, R. Ent. Soc. London, 1 (10): 31 (England). 1 9 8 4 Agrion excelsa R O S T O C K , 1 8 8 5 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 72. 1 9 8 4 Agrion exigua R O S T O C K , 1 8 8 5 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 72. 1 9 8 4 Agrion magna R O S T O C K , 1 8 8 5 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 73. 1 9 8 4 Ischnura elegans f. infuscens C A M P I O N , 1 9 0 5 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 73. 1989 Ischnura elegans elegans - ZHU, Notul. odonatol., 3 (3): 47 (China).

Distribution: Palaearctic region (except Iceland, northern Scandinavia, Corsica, Sardinia, Sicily and Malta).

elegans ebneri

SCHMIDT

1938 Ischnura elegans ebneri SCHMIDT, SitzBer. Akad. Wiss. Wien (I) 147:142. Type male: Naturhist. Mus. Wien; type-locality: Crete. 1984 Ischnura elegans eberni (misprint) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 72. 1988 Ischnura elegans ebneri- D U M O N T , DEMIRSOY & M E R T E N S , Notul. odonatol., 3 (2): 24 (Turkey).

Distribution: Crete, Turkey.

elegans marquardti

SCHMIDT

1938 Ischnura elegans marquardt SCHMIDT, SitzBer. Akad. Wiss. Wien (I) 147: 142. Type male: Naturhist. Mus., Wien; type-locality: Turkestan. Distribution: T u r k e s t a n .

269 elegans mortoni SCHMIDT 1938 Ischnura elegans mortoni SCHMIDT, SitzBer. Akad. Wiss. Wien, (I) 147: 142. Type male: Naturhist. Mus., Wien; type-locality: Spain.

Distribution: Spain. elegans pontica SCHMIDT 1938 Ischnura elegans pontica SCHMIDT, SitzBer. Akad. Wiss. Wien., (I), 147: 142. Type male: Naturhist. Mus., Wien; type-locality: Austria. 1984 Ischnura elegans pontica - STEINMANN, Fauna Hung., 160: 33 (Hungary).

Distribution: Central Europe. elegans saharensis AGUESSE 1958 Ischnura elegans saharensis AGUESSE, Rev. franc. Ent., 25: 149. Type male: unknown; typelocality: North Africa.

Distribution: North Africa. erratica CALVERT 1895 Ischnura erratica CALVERT, Proc. Cal. Acad. (2) 4: 491. Type male: Calif. Acad., type-locality: California.

Distribution: U . S . A . evansi MORTON 1919 Ischnura evansi MORTON, Entomologist's month. Mag., 55: 146. Type male: British Museum (Nat. Hist.), London; type-locality: Tigris.

Distribution: Arabia. ezoin (ASAHINA) 1952 Boninagrion ezoin ASAHINA, Mushi, 23 (6): 52. Type male: Hokkaido Univ.; type-locality: Chichi Jima. (as generotype of Boninagrion ASAHINA, 1 9 5 2 ) . 1962 Ischnura ezoin - LIEFTINCK, Insects of Micronesia, 5 (1): 41 (Bonin Island). 1984 Boninagrion ezoin - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 66.

Distribution: Bonin Island. filosa SCHMIDT 1949 Ischnura filosa SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 3Α: 23. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Ischnura filosa - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 137.

Distribution: Madagascar. fluviatilis SELYS 1876 Agrion (Ischnura) fluviatilis (BATES MS) SELYS, Synopsis des Agrionines, Add. 5: 25. Type male: Selys's Collection; type-locality: Buenos Aires. 1890 Micronympha fluviatilis - KIRBY, Syn. Cat. Neur.-Odon., London: 141. 1916 Ischnura fluviatilis - Ris, Arch. Naturg., (A) 82: 132 (Argentina). 1948 Ischnura fluviatilis - FRASER, Acta Zool. lilloana, 5: 57 (Argentina).

Distribution: Argentina.

270 forcipata

MORTON

1907 Ischnura forcipata MORTON, Trans, ent. Soc. London, 1907: 19. Type male: British Museum (Nat. Hist.), London; type-locality: Quetta. 1913 Ischnura gangetica LAIDLAW, Entomologist, 46: 235. Type male: British Museum (Nat.Hist.), London; type-locality: India. 1913 Ischnura musa BARTENEF, Rev. russ. ent., 13, 187. Type male: Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Buchara. 1916 Ischnura forcipata - LAIDLAW, Ree. Indian Mus., 12: 129. 1927 Ischnura forcipata - LIEFTINCK, Tijdschr., Ent., 70: 91 (redescription) (N. India). 1927 Ischnura gangetica - LIEFTINCK, Tijdschr., Ent., 70: 91 (proposed synonymy with forcipata MORTON, 1 9 0 7 ) .

1927 Ischnura musa - LIEFTINCK, Tijdschr., Ent., 70: 91 (proposed synonymy with forcipata MORTON, 1 9 0 7 ) .

1933 Coenagrion neeedhami NAVÄS, MUS. Heude Notes ent. chinoise, 9: 8. Type male: Heude Museum; type-locality: China. 1965 Agriocnemis nainitalensis SAHNI, Indian J. Ent., 27: 206. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: N. India. 1974 Ischnura forcipata - DUMONT, Odonatologica, 3 (3): 158. 1989 Agriocnemis nainitalensis - HÄMÄLÄINEN, Odonatologica, 18 (1): 17 (proposed synonymy with forcipata

MORTON, 1 9 0 7 ) .

Distribution: India, China. fountainei

MORTON

1905 Ischnura fountainei MORTON, Ent. Mag., 1905: 147. Type male: British Museum (Nat.Hist.), London; type-locality: Algeria. 1913 Ischnura bukharensis BARTENEF, Rev. russ. ent., 13: 186. Type male: Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Bukhara. 1961 Ischnura fountainei - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 39 (Kazakhstan). 1964 Ischnura bukharensis - AKRAMOVSKIJ, IZV. Akad. Nauk. Arm., 17 (10): 99 (proposed synonymy w i t h fountainei

MORTON, 1 9 0 5 ) .

Distribution: Central and West Asia, Algeria, Morocco, Tunisia, Libya, Egypt and Sinai to Iraq, Iran, and Saudi Arabia. gemina

(KENNEDY)

1917 Celaenura gemina KENNEDY, Proc. U.S. natn. Mus., 52: 497. Type male: U.S. natn. Mus., Washington; type-locality: California. 1984 Ischnura (Celaenura) gemina - DA VIES & TOBIN, The Dragonflies of the World, Utrecht 1: 72.

Distribution: U.S.A. genei

(RAMBUR)

1842 Agrion genei RAMBUR, Hist. nat. Ins. Neuropt., Paris: 276. Type female: Rambur's Collection; type-locality: Sicilia. 1850 Agrion (Ischnura) genei - SELYS & HÄGEN, Revue des Odonates, Paris: 187 (Sardegna). 1876 Agrion (Ischnura) genei - SELYS, Synopsis des Agrionines, Add. 5: 32 (Sicily, Sardegna, Corse). 1890 Micronympha genei - KIRBY, Syn. Cat. Neur.-Odon., London: 142. 1905 Ischnura genei - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 840. 1937 Ischnura genei - CAPRA, Boll. Soc. ent. ital., 69: 50 (Italy). 1940 Ischnura genei - NIELSEN, Mem. Soc. ent. ital., 19: 240.

Distribution: Mediterranean Islands: Corsica, Sardinia, Sicily, and Malta.

271 graellsi

(RAMBUR)

1842 Agrion graellsi RAMBUR, Hist. nat. Ins. Neuropt., Paris: 275. Type male: Graells's Collection; type-locality: Barcelona. 1 8 5 0 Agrion (Ischnura) algira SELYS & H Ä G E N , Revue des Odonates, Paris: 1 8 5 . Type: unknown; type-locality: Spain. 1876 Agrion (Ischnura) graellsi - SELYS, Synopsis des Agrioninen, Add. 5: 31 (Spain, Portugal, Algeria, Madeira). 1884 Ischnura maroccana KOLBE, Berl. ent. Z., 28: 133. Type male: Mus. Naturk. Berlin; typelocality: Morocco. 1884 Ischnura flavistyla KOLBE, Berl. ent. Z., 28:134. Type male: Mus. Naturk., Berlin; type-locality: unknown. 1890 Micronympha graellsii - KIRBY, Syn. Cat. Neur.-Odon., London: 141. 1890 Micronympha maroccana - KIRBY, Syn. Cat. Neur.-Odon., London: 141. 1 9 0 5 Ischnura graellsi - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 5 (Spain, Marocco, Algeria). 1 9 0 5 Agrion (Ischnura) algira - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 5 [proposed synonymy with graellsi ( R A M B U R , 1 8 4 2 ) ] . 1 9 0 5 Ischnura maroccana - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 5 [proposed synonymy with graellsi ( R A M B U R , 1 8 4 2 ) ] . 1 9 0 5 Ischnura flavistyla- JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 5 [proposed synonymy with graellsi ( R A M B U R , 1 8 4 2 ) ] . 1950 Ischnura graellsi - MORERA, Inst. Esp. Ent. Madrid, 92 (Spain).

Distribution: Spain, Portugal: Madeira; Algeria, Morocco, and Tunisia. haemastigma

FRASER

1927 Ischnura haemastigma FRASER, Ins. Samoa, 7: 26. Type male: British Museum (Nat. Hist.), London; type-locality: Samoa. 1953 Ischnura haemastigma - FRASER, Proc. R. ent. Soc. London, (B) 22: 122 (redescription).

Distribution: Samoa. halecarpenteri

(MUMFORD)

Coenagrion interruptum NEEDHAM (nec CHARPENTIER, 1 8 2 5 ) , Bull. Bishop Mus., 98: 1 1 1 . Type male: Bishop Museum, Honolulu; type-locality: Marquesas Island. 1942 Bedfordia hale-carpenteri MUMFORD, Ann. Mag. nat. Hist., (11) 9: 828 (as generotype of Bedfordia M U M F O R D , 1 9 4 2 ) (new name for Coenagrion interruptum NEEDHAM, 1 9 3 2 , a junior secundary homonymy of Agrion interruptum CHARPENTIER, 1825). 1984 Ischnura hale-carpentieri - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 72. 1932

Distribution: Marquesas Island. heterosticta

(BURMEISTER)

1839 Agrion heterosticta BURMEISTER, H a n d b . Ent., 2: 820. Type male: Germar's Collection; typelocality: Melbourne. 1869 Agrion distigma BRAUER, Verh. Zool.-Bot. Ges. Wien., 19:14. Type male: Hamburg Museum; type-locality: Rockhampton. 1876 Agrion (Ischnura) heterosticta - SELYS, Synopsis des Agrionines, Add. 5: 27 (Australia, New Caledonia). 1890 Micronympha heterosticta - KIRBY, Syn. Cat. Neur.-Odon., London: 141. 1890 Micronympha distigma - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1898 Ischnura heterosticta - CALVERT, Trans. Am. ent. Soc., 25: 40. 1901 Ischnura heterosticta - MARTIN, Mem. Soc. zool. France, 14: 246 (Australia, New Caledonia). 1913 Ischnura heterosticta var. tasmanica T I L L YARD, Proc. Linn. Soc. N.S. Wales, 37: 451. Male from Tasmania.

272 1913 Ischnura torresiana TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 452. Type male: Tillyard's Collection; type-locality: Queensland. 1917 Ischnura heterosticta-SIOSTEOT, Ark. Zool., Uppsala, 11 (11): 33 (New South Wales, Queensland). 1932 Ischnura heterosticta - LIEFTINCK, Nova Guinea, 15 (5): 110. 1 9 6 2 Ischnura FO/RESW/M- LIEFTINCK, Insects of Micronesia, 5 ( 1 ) : 4 1 (North Australia, New Guinea, New Hebrides, Banks Island). 1984 Ischnura heterosticta tasmanica - DAVIS & TOBIN, The Dragonflies of the World, Utrecht, 1: 74.

Distribution: Indo-Austral faunal region. hilli

PINHEY

1964 Ischnura hill PINHEY, J. ent. Soc. S.Afr., 26: 329. Type male: unknown, type-locality: Ethiopia.

Distribution: Ethiopia. inarmata

CALVERT

1898 Ischnura inarmata CALVERT, Proc. Acad. nat. Hist. Philad., 1898: 147. Type male: Acad. Sei. Philad.; type-locality: Kashmir. 1 9 0 5 Ischnura inarmata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 1 5 (Kashmir). 1916 Ischnura inarmata - LAIDLAW, Ree. Indian Mus., 12: 130. 1919 Ischnura inarmata - LAIDLAW, Ree. Indian Mus., 16: 175 (Kashmir).

Distribution: Kashmir. indefinita 1984

FRASER

Ischnura indefinita Utrecht, 1: 73.

FRASER, 1 9 4 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Argentina. indivisa (Ris) 1916 Ceratura indivisa Ris, Arch. Naturg., (A) 82: 135. Type male: not given; type-locality: Colombia. 1958 Ischnura indivisa - RACENIS, Acta biol. venezuel., 2: 182 (description of female).

Distribution: Colombia. intermedia

DUMONT

1974 Ischnura intermedia DUMONT, Odonatologica, 3 (3): 154. Type male: D u m o n t ' s Collection; type-locality: Turkey. 1 9 8 8 Ischnura intermedia - D U M O N T , DEMIRSOY & M E R T E N S , Notul. odonatol., 3 ( 2 ) : 2 4 (Turkey).

Distribution: Turkey. isoetes

LIEFTINCK

1949 Ischnura isoetes LIEFTINCK, Nova Guinea (N.S.) 5: 225. Type male: Mus. Zool., Buitenzorg; type-locality: New Guinea.

Distribution: New Guinea. japonica 1905

JACOBSON & Β Ι Α Ν Κ Ι

Ischnura japonica JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Japan.

Distribution: Japan.

814.

Type male: Zool.

273

karafutonis 1931

MATSUMURA

Ischnura karafutonis type-locality: Japan.

M A T S U M U R A , 6 0 0 0 111.

Ins. Japan:

1461.

Type male: Hokkaido Univ.;

Distribution: Japan. keilicotti

WILLIAMSON

1898 Ischnura keilicotti WILLIAMSON'S, Ent. News, 9: 209. Type male: Williamson's Collection; typelocality: Indiana.

Distribution: U.S.A. lamellata

KOLBE

1884 Ischnura lamellata KOLBE, Berl. ent. Z., 28:155. Type male: Mus. Naturk., Berlin; type-locality: Algeria. 1890 Micronympha lamellata - KIRBY, Syn. Cat. Neur.-Odon., London: 143.

Distribution: Algeria, Morocco, and Mali. nigriceps

FRASER

1924 Ischnura nigriceps FRÄSER, Ree. Indian Mus., 26: 428. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: India.

Distribution: India. ordosi

BARTENEV

1912 Ischnura ordosi BARTENEV, Ann. Mus. Zool. Acad. Imp. Sei. St. Petersbourgh, 16: 442. Type male: Zool. Inst. Mus. Acad. Sei., St. Petersburg; type-locality: Hwang-ho.

Distribution: North Korea: Hwangju. patricia

FRASER

1924 Ischnura patricia FRASER, Ree. Indian Mus., 26: 428. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: India.

Distribution: India. perparva

SELYS

1876 Agrion (Ischnura) perparva (MCLACHLAN MS) SELYS, Synopsis des Agrionines, Add. 5: 19. Type male: Selys's Collection; type-locality: Texas. 1890 Micronympha perparva - KIRBY, Syn. Cat. Neur.-Odon., London: 140. 1910 Ischnura perparva - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 69. 1915 Ischnura perparva - KENNEDY, Proc. U.S. natn. Mus., 49: 310.

Distribution: U.S.A. posita posita

(HÄGEN)

1861 Agrion positum HAGEN, Syn. Neur. N.Amer. Washington: 77. Type male: Hagen's Collection; type-locality: Savannah. 1876 Agrion (Nehalennia) posita- SELYS, Synopsis des Agrionines, Add. 5: 132 (Georgia, Washington, Philadelphia). 1890 Nehalennia posita - KIRBY, Syn. Cat. Neur.-Odon., London: 147 (U.S.A.). 1903 Ischnura posita - NEEDHAM, Bull. 68 N.Y. State Mus.: 261. 1948 Ischnura posita - ZIMMERMANN, Ins. Hawaii, 2: 380 (Hawaii). 1984 Ischnura (Ischnuridia) posita - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 73.

274 1 9 8 9 Ischnurapositaposita

- NOVELO-GUTIERREZ & PENA-OLMEDO, O d o n a t o l o g i c a , 1 8 (1): 4 4 ( U S :

Florida, Georgia, New Jersey, Carolina, Oklahoma, Tennessee).

Distribution: U.S.A. posita acicularis

DONNELLY

1965 Ischnura (Ischnuridia) acicularis DONNELLY, Fla Ent., 48 (1): 57. Type male: Donnelly's Collection; type-locality: Guatemala. 1 9 8 9 Ischnura posita acicularis - N O V E L O - G U T I E R R E Z & P E N A - O L M E D O , Odonatologica, 18 ( 1 ) : 4 5 (Mexico, Guatemala).

Distribution: Mexico, Guatemala. posita azteca 1989

NOVELO-GUTIERREZ & PENA-OLMEDO

Ischnura posita azteca N O V E L O - G U T I E R R E Z & P E N A - O L M E D O , Odontologica, 18 male: Univ. Met.-Xoch., D P A A , type-locality: Mexico.

(1): 45.

Type

Distribution: Mexico. prognata

(HÄGEN)

1861 Agrion prognatum HAGEN, Syn. Neur. N.Amer. Washington: 83. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Virginia. 1876 Agrion (Ischnura) prognatha - SELYS, Synopsis des Agrionines, Add. 5: 15. 1890 Micronympha prognatha - KIRBY, Syn. Cat. Neur.-Odon., London: 140. 1920 Anomaluraprognatha - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Anomalura KENNEDY, 1920).

1984 Ischnura (Anomalura) prognatha - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 73. 1990 Ischnura prognata - DUNKLE, Selysia, 19 (1): 1.

Distribution: U.S.A. pruinescens

(TILLYARD)

1906 Agriocnemis pruinescens TILLYARD, Proc. Linn. Ν. S. Wales Soc., 31: 191. Type male: not given, perhaps in Tillyard's Collection; type-locality: Queensland. 1913 Ischnura pruinescens -TILLYARD, Proc. Linn. N.S. Wales Soc., 37: 453. 1 9 8 4 Ischnura pruinescens - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 7 4 [as syn. with heterosticta (BURMEISTER, 1 8 3 9 ) ] .

Distribution: Australia, pumilio

(CHARPENTIER) 1825 Agrion pumilio CHARPENTIER, Hor. Ent.: 22: Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Italy. 1 8 3 6 Agrion xanthopterum STEPHENS, 111. Brit. Ent., Mand., 5 : 7 2 . Type female: British Museum (Nat. Hist.), London; type-locality: England. 1 8 3 7 Agrion aurantiaca SELYS, (nec R O S T O C K , 1 8 8 5 ) . Cat. Lep. r. Belg.: 1 2 . Type female: Mus. Nation. Hist. Nat., Paris; type-locality: Provence. 1939 Agrion rubellum CURTIS, Brit. Ent., 16: 732. Type male: British Museum (Nat.Hist.), London; type-locality: England. 1876 Agrion (Ischnura) pumilio - SELYS, Synopsis des Agrionines, Add. 5: 23 (England, Belgique, France, Italy, Hungary, Germany, Turkey, Asia Minor, Algeria, Madeira, Transcaucasica). 1885 Ischnura pumilio f. stephanella ROSTOCK, (17: 252) (emendation). 1890 Micronympha pumilio - KIRBY, Syn. Cat. Neur.-Odon., London: 140 (as generotype of Micronympha KIRBY, 1890). 1890 Agrion xanthopterum - KIRBY, Syn. Cat. Neur.-Odon., London: 140 [proposed synonymy w i t h pumilio

(CHARPENTIER, 1 8 2 5 ) ] .

275 1890 Agrion aurantiacum - KIRBY, Syn. Cat. Neur.-Odon., London: 140 [proposed synonymy with pumilio

(CHARPENTIER, 1825)].

1890 Agrion rubellum - KIRBY, Syn. Cat. Neur.-Odon., London: 140 [proposed synonymy with pumilio

(CHARPENTIER, 1825)].

1898 Ischnura pumilio - CALVERT, Trans. Am. ent. Soc., 25: 39. 1905 Ischnura pumilio - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 813 (Europe, Syria, Mesopotamia, Anatolia, Mongolia, Turkestan). 1928 Ischnura pumilio var. immaculata ZACWILICKOWSKI, Spraw. kom. fizio. Krakow: 62. Male from Poland. 1928 Ischnura pumilio var. punctata ZACWILICKOWSKI, Spraw. kom. fizio. Krakow: 68. Male from Poland. 1928 Ischnura pumilio var. striata ZACWILICKOWSKI, Spraw. kom. fizio. Krakow: 69. Male from Poland. 1928 Ischnura pumilio var. ornata ZACWILICKOWSKI, Spraw. kom. fizio. Krakow: 69. Male from Poland. 1928 Ischnura pumilio - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 25 (Egypt). 1938 Ischnura pumilio - SCHMIDT, SitzBer. Akad. Wiss. Wien., (1) 147: 142 (Lebanon). 1948 Ischnura pumilio - AKRAMOWSKI, Zool. Zborn., Moscow, 5: 152 (Armenia). 1950 Ischnura pumilio - MORERA, Inst. Esp. Ent. Madrid, 93 (Spain). 1956 Ischnura pumilio - FRÄSER, R. Ent. Soc. London, 1 (10): 31 (England). 1961 Ischnura pumilio - BELYSHEV, Fragm. Faun. Warszawa, 9 (4): 39 (Kazakhstan). 1984 Ischnura pumilio - STEINMANN, Fauna Hung., 160: 34 (Hungary).

Distribution: C. & W. Asia, Asia Minor, Europe (except Scandinavia, north Russia), North Africa.

ramburi ramburi

(SELYS)

1857 Agrion ramburi SELYS (nec CARPENTER, 1896), in RAMON, Hist. phys. Pol. Nat. Cuba, 8: 189. Type male: Selys's Collection; type-locality: Yucatan. 1857 Agrion tuberculatum SELYS (nec CHARPENTIER, 1825), Sagra Hist. Cuba Ins: 467. Type: unknown; type-locality: Cuba. 1861 Agrion iners HAGEN, Syn. Neur. N.Amer. Washington: 75. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Cuba. 1876 Agrion (Ischnura) ramburi - SELYS, Synopsis des Agrionines, Add. 5: 28. 1890 Micronypha ramburi - KIRBY, Syn. Cat. Neur.-Odon., London: 141. 1902 Ischnura ramburi - CALVERT, Biol. Centr.-Amer. London, 50: 124. 1910 Ischnura ramburi ramburi - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 70. 1910 Agrion tuberculatum - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 70 [proposed synonymy with ramburi ramburi (SELYS, 1857)]. 1910 Agrion iners - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 70 [proposed synonymy with ramburi ramburi (SELYS, 1857)]. 1919 Agrion (Ischnura) ramburi - CALVERT, Trans. Am. ent. Soc., 45: 350. 1943 Ischnura ramburi- SCHMIDT, Beitr. Fauna Perus, 2: 239 (from Canada to Paraguay and Chile).

Distribution: from Canada to Paraguay and Chile. ramburi credula

(HÄGEN)

1861 Agrion credulum HÄGEN, Syn. Neur. N.Amer. Washington: 80. Type male: Agassiz Museum, Cambridge (Mass.); type-locality: Cuba. 1890 Agrion credulum - KIRBY, Syn. Cat. Neur.-Odon., London: 141 [proposed synonymy with ramburi SELYS, 1857)]. 1910 Ischnura ramburi credula - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee 1 (1): 70. 1916 Ischnura ramburi credula - Ris, Arch. Naturg., (A), 82: 132 (Venezuela, Colombia).

276 1984 Ischnura credula - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 74 [as syn. w i t h ramburi

(SELYS, 1857)].

Distribution: Cuba, Venezuela, Colombia, rhodosoma

LIEFTINCK

1959 Ischnura rhodosoma LIEFTINCK, Nova Guinea (N.S.) 10: 220. Type male: ? Raffles Museum, Singapore; type-locality: New Guinea.

Distribution: New Guinea, rubilio

SELYS

1876 Agrion (Ischnura) rubilio SELYS, Bull. Acad. r. Belg., 41: 238. Type male: R. Mus. Brussels; type-locality: India. 1890 Micronympha rubilio - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1901 Ischnura rubilio - MARTIN, Mem. Soc. zool. France, 14: 246 (Australia). 1905 Ischnura rubilio - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 815.

Distribution: India, Australia. rufostigma rufostigma SELYS 1876 Agrion (Ischnura) rufostigma SELYS, Synopsis des Agrionines, Add. 5: 39. Type male: Selys's Collection; type-locality: India. 1890 Micronympha rufostigma - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1905 Ischnura rufostigma - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 815 (India). 1914 Ischnura rufostigma - LAIDLAW, Ree. Indian Mus., 8: 344 (Assam). 1916 Ischnura rufostigma - LAIDLAW, Ree. Indian Mus., 12: 130 (Bengal, Assam, Kashmir). 1964 Ischnura rufostigma rufostigma - SCHMIDT, Ent. Tidschr., 85: 150 (Burma).

Distribution: India, Burma, rufostigma annandalei

LAIDLAW

1919 Ischnura annandalei LAIDLAW, Ree. Indian Mus., 16: 175. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Shan States. 1964 Ischnura rufostigma annandalei - SCHMIDT, Ent. Tidschr., 85: 149 (Burma).

Distribution: Shan States, Burma. rufostigma mildredae

FRASER

1927 Ischnura mildredae FRÄSER, Ree. Indian Mus., 29: 87. Type male: British Museum (Nat.Hist.), London; type-locality: Burma. 1927 Ischnura carpentieri FRÄSER, Proc. R. ent. Soc. London (B) 15:42. Type male: R. Mus. Brussels; type-locality: Annam. 1964 Ischnura carpentieri - SCHMIDT, Ent. Tischr., 85: 150 (proposed synonymy with mildredae FRASER, 1 9 2 7 ) .

1965 Ischnura rufostigma mildredae - ASAHINA, Kontyü, 33: (4): 495 (Hong-Kong).

Distribution: East Asia. rufovittata

(BLANCHARD)

1837 Agrion rufovittata BLANCHARD, D'Orbigny Voy. Amer. Mer.: 217. Type: unknown; type-locality: Bolivia. 1890 Micronympha rufovittata - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1984 Ischnura rufovittata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 74. Distribution: Bolivia.

277 sanguinostigma FRÄSER 1953 Ischnura sanguinostigma FRÄSER, Proc. R. ent. Soc. London, (B) 22:123. Type male: Fraser's Collection; type-locality: Samoa.

Distribution: Samoa. senegalensis (RAMBUR) 1842 Agrion senegalense RAMBUR, Hist. nat. Ins. Neuropt: 276 .Type male: lost; type-locality: Senegal. 1876 Agrion (Ischnura) senegalense - SELYS, Synopsis des Agrionines, Add. 5: 29 (Senegal, Egypt, Angola, Cap, Reunion, Bombay, Java). 1890 Micronympha senegalensis - KIRBY, Syn. Cat. Neur.-Odon., London: 141. 1896 Micronympha senegalensis - KIRBY, Ann. Mag. nat. Hist., (7) 2: 244 (Pretoria, Nyasaland). 1901 Ischnura senegalensis - MARTIN, Mem. Soc. zool. France, 14: 246 (Victoria, N. Australia). 1924 Ischnura senegalensis - FRÄSER, Ree. Indian Mus., 26: 489 (W. India). 1927 Ischnura senegalensis - Ris, Zool. Meded., Leiden, 10 (1): 21 (Sumatra). 1928 Ischnura senegalensis - ANDERS, Mem. Soc. R. ent. Egyp., 3 (1): 25 (Egypt). 1934 Ischnura senegalensis - SCHMIDT, Arch. Hydrob. Suppl., 13: 339 (Java, Sumatra) («description). 1936 Ischnura senegalensis - LONGFIELD, Trans. R. ent. Soc. London, 85: 471. 1936 Ischnura senegalensis - LIEFTINCK, Revue suisse Zool., 43 (5): 136 (Bali, Lombok). 1939 Ischnura senegalensis - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 289 (Philippines). 1949 Ischnura senegalensis - FRASER, Trans. R. ent. Soc. London; 100: 138 (Mauritius). 1954 Ischnura senegalensis - LIEFTINCK, Treubia, 22 Suppl.: 74 (Siam, Malaya, Krakatau, Sumatra, Java, Borneo, Bali). 1955 Ischnura senegalensis - BRINCK, South African Animal Life, 2: 196 (Cape Prov., Basutoland, Natal, Transvaal, S. Rhodesia, S.W. Africa, Seychelles, India). 1962 Ischnura senegalensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 137 (Africa, Zanzibar, Seychelles, Mauritius, Madagascar).

Distribution: Old World tropics and temperate regions, from East Africa to Japan. sobrina SCHMIDT 1943 Ischnura sobrina SCHMIDT, Beitr. Fauna Perus, 2: 237. Type male: Mus. Naturk., Berlin; typelocality: Peru.

Distribution: Peru.. spinicauda (BRAUER) 1865 Agrion (Ischnura) spinicauda BRAUER, Verh. Zool.-Bot. Ges. Wien., 15: 511. Type male: not given; type-locality: Polynesia. 1890 Micronympha spinicauda - KIRBY, Syn. Cat. Neur.-Odon., London: 143.

Distribution: Polynesia. stueberi LIEFTINCK 1932 Ischnura stueberi LIEFTINCK, Nova Guinea, 15 (5): 111. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. taitensis SELYS 1876 Agrion (Ischnura) taitensis SELYS, Synopsis des Agrionines, Add. 5: 35. Type male: Selys's Collection; type-locality: Tahiti.

278 1890 Micronympha taitensis - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1942 Ischnura cheesmani FRÄSER, in MUMFORD, Ann. Mag. nat. Hist., (11)9: 646. Type male: British Museum (Nat.Hist.), London; type-locality: Tahiti.

Distribution: Tahiti. thelmae

LIEFTINCK

1984 Ischnura thelmae LIEFTINCK, 1966 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 74.

Distribution: Rapa Island (South Pacific Ocean). ultima Ris 1908 Ischnura ultima Ris, Dtsch. ent. Z., 1908: 518. Type male: Mus. Naturk. Berlin; type-locality: Argentina. 1916 Ischnura ultima - Ris, Arch. Naturg., (A) 82: 133. 1948 Ischnura ultima - FRASER, Acta Zool. lilloana, 5: 57 (Argentina, Chile).

Distribution: Argentina, Chile. verticalis (SAY) 1839 Agrion verticalis SAY, Jl. Acad. Sei. Philad., 8: 37. Type male: unknown; type-locality: New York. 1876 Agrion (Ischnura) verticalis - SELYS, Synopsis des Agrionines, Add. 5: 21 (New York, Washington, New Jersey, Georgia, Philadelphia, St. Louis, Illinois, Maryland, Indiana). 1890 Micronympha verticalis - KIRBY, Syn. Cat. Neur.-Odon., London: 140. 1910 Ischnura ramburii „HAGEN, 1861" - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee 1 (1): 71 [as syn. with verticalis (SAY, 1839)].

Distribution: U.S.A. vinsoni

FRASER

1949 Ischnura vinsoni FRASER, Trans. R. ent. Soc. London, 100: 140. Type male: British Museum (Nat.Hist.), London; type-locality: Mauritius. 1962 Ischnura vinsoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 137 (type male: Fraser's Collection.

Distribution: Mauritius. Genus Mesamphiagrion

KENNEDY

1920 Mesamphiagrion KENNEDY, Ohio J. Sei. Columbus, 21: 87. Type-species: Enallagma occultum Ris, 1918 (original designation).

occultum Ris 1916 Enallagma occultum Ris, Arch. Naturg., (A), 82: 118. Type male: not given, perhaps in Mus. Naturk., Berlin; type-locality: Colombia. 1920 Mesamphiagrion occultum - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Mesamphiagrion

KENNEDY, 1920).

1989 Enallagma occultum - DE MARMELS, Odontologica, 18 (3): 248.

Distribution: Colombia. Genus Millotagrion

FRASER

1953 Millotagrion FRASER, Nat. malgache, 51 (2): 159. Type-species: Millotagrion inaequistigma FRASER, 1953 (original designation). - Millotagrion - PINHELY, Publ. cult. Comp. Diam. Angola, 59: 118.

279

inaequistigma

FRÄSER

1953 Millotagrion inaequistigma FRÄSER, Nat. malgache, 5: 161. Type male: Fraser's Collection; type-locality: Madagascar (as generotype of Millotagrion FRASER, 1953).

Distribution: Madagascar. Genus Oreagrion Ris 1913 Oreagrion Ris, Nova Guinea, 9: 480. Type species: Oreagrion lorentzi Ris, 1913 (original designation).

armeniacum

LIEFTINCK

1949 Oreagrion armeniacum LIEFTINCK, Nova Guinea (N.S.) 5: 221. Type male: Raffles Museum, Singapore; type-locality: New Guinea.

Distribution: New Guinea. lorentzi Ris 1913 Oreagrion lorentzi Ris, Nova Guinea, 9: 480. Type: unknown; type-locality: New Guinea (as generotype of Oreagrion Ris, 1913).

Distribution: New Guinea. oreadum

LIEFTINCK

1949 oreadum LIEFTINCK, N o v a Guinea (N.S.) 5: 206. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. xanthocyane

LIEFTINCK

1949 Oreagrion xanthocyane LIEFTINCK, Nova Guinea (N.S.) 5: 216. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Oxyagrion 1876

SELYS

Agrion (Oxyagrion) SELYS, Synopsis des Agrionines, Add. 5: 4 6 . Type-species: Oxyagrion dissidens SELYS 1 8 7 6 (original designation). - 1 8 9 0 Oxyagrion - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 4 4 . - 1 9 1 0 Oxyagrion - M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 ( 1 ) : 5 3 (typespecies: Agrion rubidum R A M B U R , 1 8 4 2 ) . - 1 9 6 0 Oxyagrion - S T . Q U E N T I N , Beitr. neotrop. F a u n a , 2 ( 1 ) : 5 9 (key to species). - 1 9 7 4 Oxyagrion - B U L L A , Revista Mus. La Plata (Zool.), 12 ( 1 1 2 ) : 2 3 (key to species). - 1 9 7 8 Oxyagrion - C O S T A , Publ. avuls. Mus. nac. Rio de J., 61: 39 (revision, key to species).

basale

SELYS

1876 Agrion (Oxyagrion) basale SELYS, Synopsis des Agrionines, Add. 5: 59. Type male: Selys's Collection; type-locality: Brazil. 1890 Oxyagrion basale - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 4 4 . 1948 Oxyagrion basale - FRASER; Acta. Zool. lilloana, 5 : 53 (Argentina).

Distribution: Brazil, Argentina. brevistigma

SELYS

1876 Agrion (Oxyagrion) brevistigma SELYS, Synopsis des Agrionines, Add. 5: 53. Type male: Selys's Collection; type-locality: Brazil.

280 1890 Oxyagrion

brevistigma

- KIRBY,

Syn. Cat. Neur.-Odon., London:

144.

Distribution: Brazil. bruchi Naväs 1924 Oxyagrion bruchi N A V Ä S , Mem. R. Acad. Barcelona, 18 (13): 19. Type male: Mus. Cienc. Nat., Barcelona; type-locality: Argentina.

Distribution: Argentina. cardinale

FRASER

1946 Oxyagrion cardinalis FRASER, Trans. R. ent. Soc. London, 9 6 : 4 1 . Type female: Leeds Museum; type-locality: Peru. 1966 Oxyagrion cardinalis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 1 8 : 185 (type in: British Museum (Nat.Hist.), London).

Distribution: Peru. chapadense 1978

COSTA

Oxyagrion chapadense COSTA, Publ. avuls. Mus. nac. Rio de J., Rio de J.; type-locality: Brazil.

61: 92.

Type male: Mus. Nac.

Distribution: Brazil. divaricatum

CALVERT

1909 Oxyagrion divaricatum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6:181. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil. evanescens

CALVERT

1909 Oxyagrion evanescens CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 179. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil. haematinum

SELYS

1876 Agrion (Oxyagrion) haematinum 1890

SELYS,

Synopsis des Agrionines, Add. 5: 56. Type male: Selys's

Collection; type-locality: Minas Geraes. Oxyagrion haematinum - K I R B Y , Syn. Cat. Neur.-Odon., London:

144.

Distribution: Brazil. hempeli

CALVERT

1909 Oxyagrion hempeli CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 182. Type male: Carnegie Museum; type-locality: Brazil. 1948 Oxyagrion hempeli - FRASER, Acta Zool. lilloana, 5: 53 (Argentina) (redescription). 1960 Oxyagrion hempeli - S T . Q U E N T I N , Beitr. neotrop. F a u n a , 2 (1): 59.

Distribution: Brazil. impunctatum

CALVERT

1909 Oxyagrion impunctatum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 188. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil.

281 machadoi COSTA 1978 Oxyagrion machadoi COSTA, Publ. avuls. Mus. nac. Rio de J., 61: 131. Type male: Mus. Nac. Rio de J., type-locality: Brazil.

Distribution: Brazil. microstigma SELYS 1876 Agrion (Oxyagrion) microstigma SELYS, Synopsis des Agrionines, Add. 5: 54. Type male: Selys's Collection; type-locality: Brazil. 1890 Oxyagrion microstigma - KIRBY, Syn. Cat. Neur.-Odon., London: 144.

Distribution: Brazil. miniopsis SELYS 1876 Agrion (Oxyagrion) miniopsis SELYS, Synopsis des Agrionines, Add. 5: 55.Type male: McLachlan's Collection; type-locality: Bogota. 1890 Oxyagrion miniopsis - KIRBY, Syn. Cat. Neur.-Odon., London: 144.

Distribution: Brazil. pavidum HÄGEN 1876 Agrion (Oxyagrion) pavidum HAGEN, in SELYS, Synopsis des Agrionines, Add. 5: 50. Type male: Selys's Collection; type-locality: Brazil. 1890 Oxyagrion pavidum - KIRBY, Syn. Cat. Neur.-Odon., London: 144.

Distribution: Brazil. rubidum rubidum (RAMBUR) 1842 Agrion rubidum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 261. Type male: Serville's Collection; type-locality: Buenos Aires. 1876 Agrion (Oxyagrion) rubidum - SELYS, Synopsis des Agrionines, Add. 5: 57 (as generotype of Oxyagrion

SELYS, 1 8 7 6 ) .

1890 Oxyagrion rubidum - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1984 Oxyagrion rubidum rubidum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 75.

Distribution: Argentina. rubidum rufulum (HÄGEN) 1861 Agrion rufulum HÄGEN, Syn. Neur. N.Amer. Washington: 86. Type male: Hagen's Collection; type-locality: North California. 1876 Agrion (Oxyagrion) rufulum - SELYS, Synopsis des Agrionines, Add. 5: 58. 1890 Oxyagrion rufulum - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1984 Oxyagrion rubidum rufulum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 75.

Distribution: U.S.A. santosi MARTIN 1984 Oxyagrion santosi MARTIN, 1967 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 76.

Distribution: Brazil. simile COSTA 1978 Oxyagrion simile COSTA, Publ. avuls. Mus. nac. R. de J., 61: 176. Type male: Mus. Nac. Rio de J., type-locality: Venezuela.

Distribution: Venezuela.

282 sulinum

COSTA

1978 Oxyagrion sulinum COSTA, Publ. avuls. Mus. nac. R. de J., 61: 185. Type male: Mus. Nac. Rio de J., type-locality: Venezuela.

Distribution: Venezuela, terminale

SELYS

1876 Agrion (Oxyagrion) terminale SELYS, Synopsis des Agrionines, Add. 5: Type male: Selys's Collection; type-locality: Buenos Aires. 1890 Oxyagrion terminale - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1916 Oxyagrion terminale - Ris, Arch. Naturg., (A), 82: 127 (Bolivia). 1946 Oxyagrion terminale - FRÄSER, Trans. R. ent. Soc. London: 96: 40 (Argentina). 1948 Oxyagrion terminale - FRÄSER, Acta Zool. lilloana, 5: 53. 1984 Oxyagrion terminale - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 76 (Brazil).

Distribution: Bolivia, Argentina, Brazil. Genus Pacificagrion

FRÄSER

1925 Pacificagrion FRASER, Trans, ent. Soc. London, 1925: 505. Type-species: Pacificagrion rymosa FRASER, 1925 (monotypy).

dolorosa

lach-

FRASER

1953 Pacificagrion dolorosa FRASER, Proc. R. ent. Soc. London (B), 22: 119. Type male: British Museum (Nat.Hist.), London; type-locality: Samoa.

Distribution: Samoa. lachrymosa

FRASER

1925 Pacificagrion lachrymosa FRASER, Trans, ent. Soc. London, 1925: 505 (as generotype of Pacificagrion FRASER, 1925). Type male: British Museum (Nat.Hist.), London; type-locality: Samoa. 1927 Pacificagrion lachrymosa - FRASER, Ins. Samoa, 7: 33 (description of female). 1953 Pacificagrion lachrymosa - FRASER, Proc. R. ent. Soc. London, (B) 22, 120.

Distribution: Samoa.

Genus Protallagma

KENNEDY

1920 Protallagma KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type-species: Amphiagrion titicacae CALVERT, 1909 (original designation). - 1920 Oxyallagma KENNEDY, Ohio J. Sei. Columbus, 21: 87. Type-species: Agrion (Oxyagrion) dissidens SELYS, 1876 (original designation). 1958 Protallagma - FRASER, Rev. univ. Chile, 42 (2): 154 (not a valid genus). - 1984 Oxyallagma - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 76 (as including taxa).

dissidens

(SELYS)

1876 Agion (Oxyagrion) dissidens SELYS, Synopsis des Agrionines, Add. 5: 48. Type male: Selys's Collection; type-locality: Quito. 1890 Oxyagrion dissidens - KIRBY, Syn. Cat. Neur.-Odon., London: 144. 1920 Oxyagrion dissidens - KENNEDY, Ohio J. Sei. Columbus, 21: 87 (as generotype of Oxyallagma KENNEDY, 1 9 2 0 ) .

1939 Protallagma runtuni KENNEDY, Ann. ent. Soc. Amer., 32 (3): 177. Type female: unknown; type-locality: Ecuador.

283 1946 Protallagma runtunidissidens

KENNEDY, Ann. ent. Soc. Amer., 39 (3): 381. [proposed synonymy with

(SELYS, 1876)].

Distribution: Ecuador. titicacae

(CALVERT)

1909 Amphiagrion titicacae CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 203. Type male: Calvert's Collection; type-locality: Bolivia. 1909 Amphiagrion andinum FÖRSTER, Wiesbaden Jb Ver. Natk., 62: 231. Type male: Mus. Naturk., Berlin; type-locality: Bolivia. 1916 Amphiagrion titicacae - Ris, Arch. Naturg., (A) 82: 131 (Peru, Bolivia). 1920 Protallagma titicacae - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Protallagma KENNEDY, 1 9 2 0 ) .

1943 Amphiagrion 1943 Amphiagrion cacae

titicacae - SCHMIDT, Beitr. Fauna Perus, 2: 240 (Peru). andinum - SCHMIDT, Beitr. Fauna Perus, 2: 240 [proposed synonymy with titi-

(CALVERT, 1 9 0 9 ) ] .

1972 Protallagma

titicacae - BULLA, Neotropica, 18 (57): 131.

Distribution: Bolivia, Peru. Genus Rhodischnura

LAIDLAW

1919 Rhodischnura LAIDLAW, Ree. Indian Mus., 16: 177. Type-species: Ischnura nursei MORTON, 1907 (original designation).

nursei

(MORTON)

1907 Ischnura ? nursei MORTON, Trans, ent. Soc. London, 1907:19. Type male: Morton's Collection; type-locality: Deesa. 1916 Ischnura nursei - LAIDLAW, Ree. Indian Mus., 12: 131. 1919 Rhodischnura nursei - LAIDLAW, Ree. Indian Mus., 16: 177 (as generotype of Rhodischnura LAIDLAW, 1919) (description of female).

Distribution: India. Genus Skiallagma

FÖRSTER

1906 Skiallagma FÖRSTER, Insekten-Börse, 1906: 23. Type-species: Skiallagma baueri FÖRSTER, 1906 (monotypy).

baueri

FÖRSTER

1906 Skiallagma baueri FÖRSTER, Insekten-Börse, 1906: 23 (as generotype of Skiallagma FÖRSTER, 1906). Type: undesignated. 1984 Skiallagma baueri - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 76.

Distribution: Neotropical region. Genus Tigriagrion

CALVERT

1909 Tigriagrion CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 195. Type-species: Tigriagrion aurantinigrum CALVERT, 1909 (original designation). - 1960 Tigriagrion - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 54.

aurantinigrum

CALVERT

1909 Tigriagrion aurantinigrum CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 195 (as generotype of Tigriagrion CALVERT, 1909) Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil.

284

Genus Xiphiagrion

SELYS

1876 Agrion (Xiphiagrion) SELYS, Synopsis des Agrionines, Add. 5: 77 Type-species: Xiphiagrion cyanomelas SELYS, 1 8 7 6 (designated by K I R B Y , 1 8 9 0 , Syn. Cat. Neur.-Odon., London: 1 4 5 ) . - 1890 Xiphiagrion - KIRBY, Syn. Cat. Neur.-Odon., London: 145. - 1954 Xiphiagrion LIEFTINCK, Treubia, 22 Suppl.: 75.

cyanomelas

SELYS

1876 Agrion (Xiphiagrion) cyanomelas SELYS, Synopsis des Agrionines, Add. 5: 77 (as generotype of Xiphiagrion SELYS, 1876). Type male: Selys's Collection; type-locality: Moluccas. 1890 Xiphiagrion cyanomelas - KIRBY, Syn. Cat. Neur.-Odon., London: 145. 1898 Xiphiagrion karschi Ris, Ent. Nachr., 24: 326. Type male: Mus. Naturk., Berlin; type-locality: Bismarck Arch. 1934 Xiphiagrion cyanomelas - LIEFTINCK, Treubia, 14 (4): 400 (Java). 1936 Xiphiagrion cyanomelas - LIEFTINCK, Revue suisse Zool., 43 (5): 134 (Flores, Java, Celebes). 1949 Xiphiagrion cyanomelas - LIEFTINCK, Nova Guinea (N.S.) 5: 197. 1949 Xiphiagrion karschi - LIEFTINCK, Nova Guinea (N.S.) 5: 197 (proposed synonymy with cyanomelas SELYS, 1 8 7 6 ) . 1951 Xiphiagrion cyanomelas race berlandi SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 274. Male from Madagascar. 1954 Xiphiagrion cyanomelas - LIEFTINCK, Treubia, 22 Suppl.: 75 (Sumatra, Java, Borneo).

Distribution: Simalur, Engano, Java, Borneo, Palawan, Celebes, Flores, Sumba, Sula, Buru, Ceram, New Guinea, Aru, New Britain; New Ireland. truncatum

LIEFTINCK

1949 Xiphiagrion truncatum LIEFTINCK, Nova Guinea (N.S.) 5: 197. Type male: Rijksmus. Natuurl. Hist., Leiden, type-locality: New Guinea.

Distribution: New Guinea.

Genus Zoniagrion

KENNEDY

1917 Zoniagrion KENNEDY, Proc. U.S. natn. Mus., 52: 488. Type-species: Agrion SELYS, 1876 (original designation).

exclamationis

exclamationis

(SELYS)

1876 Agrion (Agrion) exclamationis SELYS, Synopsis des Agrionines, Add. 5: 141. Type male: McLachlan's Collection; type-locality: California. 1890 Coenagrion exclamationis - KIRBY, Syn. Cat. Neur.-Odon., London: 148. 1917 Zoniagrion exclamationis - KENNEDY, Proc. U.S. natn. Mus., 52: 489 (as generotype of Zoniagrion KENNEDY, 1917). Distribution: U . S . A .

Subfamily Leptobasinae Wings petiolate to distal end of De; anal vein leaves wing border at its inner end. Arc near level of distal antenodal; sectors of arc arise separately. Costal side of Dc s h o r t e r t h a n p o s t e r i o r . L a b i u m cleft f o r apical t h i r d t o half. Legs

short, spines shorter than twice separating distance. N o vulvar spine.

285 1984 Leptobasinae DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 76. Type-genus: Leptobasis

SELYS, 1 8 7 7 .

Genus Antiagrion Ris 1904 Antiagrion Ris, Hamb. Magalhaen. Reise, 7: 7. Type-species: Agrion (Erythromma) gayi SELYS, 1876 (original designation). - 1974 Antiagrion - JURZITZA, Odonatologica, 3 (4): 221.

antigone Ris 1928 Antiagrion antigone Ris, Ent. Mitt., 17: 165. Type male: unknown; type-locality: Chile.

Distribution: Chile. blanchardi SELYS 1876 Agrion (Erythromma) blanchardi SELYS, Synopsis des Agrionines, Add. 5: 199. Type male: McLachlan's Collection; type-locality: Chile. 1890 Erythromma blanchardi - KIRBY, Syn. Cat. Neur.-Odon., London: 152. 1984 Antiagrion blanchardi - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 76.

Distribution: Chile. gayi (SELYS) 1876 Agrion (Erythromma) gayi SELYS, Synopsis des Agrionines, Add. 5: 197. Type male: Mus. Nation. Hist.Nat., Paris; type-locality: Chile. 1890 Erythromma gayi - KIRBY, Syn. Cat. Neur.-Odon., London: 152. 1904 Antiagrion gayi - Ris, Hamb. Magalhaen. Reise, 7: 7 (as generotype of Antiagrion Ris, 1904). 1974 Antiagrion gayi - JURZITZA, Odonatologica, 3 (4): 222.

Distribution: Chile. grinbergi JURZITZA 1974 Antiagrion grinbergi JURZITZA, Odonatologica, 3 (4): 222. Type male: unknown; type-locality: Chile.

Distribution: Chile. Genus Chrysobasis RACENIS, 1959 Chrysobasis RACENIS, Senckenb. biol. 40: 55. Type-species: Chrysobasis buchholzi RACENIS, 1959 (original designation).

buchholzi RACENIS 1959 Chrysobasis buchholzi RACENIS, Senckenb. biol., 40: 56. Type male: Mus. nac. Caracas; typelocality: Venezuela (as generotype of Chrysobasis RACENIS, 1959).

Distribution: Venezuela. lucifer DONNELLY 1984 Chrysobasis lucifer DONNELLY, 1967 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 77.

Distribution: Guatemala.

286

Genus Diceratobasis

KENNEDY

1920 Diceratobasis KENNEDY, Ohio J. Sei. Columbus, 21: 88. Type-species: Agrion (Nehalennia) macrogaster SELYS, 1857 (original designation). - 1986 Diceratobasis - GARRISON, Odonatologica, 15 (1): 65.

macrogaster

(SELYS)

1857 Agrion (Nehalennia) macrogaster SELYS, Sagra, Hist. Cuba Ins.: 465. Type male: Selys's Collection; type-locality: Jamaica. 1877 Telebasis (Leptobasis) macrogastra - SELYS, Bull. Acad. r. Belg. (2) 43: 102. 1890 Leptobasis macrogaster - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1919 Agrion (Nehalennia) macrogaster - CALVERT, Trans. Am. ent. Soc., 45: 350 (Cuba). 1920 Diceratobasis macrogaster - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Diceratobasis KENNEDY, 1920).

Distribution: Jamaica, Cuba. melanogaster

GARRISON

1986 Diceratobasis melanogaster GARRISON, Odonatologica, 15 (1): 62. Type: unknown; type-locality: Dominica.

Distribution: Dominican Republic.

Genus Inpabasis

SANTOS

1961 Inpabasis SANTOS, Bol. Mus. Goeldi (Zool.) N.S. 34: 1. Type-species: Inpabasis eliasi SANTOS, 1961 (original designation).

eliasi

SANTOS

1961 Inpabasis eliasi SANTOS, Bol. Mus. Goeldi (Zool.) N.S. 34:2 (as generotype of Inpabasis SANTOS, 1961). Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. hubelli

SANTOS

1961 Inpabasis hubelli SANTOS, Bol. Mus. Goeldi (Zool.) N.S. 34: 5. Type male: Nac. Mus. Rio de Janeiro; type-locality: Peru.

Distribution: Peru. machadoi

SANTOS

1961 Inpabasis machadoi SANTOS, Bol. Mus. Goeldi (Zool.) N.S. 34: 4. Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. Genus Leptobasis

SELYS

1877 Telebasis (Leptobasis) SELYS, Bui. Acad. r. Belg., (2) 43:99. Type-species: Telebasis (Leptobasis) vacillans HÄGEN, 1877 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 156). 1890 Leptobasis- KIRBY, Syn. Cat. Neur.-Odon., London: 156. - 1919 Agrion (Leptobasis) - CALVERT, Trans. Am. ent. Soc., 45: 355. - 1960 Leptobasis - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 53 (key to South American species).

287 candelaria

ALAYO

1984 Leptobasis candelaria ALAYO, 1968 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the World,

Utrecht, 1: 77.

Distribution: Cuba. costalimai

SANTOS

1957 Leptobasis costalimai SANTOS, Bol. Mus. nac. Rio de Janeiro, Zool., 157: 2. Type male: Nac. Mus. Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. inversum

(SELYS)

1876 Agrion (Telagrion) inversum SELYS, Synopsis des Agrionines, Add. 5: 259. Type male: Selys's Collection; type-locality: Amazons. 1890 Telagrion inversum - KIRBY, Syn. Cat. Neur.-Odon., London: 155. 1 9 6 1 Leptobasis tuberculata SANTOS, Rev. bras. Biol., 2 1 : 1 7 1 . Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. mammilaris

CALVERT

1909 Leptobasis mammilaris CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 201. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil. vacillans vacillans

(HÄGEN)

1877 Telebasis (Leptobasis) vacillans HAGEN, in SELYS, Bull. Acad. r. Belg. (2) 43: 101. Type male: Mus. Nation. Hist. Nat., Paris: type-locality: Cuba. 1890 Leptobasis vacillans-KIRBY, Syn. Cat. Neur.-Odon., London: 156 (as gencrotype o( Leptobasis SELYS, 1877).

1909 Leptobasis vacillans race guinae CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 201. Male from Brazil. 1909 Leptobasis vacillans race ecuadorica CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 201. Male from Ecuador. 1910 Leptobasis vacillans - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee 1 (1): 64. [Type in British Museum (Nat.Hist.), London]. 1919 Agrion (Leptobasis) vacillans - CALVERT, Trans. Am. ent. Soc., 45: 355 (Cuba). 1984 Leptobasis vacillans f. guianae- DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 77. 1984 Leptobasis vacillans f. ecuadorica - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 77. 1 9 8 4 Leptobasis crocea FÖRSTER, 1 9 0 6 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 77.

Distribution: Central and South America. vacillans atrodorsum

CALVERT

1902 Leptobasis vacillans atrodorsum CALVERT, Biol. Centr.-Amer., London: 121. Type male: British Museum (Nat.Hist.), London; type-locality: Mexico. 1910 Leptobasis vacillans atrodorsum -MUTTKOWSKI, Bull. Publ. Mus. Milwaukee 1 (1): 64 (Mexico, Panama). 1958 Leptobasis atrodorsum - RACENIS, Acta biol. venezuel., 2: 194. 1984 Leptobasis vacillans f. atrodorsum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 77.

Distribution: Central America.

288 Genus Leucobasis

RACENIS

1959 Leucobasis RACENIS, Senckenb. biol. 40: 58. Type-species: Leucobasis candicans RACENIS, 1959 (original designation).

candicans

RACENIS

1959 Leucobasis candicans RACENIS, Senckenb. biol., 40: 59 (as generotype of Leucobasis 1959). Type male: Mus. Cienc. nat., Caracas; type-locality: Venezuela.

RACENIS,

Distribution: Venezuela. Genus Mesoleptobasis

SJÖSTEDT

1917 Mesoleptobasis SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 21. Type-species: Mesoleptobasis

incus

SJÖSTEDT, 1917 ( m o n o t y p y ) .

acuminata

SANTOS

1961 Mesoleptobasis acuminata SANTOS, Rev. bras. Biol., 21: 200. Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. cantralli

SANTOS

1961 Mesoleptobasis cantralli SANTOS, Rev. bras. Biol., 21: 196. Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. inca

SJÖSTEDT

1917 Mesoleptobasis incus SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 22 (as generotype of Mesoleptobasis SJÖSTEDT, 1 9 1 7 ) . Type female: Naturh. Riksmus., Stockholm; type-locality: Amazonas.

Distribution: Brazil. Genus Metaleptobasis

CALVERT

1907 Metaleptobasis CALVERT, Biol. Centr.-Amer., London: 386. Type-species: Metaleptobasis bovilla CALVERT, 1 9 0 7 (original designation). - 1 9 1 5 Metaleptobasis - WILLIAMSON, Proc. U . S . natn. Mus., 48: 601 (revision, key to species). - 1955 Metaleptobasis - RACENIS, Bol. Mus. Cienc. nat., Caracas, 1 80 (key to Venezuelan species).

amazonica

SJÖSTEDT

1917 Metaleptobasis amazonica SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 19. Type male: Naturhist. Riksmus., Stockholm; type-locality: Amazonas.

Distribution: Brazil. bicornis

(SELYS)

1877 Telebasis (Leptobasis) bicornis SELYS, Bull. Acad. r. Belg. (2) 43: 103. Type female: Selys's Collection; type-locality: Amazonas. 1890 Leptobasis bicornis - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1 9 8 4 Metaleptobasis bicornis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 7 7 .

Distribution: Brazil.

289 bovilla CALVERT 1907 Metaleptobasis bovilla CALVERT, Biol. Centr.-Amer., London: 386 (as generotype of Melaleptobasis CALVERT, 1907). Type male: Calvert's Collection; type-locality: Nicaragua. 1915 Metaleptobasis bovilla - WILLIAMSON, Proc. U.S. natn. Mus., 48: 602. 1984 Metaleptobasis bovilla - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 77 (Guatemala).

Distribution: Nicaragua, Guatemala. brysonima WILLIAMSON 1915 Metaleptobasis brysonima WILLIAMSON, Proc. U.S. natn. Mus., 48: 602. Type male: Williamson's Collection; type-locality: Trinidad. 1917 Metaleptobasis brysonima - WILLIAMSON, Ent. News, 28: 8. 1932 Metaleptobasis brysonima - GEIJSKES, Zool. Meded., Leiden, 14: 258. 1955 Metaleptobasis brysonima - RACENIS, Bol. Mus. Cienc. nat., Caracas, 1: 74 (Venezuela).

Distribution: Trinidad, Venezuela. fernandezi RACENIS 1955 Metaleptobasis fernandezi RACENIS, Bol. Mus. Cienc. nat., Caracas, 1: 75. Type male: Mus. Cienc. nat., Caracas; type-locality: Venezuela.

Distribution: Venezuela. foreli Ris 1916 Metaleptobasis foreli Ris, Arch. Naturg., (A), 82: 130. Type female: not given; type-locality: Columbia. 1955 Metaleptobasis foreli - RACENIS, Bol. Mus. Cienc. nat., Caracas, 1: 68 (description of male) (Venezuela).

Distribution: Venezuela. manicaria WILLIAMSON 1915 Metaleptobasis manicaria WILLIAMSON, Proc. U.S. natn. Mus., 48: 603. Type male: Williamson's Collection; type-locality: Trinidad. 1932 Metaleptobasis manicaria - GEIJSKES, Zool. Meded., Leiden, 14: 260 (description of female) (Trinidad). 1955 Metaleptobasis manicaria - Racenis, Bol. Mus, Cienc. nat., Caracas, 1: 73 (Venezuela).

Distribution: Trinidad, Venezuela. mauritia WILLIAMSON 1915 Metaleptobasis mauritia WILLIAMSON, Proc. U.S. natn. Mus., 48: 603. Type male: Williamson's Collection; type-locality: Trinidad. 1932 Metaleptobasis mauritia - GEIJSKES, Zool. Meded., Leiden, 14: 258 (redescription).

Distribution: Trinidad. quadricornis (SELYS) 1877 Telebasis (Leptobasis) quadricornis SELYS, Bull. Acad. r. Belg., (2) 43: 104. Type female: Selys's Collection; type-locality: Para. 1890 Leptobasis quadricornis - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1984 Metaleptobasis quadricornis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 78.

Distribution: Brazil.

290 rosea (SELYS) 1877 Telebasis (Leptobasis) rosea SELYS, Bull. Acad. r. Belg. (2), 43: 105. Type male: Selys's Collection; type-locality: Amazonas. 1890 Leptobasis rosea - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 5 6 . 1 9 8 4 Metaleptobasis rosea - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 7 8 .

Distribution: Brazil. selysi SANTOS 1956 Metaleptobasis selysi type-locality: Brazil.

SANTOS,

Rev. bras. Biol., 16: 381. Type male: Nac. Mus., Rio de Janeiro;

Distribution: Brazil. tetragena CALVERT 1948 Metaleptobasis tetragena locality: British Guiana.

CALVERT,

Zoologica, 33: 60. Type male: Calvert's Collection; type-

Distribution: British Guyana. weibezahni RACENIS 1955 Metaleptobasis weibezahni R A C E N I S , Bol. Mus. Cienc. nat. Caracas, 1: 77. Type male: Mus. Cienc. nat., Caracas, type-locality: Venezuela.

Distribution: Venezuela. westfalli CUMMING 1954 Metaleptobasis westfalli cality: Panama.

CUMMING,

Florida Ent., 37: 25. Type male: Univ. Florida; type-lo-

Distribution: Panama.

Subfamily Pseudagriinae Wings petiolate barely to Ac. Anal vein arising approximately at level of Ac. Labium cleft to apical quarter - third. Claw hooks present. N o vulvar spines. 1913 Pseudagriinae T I L L Y A R D , Proc. Linn. Soc. N.S. Wales, 3 7 : 472 (emendation). - 1957 Pseudagriinae- FRÄSER, R. Zool. N . S . Wales, 1 9 5 7 : 48. Type genus: Pseudagrion SELYS, 1876 (with genera: Pseudagrion, Caliagrion, Ceriagrion, Xanthagrion, Megalagrion, Anisagrion, Erythagrion, Telagrion, Leptagrion, Argiagrion). - 1957 Amphicneminae FRÄSER, R. Zool. Soc. N.S. Wales, 1 9 5 7 : 4 8 . Type-genus: Amphicnemis SELYS, 1863 (with genera: Amphicnemis, Archibasis, Teinobasis, Papuagrion, Plagulibasis, Pericnemis, Nesobasis, Leptobasis, Antiagrion).

Genus Aeolagrion WILLIAMSON 1906 Hylaeagrion FÖRSTER, Insekten-Börse, 2 3 : 15. Type-species: Hylaeagrion argenteolineatum FÖRSTER, 1906 (monotypy). - 1 9 1 7 Aeolagrion WILLIAMSON, Ent. News, 2 8 : 2 4 2 . Type-species: Agrion dorsale BURMEISTER, 1839 (original designation). - 1 9 8 4 Hylaeagrion - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 78 (as syn. with Aeolagrion WILLIAMSON, 1917).

dorsale (BURMEISTER) 1839 Agrion dorsale BURMEISTER, Handb. Ent., 2 819. Type male: Sommer's Collection; type-locality: Brazil. 1876 Agrion (Leptagrion) dorsale - SELYS, Synopsis des Agrionines, Add. 5: 271.

291 1890 Leptagrion dorsale - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1898 Leptagrion dorsale - CALVERT, Trans. Am. ent. Soc., 25: 37. 1906 Hylaeagrion argenteolineatum FÖRSTER, Insekten-Börse, 23:16 (as generotype of Hylaeagrion FÖRSTER, 1906). Type male: Förster's Collection; type-locality: Brazil. 1917 Leptagrion dorsale - SJÖSTEDT, Ark. Zool. Uppsala, 11 (15): 15 (Amazonas). 1 9 1 7 Aeolagrion dorsale - WILLIAMSON, Ent. News, 28: 2 4 2 (as generotype of Aeolagrion W I L L I A M SON, 1 9 1 7 ) .

1917 Hylaeagrion argenteolineatum - WILLIAMSON, Ent. News, 28: 242 [proposed synonymy with dorsale (BURMEISTER, 1839)].

Distribution: Brazil. demerarum

WILLIAMSON

1917 Aeolagrion demerarum WILLIAMSON, Ent. News, 28: 244. Type male: Williamson's Collection; type-locality: Georgetown.

Distribution: Guyana. flammeum

(SELYS)

1876 Agrion (Leptagrion) flammeum (BATES MS) SELYS, Synopsis des Agrionines, Add. 5: 276. Type male: Selys's Collection; type-locality: Amazonas. 1890 Leptagrion flammeum - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1917 Leptagrion flammeum - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 13 (Manaos). 1946 Aeolagrion flammeum - FRASER, Trans. R. ent. Soc. London; 97: 458 (Peru).

Distribution: Peru, Brazil. foliaceum

(SJÖSTEDT)

1917 Leptagrion foliaceum SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 13. Type male: Naturhist. Riksmus., Stockholm; type-locality: Amazonas. 1984 Aeolagrion foliaceum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 78 (S. America).

Distribution: Brazil. fulvum

NEEDHAM

1933 Aeolagrion fulvum NEEDHAM, Amer. Mus. Novit., 664: 5. Type male: Needham's Collection; type-locality: Venezuela.

Distribution: Venezuela. inalatum

CALVERT

1961 Aeolagrion inalatum CALVERT, Acad. nat. Sei. Philad., 113: 8. Type female: Acad. nat. Sei. Philad.; type-locality: Peru.

Distribution: Peru. inca

(SELYS)

1876 Agrion (Leptagrion) inca SELYS, Synopsis des Agrionines, Add. 5: 272. Type male: Selys's Collection; type-locality: Peru. 1890 Leptagrion inca - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1917 Leptagrion inca - WILLIAMSON, Ent. News, 28: 247. 1943 Leptagrion inca - SCHMIDT, Beitr. F a u n a Perus, 2: 240 (Peru). 1958 Aeolagrion inca - RACENIS, Acta biol. venezuel., 2: 474.

Distribution: Peru.

292

Genus Amphicnemis SELYS 1863 Amphicnemis (Amphicnemis) SELYS, Synopsis des Agrionines, Add. 4: 6. Type-species: Amphicnemis wallacei SELYS, 1 8 6 3 (monotypy). - 1 8 7 7 Telebasis (Amphicnemis) - SELYS, Bull. Acad. r. Belg., (2) 43: 126 (revision). - 1890 Amphicnemis - KIRBY, Syn. Cat. Neur.-Odon., London: 157. - 1939 Amphicnemis - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 362 (key to Malaysian species). - 1954 Amphicnemis - LIEFTINCK, Treubia, 22 Suppl.: 65.

amabilis

LIEFTINCK

1940 Amphicnemis amabilis LIEFTINCK, Treubia, 17: 368. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Amphicnemis amabilis - LIEFTINCK; Treubia, 22 Suppl.: 64 (Borneo).

Distribution: Borneo. annae

LIEFTINCK

1940 Amphicnemis annae LIEFTINCK, Treubia, 17: 370. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Amphicnemis annae - LIEFTINCK; Treubia, 22 Suppl.: 65 (Borneo).

Distribution: Borneo. bicolor (MARTIN) 1898 Teinobasis bicolor MARTIN, Ann. Soc. ent. France, 66: 593. Type male: Martin's Collection; type-locality: Banguey Island. 1940 Amphicnemis bicolor - LIEFTINCK, Treubia, 17: 368. 1954 Amphicnemis bicolor - LIEFTINCK, Treubia, 22 Suppl.: 65 (Borneo).

Distribution: Borneo. billitonis

LIEFTINCK

1940 Amphicnemis billitonis LIEFTINCK, Treubia, 17: 374. Type male: Buitenzorg Museum; typelocality: Billiton Island. 1954 Amphicnemis billitonis - LIEFTINCK, Treubia, 22 Suppl. 65.

Distribution: Billiton Island. bonita (NEEDHAM & GYGER) 1939 Pericnemis bonita NEEDHAM & G Y G E R Philipp. J . Sei., 70 (3): 292. Type male: not given; type-locality: Luzon. 1940 Amphicnemis bonita - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands. cantuga (NEEDHAM & GYGER) Pericnemis cantuga NEEDHAM & G Y G E R , Philipp. J . Sei., type-locality: M i n d a n a o . 1940 Amphicnemis cantuga - LIEFTINCK, Treubia, 17: 362. 1939

7 0 (3): 2 9 6 .

Type male: not given;

Distribution: Philippine Islands. circularis

LIEFTINCK

1974 Amphicnemis circularis LIEFTINCK, Steenstrupia, 3 (12): 134. Type male: Rijksmus. Natuurl. Hist., Leiden, type-locality: Tawi Tawi.

Distribution: Pacific Ocean Islands.

293 dactylostyla LIEFTINCK 1953 Amphicnemis dactylostyla LIEFTINCK, Treubia, 22: 386. Type male: Rijksmus. Natuurl. Hist., Leiden, type-locality: Borneo. 1954 Amphicnemis dactylostyla - LIEFTINCK, Treubia, 22 Suppl.: 65.

Distribution: Borneo. dentifer (NEEDHAM & GYGER) 1939 Teinobasis dentifer NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 300. Type female: not given; type-locality: Mindanao. 1940 Amphicnemis dentifer - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands. ecornuta SELYS 1889 Amphicnemis ecornuta SELYS, Ann. Mus. civ. Stor. nat. Genova, 27: 482. Type male: Mus. civ. Stor. nat. Genova; type-locality: Sumatra. 1898 Amphicnemis ecornuta - KRÜGER; Stettiner ent. Ztg. 59: 123 (Sumatra). 1913 Amphicnemis ecornuta - LAIDLAW, Proc. Zool. Soc. London, 1913: 74. 1940 Amphicnemis ecornuta - LIEFTINCK, Treubia, 17: 363. 1954 Amphicnemis ecornuta - LIEFTINCK, Treubia, 22 Suppl.: 65 (Sumatra).

Distribution: Sumatra. erminea LIEFTINCK 1953 Amphicnemis erminea LIEFTINCK, Treubia, 22: 247. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Amphicnemis erminea - LIEFTINCK, Treubia, 22 Suppl.: 66 (Borneo).

Distribution: Borneo, flavicornis

(NEEDHAM & GYGER )

1939 Pericnemis flavicornis NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 294. Type male: not given; type-locality: Luzon. 1940 Amphicnemis flavicornis - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands. furcata BRAUER 1868 Amphicnemis furcata BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 543. Type male: Semper's Collection; type-locality: Philippines. 1877 Telebasis (Amphicnemis) furcata - SELYS, Bull. Acad. r. Belg., (2) 43: 127 (redescription). 1939 Amphicnemis furcata - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 298. 1957 Amphicnemis furcata - LIEFTINCK, Zool. Meded., Leiden, 35: 165.

Distribution: Philippine Islands. glauca BRAUER 1868 Amphicnemis glauca BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 542. Type female: Semper's Collection; type-locality: Luzon. 1877 Telebasis (Telebasis) glauca - SELYS, Bull. Acad r. Belg., (2), 43: 121 (Luzon). 1890 Teinobasis glauca - KIRBY, Syn. Cat. Neur.-Odon, London: 157. 1939 Pericnemis glauca - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 297. 1940 Amphicnemis glauca - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands.

294 gracilis KRÜGER 1898 Amphicnemis type-locality: 1 9 3 5 Amphicnemis 1940 Amphicnemis 1954 Amphicnemis

gracilis KRÜGER, Stettiner ent. Ztg, 59: 121. Type male: Mus. Naturk., Berlin; Sumatra. gracilis - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 1 0 . gracilis - LIEFTINCK, Treubia, 17: 365. gracilis - LIEFTINCK, Treubia, 22 Suppl.: 66 (Sumatra).

Distribution: Sumatra. incallida (NEEDHAM & GYGER) 1939 Pericnemis incallida NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 291. Type male: not given; type-locality: Luzon. 1940 Amphicnemis incallida - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands. kuiperi LIEFTINCK 1937 Amphicnemis kuiperi LIEFTINCK, Treubia, 16 (1): 99. Type male: Buitenzorg Museum; typelocality: Billiton. 1940 Amphicnemis kuiperi - LIEFTINCK, Treubia, 17: 363. 1954 Amphicnemis kuiperi - LIEFTINCK, Treubia, 22 Suppl.: 66.

Distribution: Billiton Island. lestoides BRAUER 1868 Amphicnemis lestoides BRAUER, Verh. Zool.-Bot Ges. Wien, 8: 541. Type male: Semper's Collection; type-locality: Philippines. 1877 Telebasis (Amphicnemis) lestoides - SELYS, Bull. Acad. r. Belg. (2) 43: 130 (redescription) (Mindanao). 1939 Pericnemis lestoides - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 297 (redescription). 1940 Amphicnemis lestoides - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands. madelenae LAIDLAW 1913 Amphicnemis madelenae LAIDLAW, Proc. Zool. Soc. London, 1913: 71. Type male: Laidlaw's Collection; type-locality: Borneo. 1940 Amphicnemis madelenae - LIEFTINCK, Treubia, 17: 376 (redescription). 1954 Amphicnemis madelenae - LIEFTINCK, Treubia, 22 Suppl.: 66 (Borneo).

Distribution: Borneo. mariae LIEFTINCK 1940 Amphicnemis mariae LIEFTINCK, Treubia, 17: 374. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Amphicnemis mariae - LIEFTINCK, Treubia, 22 Suppl.: 66 (Borneo).

Distribution: Borneo. martini Ris 1911 Amphicnemis type-locality: 1913 Amphicnemis 1940 Amphicnemis 1954 Amphicnemis

martini Ris, Ann. Soc. ent. Belg., 55: 237. Type male: Mus. Naturk. Berlin; Borneo. martini - LAIDLAW, Proc. Zool. Soc. London, 1913: 72. martini - LIEFTINCK, Treubia, 17: 362. mariae - LIEFTINCK, Treubia, 22 Suppl.: 67 (Borneo).

Distribution: Borneo.

295 mcgregori

(NEEDHAM & G Y G E R )

1939 Pericnemis mcgregori NEEDHAM & GYGER, Philipp. J. Sei, 70 (3): 293. Type male: not given; type-locality: Luzon. 1940 Amphicnemis mcgregori - LIEFTINCK, Treubia, 17: 362.

Distribution: Philippine Islands. pandanicola

LIEFTINCK

1953 Amphicnemis pandanicola LIEFTINCK, Treubia, 22: 388. Type male: Buitenzorg Museum; typelocality: Borneo. 1954 Amphicnemis

pandanicola

- LIEFTINCK, Treubia, 22 Suppl.: 67 (Borneo).

Distribution: Borneo. platystyla

LIEFTINCK

1953 Amphicnemis platystyla LIEFTINCK, Treubia, 22: 246. Type male: Buitenzorg Museum; typelocality: Borneo. 1954 Amphicnemis platystyla - LIEFTINCK, Treubia, 22: Suppl.: 67 (Borneo).

Distribution: Borneo. remiger

LAIDLAW

1912 Amphicnemis remiger lection; type-locality: 1913 Amphicnemis remiger 1954 Amphicnemis remiger

LAIDLAW, J. Str. Br. R. Asiatic Soc., 63: 96. Type male: Laidlaw's ColBorneo. - LAIDLAW, Proc. Zool. Soc. London: 1913: 77. - LIEFTINCK, Treubia, 22 Suppl.: 67 (Borneo).

Distribution: Borneo. smedleyi

LAIDLAW

1926 Amphicnemis lousiae smedleyi LAIDLAW, J. Malay. Br. R. Asiatic Soc., 4: 219. Type male: Laidlaw's Collection; type-locality: Mentawei Island. 1940 Amphicnemis smedleyi - LIEFTINCK, Treubia, 17: 364. 1954 Amphicnemis smedleyi - LIEFTINCK, Treubia, 22 Suppl.: 67.

Distribution: Mentawei Island. wallacei

SELYS

1863 Amphicnemis (Amphicnemis) wallacii SELYS, Synopsis des Agrionines, A d d . 4: 9 (as generotype of Amphicnemis SELYS, 1863). Type male: Selys's Collection; type-locality: Borneo. 1890 Amphicnemis wallacei - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1913 Amphicnemis lousiae LAIDLAW, Proc. Zool. Soc. London, 1913: 71. Type male: Laidlaw's Collection; type-locality: Borneo. 1940 Amphicnemis lousiae - LIEFTINCK, Treubia, 17: 371 (proposed synonymy with wallacei SELYS, 1863).

1954 Amphicnemis

wallacii - LIEFTINCK, Treubia, 22 Suppl.: 67 (Borneo).

Distribution: Borneo.

Genus Anisagrion

SELYS

1876 Agrion (Anisagrion) SELYS, Synopsis des Agrionines, Add. 5: 242. Type-species: Anisagrion allopterum SELYS, 1876 (original designation). - 1890 Anisagrion - KIRBY, Syn. Cat. Neur.Odon., London: 154. - 1960 Anisagrion - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 56 (key to species).

296 allopterum

SELYS

1876 Agrion (Anisagrion) allopterum SELYS, Synopsis des Agrionines, Add. 5: 243. (as generotype of Anisagrion SELYS, 1876). Type male: Selys's Collection; type-locality: Guatemala. 1876 Agrion (Anisagrion) allopterum race rubicaudum SELYS, Synopis des Agrionines, Add. 5: 244. Male from Mexico. 1890 Anisagrion allopterum - KIRBY, Syn. Cat. Neur.-Odon., London: 154 (Mexico, Guatemala). 1890 Anisagrion rubicaudum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1916 Anisagrion allopterum - Ris, Arch. Naturg., (A) 82: 117 (Costa Rica).

Distribution: Central America. inornatum

(SELYS)

1876 Agrion (Leptagrion) inornatum SELYS, Synopsis des Agrionines, Add. 5: 273. Type female: Mus. Tierk., Dresden; type-locality: unknown. 1890 Leptagrion inornatum - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1953 Anisagrion meridionale RACENIS, Opusc. ent., 18:181. Type: not given; type-locality: Venezuela. 1984 Anisagrion meridionale - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 79 [(proposed synonymy with inornatum (SELYS, 1876)].

Distribution: Venezuela. kennedyi

LEONARD

1937 Anisagrion kennedyi LEONARD, OCC. Pap. Mus. Zool. Univ. Mich., 354: 2. Type male: Zool. Univ. Michigan; type-locality: Panama.

Distribution: Panama. truncatipennis

CALVERT

1902 Anisagrion truncatipennis CALVERT, Biol. Centr.-Amer. Neur.: 106. Type male: Calvert's Collection; type-locality: Guatemala.

Distribution: Guatemala.

Genus Apanisagrion

KENNEDY

1902 Apanisagrion KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type-species: Agrion (Nehalennia) lais SELYS, 1876 (original designation). 1960 Apanisagrion - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 56.

lais

(SELYS)

1876 Agrion (Nehalennia) lais SELYS, Synopsis des Agrionines, Add. 5: 282. Type male: Naturhist. Mus. Wien; type-locality: Mexico. 1890 Nehalennia lais - KIRBY, Syn. Cat. Neur.-Odon., London: 147. 1902 Anisagrion lais - CALVERT, Biol. Centr.-Amer., London: 106 (redescription) (Honduras, Mexico). 1920 Apanisagrion lais - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Apanisagrion KENNEDY, 1 9 2 0 ) .

Distribution: Central America.

Genus Archibasis

KIRBY

1877 Telebasis (Stenobasis) SELYS, Bull. Acad. r. Belg. (2): 43:107. Type-species: Stenobasis oscillans SELYS, 1877 (original designation). - 1890 Archibasis KIRBY, Syn. Cat. Neur.-Odon., London: 156 ( a n e w n a m e f o r Stenobasis

SELYS, 1 8 7 7 , a j u n i o r p r i m a r y h o m o n y m y o f Stenobasis

AGAS-

297 siz, 1846 in Diptera). - 1930 Archibasis - Ris, Ark. Zool. Uppsala, 21A (31): 15. - 1949 Archlbasis LIEFTINCK, Nova Guinea (N.S.) 5 : 1 8 4 (key to species). - 1 9 5 4 Archibasis - LIEFTINCK, Treubia, 22 Suppl.: 60 (generotype: Stenobasis melanocyana SELYS, 1 8 7 7 ) .

crucigera

LIEFTINCK

1949 Archibasis crucigera LIEFTINCK, N o v a Guinea (N.S.) 5: 192. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. incisura

LIEFTINCK

1949 Archibasis incisura LIEFTINCK, N o v a Guinea (N.S.) 5: 186. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1954 Archibasis incisura - LIEFTINCK, Treubia, 22 Suppl.: 60 (Borneo).

Distribution: Borneo. melanocyana

(SELYS)

1877 Telebasis (Stenobasis) melanocyana SELYS, (nec Ris, 1930), Bull. Acad. r. Belg. (2) 43: 110. Type male: Selys's Collection; type-locality: Malacca. 1890 Archibasis melanocyana - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1927 Archibasis melanocyana - Ris, Zool. Meded., Leiden, 10: (1): 25 (Sumatra). 1934 Archibasis melanocyana - LIEFTINCK, Treubia, 14 (4): 397 (Java). 1942 Archibasis melanocyana - FRÄSER, Proc. R. ent. Soc. London, (B) 11: 97. 1949 Archibasis melanocyana - LIEFTINCK, N o v a Guinea (N.S.) 5: 184. 1954 Archibasis melanocyana - LIEFTINCK, Treubia, 22 Suppl.: 60 (Sumatra, Borneo, Malaya) (as generotype of Archibasis KIRBY, 1890).

Distribution: Malaysia, Borneo, Sumatra, Java. mimetes

(TILLYARD)

1913 Stenobasis mimetes TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 472. Type male: Tillyard's Colletion; type-locality: Queensland. 1930 Archibasis mimetes - Ris, Ark. Zool. Uppsala, 21A (31): 17 (redescription). 1931 Archibasis mimetes - FRÄSER, Ree. Indian Mus., 33: 465.

Distribution: Australia. oscillans

(SELYS)

1877 Telebasis (Stenobasis) oscillans SELYS, Bull. Acad. r. Belg. (2): 43:108. Type male: Termeszett. Muz., Budapest; type-locality: Siam. 1890 Archibasis oscillans - KIRBY, Syn. Cat. Neur.-Odon., London: (as generotype of Archibasis KIRBY, 1890).

1914 Archibasis oscillans - LAIDLAW, Ree. Indian Mus., 8 343. 1919 Archibasis oscillans - LAIDLAW, Ree. Indian Mus., 16: 194 (India). 1924 Pseudagrion praeclavum - FRÄSER, Ree. Indian Mus., 26: 493. Type male: British Museum (Nat.Hist.), London; type-locality: West India. 1931 Archibasis mimetes praeclavum - FRÄSER, Ree. Indian. Mus., 33: 465. 1949 Pseudagrion praeclavum - LIEFTINCK, Nova Guinea (N.S.), 5: 182 [proposed synonymy with oscillans (SELYS, 1 8 7 7 ) ] . 1954 Archibasis oscillans - LIEFTINCK, Treubia, 22 Suppl.: 60 (Siam, Sumatra, Java).

Distribution: Oriental region.

298 tenella LIEFTINCK 1949 Archibasis tenella LIEFTINCK, N o v a Guinea (N.S.) 5:186. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1954 Archibasis tenella - LIEFTINCK, Treubia, 22 Suppl.: 61 (Borneo).

Distribution: Borneo. viola

LIEFTINCK

1927 Archibasis melanocyana Ris (nec SELYS, 1877), Zool. Meded., Leiden, 10: 25. Type male: unknown; type-locality: Sumatra. 1948 Archibasis viola LIEFTINCK, (nom.nud.), Treubia, 19: 285. 1949 Archibasis viola LIEFTINCK, N o v a Guinea (N.S.) 5:185. (new name for Archibasis melanocyana Ris, 1927, a junior secondary homonymy of Archibasis melanocyana SELYS, 1877). 1954 Archibasis viola-LIEFTINCK, Treubia, 22 Suppl.: 61 (Malaya, Engano, Sumatra, Java, Borneo).

Distribution: Oriental region. Genus Argiagrion SELYS 1876 Agrion (Argiagrion) SELYS, Synopsis des Agrionines, Add. 5: 240. Type-species: Argiagrion leonium SELYS, 1 8 7 6 (original designation). - 1 8 9 0 Argiagrion - K I R B Y , Syn. Cat. Neur.-Odon., London: 154. - 1962 Argiagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 121.

leoninum

SELYS

1876 Agrion (Argiagrion) leoninum SELYS, Synopsis des Agrionines, Add. 5: 240 (as generotype of Argiagrion SELYS, 1876). Type female: McLachlan's Collection; type-locality: Sierra Leone. 1890 Argiagrion leoninum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1962 Argiagrion leoninum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 121.

Distribution: Sierra Leone. Genus Caliagrion

TILLYARD

1913 Caliagrion TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 468. Type-species: Pseudagrion hursti M A R T I N , 1 9 0 1 (original designation).

billing-

billinghursti (MARTIN) 1901 Pseudagrion billinghursti MARTIN, Mem. Soc. Zool. France, 14: 246. Type male: Martin's Collection; type-locality: Victoria. 1913 Caliagrion billinghursti - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 468 (as generotype of Caliagrion

TILLYARD, 1 9 1 3 ) .

Distribution: Australia. Genus Ceriagrion SELYS 1876 Agrion (Ceriagrion) SELYS, Synopsis des Agrionines, Add. 5: 235. Type-species: Agrion rhosoma)

cerinorubellum

BRAUER, 1 8 6 5 ( d e s i g n a t e d b y K I R B Y , 1 8 9 0 , S y n . C a t .

(Pyr-

Neur.-Odon.,

London: 154). - 1869 Brachybasis SELYS, A n n . Soc. ent. Belg., 12: 95 (nom.nud.) - 1898 Brachybasis - KIRBY, A n n . Mag. nat. Hist., (7) 2: 245. - 1908 Brachybasis - Ris, Jenaische Denkschr. med. nat. Gesch., 13: 303 (type-species: Agrion rhomboidale BEAUVOIS, 1805). 1905 Ceriagrion - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 831 (key to species). 1919 Ceriagrion - LAIDLAW, Ree. Indian Mus., 16: 187 (key to species, revision). - 1920 P a laeobasis K E N N E D Y , Ohio J . Sei. Columbus, 2 1 : 88. Type-species: Libellula tenella VILLIERS, 1789 (original designation). - 1921 Ceriagrion - Ris, Annls. S. Afr. Mus., 18: 314 (key to African species). - 1 9 3 2 Ceriagrion - LIEFTINCK, N o v a Guinea, 1 5 ( 5 ) : 1 0 8 . - 1 9 3 5 Palaeobasis

299 C O W L E Y , Entomologist, 6 8 : 1 5 4 (proposed synonymy with Ceriagrion SELYS, 1 8 7 6 ) . 1941 Ceriagrion - FRÄSER, Proc. R. Ent. Soc. London, (B) 10: 62 (key to African species). - 1951 Ceriagrion - SCHMIDT, Arq. Mus. Bocage, 20: 157 (key to African and Malagasy species). - 1953 Ceriagrion - SCHMIDT, Mitt, münch. ent. Ges., 43: 5 (key to W. Palaearctic species). - 1954 Ceriagrion - LIEFTINCK, Treubia, 22 Suppl.: 54. - 1956 Ceriagrion - FRASER, R. Ent. Soc. London; 1 (10): 30. - 1 9 6 2 Ceriagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 118. - 1962 Brachybasis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 118 (proposed synonymy with Ceriagrion SELYS, 1876). - 1962 Nepocnemis FRASER (nom.nud.) - loc. cit. PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120 (as Nepocnemis platystyla FRASER (ined.). Type in C.B. (Tervuren Museum), as syn. with Ceriagrion SELYS, 1876. -

aeruginosum

(BRAUER)

1869 Agrion aeruginosum BRAUER, Verh. Zool.-Bot. Ges. Wien, 19: 13. Type male: Zool. Inst. Mus. Univ., H a m b u r g ; type-locality: R o c k h a m p t o n . 1890 Micronympha aeruginosum - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1891 Ceriagrion ceromandelinanum race erubescens SELYS, Ann. Mus. civ. Stor. Genova, 10 (30): 516. Male from Queensland. 1901 Ischnura aeruginosum - MARTIN, Mem. Soc. zool. France, 14: 246 (Queensland). 1927 Ceriagrion erubescens - Ris, Zool. Meded., Leiden, 10 (1): 25 (Sumatra). 1931 Ceriagrion erubescens - SCHMIDT, Konowia, 10 (3): 183 (China: Shanghai). 1933 Ceriagrion erubescens - SJÖSTEDT, Ark. Zool. Uppsala, 25A (5): 21 (Szechwan). 1934 Ceriagrion erubescens - SCHMIDT, Arch. Hydrob. Suppl., 13: 353 (Java). 1949 Ceriagrion ? erubescens - LIEFTINCK, Treubia, 20 (2): 346 (Solomon Islands). 1951 Ceriagrion erubescens - LIEFTINCK, Treubia, 21 (1): 196 (Australo-Papuan region). 1954 Ceriagrion erubescens - LIEFTINCK, Treubia, 22 Suppl.: 55 (proposed synonymy with lactericum LIEFTINCK, 1 9 5 1 ) .

1959 Ceriagrion aeruginosum - LIEFTINCK, Ent. Mitt. Zool. Staatsinst. Zool. Mus. Hamburg, 21: 2. 1 9 5 9 Ceriagrion ceromandelianum race erubescens - LIEFTINCK Ent. Mitt. Zool. Staatsinst. Zool. Mus. Hamburg, 21: 2 [proposed synonymy with aeruginosum ( B R A U E R , 1 8 6 9 ) ] .

Distribution: Indo-austral region. annulatum

FRASER

1955 Ceriagrion annulatum FRASER, Revue Zool.-Bot. afr., 51: 35. Type male: Mus. r. Afr. centr. Tervuren; type-locality: Belgian Congo. 1962 Ceriagrion annulatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 119.

Distribution: Congo Republic. annulosum

LIEFTINCK

1934 Ceriagrion annulosum LIEFTINCK, Stylops, 3: 11. Type male: Fruhstorfer's Collection; typelocality: Java. 1934 Ceriagrion annulosum - LIEFTINCK, Treubia, 14 (4): 394 (Java). 1954 Ceriagrion annulosum - LIEFTINCK, Treubia, 22 Suppl.: 54 (Sumatra, Java).

Distribution: Sumatra, Java. auranticum

FRASER

1922 Ceriagrion auranticum FRASER, J. nat. Hist. Soc. Siam, 4: 236. Type male: British Museum (Nat.Hist.), London; type-locality: Bangkok.

Distribution: Thailand.

300 auritum

FRÄSER

1951 Ceriagrion auritum FRÄSER, Mem. Inst. sei. Madagascar, 6A: 412. Type male: Inst. Rech. Sei. Madagascar; type-locality: Madagascar. 1962 Ceriagrion auritum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 119.

Distribution: Madagascar. azureum

(SELYS)

1892 Pseudagrion azureum SELYS, Ann. Mus. civ. Stor. nat. Genova, (2) 10: 513. Type male: Mus. civ. Stor. nat., Genova, type-locality: Burma. 1916 Pseudagrion azureum - LAIDLAW, Ree. Indian Mus., 12: 25 (Burma). 1 9 8 4 Ceriagrion azureum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 0 (India, China).

Distribution: India, Burma, China, Laos. bakeri

FRASER

1941 Ceriagrion bakeri FRÄSER, Proc. R. ent. Soc. London, (B) 10: 62. Type male: British Museum (Nat.Hist.), London; type-locality: Uganda. 1951 Ceriagrion villiersi FRASER, Bull. Inst, franc. Afr. noire, 13: 1086. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Togo. 1962 Ceriagrion bakeri - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 119 (Uganda, Nigeria). 1962 Ceriagrion villiersi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 121. 1 9 8 4 Ceriagrion villiersi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 8 1 (proposed synonymy with bakeri FRASER, 1 9 4 1 ) .

Distribution: Central Africa. batjanum 1984

ASAHINA

Ceriagrion batjanum Utrecht, 1: 80.

ASAHINA, 1 9 6 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Batjan. bellona bellona

LAIDLAW

1915 Ceriagrion bellona locality: Borneo. 1920 Ceriagrion bellona 1934 Ceriagrion bellona 1954 Ceriagrion bellona 1984 Ceriagrion bellona

LAIDLAW, J. Sarawak Mus., 2: 274. Type male: Sarawak Museum; type- LAIDLAW, Proc. Zool. Soc. London, 1920: 335 (Borneo). - LAIDLAW, J. Fed. Mai. States Mus., 17: 551. - LIEFTINCK, Treubia, 22 Suppl.: 54 (Borneo). bellona - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 80.

Distribution: Borneo. bellona hoogerwerfi

LIEFTINCK

1940 Ceriagrion hoogerwerfi LIEFTINCK, Treubia, 17: 260. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Sumatra. 1954 Ceriagrion hoogerwerfi - LIEFTINCK, Treubia, 22 Suppl.: 55 (Sumatra). 1984 Ceriagrion bellona hoogerwerfi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 80.

Distribution: Sumatra. calamineum

LIEFTINCK

1951 Ceriagrion calamineum LIEFTINCK, Treubia, 21 (1): 189. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Malay Peninsula.

301 1954 Ceriagrion calamineum - LIEFTINCK, Treubia, 22 Suppl.: 54 (Penang, Sumatra, Java).

Distribution: Sumatra, Java, Malay Peninsula, and Cambodia. cerinorubellum

(BRAUER)

1865 Agrion (Pyrrhosoma) cerino-rubellum BRAUER, Verh. Zool.-Bot Ges. Wien, 15: 511. Type male: not given; type-locality: Ceylon. 1876 Agrion (Ceriagrion) cerinorubellum - SELYS, Synopsis des Agrionines, Add. 5: 236 (Malacca, Borneo, Ceylon). 1890 Ceriagrion cerinorubellum - KIRBY, Syn. Cat. Neur.-Odon., London: 154 (as generotype of Ceriagrion

SELYS, 1 8 7 6 ) .

1891 Ceriagrion cerinorubellum - KARSCH, Ent. Nachr., 17 (16): 243 (Sumatra). 1916 Ceriagrion cerinorubellum - LAIDLAW, Ree. Indian Mus., 12: 133 (Ceylon, India, Burma, Malaysia, Sumatra, Borneo). 1924 Ceriagrion cerinorubellum - FRÄSER, Ree. Indian Mus., 26: 492 (W. India). 1927 Ceriagrion cerinorubellum - Ris, Zool. Meded., Leiden, 10 (1): 25 (Sumatra). 1934 Ceriagrion cerinorubellum - LIEFTINCK, Treubia, 14 (4): 394 (Java). 1935 Ceriagrion cerinorubellum - LIEFTINCK, Misc. Zool. Sum., 92-93: 20 (Sumatra). 1948 Ceriagrion cerinorubellum - LIEFTINCK, Treubia, 19: 285. 1954 Ceriagrion cerinorubellum - LIEFTINCK, Treubia, 22 Suppl.: 55 (Siam, Malaya, Mentawei Island, Sumatra, Java, Borneo).

Distribution: Oriental region. chaoi

SCHMIDT

1964 Ceriagrion chaoi SCHMIDT, Ent. Tidschr., 85: 152. Type male: Schmidt's Collection; type-locality: Burma.

Distribution: Burma. citrinum

CAMPION

1914 Ceriagrion citrinum CAMPION, Ann. Mag. nat. Hist., (8) 14: 278. Type male: British Museum (Nat.Hist.), London; type-locality: S. Nigeria. 1941 Ceriagrion citrinum - ST. QUENTIN, Annls. Naturh. Mus. Wien, 52: 108 (Belg. Congo). 1962 Ceriagrion citrinum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 119 (Nigeria, Belgian Congo).

Distribution: West Africa, coeruleum

LAIDLAW

1919 Ceriagrion coeruleum LAIDLAW, Ree. Indian Mus., 16: 189. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Himalaya.

Distribution: Himalaya, coraliinum

CAMPION

1914 Ceriagrion coraliinum CAMPION, Ann. Mag. nat. Hist., (8) 14: 279. Type male: British Museum (Nat.Hist.), London; type-locality: Sierra Leone. 1941 Ceriagrion bidentatum FRÄSER, Proc. R. ent. Soc. London, (B) 10: 64. Type male: British Museum (Nat.Hist.), London; type-locality: Uganda. 1952 Ceriagrion coraliinum - LONGFIELD, Proc. R. ent. Soc. London; (B) 21: 43 (redescription). 1955 Ceriagrion coraliinum - FRASER, Revue Zool.-Bot. afr., 51: 37 (Belgian Congo). 1962 Ceriagrion coraliinum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 119 (Angola, Belgian Congo, Cameroon, Nigeria, Sierra Leone). 1962 Ceriagrion bidentatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 119.

302 1984 Ceriagrion bidentatum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 80 (proposed synonymy with corallinum C A M P I O N , 1 9 1 4 ) .

Distribution: West Africa, coromandelianum

(FABRICIUS) 1798 Agrion coromandelianum FABRICIUS, Ent. Syst. Suppl.: 287. Type male: Univ. zool. Mus., Kobenhavn, type-locality: India. 1842 Agrion cerinum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 279. Type male: Guerin's Collection; type-locality: Senegal. 1876 Agrion (Ceriagrion) coromandelianum - SELYS, Synopsis des Agrionines, Add. 5: 238. 1876 Agrion cerinum - SELYS, Synopsis des Agrionines, Add. 5: 238 [proposed synonymy with coromandelianum (FABRICIUS 1 7 9 8 ) ] . 1890 Ceriagrion coromandelianum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1916 Ceriagrion coromandelianum - LAIDLAW, Ree. Indian Mus., 12: 132 (Ceylon, India, Burma, Indo-China, Sunda, Celebes). 1951 Ceriagrion coromandelianum - LIEFTINCK, Treubia, 21 (1): 187 (redescription) (India, Ceylon, Assam). 1955 Ceriagrion coromandelianum - BAIJAL & A G A R W A L , Agra Univ. J. Res., 4 (2): 470 (India, Ceylon, Burma, Malaysia, Indo-China, South China).

Distribution: Indo-Austral region. fallax fallax Ris 1914 1919 1955 1956 1984

Ceriagrion fallax Ris, Ent. Mitt., Berlin, 3:47. Type male: Ris's Collection; type-locality: China. Ceriagrion fallax - LAIDLAW, Ree. Indian Mus., 16: 191 (Burma). Ceriagrion fallax - SANTOCK, Agra Univ. J. Res. 4 Suppl.: 747 (Himalayas). Ceriagrion fallax - ASAHINA, Ent. Medd., 27 (4-5): 219 (South China). Ceriagrion fallax fallax - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 80.

Distribution: China, Burma, India. fallax cerinomelas

LIEFTINCK

1927 Ceriagrion cerinomelas LIEFTINCK, Ent. Tijdschr., 70: 88. Type male: R. Mus., Amsterdam; type-locality: North India. 1928 Ceriagrion cerinomelas - MORTON, Trans, ent. Soc. London, 76: 112 (Yunnan). 1933 Ceriagrion cerinomelas - FRASER, Fauna brit. India, Odon., 1: 321 (proposed synonymy with fallax Ris, 1914). 1984 Ceriagrion fallax cerinomelas - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 80.

Distribution: India, and South China, fallax pendleburyi

LAIDLAW 1931 Ceriagrion pendleburyi L A I D L A W , I. Fed. Mai. States Mus., 16: 198. Type male: Raffles Museum; Singapore; type-locality: Perak. 1954 Cerinomelas pendleburyi - LIEFTINCK, Treubia, 22 Suppl.: 56 (Malaysia). 1984 Ceriagrion fallax pendleburyi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 81 (India, Malaya).

Distribution: Malaysia, Thailand, Burma, and India. glabrum glabrum

(BURMEISTER)

1839 Agrion glabrum BURMEISTER, Handb. Ent., 2: 821. Type male: Halle Museum; type-locality: Cape of Good Hope. 1805 Agrion rhomboidalis BEAUVOIS, Ins. Afr. Amer.: 85 (nom.nud.). 1842 Agrion ferruginueum RAMBUR, Hist. nat. Ins. Neuropt. Paris: 280. Type male: Guerin's Collection; type-locality: Madagascar.

303 1876 Agrion (Ceriagrion) glabrum SELYS, Synopsis des Agrionines, Add. 5: 237 (Tropical Africa, Madagascar, Mauritius, Seychelles). 1876 Agrion ferrugineum - SELYS, Synopsis des Agrionines, Add. 5: 237 [proposed synonymy with glabrum (BURMEISTER, 1 8 3 9 ) ] . 1890 Ceriagrion glabrum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1898 Brachybasis rhomboidalis - KIRBY, Ann. Mag. nat. Hist., (7) 2: 245 (as generotype of Brachybasis KIRBY, 1898). 1913 Ceriagrion glabrum - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 471 (Queensland). 1928 Ceragrion (misprint) glabrum - ANDERS, Mem. Soc. R. ent. Egypt, 3: (1): 25 (Egypt). 1949 Agrion rhomboidalis - FRASER, Trans. R. ent. Soc. London, 100: 137 [proposed synonymy with glabrum (BURMEISTER), 1 8 3 9 ) ] . 1958 Ceriagrion glabrum - PINHEY, Stuttg. Beitr. Naturk. 10: 1 (Kilimandjaro).

Distribution: tropical Africa, Mauritius, Madagascar, Seychelles, ? Australia, glabrum longispinum PINHEY 1963 Ceriagrion glabrum longispinum PINHEY, Ann. Mag. Nat. Hist., (13), 6: 18. Type: unknown; type-locality: Congo.

Distribution: Congo Republic. georgfreyi

SCHMIDT

1953 Ceriagrion Georg-Freyi SCHMIDT, Mitt, miinch. ent. Ges., 43: 1. Type male: Schmidt's Collection; type-locality: Turkey.

Distribution: Turkey. hamoni FRASER 1955 Ceriagrion hamoni FRASER, Rev. franc. Ent., 22: 237. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Fr. W. Africa. 1962 Ceriagrion hamoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120.

Distribution: West Africa, ignitum CAMPION 1914 Ceriagrion ignitum CAMPION, Ann. Mag. nat. Hist., (8) 14: 281. Type male: British Museum (Nat.Hist.), London; type-locality: Gold Coast. 1962 Ceriagrion ignitum - PINHEY, Publ. cult. Comp. Diam. Angola, 59:120 (Gold Coast, Liberia). 1984 Ceriagrion ignitum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 80 (Ghana).

Distribution: West Africa. inaequale

LIEFTINCK

1932 Ceriagrion inaequale LIEFTINCK, Nova Guinea, 15 (5): 108. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. indochinense ASAHINA 1984 Ceriagrion indochinense ASAHINA, 1967 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the

World, Utrecht, 1: 80.

Distribution: Vietnam.

304 katambovae 1984

PINHEY

Ceriagrion katambovae Utrecht, 1: 81.

PINHEY, 1 9 6 1

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: : Zambia. kordofanicum Ris 1924 Ceriagrion kordofanicum Ris, in ALFKEN, Denkschr. Akad. Wiss., 99: 279. Type male: Naturhist. Mus. Wien; type-locality: Tonga. 1941 Ceriagrion kordofanicum - FRÄSER, Proc. R. ent. Soc. London, (B) 10: 66 (redescription) (Uganda). 1962 Ceriagrion kordofanicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 170 (with Ceriagrion rubicundum FRASER (ined.) from Kenya - nom.nud.).

Distribution: Sudan: Tonga, Uganda, and Zaire. latericium latericium

LIEFTINCK

1951 Ceriagrion latericium LIEFTINCK, Treubia, 21 (1): 193. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Sumatra. 1953 Ceriagrion latericium - LIEFTINCK, Verh. naturg. Ges. Basel, 64: 155. (Malaya, Sumatra, Java, Sumba). 1987 Ceriagrion latericium - HÄMÄLÄINEN, Odonatologica, 16 (2): 184 (proposed synonymy with auranticum FRASER, 1 9 2 2 ) .

Distribution: Sumatra, Malaysia, Java, Sumba. latericium ryukyuanum 1984

ASAHINA

Ceriagrion latericium ryukyuanum of the World, Utrecht, 1: 81.

ASAHINA, 1 9 6 7

- loc. cit.

D A V I E S & TOBIN,

The Dragonflies

Distribution: East Asia. lieftincki 1984

ASAHINA

Ceriagrion lieftincki Utrecht, 1: 82.

ASAHINA, 1 9 6 7

- loc. cit

D A V I E S & TOBIN,

The Dragonflies of the World,

Distribution: Philippine Islands. madagazureum

FRASER

1949 Ceriagrion madagazureum FRASER, Mem Inst. sei. Madagascar, 3Α: 26. Type male: Mus. N a tion. Hist. Nat., Paris; type-locality: Madagascar. 1962 Ceriagrion madegazureum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120.

Distribution: Madagascar. malaisei

SCHMIDT

1964 Ceriagrion malaisei SCHMIDT, Ent. Tidskr., 85: 152. Type male: Schmidt's Collection; typelocality: Burma.

Distribution: Burma, Laos, Thailand, and Vietnam. melanurum

SELYS

1876 Agrion (Ceriagrion) coromandelianum race melanurum SELYS, Synopsis des Agrionines, A d d . 5: 239. Type male: Selys's Collection; type-locality: Japan. 1890 Ceriagrion melanurum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1919 Ceriagrion melanurum - L A I D L A W , Ree. Indian Mus., 16: 191 (China, Sumatra, Japan).

305 1933 Ceriagrion melanurum - SJÖSTEDT, Ark. Zool., Uppsala, 25A (5): 20 (Szechwan).

Distribution: from Japan to South China, ? Sumatra. moorei

LONGFIELD

1952 Ceriagrion moorei LONGFIELD, Proc. R. ent. Soc. London, (B) 21: 44. Type male: British Museum (Nat.Hist.), London; type-locality: Gold Coast. 1962 Ceriagrion moorei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120.

Distribution: Ghana. mourae PINHEY 1984

Ceriagrion mourae Utrecht, 1: 81.

PINHEY, 1 9 6 6

- loc. cit

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Mozambique. nigroflavum FRASER 1933 Ceriagrion nigroflavum FRASER, Fauna brit. India, Odon., 1: 323. Type male: British Museum (Nat.Hist.), London; type-locality: Burma. 1984 Ceriagrion nigroflavum — DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 81 (India, Indochina).

Distribution: India, Vietnam, Burma, nigrolineatum

SCHMIDT

1951 Ceriagrion nigrolineatum SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 271. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Ceriagrion nigrolineatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120.

Distribution: Madagascar, nipponicum 1984

ASAHINA

Ceriagrion nipponicum World, Utrecht, 1: 81.

ASAHINA, 1 9 6 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: China, Japan. oblungulum SCHMIDT 1951 Ceriagrion oblungulum SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 268. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Ceriagrion oblungulum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120.

Distribution: Madagascar. olivaceum olivaceum

LAIDLAW

1914 Ceriagrion olivaceum LAIDLAW, Ree. Indian Mus., 8: 345. Type male: Nation. Coll. zool. Surv. India, Calcutta, type-locality: Burma. 1916 Ceriagrion olivaceum - LAIDLAW, Ree. Indian Mus., 12: 133 (Assam, Burma). 1924 Ceriagrion olivaceum - FRASER, Ree. Indian Mus., 26: 492 (West India). 1933 Ceriagrion olivaceum - SJÖSTEDT, Ark. Zool. Uppsala, 25A (5): 20 (Szechwan). 1 9 5 5 Ceriagrion olivaceum - BAIJAL & A G A R W A L , Agra Univ. J . res., 4 ( 2 ) : 4 7 0 (India, Burma). 1 9 8 4 Ceriagrion olivaceum olivaceum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1 : 8 1 .

Distribution: China, India, Burma.

306 olivaceum aurantiacum

FRÄSER

1924 Ceriagrion aurantiacum FRÄSER, Ree. Indian Mus., 26: 492. Type male: British Museum (Nat.Hist.), London; type-locality: Nilgiris. 1966 Ceriagrion olivaceum aurantiacum - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 180.

Distribution: India, pallidum

FRASER

1933 Ceriagrion pallidum FRÄSER, J. Siam Soc. nat. Hist., Suppl.: 9:118. Type male: British Museum (Nat.Hist.), London; type-locality: Laos.

Distribution: Laos.

platystigma

FRASER

1941 Ceriagrion platystigma FRASER, Proc. R. ent. Soc. London, (B) 10: 63. Type male: British Museum (Nat.Hist.), London; type-locality: U g a n d a . 1955 Ceriagrion platystigma - FRASER, Revue Zool. Bot. afr., 51: 36 (Belgian Congo). 1 9 5 9 Ceriagrionplatystigma-hoNGFiELO, Publ.c. cult. Comp. Diam. Angola, 4 5 : 2 6 (Angola, Uganda). 1962 Ceriagrion platystigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120 (Uganda, Cameroon, Belgian Congo, Rhodesia, Transvaal, S. Nigeria, Angola) + Nepocnemis platystyla FRASER (ined.) (Tervuren Museum as a form of platystigma FRASER, 1941).

Distribution: tropical Africa. praetermissum

LIEFTINCK

1929 Ceriagrion praetermissum LIEFTINCK, Tijdschr. Ent., 72: 115. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Java. 1934 Ceriagrion praetermissum - SCHMIDT, Arch. Hydrob. Suppl.: 13: 353. 1934 Ceriagrion praetermissum - LIEFTINCK, Treubia, 14 (4): 395 (Java). 1954 Ceriagrion praetermissum

- LIEFTINCK, Treubia, 22 Suppl.: 56 (Sumatra, Java).

Distribution: Java, Sumatra. rubellocerinum

FRASER

1947 Ceriagrion rubellocerinum FRASER, Trans. R. ent. Soc. London, 98:28. Type male: Mus. Nation. Hist.Nat., Paris; type-locality: Ivory Coast. 1962 Ceriagrion rubellocerinum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120 (Ivory Coast, Liberia).

Distribution: West Africa. rubiae

LAIDLAW

1916 Ceriagrion rubiae LAIDLAW, Ree. Indian Mus., 12: 132. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Conchin. 1919 Ceriagrion rubiae - LAIDLAW, Ree. Indian Mus., 16: 190 (India). 1924 Ceriagrion rubiae - FRASER, Ree. Indian Mus., 26: 491 (West India).

Distribution: India. sakejii

PINHEY

1963 Ceriagrion sakejii PINHEY, Ann. Mag. nat. Hist., (13) 5: 21. Type male: British Museum (Nat.Hist.), London; type-locality: Rhodesia.

Distribution: Rhodesia.

307 sanguinostigma

FRÄSER

1955 Ceriagrion sanguinostigma FRÄSER, Revue Zool. Bot. afr., 51: 33. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1962 Ceriagrion sanguinostigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120.

Distribution: Congo Republic. sinense 1984

ASAHINA

Ceriagrion sinense Utrecht, 1: 81.

ASAHINA, 1 9 6 7

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: East China. suave Ris 1921 Ceriagrion suave Ris, Annls. S. Afr. Mus., 18: 316. Type male: Mus. r. Afr. centr., Tervuren, type-locality: Katanga. 1945 Ceriagrion suave - LONGFIELD, Arq. Mus. Bocage, 16: 28 (Angola). 1962 Ceriagrion suave - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 120 (Rhodesia, Belgian Congo, Angola, Kenya, Nigeria).

Distribution: tropical Africa. tenellum tenellum

(VILLIERS)

1789 Libellula tenella VILLIERS, Linn. Ent., 3: 15. Type: unknown; type-locality: France. 1823 Agrion rubella VAN DER LINDEN, Opusc. Sei., 4:105. Type female: unknown; type-locality: Italy. 1850 Agrion (Pyrrhosoma) tenellum - SELYS & H Ä G E N , Revue des Odonates, Paris: 180 (France, England, Italy, Switzerland, Greece, Algeria). 1876 Agrion (Pyrrhosoma) tenellum - SELYS, Synopsis des Agrionines, Add. 5: 184 (France, Belgium, Italy, Creta, Algeria). 1890 Pyrrhosoma tenellum - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1916 Ceriagrion tenellum - Ris, Mitt. Ent. Zürich, 2: 108. 1920 Palaeobasis tenellum - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Palaeobasis KENNEDY, 1920).

1935 Ceriagrion tenellum - COWLEY, Entomologist, 68: 154. 1950 Pyrrhosoma tenellum - MOREIEA, Inst. Esp. Ent. Madrid: 82 (Spain). 1 9 5 6 Ceriagrion tenellum - FRÄSER, R . Ent. Soc. London, 1 ( 1 0 ) : 3 0 (England).

Distribution: Europa, Algeria. tenellum nielseni

SCHMIDT

1953 Ceriagrion tenellum nielseni SCHMIDT, Mitt, münch. ent. Ges., 43: 6. Type male: Schmidt's Collection; type-locality: Sicily.

Distribution: Sicily. tricrenaticeps 1984

LEGRAND

Ceriagrion tricrenaticeps World, Utrecht, 1: 81.

LEGRAND, 1 9 8 4

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Gabon. varians

(MARTIN)

1908 Aciagrion varians MARTIN, Ann. Mus., civ. Stor. nat. Genova, 43: 660. Type male: Mus., civ. Stor. nat. Genova, type-locality: Guinea. 1962 Aciagrion varians - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 133 (Portuguese Guinea, Uganda).

308 1984 Ceriagrion varians - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 81.

Distribution: West Africa. whellani

LONGFIELD

1952 Ceriagrion whellani LONGFIELD, Proc. R. ent. Soc. London (B) 21: 42. Type male: British Museum (Nat.Hist.), London; type-locality: South Rhodesia. 1955 Ceriagrion whellani - FRASER, Revue Zool. Bot. afr., 51: 37 (Belgian Congo). 1962 Ceriagrion whellani - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 121 (Rhodesia, Belgian Congo, Sierra Leone).

Distribution: tropical Africa.

Genus Hylaeonympha

RACENIS

1968 Hylaeonympha RACENIS, Publ. ocas. Mus. Ci. nat. Caracas (Zool.), 11: 5. Type-species: Hylaeonympha magoi RACENIS, 1968 (original designation).

magoi

RACENIS

1968 Hylaeonympha magoi RACENIS, Publ. ocas. Mus. Ci. nat. Caracas (Zool.), 11: 7 (as generotype of Hylaeonympha RACENIS, 1968). Type male: Caracas Museum; type-locality: Venezuela.

Genus Leptagrion SELYS 1876 Agrion (Leptagrion) SELYS, Synopsis des Agrionines, Add. 5:, 264. Type-species: Agrion macrurum BURMEISTER, 1839 (original designation) - 1890 Leptagrion - KIRBY, Syn. Cat. Neur.Odon., London: 155. - 1960 Leptagrion - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 46 (key to South American species). - 1962 Leptagrion - DIAS DOS SANTOS, Bol. Mus. nac. Rio de Janeiro (N.S.) (Zool.) 233: 5 (key to species).

acuta SANTOS 1961 Leptagrion acuta SANTOS, Rev. bras. Biol., 21: 359. Type male: Santos's Collection; typelocality: Brazil.

Distribution: Brazil. aculeatum

SANTOS

1984 Leptagrion aculeatum SANTOS, 1965 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 82.

Distribution: Brazil. andromache SELYS 1876 Agrion (Leptagrion) andromache (HÄGEN without description) SELYS, Synopsis des Agrionines, Add. 5: 268. Type male: Hagen's Collection; type-locality: Brazil. 1890 Leptagrion andromache - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. auriceps

ST. QUENTIN

I960 Leptagrion auriceps ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 47. Type male: Naturhist. Mus., Wien; type-locality: Brazil.

Distribution: Brazil.

309 autazensis SJÖSTEDT 1917 Leptagrion autazensis SJÖSTEDT, Ark. Zool. Uppsala, 11 (15): 11. Type male: Naturhist. Riksmus., Stockholm; type-locality: Amazonas.

Distribution: Brazil. beebeanum CALVERT 1948 Leptagrion beebeanum CALVERT, Zoologica, 33: 58. Type male: Calvert's Collection; type-locality: British Guiana.

Distribution: British Guyana. bocainense SANTOS 1984

Leptagrion bocainense Utrecht, 1: 82.

SANTOS, 1 9 7 9

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Brazil. capixabae SANTOS 1984

Leptagrion capixabae Utrecht, 1: 81.

SANTOS, 1 9 6 5

Distribution: Brazil. croceum (BURMEISTER) 1839 Agrion croceum BURMEISTER, Handb. Ent., 2: 819. Type male: Sommer's Collection; type-locality: Suriname. 1898 Leptagrion croceum - CALVERT, Trans. Am. ent. Soc., 25: 37.

Distribution: Suriname. dardanoi SANTOS 1984

Leptagrion dardanoi Utrecht, 1: 82.

SANTOS, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Brazil. dispar SELYS 1876 Agrion (Leptagrion) elongatum race dispar (BATES MS) SELYS, Synopsis des Agrionines, Add. 5: 271. Type male: Selys's Collection; type-locality: Brazil. 1890 Leptagrion dispar - KIRB»1, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. elongatum SELYS 1876 Agrion (Leptagrion) elongatum SELYS, Synopsis des Agrionines, Add. 5: 269. Type male: Selys's Collection; type-locality: Brazil. 1890 Leptagrion elongatum - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. fernandezianum RACENIS 1958 Leptagrion fernandezianum RACENIS, Acta biol. venezuel., 2:195. Type male: Caracas Museum; type-locality: Venezuela.

Distribution: Venezuela.

310 garbei

SANTOS

1961 Leptagrion garbei SANTOS, Rev. bras. Biol., 21: 360. Type: unknown; type-locality: Brazil.

Distribution: Brazil. macrurum

(BURMEISTER)

1839 Agrion macrurum BURMEISTER, H a n d b . Ent., 2: 819. Type male: Sommer's Collection; typelocality: Brazil. 1876 Agrion (Leptagrion) macrurum - SELYS, Synopsis des Agrionines, Add. 5: 267 (as generotype o f Leptagrion

SELYS, 1 8 7 6 ) .

1890 Leptagrion macrurum - KIRBY, Syn. Cat. Neur.-Odon., London: 155. 1898 Leptagrion macrurum - CALVERT, Trans. Am. ent. Soc., 25: 37.

Distribution: Brazil. obsoletum

SELYS

1876 Agrion (Leptagrion) obsoletum SELYS, Synopsis des Agrionines, Add. 5: 275. Type male: McLachlan's Collection; type-locality: Amazonas. 1890 Leptagrion obsoletum - KIRBY, Syn. Cat. Neur.-Odon., London: 156.

Distribution: Brazil, perlongum

CALVERT

1 9 8 4 L e p t a g r i o n p e r l o n g u m CALVERT, 1 9 0 9 - l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 82.

Distribution: Argentina. prothoracicum

(KIMMINS)

1945 Telagrion prothoracicum KIMMINS, Ann. Mag. nat. Hist., (11) 12: 187. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador. 1984 Leptagrion prothoracicum - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 82.

Distribution: Ecuador. porrectum

SELYS

1876 Agrion (Leptagrion) porrectum SELYS, Synopsis des Agrionines, Add. 5: 265. Type male: Selys's Collection; type-locality: Brazil. 1890 Leptagrion porrectum - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. rufum

SELYS

1876 Agrion (Leptagrion) rufum SELYS, Synopsis des Agrionines, Add. 5: 277. Type male: Mus. Tierk., Dresden; type-locality: unknown. 1890 Leptagrion rufum - KIRBY, Syn. Cat. Neur.-Odon., London: 156.

Distribution: ? South America, siqueirai

SANTOS

1 9 8 4 L e p t a g r i o n siqueirai SANTOS, 1 9 6 8 - l o c . c i t . DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 82.

Distribution: Brazil.

311 vriesianum SANTOS 1984

Leptagrion vriesianum Utrecht, 1: 82.

SANTOS, 1 9 7 8

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Brazil. Genus Megalagrion M C L A C H L A N 1883 Megalagrion MCLACHLAN, Ann. Mag. nat. Hist., (5) 7: 237. Type-species: Megalagrion blackburni MCLACHLAN, 1883 (original designation). - 1920 Hawaiiagrion KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type-species: Agrion (Agrion) xanthomelas SELYS, 1876 (original designation). - 1920 Kilauagrion KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type-species: Agrion nesiotes PERKINS, 1899 (original designation). - 1920 Oahuagrion KENNEDY, Ohio J. Sei. Columbus, 21: 86. Type-species: Agrion oahuensis BLACKBURN, 1884 (original designation) 1948 Hawaiiagrion - ZIMMERMANN, Ins. Hawaii, 2: 342 (proposed synonymy with Megalagrion MCLACHLAN, 1883). - 1948 Kilauagrion - ZIMMERMANN, Ins. Hawaii, 2: 342 (proposed synonymy with Megalagrion MCLACHLAN, 1883). - 1948 Oahuagrion - ZIMMERMANN, Ins. Hawaii, 2: 342 (proposed synonymy with Megalagrion MCLACHLAN, 1883).

adytum PERKINS 1899 Agrion adytum PERKINS, Fauna Hawaiiensis, 2:69. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1899 Agrion adytum var. tillyardi PERKINS, Fauna Hawaiiensis, 2: 69. Male from Hawaii. 1917 Megalagrion adytum - KENNEDY, Ent. News, 28 (1): 11. 1948 Megalagrion adytum - ZIMMERMANN, Ins. Hawaii, 2: 359 (Hawaii).

Distribution: Hawaii. amaurodytum amaurodytum (PERKINS) 1899 Agrion amaurodytum PERKINS, Fauna Hawaiiensis, 2: 66. Type male: B.P. Bishop Mus., Honolulu; type-locality: Hawaii. 1917 Megalagrion amaurodytum amaurodytum - KENNEDY, Ent. News, 28 (1): 11. 1948 Megalagrion amaurodytum amaurodytum - ZIMMERMANN, Ins. Hawaii, 2: 359 (Hawaii).

Distribution: Hawaii. amaurodytum fallax (PERKINS) 1899 Agrion amaurodytum var. fallax PERKINS, Fauna Hawaiiensis, 2: 67. Male from Hawaii. 1948 Megalagrion amaurodytum fallax - ZIMMERMANN, Ins. Hawaii, 2: 359 (Hawaii).

Distribution: Hawaii. amaurodytum peles (PERKINS) 1899 Agrion amaurodytum race peles PERKINS, Fauna Hawaii, 2: 67. Male - type-locality: not specified by Perkins. 1917 Megalagrion amaurodytum race peles - KENNEDY, Ent. News, 28 (1): 11. 1948 Megalagrion amaurodytum peles - ZIMMERMANN, Ins. Hawaii, 2: 360 (Hawaii).

Distribution: Hawaii. amaurodytum wainaeanum (PERKINS) 1899 Agrion amaurodytum race wainaeanum PERKINS, Fauna Hawaii, 2: 67. Male from Oahu. 1938 Megalagrion amaurodytum wainaeanum - SCHMIDT, Ann. Ent. Soc. Amer., 31 (3): 326. 1948 Megalagrion amaurodytum wainaeanum - ZIMMERMANN, Ins. Hawaii, 2: 361 (Hawaii).

Distribution: Hawaii.

312 blackburni

MCLACHLAN

1882 Megalagrion blackburni MCLACHLAN, Ann. Mag. nat. Hist., (5), 12: 237 (as generotype of Megalagrion McLachlan, 1883). Type male: British Museum (Nat.Hist.), London; type-locality: Hawaii. 1899 Arion blackburni - PERKINS, F a u n a Hawaii, 2: 76. 1917 Arion blackburni - KENNEDY, Ent. News, 28 (1): 11. 1948 Megalagrion blackburni - ZIMMERMANN, Ins. Hawaii, 2: 361 (Hawaii).

Distribution: Hawaii. calliphya calliphya

MCLACHLAN

1883 Megalagrion calliphya MCLACHLAN, Ann. Mag. nat. Hist., (5) 12: 236. Type male: British Museum (Nat.Hist.), London; type-locality: Hawaii. 1884 Agrion satelles BLACKBURN, Ann. Mag. nat. Hist., (5) 14:414. Type male: B.P. Bishop Museum; type-locality: Hawaii. 1890 Coenagrion safeties - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1917 Megalagrion calliphya calliphya - KENNEDY, Ent. News, 28: (1): 11. 1920 Hawaiiagrion calliphya - KENNEDY, Ohio J. Sei. Columbus, 21: 86. 1948 Megalagrion calliphya calliphya - ZIMMERMANN, Ins. Hawaii, 2: 361 (Hawaii).

Distribution: Hawaii. calliphya microdemas

(PERKINS)

1899 Agrion calliphya race microdemas locality: given by Perkins. 1917 Megalagrion calliphya microdemas 1920 Hawaiiagrion calliphya microdemas 1948 Megalagrion calliphya microdemas

PERKINS, Fauna Hawaii, 2: 71. Male - no definite; type- KENNEDY, Ent. News, 28: 12. - KENNEDY, Ohio J. Sei. Columbus, 21: 86. - ZIMMERMANN, Ins. Hawaii, 2: 362 (Hawaii).

Distribution: Hawaii. hawaiense

(MCLACHLAN)

1883 Agrion hawaiense MCLACHLAN, Ann. Mag. nat. Hist., (5) 12: 232. Type male: British Museum (Nat.Hist.), London; type-locality: Hawaii. 1883 Agrion deceptor MCLACHLAN, Ann. Mag. nat. Hist., (5) 12: 235. Type male: British Museum (Nat.Hist.), London; type-locality: Hawaii. 1890 Coenagrion hawaiense - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1890 Coenagrion deceptor - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1899 Agrion calverti PERKINS, Fauna Hawaiiensis, 2: 694. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1936 Megalagrion hawaiense - WILLIAMS, Proc. Haw. ent. Soc., 9: 324 (redescription). 1936 Agrion calverti - WILLIAMS, Proc. Haw. ent. Soc., 9: 324 [proposed synonymy with hawaiense (MCLACHLAN,

1883)].

1948 Agrion deceptor - ZIMMERMANN, Ins. Hawaii, 2: 363 [proposed synonymy with hawaiense (MCLACHLAN,

1883)].

Distribution: Hawaii. eudytum

(PERKINS)

1899 Agrion eudytum PERKINS, F a u n a Hawaiiensis, 2: 68. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion eudytum - KENNEDY, Ent. News, 28: 12. 1948 Megalagrion eudytum - ZIMMERMANN, Ins. Hawaii, 2: 362 (Hawaii).

Distribution: Hawaii.

313 heterogamias

(PERKINS)

1899 Agrion heterogamias PERKINS, F a u n a Hawaiiensis, 2: 77. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion heterogamias - KENNEDY, Ent. News, 28: 13. 1948 Megalagrion heterogamias - ZIMMERMANN, Ins. Hawaii, 2: 365 (Hawaii).

Distribution: Hawaii. jugorum

(PERKINS)

1899 Agrion jugorum PERKINS, F a u n a Hawaiiensis, 2: 72. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion jugorum - KENNEDY, Ent. News, 28: 11. 1948 Megalagrion jugorum - ZIMMERMANN, Ins. Hawaii, 2: 365 (Hawaii).

Distribution: Hawaii. kauaiense

(PERKINS)

1899 Agrion kauaiense PERKINS, F a u n a Hawaiiensis, 2: 75. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion kauaiense - KENNEDY, Ent. News, 28: 11. 1948 Megalagrion kauaiense - ZIMMERMANN, Ins. Hawaii, 2: 365 (Hawaii).

Distribution: Hawaii. koelanse

(BLACKBURN)

1884 Agrion koelense BLACKBURN, Ann. Mag. nat. Hist., (5) 14: 417. Type male: B.P. Bishop M u seum, Honolulu; type-locality: Hawaii. 1890 Coenagrion koelense - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1899 Agrion asteliae PERKINS, F a u n a Hawaiiensis, 2:66. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1948 Megalagrion koelense - ZIMMERMANN, Ins. Hawaii, 2: 366. 1948 Agrion asteliae - ZIMMERMANN, Ins. Hawaii, 2 : 366 [proposed synonymy with koelense ( B L A C K BURN, 1884)].

Distribution: Hawaii. leptodemas

(PERKINS)

1899 Agrion leptodemas PERKINS, F a u n a Haiwaiiensis, 2: 70. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion leptodemas - KENNEDY, Ent. News, 28. 12. 1936 Megalagrion leptodemas - WILLIAMS, Proc. Haw. ent. Soc., 9: 304 (redescription) (Hawaii). 1948 Megalagrion leptodemas - ZIMMERMANN, Ins. Hawaii, 2: 368 (Hawaii).

Distribution: Hawaii. molokaiense

(PERKINS)

1899 Agrion molokaiense PERKINS, F a u n a Hawaiiensis, 2: 73. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion molokaiense - KENNEDY, Ent. News, 28: 11. 1920 Hawaiiagrion molokaiense - KENNEDY, Ohio J. Sei. Columbus, 21: 86. 1948 Megalagrion molokaiense - ZIMMERMANN, Ins. Hawaii, 2: 370 (Hawaii).

Distribution: Hawaii.

314 nesiotes

(PERKINS)

1899 Agrion nesiotes PERKINS, F a u n a Hawaiiensis, 2:72. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1920 Kilauagrion nesiotes - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Kilauagrion KENNEDY, 1 9 2 0 ) .

1934 Kilauagrion dinesiotes KENNEDY, Ann. Ent. Soc. Amer., 27 (2): 343. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1948 Megalagrion nesiotes - ZIMMERMANN, Ins. Hawaii, 2: 370. 1948 Kilauagrion dinesiotes - ZIMMERMANN, Ins. Hawaii, 2: 370 [proposed synonymy with nesiotes (PERKINS, 1 8 9 9 ) ] .

Distribution: Hawaii. nigrohamatum nigrohamatum

(BLACKBURN)

1884 Agrion nigrohamatum BLACKBURN, Ann. Mag. nat. Hist., (5) 14. 416. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1890 Coenagrion nigrohamatum - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1917 Megalagrion nigrohamatum - KENNEDY, Ent. News, 28: 13. 1920 Hawaiiagrion nigrohamatum - KENNEDY, Ohio J. Sei. Columbus, 21: 86. 1936 Megalagrion nigrohamatum -WILLIAMS, Proc. Haw. ent. Soc., 9: 318 (redescription) (Hawaii). 1948 Megalagrion nigrohamatum nigrohamatum - ZIMMERMANN, Ins. Hawaii, 2: 370 (Hawaii).

Distribution: Hawaii. nigrohamatum nigrolineatum

(PERKINS)

1899 Agrion nigrohamatum race nigrolineatum PERKINS, F a u n a Hawaiiensis, 2: 65. Male from Hawaii. 1917 Megalagrion nigrohamatum nigrolineatum - KENNEDY, Ent. News, 28: 13. 1920 Hawaiiagrion nigrohamatum nigrolineatum - KENNEDY, Ohio J. Sei. Columbus, 21: 86. 1936 Megalagrion nigrohamatum race nigrolineatum - WILLIAMS, Proc. Haw. ent. Soc., 9: 318. 1948 Megalagrion nigrohamatum nigrolineatum - ZIMMERMANN, Ins. Hawaii, 2: 372 (Hawaii).

Distribution: Hawaii, oahuense

(BLACKBURN)

1884 Agrion oahuense BLACKBURN, Ann. Mag. nat. Hist., (5) 14: 415. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1890 Coenagrion oahuense - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1899 Agrion oahuense - PERKINS, Fauna Hawaiiensis, 2: 74 (description of female) (Hawaii). 1920 Oahuagrion oahuense - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Oahuagrion KENNEDY, 1920).

1936 Megalagrion 1948 Megalagrion

oahuense - WILLIAMS, Proc. Haw. ent. Soc., 9: 339 (redescription) (Hawaii). oahuense - ZIMMERMANN, Ins. Hawaii, 2: 372.

Distribution: Hawaii. oceanicum

MCLACHLAN

1882 Megalagrion oceanicum MCLACHLAN, Ann. Mag. nat. Hist., (5) 12: 239. Type male: British Museum (Nat.Hist.), London; type-locality: Hawaii. 1899 Agrion oceanicum - PERKINS, F a u n a Hawaiiensis, 2: 76. 1936 Megalagrion oceanicum - WILLIAMS, Proc. Haw. ent. Soc., 9: 320 (redescription) (Hawaii). 1948 Megalagrion oceanicum - ZIMMERMANN, Ins. Hawaii, 2: 76.

Distribution: Hawaii.

315 oresitrophum

(PERKINS)

1899 Agrion oresitrophum PERKINS, F a u n a Hawaiiensis, 2: 69. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion oresitrophum - KENNEDY, Ent. News, 28: 10. 1948 Megalagrion oresitrophum - ZIMMERMANN, Ins. Hawaii, 2: 377 (Hawaii).

Distribution: Hawaii. orobates

(PERKINS)

1899 Agrion orobates PERKINS, F a u n a Hawaiiensis, 2: 70. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion orobates - Kennedy, Ent. News, 28: 10. 1948 Megalagrion orobates - ZIMMERMANN, Ins. Hawaii, 2: 377 (Hawaii).

Distribution: Hawaii. pacificum

(MCLACHLAN)

1883 Agrion pacificum MCLACHLAN, Ann. Mag. nat. Hist., (5) 12: Type male: British Museum (Nat.Hist.), London; type-locality: Hawaii. 1890 Coenagrion pacificum - KIRBY, Syn. Cat. Neur.-Odon., London: 151. 1899 Agrion pacificum - PERKINS, F a u n a Hawaiiensis, 2: 64. 1917 Megalagrion pacificum - KENNEDY, Ent. News, 28: 12. 1948 Megalagrion pacificum - ZIMMERMANN, Ins. Hawaii, 2: 377 (Hawaii).

Distribution: Hawaii. paludicola

MACIOLEK &

HOWARTH

1 9 8 4 M e g a l a g r i o n paludicola MACIOLEK & HOWARTH, 1 9 7 9 - l o c . cit. D A VIES & TOBIN, T h e D r a -

gonflies of the World, Utrecht, 1: 83.

Distribution: Hawaii. vagabundum

(PERKINS)

1899 Agrion vagabundum PERKINS, F a u n a Hawaiiensis, 2: 75. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion vagabundum - KENNEDY, Ent. News, 28: 13. 1948 Megalagrion vagabundum - ZIMMERMANN, Ins. Hawaii, 2: 378 (Hawaii).

Distribution: Hawaii. williamsoni

(PERKINS)

1899 Agrion williamsoni PERKINS, F a u n a Hawaiiensis, 2: 69. Type male: B.P. Bishop Museum, Honolulu; type-locality: Hawaii. 1917 Megalagrion williamsoni - KENNEDY, Ent. News, 28: 10. 1948 Megalagrion williamsoni - ZIMMERMANN, Ins. Hawaii, 2: 379.

Distribution: Hawaii. xanthomelas

(SELYS)

1876 Agrion (Agrion) xanthomelas SELYS, Synopsis des Agrionines, Add. 5: 174. Type male: Selys's Collection; type-locality: Sandwich Islands. 1890 Coenagrion xanthomelas - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1920 Hawaiiagrion xanthomelas - KENNEDY, Ohio J. Sei. Columbus, 21: 86 (as generotype of Hawaiiagrion

KENNEDY, 1 9 2 0 ) .

316 1936 Megalagrion xanthomelas - WILLIAMS, Proc. Haw. ent. Soc., 9: 310 (redescription) (Hawaii). 1948 Megalagrion xanthomelas - ZIMMERMANN, Ins. Hawaii, 2: 379 (Hawaii).

Distribution: Hawaii.

Genus Melanesobasis DONNELLY 1984

Melanesobasis D O N N E L L Y , 1 9 8 4 . Type-species: Nesobasis corniculata T I L L Y A R D , 1 9 2 3 (original designation) - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 83.

annulata (BRAUER) 1969 Pericnemis annulata BRAUER, Verh. Zool.-Bot. Ges. Wien, 19: 10. Type male: Zool. Inst. Mus. Univ. Hamburg; type-locality: Samoa. 1959 Nesobasis annulata - LIEFTINCK, Ent. Mitt. Zool. Staatsinst. Zool. Mus. Hamburg, 21: 1. 1984 Melanesobasis annulata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 83.

Distribution: Samoa. bicellulare DONNELLY 1984 Melanesobasis bicellulare World, Utrecht, 1: 83.

DONNELLY,

1984 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Vanuatu. corniculata corniculata (TILLYARD) 1923 Nesobasis corniculata TILLYARD, Trans, ent. Soc. London, 1923: 319. Type male: Cawthron Inst. Coll., type-locality: Fiji. 1 9 8 4 Melanesobasis corniculata corniculata - D O N N E L L Y , 1 9 8 4 (as generotype of Melanesobasis D O N N E L L Y , 1 9 8 4 ) . - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 3 .

Distribution: Fiji. corniculata marginata DONNELLY 1984 Melanesobasis corniculata marginata D O N N E L L Y , 1984 - loc. cit. gonflies of the World, Utrecht, 1: 83.

DAVIES & TOBIN,

The Dra-

Distribution: Fiji. flavilabris (SELYS) 1891 Agrion (Nesobasis) flavilabris SELYS C.R. ent. Belg., 35: 55. Type male: Godeflroy Museum, Hamburg; type-locality: Fiji. 1923 Nesobasis flavilabris - TILLYARD, Trans, ent. Soc. London, 1923: 319 (Fiji). 1 9 8 4 Melanesobasis flavilabris - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 4 [incorrect synonymy with annulata ( B R A U E R , 1 8 6 9 ) ] .

Distribution: Fiji. maculosa DONNELLY 1984 Melanesobasis maculosa Utrecht, 1: 84.

DONNELLY,

1984 - cit. DAVIES &

TOBIN,

The Dragonflies of the World,

Distribution: Fiji. mcleani DONNELLY 1984 Melanesobasis mcleani World, Utrecht, 1: 84.

Distribution: Fiji.

DONNELLY,

1984 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

317 prolixa DONNELLY 1984 Melanesobasis prolixa DONNELLY, 1984 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 84.

Distribution: Fiji. simmondsi (TILLYARD) 1923 Nesobasis simmondsi T I L L YARD, Trans, ent. Soc. London, 1923: 320. Type male: Cawthron Inst. Coll.; type-locality: Fiji. 1984 Melanesobasis simmondsi - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 84.

Distribution: Fiji. Genus Minagrion SANTOS 1965 Minagrion SANTOS, Atas Soc. Biol. Rio de Janeiro, 9: 9. Type-species: Agrion mecistogastrum SELYS, 1876 (original designation).

(Telagrion)

caldense SANTOS 1965 Minagrion caldense SANTOS, Atas Soc. Biol. Rio de Janeiro, 9: 10. Type male: Inst. Biol. Def. Agr., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. canaanensis SANTOS 1967 Minagrion canaanensis SANTOS, Atas Soc. Biol. Rio de Janeiro, 11: 7. Type male: Inst. Biol. Def. Agr., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. mecistogastrum (SELYS) 1876 Agrion (Telagrion) mecistogastrum SELYS, Synopsis des Agrionines, Add. 5: 262. Type male: Selys's Collection; type-locality: Brazil. 1965 Minagrion mecistogastrum - SANTOS, Atas Soc. Biol. Rio de Janeiro, 9: 9 (as generotype of Minagrion SANTOS, 1965).

Distribution: Brazil. ribeiroi SANTOS 1956 Minagrion ribeiroi SANTOS, Rev. bras. Biol., 16: 369. Type male: Santos's Collection; typelocality: Brazil.

Distribution: Brazil. waltheri (SELYS) 1876 Agrion (Agrion) waltheri SELYS, Synopsis des Agrionines, Add. 5: 176. Type male: Selys's Collection; type-locality: Minas Geraes. 1890 Coenagrion waltheri - KIRBY, Syn. Cat. Neur.-Odon., London: 150. 1956 Telagrion serracipoensis SANTOS, Rev. bras. Biol., 16: 369. Type male: Santos's Collection; type-locality: Brazil. 1984 Minagrion waltheri - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 84. 1984 Teleagrion serracipoensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 84 [proposed synonymy with waltheri (SELYS, 1 8 7 6 ) ] .

Distribution: Brazil.

318 Genus Nesobasis

SELYS

1891 Agrion (Nesobasis) SELYS, C.R. ent. Belg., 35: 51. Type-species: Agrion (Nesobasis) erythrops SELYS, 1891 (original designation). - 1923 Nesobasis - T I L L Y A R D , Trans, ent. Soc. London, 1923: 313 (revision, key to species).

angulicollis

TILL YARD

1923 Nesobasis angulicollis T I L L Y A R D , Trans, ent. Soc. London, 1923: 322. Type male: Cawthron Inst. Coll., type-locality: Fiji.

Distribution: Fiji. aurantiaca

TILLYARD

1923 Nesobasis aurantiaca T I L L Y A R D , Trans, ent. Soc. London, 1923: 332. Type male: Cawthron Inst. Coll; type-locality: Fiji.

Distribution: Fiji. bidens 1958

KIMMINS

Nesobasis bidens KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), (Nat.Hist.), London; type-locality: New Hebrides.

6: 2 3 9 .

Type male: British Museum

Distribution: New Hebrides. brachycerca

TILLYARD

1923 Nesobasis brachycerca T I L L Y A R D , Trans, ent. Soc. London, 1923: 332. Type male: Cawthron Inst. Coll., type-locality: Fiji.

Distribution: Fiji. campioni

TILLYARD

1923 Nesobasis campioni T I L L Y A R D , Trans, ent. Soc. London, 1923: 329. Type male: Cawthron Inst. Coll. type-locality: Fiji.

Distribution: Fiji. comosa

TILLYARD

1923 Nesobasis comosa T I L L Y A R D , Trans, ent. Soc. London, 1923: 321. Type male: Cawthron Inst. Coll.; type-locality: Fiji.

Distribution: Fiji. erythrops

SELYS

1891 Agrion (Nesobasis) erythrops SELYS, C.R. ent. Belg., 35: 53 (as generotype of Nesobasis 1891). Type male: Godeffroy Museum, Hamburg; type-locality: Fiji. 1923 Nesobasis erythrops - T I L L Y A R D , Trans, ent. Soc. London, 1 9 2 3 : 317 (Fiji).

SELYS,

Distribution: Fiji. heteroneura

TILLYARD

1923 Nesobasis heteroneura T I L L Y A R D , Trans, ent. Soc. London, 1923: 333. Type male: Cawthron Inst. Coll.; type-locality: Fiji.

Distribution: Fiji.

319 leveri

KIMMINS

1943 Nesobasis leveri KIMMINS, Ann. Mag. nat. Hist., (11) 10: 699. Type male: British Museum (Nat.Hist.), London; type-locality: Fiji.

Distribution: Fiji, longistyla

SELYS

1891 Agrion (Nesobasis) longistyla SELYS, C.R. ent. Belg., 35: 57. Type male: Godeffroy Museum, Hamburg; type-locality: Fiji. 1923 Nesobasis longistyla - TILL YARD, Trans, ent. Soc. London; 1923: 334 (Fiji).

Distribution: Fiji. malekulana

KIMMINS

1936 Nesobasis malekulana KIMMINS, Ann. Mag. nat. Hist., (10) 18: 72. Type male: British Museum (Nat.Hist.), London; type-locality: New Hebrides.

Distribution: New Hebrides. nigrostigma

SELYS

1891 Agrion (Nesobasis) nigrostigma SELYS, C.R. ent. Belg., 35: 56. Type male: Godeffroy Museum, Hamburg; type-locality: Fiji. 1923 Nesobasis nigrostigma - TILLYARD, Trans, ent. Soc. London, 1923: 322 (Fiji).

Distribution: Fiji. selysi

TILLYARD

1923 Nesobasis selysi TILLYARD, Trans, ent. Soc. London, 1923: 327. Type male: Cawthron Inst. Coll.; type-locality: Fiji.

Distribution: Fiji. subhumeralis

TILLYARD

1923 Nesobasis subhumeralis TILLYARD, Trans, ent. Soc. London, 1923: 325; type male: Cawthron Inst. Coll.; type-locality: Fiji.

Distribution: Fiji. telegastrum

SELYS

1891 Agrion (Nesobasis) telegastrum SELYS, C.R. ent. Belg., 35: 56. Type male: Godeffroy Museum, H a m b u r g ; type-locality: Fiji. 1923 Nesobasis telegastrum - TILLYARD, Trans, ent. Soc. London, 1923: 317 (Fiji).

Distribution: Fiji. Genus Papuagrion Ris 1913 Papuagrion Ris, Nova Guinea, 9 (3): 482. Type-species: Papuagrion reductum Ris, 1913 (original designation). - 1935 Papuagrion LIEFTINCK, N o v a Guinea, 17: 234 (revision, key to New Guinean species). - 1937 Papuagrion - LIEFTINCK, Nova Guinea (N.S.) 1: 307 (key to species). - 1984 Papuagrion - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 84 [type-species: Teinobasis (Stenobasis) occipitalis SELYS, 1877].

320 auriculatum

LIEFTINCK

1937 Papuagrion auriculatum LIEFTINCK, Nova Guinea (N.S.) 1: 321. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. corruptum

LIEFTINCK

1938 Papuagrion corruptum LIEFTINCK, Nova Guinea (N.S.) 2: 105. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. degeneratum

LIEFTINCK

Papuagrion magnanimum LIEFTINCK (nec SELYS, 1 8 7 6 ) , N o v a Guinea, 17: 2 4 5 . Type female: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea. 1937 Papuagrion degeneratum LIEFTINCK, Nova Guinea (N.S.) 1: 316 (new name for Papuagrion magnanimum LIEFTINCK, 1935, a junior secondary homonymy of Agrion (Pseudagrion) magnanimum SELYS, 1876). 1935

Distribution: New Guinea. digitiferum

LIEFTINCK

1949 Papuagrion digitiferum LIEFTINCK, Nova Guinea (N.S.) 5:155. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. ekari

LIEFTINCK

1949 Papuagrion ekari LIEFTINCK, N o v a Guinea (N.S.) Leiden; type-locality: New Guinea.

5:148.

Type male: Rijksmus. Natuurl. Hist.,

Distribution: New Guinea, flavipedum LIEFTINCK 1949 Papuagrion flavipedum LIEFTINCK, Nova Guinea (N.S.) 5: 156. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, flavithorax

(SELYS)

1878 Pseudagrion flavithorax SELYS, Mitt. Zool. Mus. Dresden, 3: 322. Type male: R. Mus., Brussels; type-locality: Papua. 1937 Papuagrion flavithorax - LIEFTINCK, Nova Guinea (N.S.) 1: 310 («description).

Distribution: New Guinea. fraterculum LIEFTINCK 1937 Papuagrion fraterculum LIEFTINCK, Nova Guinea (N.S.) 1: 323. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, gurneyi

LIEFTINCK

1949 Papuagrion gurneyi LIEFTINCK, Treubia, 20 (2): 341. Type male: U S Nation. Mus., Washington; type-locality: Bougainville. Distribution: Bougainville.

321 insulare

LIEFTINCK

1949 Papuagrion insulare LIEFTINCK, Nova Guinea (N.S.) 5: 160. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Biak Island.

Distribution: Indonesia: Biak Island. laminatum

LIEFTINCK

1937 Papuagrion laminatum LIEFTINCK, Nova Guinea (N.S.) 1: 319. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. magnanimum

(SELYS)

1876 Agrion (Pseudagrion) magnanimum SELYS (nec LIEFTINCK, 1935), Synopsis des Agrionines, Add. 5: 226. Type male: McLachlan's Collection; type-locality: Aru. 1 8 9 0 Pseudagrion magnanimum - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 5 3 . 1 9 8 4 Papuagrion magnanimum - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 5 .

Distribution: Aru. occipitale

(SELYS)

1877 Telebasis (Stenobasis) occipitalis SELYS, Bull. Acad. r. Belg. (2) 43: 111. Type male: Selys's Collection; type-locality: New Guinea. 1890 Archibasis (n.n.) occipitalis - K I R B Y , Syn. Cat. Neur.-Odon., London: 156. 1908 Stenobasis occipitalis - M A R T I N , Bull. Soc. ent. Ital., 6 0 : 206. 1935 Papuagrion occipitale - LIEFTINCK, Nova Guinea 1 7 : 238 (redescription). 1984 Papuagrion occipitale - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 85 (as generotype of Papuagrion Ris, 1913).

Distribution: New Guinea. oppositum

LIEFTINCK

1949 Papuagrion oppositum LIEFTINCK, Nova Guinea (N.S.) 5: 146. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. pandanicolum

LIEFTINCK

1949 Papuagrion pandanicolum LIEFTINCK, N o v a Guinea (N.S.) 5: 153. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, parameles

LIEFTINCK

1949 Papuagrion parameles LIEFTINCK, Nova Guinea (N.S.) 5: 159. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, pesechem pesechem

LIEFTINCK

1949 Papuagrion pesechem pesechem LIEFTINCK, Nova Guinea (N.S.) 5: 152. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea. Distribution: New Guinea.

322

pesechem corniculatum

LIEFTINCK

1949 Papuagrion pesechem corniculatum LIEFTINCK, Nova Guinea (N.S.) 5: 152. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, prothoracale

LIEFTINCK

1935 Papuagrion prothoracel LIEFTINCK, Nova Guinea, 17: 242. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. rectangulare

LIEFTINCK

1937 Papuagrion rectangulare LIEFTINCK, Nova Guinea (N.S.) 5: 313. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, reductum Ris 1913 Papuagrion reductum Ris, N o v a Guinea, 9 (3): 483. Type male: Inst. Tax. Zoöl. Amsterdam; type-locality: New Guinea (as generotype of Papuagrion Ris, 1913). 1935 Papuagrion reductum - LIEFTINCK, Nova Guinea, 17: 248.

Distribution: New Guinea. rufipedum

LIEFTINCK

1937 Papuagrion rufipedum LIEFTINCK, Nova Guinea (N.S.) 1: 317. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, spinicaudum

LIEFTINCK

1937 Papuagrion spinicaudum LIEFTINCK, Nova Guinea (N.S.) 5: 311. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Pericnemis

HÄGEN

1863 Amphicnemis (Pericnemis) HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 7. Type-species: Pericnemisstictica SELYS, 1 8 6 3 (monotypy). - 1 8 7 7 Telebasis (Pericnemis) - SELYS, Bull. Acad, r. Belg. (2): 43: 131 (revision). - 1890 Pericnemis - KIRBY, Syn. Cat. Neur.-Odon., London: 1 5 8 . - 1 9 3 9 Pericnemis - NEEDHAM & G Y G E R , Philipp. J . Sei., 7 0 ( 3 ) : 2 9 1 (key to species). 1954 Pericnemis - LIEFTINCK, Treubia, 22 Suppl.: 63.

stictica

SELYS

1863 Amphicnemis (Pericnemis) stictica SELYS, Synopsis des Agrionines, Add. 4:8. Type male: Musee de la East India House, London (as generotype of Pericnemis HÄGEN, 1863); type-locality: Java. 1877 Amphicnemis (Pericnemis) sticia - SELYS, Bull. Acad. r. Belg. (2): 43: 132 (redescription) (type in Mus. N a t u r k . Berlin). 1890 Pericnemis stictica - KIRBY, Syn. Cat. Neur.-Odon., London: 158. 1927 Pericnemis stictica - Ris, Zool. Meded., Leiden, 10 (1): 25 (Sumatra). 1934 Pericnemis stictica - LIEFTINCK, Treubia, 14 (4): 398 (Java, Sumatra, Borneo, Malay Archipelago).

Distribution: from Malaysia to Borneo.

323 triangularis

LAIDLAW

1931 Pericnemis triangularis LAIDLAW, J. Fed. Mai. States Mus., 16: 248. Type female: Laidlaw's Collection; type-locality: Borneo. 1954 Pericnemis triangularis - LIEFTINCK, Treubia, 22 Suppl.: 64 (Borneo).

Distribution: Borneo. Genus Plagulibasis

LIEFTINCK

1949 Plagulibasis LIEFTINCK, Nova Guinea (N.S.) 5: 176. Type-species: Nesolabis ciliata Ris, 1913 (original designation).

ciliata (Ris) 1913 Nesobasis ciliata Ris, Nova Guinea, 9: 485. Type male: unknown; type-locality: New Guinea. 1949 Plagulibasis ciliata - LIEFTINCK, Nova Guinea (N.S.) 5: 176 (as generotype of Plagulibasis LIEFTINCK, 1949).

Distribution: New Guinea. Genus Pseudagrion

SELYS

1876 Agrion (Pseudagrion) SELYS, Synopsis des Agrionines, Add. 5: 200. Type-species: Agrion fuscigerum RAMBUR, 1842 (original designation). - 1890 Pseudagrion - KIRBY, Syn. Cat. Neur.Odon., London: 153. - 1905 Pseudagrion - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 830 (key to species). - 1916 Pseudagrion - LAIDLAW, Ree. Indian Mus., 12: 21. (revision, key to species). - 1921 Pseudagrion - Ris, Annls. S. Afr. Mus., 18: 297 (key to species). 1931 Pseudagrion - CHUJO, Trans, nat. Hist. Soc. Formosa, 21 (112): 40. - 1934 Pseudagrion - SCHMIDT, Arch. Hydrobiol. Suppl.: 13: 346. - 1936 Pseudagrion - SCHMIDT, Abh. senckenb. naturf. Ges., 433:1 (revision, key to African species). - 1936 Pseudagrion - LONGFIELD, Trans. R. ent. Soc. London, 85: 472. - 1939 Pseudagrion - NEEDHAM & GYGER, Philipp. J. Sei. 70 (3): 281 (key to species). - 1941 Paulinagrion FRÄSER, Proc. R. ent. Soc. London, (B) 10: 37. Type-species: Paulinagrion platystictoides FRASER 1941 (original designation). - 1954 Pseudagrion LIEFTINCK, Treubia, 22 Suppl.: 56. - 1962 Pseudagrion LIEFTINCK, Insects of Micronesia, 5 (1): 36. - 1962 Pseudagrion - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 121 (type-species: Agrion caffrum BURMEISTER, 1839). - 1973 Pseudagrion - DUMONT, Israel J. Zool., 22 (2-4): 192 (key to regional species).

acaciae

FÖRSTER

1906 Pseudagrion acaciae FÖRSTER, Mannheim. Jb. Ver. Natk., 71-72: 56. Type male: Förster's Collection; type-locality: Transvaal. 1921 Pseudagrion acaciae - RIS, Annls. A. Afr. Mus., 18: 308 (Zululand, Nyassaland, Rhodesia) (redescription). 1928 Pseudagrion acaciae - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 22 (Egypt). 1936 Pseudagrion acaciae - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 53 (Rhodesia, Nyassaland, Egypt, Abyssinia, Sudan, Lebanon). 1962 Pseudagrion acaciae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 123 (from Egypt to South Africa, and Palestine).

Distribution: from Egypt to Natal, Israel, Lebanon, aguessei

PINHEY

1964 Pseudagrion aguessei PINHEY, Bull. Inst, franc. Afr. noire, (A) 26: 1146. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Sierra Leone.

Distribution: Sierra Leone.

324 alcicorne alcicorne

FÖRSTER

1906 Pseudagrion alcicorne FÖRSTER, Mannheim. Jb. Ver. Natk., 7 1 - 7 2 . 64. Type male: A n n A r b o r Museum; type-locality: Madagascar. 1936 Pseudagrion alcicorne - SCHMIDT, Abh. senckenb. naturf. Ges., 43: 42. 1962 Pseudagrion alcicorne alcicorne - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. alcicorne digitatum

SCHMIDT

1951 Pseudagrion alcicorne digitatum SCHMIDT, Mem. Inst. sei. Madagscar, 6A: 237. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion alcicorne digitatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. ambatoroae 1984

AGUESSE

Pseudagrion ambatoroae World, Utrecht, 1: 85.

AGUESSE, 1 9 6 8

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: Madagascar. ampolomitae 1984

AGUESSE

Pseudagrion ampolomitae World, Utrecht, 1: 85.

AGUESSE, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Madagascar. andamanicum

FRÄSER

1924 Pseudagrion andamanicum FRÄSER, Ree. Indian Mus., 26: 411. Type male: British Museum (Nat.Hist.), London; type-locality: A n d a m a n Island.

Distribution: Andaman Island. angelicum

FRASER

1947 Pseudagrion angelicum FRASER, Trans. R. ent. Soc. London, 98: 25. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Ivory Coast. 1962 Pseudagrion angelicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 123 (Ivory Coast, Nigeria).

Distribution: West Africa. angolense

SELYS

1876 Agrion (Pseudagrion) angolense SELYS, Synopsis des Agrionines, Add. 5: 203. Type male: McLachlan's Collection; type-locality: Angola. 1890 Pseudagrion angolense - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1821 Pseudagrion angolense - Ris, Annls. S. Afr. Mus., 18: 302 (S.Africa, Angola). 1936 Pseudagrion angolense - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 39. 1936 Pseudagrion angolense - LONGFIELD, Trans. R. ent. Soc. London, 85: 472 (Uganda, Congo). 1947 Pseudagrion angolense - FRASER, Trans. R. ent. Soc. London, 98: 25 (Ivory Coast). 1947 Pseudagrion monardi LONGFIELD, Arq. Mus. Bocage, 16: 12. Type male: British Museum (Nat.Hist.), London; type-locality: Angola. 1959 Pseudagrion angolense - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 17 (Tanganyika, Kenya, U g a n d a , Congo, Angola). 1962 Pseudagrion hageni- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 123 (incorrect synonymy with angolense SELYS, 1 8 7 6 ) .

325 1962 Pseudagrion monardi - KIMMINS, Ann. Mag. nat. Hist., (13) 5: 331 (proposed synonymy with angolense

SELYS, 1 8 7 6 ) .

1 9 8 4 P s e u d a g r i o n f i l i f o r m e R i s & SCHMIDT, 1 9 3 6 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f

the World, Utrecht, 1: 87 (proposed synonymy with angolense SELYS, 1876).

Distribution: tropical Africa. apicale

SCHMIDT

1949 Pseudagrion apicale SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 3A: 23. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion apicale - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. approximatum

SCHMIDT

1951 Pseudagrion approximatum SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 230. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion approximatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. assegaii

PINHEY

1950 Pseudagrion assegaii PINHEY, Ann. Transvaal Mus., 21: 261. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Transvaal. 1962 Pseudagrion assegaii - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 123 (Transvaal, S. Rhodesia).

Distribution: South Africa. aureofrons

TILLYARD

1906 Pseudagrion aureofrons TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 189. Type male: Tillyard's Collection; type-locality: Queensland. 1913 Pseudagrion aureofrons - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 470 (Queensland, New South Wales). 1917 Pseudagrion aureofrons - SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 38 («description).

Distribution: Australia. australasiae

SELYS

1876 Agrion (Pseudagrion) australasiae SELYS, Synopsis des Agrionines, Add. 5: 216. Type female: R. Mus., Brussels; type-locality: Queensland. 1890 Pseudagrion australasiae - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1901 Pseudagrion australasiae - MARTIN, Mem. Soc. zool. France, 14: 246 (Victoria). 1913 Pseudagrion australasiae - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 469 (Queensland, New South Wales). 1916 Pseudagrion australasiae - LAIDLAW, Ree. Indian Mus., 12: 23 (India to Australia). 1919 Pseudagrion bengalense LAIDLAW, Ree. Indian Mus., 16: 192 [as new name for Agrion (Pseudagrion) australasiae SELYS, 1 8 7 6 ] . 1936 Pseudagrion australasiae - LIEFTINCK, Konowia, 15 ( 3 - 4 ) : 167 (redescription). 1954 Pseudagrion australasiae - LIEFTINCK, Treubia, 22 Suppl.: 56 (Siam, Malaya, Sumatra, Java). 1954 Pseudagrion bengalense - LIEFTINCK, Treubia, 22 Suppl.: 56 (proposed synonymy with australasiae

SELYS, 1 8 7 6 ) .

Distribution: from India to Australia.

326 azureum

NEEDHAM & GYGER

1939 Pseudagrion azureum NEEDHAM & G Y G E R , Philipp. J. Sei. 70 (3): 283. Type male: not given; type-locality: Manila.

Distribution: Philippine Islands, basicornu

SCHMIDT

1934 Pseudagrion basicornu NIELSEN, Boll. Lab. Ent. R. Ist. Agr. Bologna, 7: 183 (emendation). 1 9 3 6 Pseudagrion basicornu SCHMIDT (new name for Pseudagrion basicornu NIELSEN, 1 9 3 4 ) , Abh. senckenb. naturf. ges., 433: 36. Type male: Martin's Collection; type-locality: Congo Republic. 1962 Pseudagrion basicornu - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 123 (Belgian Congo, Ivory Coast, Angola, Guinea, Dahomey, Liberia, Nigeria).

Distribution: West Africa. beadlei

PINHEY

1961 Pseudagrion beadlei PINHEY, Publ. Mus. Brit. (Nat.Hist.), Ent.: 33. Type male: British Museum (Nat.Hist.), London; type-locality: Lake Victoria. 1962 Pseudagrion beadlei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 123. 1 9 8 4 Pseudagrion beadlei - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 6 (as form of sjöstedti FÖRSTER, 1 9 0 6 ) . 1984 Pseudagrion beadlei - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 91 (as subspecies of sjöstedti FÖRSTER, 1 9 0 6 ) .

Distribution: Lake Victoria. bicoerulans

MARTIN

1906 Pseudagrion bicoerulans MARTIN, Bull. Mus. Hist. nat. Paris, 1906: 510. Type male: unknown; type-locality: Afr. Orientale. 1909 Pseudagrion grande SJÖSTEDT, Wiss. Ergebn. Exp. Kilimandjaro-Meru, 14: 45. Type male: Naturhist. Riksmus., Stockholm; type-locality: Kilimandjaro. 1915 Pseudagrion stellatum MARTIN, Voy. Alluaud Jeannel Afr. orient., Odon: 46. Type male: British Museum (Nat.Hist.), London; type-locality: Tanzania. 1915 Pseudagrion stellatum form major MARTIN, Voy. Alluaud Jeannel Afr. orient., Odon: 46. Male from Kilimandjaro. 1936 Pseudagrion bicoerulans - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 36. 1936 Pseudagrion stellatum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 36 (proposed synonymy with bicoerulans M A R T I N , 1 9 0 6 ) .

Distribution: Kenya, Tanzania. bidentatum

MORTON

1907 Pseudagrion bidentatum MORTON, Trans, ent. Soc. London, 1907: 308. Type male: British Museum (Nat.Hist.), London; type-locality: Deesa.

Distribution: India: Deesa. caffrum

(BURMEISTER)

1839 Agrion caffrum BURMEISTER, H a n d b . Ent., 2: 821. Type male: Agassiz Museum, Cambridge, (Mass.); type-locality: Port Natal. 1 8 7 6 Agrion palliatum ( K L U G & H A G E N M S S ) SELYS, Bull. Acad. r. Belg. ( 2 ) : 4 3 : 4 9 7 . Type male: Agassiz Museum, Cambridge, (Mass.); type-locality: Cape. 1890 Pseudagrion caffrum - KIRBY, Syn. Cat. Neur.-Odon., London: 187 [incorrect synonymy with Brachybasis rhomboidalis (BEAUVOIS, 1 8 0 5 ) ] . 1898 Pseudagrion caffrum - CALVERT, Trans. Amer. ent. Soc., 25: 42 (redescription of holotype). 1921 Pseudagrion caffrum - Ris, Annls. S. Afr. Mus., 18: 301 (South Africa).

327 1962 Pseudagrion Pseudagrion

caffrum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124 (as generotype of SELYS, 1 8 7 6 ) .

1962 Agrion palliatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124 [proposed synonymy with caffrum (BURMEISTER, 1 8 3 9 ) ] .

Distribution: South Africa. calosomum

LIEFTINCK

1936 Pseudagrion calosomum LIEFTINCK, Revue suisse Zool., 43 (5): 124. Type male: Buitenzorg Museum; type-locality: Sumba. 1953 Pseudagrion calosomum - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 155 (redescription).

Distribution: Sumba. celebense

LIEFTINCK

1937 Pseudagrion celebense LIEFTINCK, Treubia, 16 (1): 90. Type male: Buitenzorg Museum; typelocality: Celebes.

Distribution: Celebes. cheliferum

FRASER

1949 Pseudagrion cheliferum FRASER, Mem. Inst. sei. Madagascar, 3A: 29. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Pseudagrion cheliferum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. chloroceps

FRASER

Pseudagrion igniceps FRASER (nec FRASER, 1 9 5 3 ) , M e m Inst. sei. Madagascar, ( E ) 3 : 2 9 . Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1955 Pseudagrion chloroceps FRASER, Nat. malgache, 5: 163 (new name for Pseudagrion igniceps FRASER, 1953, a junior secondary homonymy of Pseudagrion igniceps FRASER, 1953 -» see igniceps). 1962 Pseudagrion chloroceps - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129. 1953

Distribution: Madagascar. cingillum

(BRAUER)

1869 Agrion cingillum BRAUER, Verh. Zool.-Bot. Ges. Wien, 19: 11. Type male: H a m b u r g Museum; type-locality: R o c k h a m p t o n . 1890 Micronympha cingillum - KIRBY, Syn. Cat. Neur.-Odon., London: 143. 1926 Pseudagrion papuense TILLYARD, Ree. Australian Mus., 15: 159. Type male: Tillyard's Collection; type-locality: New Guinea. 1959 Pseudagrion cingillum - LIEFTINCK, Ent. Mitt. Zool. Staatsinst. Zool. Mus. Hamburg, 21: 1. 1959 Pseudagrion papuense - LIEFTINCK, Ent. Mitt. Zool. Staatsinst. Zool. Mus. H a m b u r g , 21: 1 [proposed synonymy with cingillum (BRAUER, 1 8 6 9 ) ] .

Distribution: Australia, New Guinea. citricola

BARNARD

1937 Pseudagrion citricola BARNARD, Annls. S. Afr. Mus., 32: 212. Type male: South African Museum; type-locality: Cape. 1962 Pseudagrion citricola - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124 (Cape Province).

Distribution: South Africa.

328 civicum

LIEFTINCK

1932 Pseudagrion civicum LIEFTINCK, N o v a Guinea, 15 (5): 86. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. coarctatum

LIEFTINCK

1932 Pseudagrion coarctatum LIEFTINCK, Nova Guinea, 15 (5): 94. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. coeleste coeleste

LONGFIELD

1947 Pseudagrion coelestis LONGFIELD, Arq. Mus. Bocage, 16: 9. Type male: British Museum (Nat.Hist.), London; type-locality: Angola. 1962 Pseudagrion coeleste - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124 (Angola, Zambezi). 1984 Pseudagrion coeleste coeleste - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 86.

Distribution: West Africa. coeleste samfyae

PINHEY

1984 Pseudagrion coeleste samfyae the World, Utrecht, 1: 90.

PINHEY,

1964 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of

Distribution: Rhodesia. coeruleiceps

LONGFIELD

1959 Pseudagrion coeruleiceps LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 19. Type male: British Museum (Nat.Hist.), London; type-locality: Nigeria. 1962 Pseudagrion coeruleiceps - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124.

Distribution: Nigeria. coeruleipunctum

PINHEY

1964 Pseudagrion coeruleipunctum PINHEY, Publ. cult. Comp. Diam. Angola, 63: 103. Type female: Nat. Mus., Bulawayo; type-locality: Angola.

Distribution: Angola. commoniae commoniae

(FÖRSTER)

1902 Erythromma commoniae FÖRSTER, Mitt. Ver. Karlsruhe 15: 75. Type male: A n n A r b o r Museum; type-locality: Erythrea. 1 9 0 5 Erythromma commoniae - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 0 (Africa). 1962 Erythromma commoniae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 131 (Eritrea). 1984 Pseudagrion commoniae commoniae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 8 6 .

Distribution: Eritrea. commoniae nigerrimum

PINHEY

1950 Pseudagrion nigerrimum PINHEY, Ann. Transvaal Mus., 21:265. Type male: Transvaal Museum; type-locality: South Rhodesia. 1962 Pseudagrion nigerrimum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127 (Rhodesia, M o zambique, Kenya).

329 1984 Pseudagrion commoniae nigerrimum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 86.

Distribution: Rhodesia, Mozambique, Kenya. conspicuum

FRASER

1947 Pseudagrion conspicuum FRASER, Trans. R. ent. Soc. London; 98:27. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Ivory Coast. 1962 Pseudagrion conspicuum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124 (Ivory Coast, Belgian Congo, South Nigeria).

Distribution: West Africa. coomansi

LIEFTINCK

1937 Pseudagrion coomansi LIEFTINCK, Treubia, 16 (1): 86. Type male: Buitenzorg Museum; typelocality: Borneo. 1954 Pseudagrion coomansi - LIEFTINCK, Treubia, 22 Suppl.: 57 (Borneo, Malaya).

Distribution: Malaysia, Borneo. coriaceum

SELYS

1876 Agrion (Pseudagrion) coriaceum SELYS, Synopsis des Agrionines, Add. 5: 223. Type male: Naturhist. Mus. Wien; type-locality: Amboine. 1890 Pseudagrion coriaceum - KIRBY, Syn. Cat. Neur.-Odon., London: 153.

Distribution: Malaysia. crocops

SELYS

1876 Agrion (Pseudagrion) crocops SELYS, Synopsis des Agrionines, Add. 5: 222. Type male: McLachlan's Collection; type-locality: Celebes. 1890 Pseudagrion crocops - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1896 Pseudagrion semicolon FÖRSTER, Ent. Nachr., 22: 325. Type male: Mus. Naturk., Berlin; typelocality: Celebes. 1915 Pseudagrion crocops - Ris, Nova Guinea, 13: 97. 1939 Pseudagrion crocops - NEEDHAM & G Y G E R , Philipp. J. Sei. 70 (3): 284.

Distribution: Celebes. cyathiforme

PINHEY

1973 Pseudagrion cyathiforme PINHEY, Odonatologica, 2 (4): 317. Type male: Pinhey's Collection; type-locality: Liberia.

Distribution: Liberia. deconcertans 1984

AGUESSE

Pseudagrion deconcertans World, Utrecht, 1: 87.

AGUESSE, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Madagascar. decorum

(RAMBUR)

1842 Agrion decorum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 258. Type male: Serville's Collection; type-locality: Bombay. 1876 Agrion (Pseudagrion) decorum - SELYS, Synopsis des Agrionines, Add. 5: 212. 1890 Pseudagrion decorum - KIRBY, Syn. Cat. Neur.-Odon., London: 153.

330 1916 Pseudagrion decorum - LAIDLAW, Ree. Indian Mus., 12: 24 (India, Sumatra). 1924 Pseudagrion decorum - FRÄSER, Ree. Indian Mus., 26: 497 (West India).

Distribution: India, Sumatra. demorsum

NEEDHAM

1935 Pseudagrion demorsum NEEDHAM, Bull. Bishop Mus., 114:71. Type male: B.R Bishop Museum, Honolulu; type-locality: Marquesas Island.

Distribution: Marquesas Island, deningi PINHEY 1984 Pseudagrion deningi PINHEY, 1961 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the World,

Utrecht, 1: 87.

Distribution: North Rhodesia. digitatum

SCHMIDT

1949 Pseudagrion digitatum SCHMIDT, in FRASER, Mem Inst. sei. Madagascar, 3A: 22. Type male: Inst. Rech. sei. Madagascar; type-locality: Madagascar.

Distribution: Madagascar, dispar SCHMIDT 1949 Pseudagrion dispar SCHMIDT, in FRASER, Mem Inst. sei. Madagascar, 3A: 22. Type male: Inst. Rech. sei. Madagascar; type-locality: Madagascar. 1962 Pseudagrion dispar - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. divaricatum SCHMIDT 1951 Pseudagrion divaricatum SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 223. Type male: Inst. Rech. sei. Madagascar; type-locality: Madagascar. 1962 Pseudagrion divaricatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. dundoense

LONGFIELD

1959 Pseudagrion dundoense LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 24. Type male: not given; type-locality: Angola. 1962 Pseudagrion dundoense - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124.

Distribution: Angola, elongatum

NEEDHAM

1930 Pseudagrion elongatum NEEDHAM, Zool. Sinica, (A) 11: 261. Type male: ? Peking Museum; type-locality: China.

Distribution: China, epiphonematicum KARSCH 1891 Pseudagrion epiphonematicum KARSCH, Ent. Nachr., 17 (5): 68. Type male: Mus. Naturk., Berlin; type-locality: Cameroon. 1908 Pseudagrion epiphonematicum - MARTIN, Annali. Mus. civ. Stor. nat. Genova, 43: 654 (Fernando Po). 1936 Pseudagrion epiphonematicum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 38. 1 9 3 6 Pseudagrion epiphonematicum — L O N G F I E L D , Trans. R . ent. Soc. L o n d o n , 85: 473 (Ruwenzori).

331 1961 Pseudagrion epiphonematicum - PINHEY, Entomologist's month. Mag., 97: 103 (description of allotype). 1 9 6 2 Pseudagrion epiphonematicum - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 2 4 (Cameroon, Spanish Guinea, Portuguese Guinea, Belgian Congo, Uganda).

Distribution: tropical Africa. estesi

PINHEY

1971 Pseudagrion estesi PINHEY, Arnoldia, Rhodesia, 5 (6): 2. Type male: Pinhey's Collection; typelocality: Angola.

Distribution: Angola. evidanum

NEEDHAM &

GYGER

1939 Pseudagrion evidanum NEEDHAM & G Y G E R , Philipp. J. Sei. 70 (3): 286. Type male: not given; type-locality: Luzon.

Distribution: Philippine Islands. farinicolle

LIEFTINCK

1932 Pseudagrion farinicolle LIEFTINCK, N o v a Guinea, 15 (5): 88. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. fisheri 1984

PINHEY

Pseudagrion fisheri PINHEY, 1 9 6 1 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 87.

Distribution: North Rhodesia. flavifrons

NEEDHAM &

GYGER

1939 Pseudagrion flavifrons NEEDHAM & G Y G E R , Philipp. J. Sei. 70 (3): 285. Type male: not given; type-locality: Luzon.

Distribution: Philippine Islands. flavipes flavipes

SJÖSTEDT

1899 Pseudagrion hemicolon KARSCH, Ent. Nachr., 25 (11): 8. Type male: lost; type-locality: Cameroon. 1900 Pseudagrion flavipes SJÖSTEDT, Bih. Svenska Akad., 25 (2): 59. Type male: Riksmus. Naturhist., Stockholm; type-locality: Cameroon. 1936 Pseudagrion flavipes - SCHMIDT., Abh. senckenb. naturf. Ges., 433: 22. 1936 Pseudagrion hemicolon - SCHMIDT., Abh. senckenb. naturf. Ges., 433: 22 (proposed synonymy with flavipes (SJÖSTEDT, 1900). 1962 Pseudagrion flavipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124. 1984 Pseudagrion hemicolon - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 88 (as good species).

Distribution: Cameroon, Nigeria, Togo, Ghana. flavipes leonense

PINHEY

1964 Pseudagrion flavipes leonensis PINHEY, Bull. Inst, franc. Afr. noire, (A) 26: 1147. Type male: Mus. r. Afr. centr. Tervuren; type-locality: Sierra Leone.

Distribution: Sierra Leone.

332 furcigerum

(RAMBUR)

1842 Agrion furcigerum RAMBUR, Hist. nat. Ins. Neuropt. Paris: 261. Type male: R. Mus., Brussels; type-locality: Cape. 1876 Agrion (Pseudagrion J furcigerum - SELYS, Synopsis des Agrionines, Add. 5: 206 (as generotype of Pseudagrion SELYS, 1876) (Cape of G o o d Hope). 1890 Pseudagrion furcigerum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1921 Pseudagrion furcigerum - Ris, Annls. S. Afr. Mus., 18: 300 (Cap). 1936 Pseudagrion furcigerum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 17. 1962 Pseudagrion furcigerum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124.

Distribution: South Africa. gamblesi

PINHEY

1984 Pseudagrion gamblesi PINHEY, 1978 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 87.

Distribution: Zimbabwe. giganteum

SCHMIDT

1951 Pseudagrion giganteum SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 6A: 239. Type female: Schmidt's Collection, type-locality: Madagascar. 1962 Pseudagrion gigantuem - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar.

gigas

SCHMIDT

1936 Pseudagrion gigas SCHMIDT, Abh. senckenb. naturf. Ges., 433: 33. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Sikasso. 1962 Pseudagrion gigas - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 125 (Transvaal, Rhodesia, Mozambique, Kenya, French Guinea, North Nigeria).

Distribution: tropical Africa. glaucescens glaucescens

SELYS

1876 Agrion (Pseudagrion) glaucescens SELYS, Synopsis des Agrionines, Add. 5: 208.Type: not designated, series of both sexes in R. Mus. Brussels; type-locality: Sierra Leone. 1890 Pseudagrion glaucescens - KIRBY, Syn. Cat. Neur.-Odon. London: 153. 1917 Pseudagrion zumbense NAVÄS, Broteria, 15: 72. Type male: unknown; type-locality: Mozambique. 1936 Pseudagrion glaucescens - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 61. 1936 Pseudagrion zumbense - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 61 (proposed synonymy w i t h glaucescens

SELYS, 1 8 7 6 ) .

1962 Pseudagrion glaucescens - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 125 (French Equatorial Africa, Cameroons, Rhodesia, Mozambique, Belgian Congo, Tanganyika, Uganda, Sierra Leone, Nigeria).

Distribution: tropical Africa. glaucescens mozambicensis

PINHEY

1962 Pseudagrion mozambicensis PINHEY, Occ. Pap. nat. Mus. Rhodesia, (Β) 3: 894. Type: unknown; type-locality: Mozambique.

Distribution: Mozambique.

333 glaucescens zambesiensis 1984

PINHEY

Pseudagrion glaucescens zambesiesis of the World, Utrecht, 1: 92.

PINHEY, 1 9 6 4

- loc. cit.

D A VIES & TOBIN,

The Dragonflies

Distribution: Mozambique: Zambezi. glaucoideum

SCHMIDT

1936 Pseudagrion glaucoideum SCHMIDT, Abh. senckenb. naturf. Ges., 433: 66. Type male: Mus. Nation. Hist. N a t . Paris; type-locality: Congo. 1962 Pseudagrion glaucoideum -PINHEY, Publ. cult. Comp. Diam. Angola, 59:125 (Belgian Congo, Liberia). 1971 Pseudagrion isidromorai - PINHEY, Arnoldia, Rhodesia, 5 (6): 1 (proposed synonymy with glaucoideum

SCHMIDT, 1 9 3 6 ) .

1973 Pseudagrion glaucoideum - PINHEY, Odonatologica, 2 (4): 318. 1984 Pseudagrion isidromorai SART, 1967 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e W o r l d ,

Utrecht, 1: 88 from West Africa.

Distribution: West Africa. greeni I960

PINHEY

Pseudagrion greeni PINHEY, Entomologist's month. Mag., 96: lawayo; type-locality: South Africa.

260.

Type male: Nat. Mus. Bu-

Distribution: South Africa. grilloti

LEGRAND

1987 Pseudagrion grilloti LEGRAND, Rev. Fr. Entomol., (N.S.) 9 (2): 77. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Congo.

Distribution: Congo Republic. guichardi

KIMMINS

1958 Pseudagrion guichardi KIMMINS, Bull. Brit. Mus. (Nat.Hist.) Ent., 7: 351. Type male: British Museum (Nat.Hist.), London; type-locality: Ethiopia. 1962 Pseudagrion guichardi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 125.

Distribution: Ethiopia. hageni hageni

KARSCH

1893 Pseudagrion hageni KARSCH, Berl. ent. Z., 38: 38. Type male: Mus. Naturk., Berlin; typelocality: D.O. Africa. 1936 Pseudagrion hageni - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 39 (incorrect synonymy with angolense SELYS, 1876). 1963 Pseudagrion angolense rubridorsum BALINSKY, J. ent. Soc. S. Afr., 26: 245. Type male: unknown; type-locality: Cape.

Distribution: East and South Africa. hageni tropicanum 1984

PINHEY

Pseudagrion hageni tropicanum the World, Utrecht, 1: 91.

PINHEY, 1 9 6 6

Distribution: Central Africa.

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of

334 hamoni

FRÄSER

1955 Pseudagrion hamoni FRÄSER, Rev. franc. Ent., (22) 3: 239. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Fr. W. Africa. 1956 Pseudagrion whellani PINHEY, Occ. Pap. Coryndon Mus., 4: 18. Type male: British Museum (Nat.Hist.) London; type-locality: E. & W. Africa. 1962 Pseudagrion hamoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 125 (French Congo). 1962 Pseudagrion whellani - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129 (Rhodesia, Mozambique, Tanganyika, Kenya, Belgian Congo, Nigeria, Sudan). 1984 Pseudagrion hamoni - RAYUK & ZUBEIR, Entomologist's month, Mag., 120: 157 (Sudan).

Distribution: Central Africa. hamulum

SCHMIDT

1951 Pseudagrion hamulus SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 6A: 220. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion hamulus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. helenae helenae

BALINSKY

1964 Pseudagrion helenae BALINKSY, Novos Taxa ent., 36: 3. Type: unknown; type-locality: Bechuanaland.

Distribution: South Africa, helenae planivarum

PINHEY

1984 Pseudagrion helenae planivarum PINHEY, 1964 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of

the World, Utrecht, 1: 90.

Distribution: Zambia. hypermelas

SELYS

1876 Agrion (Pseudagrion) hypermelas SELYS, Synopsis des Agrionines, Add. 5: 229. Type male: Selys's Collection; type-locality: India. 1890 Pseudagrion hypermelas - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1 9 0 5 Pseudagrion hypermelan (misprint) - JACOBSON & BIANKI, Pryam. lozhonos. Ross, imp.: 8 3 0 . 1916 Pseudagrion hypermelas - LAIDLAW, Ree. Indian Mus., 12: 25 (India, Sumatra). 1924 Pseudagrion hypermelas - FRÄSER, Ree. Indian Mus., 26: 497 (West India). 1935 Pseudagrion hypermelas - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 11 (Sumatra).

Distribution: India, Sumatra. igniceps

FRASER

1953 Pseudagrion igniceps FRASER (nec FRASER, 1953), Nat. malgache, 5 : 163. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Pseudagrion igniceps - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Madagascar. ignifer

TILLYARD

1906 Pseudagrion ignifer TILLYARD, Proc. Linn. N. S. Wales, 31: 188. Type male: Tillyard's Collection, type-locality: Queensland. 1913 Pseudagrion ignifer - TILLYARD, Proc. Linn. N. S. Wales, 37: 469. 1917 Pseudagrion ignifer - SJÖSTEDT, Ark. Zool., Uppsala, 11 (11): 37. Distribution: A u s t r a l i a .

335 incisurum

LIEFTINCK

Pseudagrion incisurum LIEFTINCK, Treubia, 20 ( 2 ) : 3 3 6 . Type male: Rijksmus. Natuurl. Hist., Leiden, type-locality: Guadalcanal. 1984 Pseudagrion incisurum - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 88 (Solomon Island). 1949

Distribution: Pacific area. inconspicuum inconspicuum Ris 1931 Pseudagrion inconspicuum Ris, Revue Suisse zool., 38: 98. Type male: Senckenberg Musuem; type-locality: Angola. 1 9 3 6 Pseudagrion inconspicuum - S C H M I D T , A b h . senckenb. naturf. Ges., 433: 2 4 . 1962 Pseudagrion inconspicuum - PINHEY, Publ. cult. Comp. Diam. Angola, 59:125 (Cape Province, N. Rhodesia, Angola).

Distribution: Central and South Africa. inconspicuum arabicum

WATERSTON

1984 Pseudagrion inconspicuum arabicum WATERSTON, 1980 - loc. cit. gonflies of the World, Utrecht, 1: 85.

D A VIES & TOBIN,

The Dra-

Distribution: Arabia. indicum

FRASER

1924 Pseudagrion indicum FRÄSER, Ree. Indian Mus., 26: 495. Type male: British Museum (Nat.Hist.), London; type-locality: West India.

Distribution: West India. inopinatum 1984

BALINSKY

Pseudagrion inopinatum World, Utrecht, 1: 88.

BALINSKY, 1 9 7 1

- loc. cit.

D A V I E S & TOBIN,

The Dragonflies of the

D A V I E S & TOBIN,

The Dragonflies of the

Distribution: Transvaal. kaffinum 1984

CONSIGLIO

Pseudagrion kaffinum CONSIGLIO, World, Utrecht, 1: 88.

1978

- loc. cit.

Distribution: Ethiopia. kersteni

(GERSTAECKER)

1869 Agrion kersteni GERTAECKER, Arch. f. nat., 35 (1): 222. Type male: Mus. N a t u r k . Berlin; type-locality: Cape of G o o d Hope. 1869 Agrion deckeni GERSTAECKER, Arch. f. nat., 35 (1): 222. Type male: Mus. Naturk. Berlin; type-locality: Togo. 1 8 7 6 Agrion (Pseudagrion) praetextatum SELYS ( H Ä G E N M S ) , Synopsis des Agrionines, Add. 5 : 204. Type male: Selys's Collection; type-locality: Zanzibar. 1890 Coenagrion kersteni - K I R B Y , Syn. Cat. Neur.-Odon., London: 151. 1890 Coenagrion deckeni - K I R B Y , Syn. Cat. Neur.-Odon., London: 150. 1890 Pseudagrion praetextatum - K I R B Y , Syn. Cat. Neur.-Odon., London: 153. 1898 Pseudagrion deckeni - K I R B Y , Ann. Mag. nat. Hist., (7) 2: 244 (Pretoria, Nyasaland). 1 9 0 5 Pseudagrion praetextatum praetextatum - JACOBSON & BIANKI - Pryam. lozhos. Ross, imp.: 830.

1906 Pseudagrion erythraceum

FÖRSTER,

Jb. Ver. Natk. Mannheim,

71-72: 1

(nom.nud.) - lost.

336 1909 Pseudagrion kilimandjaricum SJÖSTEDT, Kilimandjaro-Meru Exp., 14: 47. Type male: Naturhist. Riksmus., Stockholm; type-locality: Tanzania. 1912 Pseudagrion kersteni - M A R T I N , Fenill. jeun. Nat., ( 5 ) 4 2 : 99. 1921 Pseudagrion praetextatum - Ris, Annls S. Afr. Mus., 18: 303 (Zululand, Transvaal, Nyassa, Natal). 1936 Pseudagrion kersteni - S C H M I D T , Abh. senckenb. naturf. Ges., 4 3 3 : 18. 1936 Agrion deckeni - S C H M I D T , Abh. senckenb. naturf. Ges., 433: 18 [proposed synonymy with kersteni (GERSTAECKER, 1869)]. 1936 Agrion (Pseudagrion) praetextatum - S C H M I D T , Abh. senckenb. naturf. Ges., 433:18 [proposed synonymy with kersteni (GERSTAECKER, 1869)]. 1 9 3 6 Pseudagrion erythraceum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 1 8 [proposed synonymy with kersteni (GERSTAECKER, 1 8 6 9 ) ] . 1 9 3 6 Pseudagrion kilimanjaricum - S C H M I D T , Abh. senckenb. naturf. Ges., 433: 1 8 [proposed synonymy with kersteni (GERSTAECKER, 1 8 6 9 ) ] .

Distribution: East Africa, South African Republic. kibalense

LONGFIELD

1959 Pseudagrion kibalense LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 22. Type male: not given; type-locality: Uganda. 1 9 6 2 Pseudagrion kibalense - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 2 6 .

Distribution: Uganda. laidlawi

FRASER

1922 Pseudagrion laidlawi FRASER, Mem. Dept. Agr. India, 7: 48. Type male: originally in Fraser's Collection, now unknown; type-locality: India.

Distribution: India. lindicum

GRÜNBERG

1902 Pseudagrion lindicum G R Ü N B E R G , Sber Ges. Naturf. Freunde Berl., 9: 230. Type male: Mus. Naturk., Berlin; type-locality: Lindi. 1936 Pseudagrion lindicum - S C H M I D T , Abh. senckenb. naturf. Ges., 433: 64 (Zanzibar). 1962 Pseudagrion lindicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 126 (Belgian Congo, Tanganyika, Kenya, Somalia).

Distribution: tropical Africa. lucidum

SCHMIDT

1951 Pseudagrion lucidum SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 6A: 255. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion lucidum - P I N H E Y , Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. macrolucidum

AGUESE

1984 Pseudagrion macrolucidum World, Utrecht, 1: 89.

AGUESE,

1968 - loc. cit.

D A V I E S & TOBIN,

The Dragonflies of the

Distribution: Madagascar. makabusiense

PINHEY

1950 Pseudagrion makabusiensis P I N H E Y , Ann. Transvaal Mus., 21: 263. Type male: Transvaal Museum; type-locality: South Rhodesia.

337 1962 Pseudagrion makabusiensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 126. 1987 Pseudagrion makabusiensis - LEGRAND, Rev. Ft. Entomol. (N.S.) 9 (2): 79.

Distribution: Rhodesia. malabaricum

FRASER

1924 Pseudagrion malabaricum FRÄSER, Ree. Indian Mus., 26: 494. Type male: British Museum (Nat.Hist.), London; type-locality: West India. 1955 Pseudagrion malabaricum - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 73 (Ceylon).

Distribution: Malabar, Sri Lanka. malagasoides

PINHEY

1973 Pseudagrion malagasoides PINHEY, Odonatologica, 2 (4): 321. Type male: unknown; type-locality: Liberia.

Distribution: Liberia, malgassicum

SCHMIDT

1949 Pseudagrion malgassicum SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 3A: 22. Type male: not given; type-locality: Madagascar. 1962 Pseudagrion malgassicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. massaicum

SJÖSTEDT

1909 Pseudagrion punctatum massaicum SJÖSTEDT, Wiss. Ergebn. Exp. Kilimandjaro-Meru, Odon.: 48. Type male: Naturhist. Riksmus., Stockholm; type-locality: Tanzania. 1915 Pseudagrion massaicum - Ris, Ent. Mitteil., 4: 141. 1921 Pseudagrion massaicum - Ris, Annls S. Afr. Mus., 18: 310 (Zuzuland, Caffraria) (redescription). 1936 Pseudagrion massaicum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 50. 1936 Pseudagrion massaicum - LONGFIELD, Trans. R. ent. Soc. London, 85: 473 (Uganda, Belgian Congo). 1937 Pseudagrion massaicum var. cogmani BARNARD, Ann. S. Afr. Mus., 32: 215. Male from Cape. 1941 Pseudagrion massaicum - ST. QUENTIN, Ann. Naturh. Mus., Wien, 52: 109 (Kilimandjaro). 1951 Pseudagrion massaicum - FRASER, Mem. Inst. sei. Madagascar, 6A: 411 (Madagascar). 1962 Pseudagrion massaicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 126 (Cape to Kenya, Uganda, Angola, Belgian Congo, Sudan, Guinea, Comoro Island). 1 9 8 4 Pseudagrion chomae PINHEY, 1 9 6 4 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 86. 1984 Pseudagrion cogmani - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 86.

Distribution: tropical Africa, Madagascar, and Comoro Island. melanicterum

SELYS

1876 Agrion (Pseudagrion) melanicterum SELYS, Synopsis des Agrionines, Add. 5: 202. Type: not designated, males and females series in R. Mus. Brussels; type-locality: Sierra Leone. 1890 Pseudagrion melanicterum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1893 Pseudagrion emarginatum KARSCH, Berl. ent. Z., 38: 39. Type male: Mus. Naturk., Berlin; type-locality: Togo. 1908 Pseudagrion melanicterum - MARTIN, Ann. Mus. civ. Stor. nat. Genova, 43:654 (Fernando Po). 1936 Pseudagrion melanicterum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 43. 1936 Pseudagrion emarginatum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 43 [proposed synonymy with melanicterum (SELYS, 1876)]. 1959 Pseudagrion melanicterum - LONGFIELD, Publ. cult. Comp. Diam. Angola, 45: 17 (Angola).

338 1962 Pseudagrion melanicterum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 126 (Rhodesia, Belgian Congo, Uganda, Cameroon, Nigeria, Liberia, French Equatorial Guinea, Togo, Dahomey, Sierra Leone, Dakar. 1987 Pseudagrion melanicterum

- LEGRAND, Rev. Fr. Entomol., (N.S.) 9 (2): 81.

Distribution: tropical Africa. mellisi

SCHMIDT

1951 Pseudagrion mellisi SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 6A: 231. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion mellisi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. merina

SCHMIDT

1949 Pseudagrion merina SCHMIDT, in FRASER, Mem. Inst. sei. Madagascar, 3A: 22. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion merina - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. microcephalum

(RAMBUR)

1842 Agrion microcephalum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 259. Type male: Guerin's Collection; type-locality: Bombay. 1876 Agrion (Pseudagrion) microcephalum - SELYS, Synopsis des Agrionines, Add. 5: 214 (Bombay, Singapore, Luzon, Java). 1890 Pseudagrion microcephalum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1916 Pseudagrion microcephalum - Ris, Suppl. Ent., 5: 40 (Queensland). 1916 Pseudagrion microcephalum - LAIDLAW, Ree. Indian Mus., 12: 23 (India, Burma, Tonkin to the Bismarck Archipelago). 1917 Pseudagrion microcephalum - SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 37 (Queensland). 1924 Pseudagrion microcephalum - FRASER, Ree. Indian Mus., 26: 497 (West India). 1931 Pseudagrion microcephalum - CHUJO, Trans, nat. Hist. Soc. Formosa, 21 (112): 40 (Formosa). 1934 Pseudagrion microcephalum - SCHMIDT, Arch. Hydrob. Suppl.: 13: 349 (Java, Sumatra). 1934 Pseudagrion microcephalum - LIEFTINCK, Treubia, 14 (4): 396 (Java). 1935 Pseudagrion microcephalum - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 1 1 . 1936 Pseudagrion microcephalum - LIEFTINCK, Revue suisse Zool., 43 (5): 122. 1939 Pseudagrion microcephalum - NEEDHAM & G Y G E R , Philipp. J. Sei. 70 (3): 284 (Philippines). 1949 Pseudagrion microcephalum - LIEFTINCK, Treubia, 20 (2): 338 (redescription) (Solomon Islands). 1954 Pseudagrion microcephalum - LIEFTINCK, Treubia, 22 Suppl.: 57 (Malaya, Sumatra, Krakatau, Java, Bali, Borneo).

Distribution: Oriental region, and Queensland. mohelii 1984

AGUESE

Pseudagrion mohelii Utrecht, 1: 89.

AGUESE, 1 9 6 8

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Madagascar. newtoni

PINHEY

1962 Pseudagrion newtoni PINHEY, Oec. Pap. nat. Mus. S. Rhodesia, (B) 3: 895. Type male: S. Rhodesia Museum; type-locality: Central Africa.

Distribution: Rhodesia.

339 nigripes

SCHMIDT

1951 Pseudagrion nigripes SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 242. Type male: Schmidt's Colletion; type-locality: Madagascar. 1962 Pseudagrion nigripes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. nigrofasciatum

LIEFTINCK

1934 Pseudagrion nigrofasciatum LIEFTINCK, Stylops, 3: 6. Type male: Buitenzorg Museum; typelocality: Java. 1934 Pseudagrion infracavum SCHMIDT, Arch. Hydrob. Suppl.: 13: 350. Type male: Mus. Naturk., Berlin; type-locality: Java. 1954 Pseudagrion nigrofasciatum - LIEFTINCK, Treubia, 22 Suppl.: 58.

Distribution: Java. niloticum 1984

DUMONT

Pseudagrion niloticum Mag., 120: 1 5 7 .

DUMONT, 1978

- loc. cit.

R A Y A H & ZUBEIR,

Entomologist's month.

Distribution: Sudan. nubicum

SELYS

1876 Agrion (Pseudagrion) nubicum SELYS, Synopsis des Agrionines, Add. 5: 211. Type: not designates, 2 males in R. Mus. Brussels; type-locality: Nubia. 1890 Pseudagrion nubicum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1900 Coenagrion glaucum SJÖSTEDT, Bih. Svenska Akad., 25 (2): 61. Type male: Naturhist. Riksmus., Stockholm; type-locality: Cameroon. 1 9 0 5 Pseudagrion nubicum - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 0 (Nubia). 1928 Pseudagrion nubicum - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 23 (Egypt, Sudan). 1936 Pseudagrion nubicum - LONGFIELD, Trans. R. ent. Soc. London, 85: 473 (Congo). 1962 Pseudagrion nubicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127 (Congo, Rhodesia, Nyasaland, Kenya, Sudan, Cameroon, French Guinea, Sierra Leone, Egypt). 1962 Coenagrion glaucum — PINHEY, Publ. cult. Comp. Diam. Angola, 59:127 (proposed synonymy with nubicum SELYS, 1 8 7 6 ) .

Distribution: from Egypt to Rhodesia. olsufieffi

SCHMIDT

1951 Pseudagrion olsufieffi SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 223. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion olsufieffi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. orientate 1984

BELYSHEV

Pseudagrion Orientale BELYSHEV, World, Utrecht, 1: 89.

1973

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Siberia. pacificum

T I L L YARD

1923 Pseudagrion pacificum TILLYARD, Trans, ent. Soc. London, 1923: 311. Type male: Cawthron Inst. Coll.; type-locality: Fiji.

Distribution: Fiji.

340 palauense LIEFTINCK 1962 Pseudagrion palauense LIEFTINCK, Insects of Micronesia, 5 (1): 36. Type male: Kyushu Univ.; type-locality: Eastern Caroline Island.

Distribution: Caroline Island. pelecotomum LIEFTINCK 1932 Pseudagrion pelecotomum LIEFTINCK, Nova Guinea, 15 (5): 93. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. perfuscatum LIEFTINCK 1937 Pseudagrion perfuscatum LIEFTINCK, Treubia, 16 (1): 94. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Pseudagrion perfuscatum - LIEFTINCK, Treubia, 22 Suppl.: 58 (Borneo).

Distribution: Borneo. pilidorsum pilidorsum (BRAUER) 1868 Agrion pilidorsum BRAUER, Verh. Zool.-Bot. Ges., Wien, 18: 553, Type male: ? Naturhist. Mus., Wien, type-locality: Luzon. 1876 Agrion (Pseudagrion) pilidorsum - SELYS Synopsis des Agrionines, Add. 5: 224 (Luzon, Manila). 1890 Pseudagrion pilidorsum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1916 Pseudagrion pilidorsum - Ris, Suppl. ent. Berlin, 5: 43 (Formosa). 1936 Pseudagrion pilidorsum pilidorsum - LIEFTINCK, Revue suisse Zool., 43 (5): 127 (key to subspecies) (Philippine Islands, Formosa). 1939 Pseudagrion pilidorsum - NEEDHAM & G Y G E R , Philipp. J. Sei. 70 (3). 281. 1954 Pseudagrion pilidorsum pilidorsum - LIEFTINCK, Treubia, 22 Suppl.: 58 (Luzon: Manila, Mindanao).

Distribution: Philippine Islands, Taiwan. pilidorsum declaratum LIEFTINCK 1936 Pseudagrion pilidorsum declaratum LIEFTINCK, Revue suisse Zool., 43 (5): 132. Type male: Buitenzorg Museum; type-locality: Lombok.

Distribution: Lombok. pilidorsum deflexum LIEFTINCK 1936 Pseudagrion pilidorsum deflexum LIEFTINCK, Revue suisse Zool., 43 (5). 134. Type male: Buitenzorg Museum; type-locality: Timor.

Distribution: Timor. pilidorsum enganoense LIEFTINCK 1948 Pseudagrion enganoense LIEFTINCK, Treubia, 19 (2): 290. Type male: ? Buitenzorg Museum; type-locality: Engano Island. 1954 Pseudagrion pilidorsum enganoense - LIEFTINCK, Treubia, 22 Suppl.: 58.

Distribution: Engano Island. pilidorsum obscurum LIEFTINCK 1936 Pseudagrion pilidorsum obscurum LIEFTINCK, Revue suisse Zool., 43 (5): 131. Type male: Buitenzorg Museum; type-locality: Nias.

341 1948 Pseudagrion pilidorsum 1954 Pseudagrion pilidorsum

obscurum - LIEFTINCK, Treubia, 19: 284. obscurum - LIEFTINCK, Treubia, 22 Suppl.: 58 (Nias).

Distribution: Indonesia: Nias Island. pilidorsum simalurum

LIEFTINCK

1948 Pseudagrion pilidorsum simalurum LIEFTINCK, Treubia, 19 (2): 289. Type male: ? Buitenzorg Museum; type-locality: Simalur. 1954 Pseudagrion pilidorsum simalurum - LIEFTINCK, Treubia, 22 Suppl.: 59 (Simalur, Mentawei I.).

Distribution: Simalur, Mentawei Island. pontogenes Ris 1915 Pseudagrion pontogenes Ris, Ent. Mitt., 4:140. Type male: A n n A r b o r Museum; type-locality: Mayotte. 1936 Pseudagrion pontogenes - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 23. 1962 Pseudagrion pontogenes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Comoro Island. pruinosum pruinosum

(BURMEISTER)

1839 Agrion pruinosum (DE H A A N MS) BURMEISTER, H a n d b . Ent., 2: 821. Type male: Hoffmannsegg's Collection; type-locality: Java. 1876 Agrion (Pseudagrion) pruinosum - SELYS, Synopsis des Agrionines, Add. 5: 228. 1890 Pseudagrion pruinosum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1898 Agrion pruinosum - CALVERT, Trans. Am. ent. Soc., 25: 41. 1927 Pseudagrion pruinosum - Ris, Zool. Meded., Leiden, 10 (1): 23 (Sumatra). 1934 Pseudagrion pruinosum pruinosum - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 347 (key to subspecies). 1935 Pseudagrion pruinosum pruinosum - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 11 (Sumatra). 1954 Pseudagrion pruinosum pruinosum - LIEFTINCK, Treubia, 22 Suppl.: 59 (Sumatra, Java, Bali).

Distribution: Java, Sumatra, Bali. pruinosum ranauense

SCHMIDT

1934 Pseudagrion pruinosum ranauense SCHMIDT, Arch. Hydrobiol. Suppl., 13: 348. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1934 Pseudagrion pruinosum fraseri SCHMIDT, Arch. Hydrobiol. Suppl., 13: 348. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1 9 3 5 Pseudagrion pruinosum fraseri - LIEFTINCK Misc. Zool. Sum., 9 2 - 9 3 : 11 (roposed synonymy with pruinosum ranauense SCHMIDT, 1 9 3 4 ) . 1954 Pseudagrion pruinosum ranauense LIEFTINCK, Treubia, 22 Suppl.: 59 (Siam, Malaya, Sumatra).

Distribution: from Thailand to Sumatra. pterauratum 1984

AGUESSE

Pseudagrion pterauratum World, Utrecht, 1: 90.

AGUESSE, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Madagascar. punctum 1842 1862 1876 1890 1905

(RAMBUR)

Agrion punctum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 260. Type, sex and locality not given. Agrion punctum - SELYS, in MAILLARD, Reunion, 2: 33 (Reunion, Mauritius, Cap). Agrion (Agrion) punctum - SELYS, Synopsis des Agrionines, Add. 5: 181 (Mauritius). Coenagrion punctum - KIRBY, Syn. Cat. Neur.-Odon., London: 150. Pseudagrion punctum - KIRBY, Ann. Mag. nat. Hist., (7) 15: 191 (Salibury).

342 1936 Pseudagrion punctum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 47. 1949 Pseudagrion punctum - FRÄSER, Trans. R. ent. Soc. London, 100: 138 (Mauritius). 1962 Pseudagrion punctum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130. (type in: R. Mus., Brussels) (Madagascar, Mauritius, Reunion).

Distribution: Madagascar, Mauritius, Reunion, ? Cap, and Zimbabwe. quadrioculatum 1984

PINHEY

Pseudagrion quadrioculatum World, Utrecht, 1: 90.

PINHEY, 1 9 6 4

- loc. cit

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Congo. renaudi

FRASER

1953 Pseudagrion renaudi FRASER, Nat. malgache, 5: 162. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Pseudagrion renaudi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. risi

SCHMIDT

1936 Pseudagrion risi SCHMIDT, Abh. senckenb. naturf. Ges., 433: 34. Type male: Dahlem Museum; type-locality: Cameroon. 1941 Paulinagrion platystictoides FRASER, Proc. R. ent. Soc. London (B) 10: 37. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Cameroon (as generotype of Paulinagrion FRASER, 1941). 1962 Pseudagrion risi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127. 1962 Pseudagrion platystictoides - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 113.

Distribution: West Africa. rubiceps rubiceps

SELYS

1876 Pseudagrion rubiceps locality: Java. 1916 Pseudagrion rubiceps 1919 Pseudagrion rubiceps 1924 Pseudagrion rubiceps 1930 Pseudagrion rubiceps 1936 Pseudagrion rubiceps 1954 Pseudagrion rubiceps 1965 Pseudagrion rubiceps

SELYS, Bull. Acad. r. Belg. 42: 510. Type male: Selys's Collection; type-

LAIDLAW, Ree. Indian Mus., 12: 24 (India, Burma, Java). LAIDLAW, Ree. Indian Mus., 16: 193. FRASER, Ree. Indian Mus., 26: 497 (West India). Ris, Ark. Zool. Uppsala, 21A (31): 20 (India, Sumatra). LIEFTINCK, Revue suisse Zool., 43 (5): 122 (Flores). LIEFTINCK, Treubia, 22 Suppl.: 60 (Malaya, Sumatra, Java). ASAHINA, Kontyü, 33 (4): 495 (Hong-Kong).

Distribution: from Philippine Islands to Java. rubiceps ceylonicum

KIRBY

1891 Pseudagrion ceylonica, KIRBY, Proc. Zool. Soc. London, 1891: 205. Type female: British M u seum (Nat.Hist.), London; type-locality: Ceylon. 1919 Archibasis ceylonica - LAIDLAW, Ree. Indian Mus., 16: 194. 1930 Archibasis ceylonica - Ris, Ark. Zool. Uppsala, 21A (31): 30 (Ceylon). 1955 Pseudagrion rubiceps ceylonicum - LIEFTINCK, Zool. Meded., Leiden,34 (5): 74 (Ceylon).

Distribution: Sri Lanka. rufocinctum 1956

PINHEY

Pseudagrion rufocinctum, PINHEY, OCC. Pap. Cornyndon Mus., 4: seum (Nat.Hist.), London; type-locality: Uganda.

24.

Type male: British M u -

343 1962 Pseudagrion rufocinctum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127.

Distribution: Uganda. rufostigma

LONGFIELD

1947 Pseudagrion rufostigma LONGFIELD, Arq. Mus. Bocage, 16: 11. Type male: British Museum (Nat.Hist.), London; type-locality: Angola. 1960 Pseudagrion rufostigma - PINHEY, Entomologist's month. Mag., 96: 260. 1962 Pseudagrion rufostigma - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127 (Angola, N. Rhodesia).

Distribution: North Rhodesia, Angola. salisburyense Ris 1921 Pseudagrion salisburyense Ris, Annls S. Afr. Mus., 18: 306. Type male: Cape Town Museum; type-locality: South Africa. 1936 Pseudagrion salisburyense - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 25. 1 9 6 2 Pseudagrion salisburyense - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 2 8 (Cape Province, Belgian Congo, Angola, Somalia). 1 9 8 4 Pseudagrion salisburyense f. chongwe PINHEY, 1 9 6 1 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 86 - from North Rhodesia.

Distribution: tropical Africa. samoense

FRASER

1924 Pseudagrion samoensis FRASER, Trans, ent. Soc. London; 1924:430. Type male: British Museum (Nat.Hist.), London; type-locality: Samoa.

Distribution: Samoa. schmidtianum

LIEFTINCK

1936 Pseudagrion schmidtianum LIEFTINCK, Revue suisse zool., 43 (5): 122. Type male: Mus. Naturk., Berlin; type-locality: Timor.

Distribution: Timor. serrulatum

KARSCH

1894 Pseudagrion serrulatum K A R S C H Berlin ent. Z., 39 (1): 16. Type male: Mus. Naturk., Berlin; type-locality: Cameroon. 1936 Pseudagrion serrulatum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 46. 1956 Pseudagrion superbum FRASER, Revue Zool.-Bot. afr., 54: 382. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1962 Pseudagrion serrulatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59:128 (Cameroon, Uganda, Belgian Congo).

Distribution: Cameroon, Belgian Congo, and Uganda, seyrigi

SCHMIDT 1951 Pseudagrion seyrigi SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 260. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion seyrigi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar.

344 siamense FRÄSER 1922 Pseudagrion siamensis FRÄSER, J . nat. Hist. Soc. Siam, 4: 235. Type: originally in Coll. Fraser, now unknown; type-locality: Bangkok.

Distribution: Thailand. silaceum LIEFTINCK 1932 Pseudagrion silaceum LIEFTINCK, Nova Guinea, 15 (5): 84. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea. simile SCHMIDT 1951 Pseudagrion simile SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 233. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion simile - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. simonae LEGRAND 1987 Pseudagrion simonae LEGRAND, Rev. Fr. Entomol., (N.S.) 9 (2): 79. Type male: Legrand's Collection; type-locality: Congo.

Distribution: Congo Republic. sjoestedti sjoestedti FÖRSTER 1906 Pseudagrion sjöstedti FÖRSTER, Jb Mannheim Ver. Natk., 71-72: 62. Type male: Förster's Collection; type-locality: Cameroon. 1915 Pseudagrion sjöstedti - Roi, Zentr. Afr. Exp., Zool.: 337 (Sudan). 1921 Pseudagrion sjöstedti - Ris, Annls S. Afr. Mus., 18: 312 (Ivory Coast, Congo). 1936 Pseudagrion sjöstedti - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 57. 1960 Pseudagrion greigi PINHEY, Entomologist's month. Mag. 96: 259. Type male: Nat. Mus., Bulawayo; type-locality: South Africa. 1974 Pseudagrion sjoestedti - GAMBLES, Entomologist's month. Mag., 110: 117 (allotype designated).

Distribution: tropical Africa. sjoestedti jacksoni PINHEY 1961 Pseudagrion jacksoni PINHEY, Publ. Brit. Mus., (Nat.Hist.), Ent: 37. Type male: British Museum (Nat.Hist.), London; type-locality: Uganda. 1962 Pseudagrion sjoestedti jacksoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 125.

Distribution: Uganda. sjoestedti nigeriense GAMBLES 1974 Pseudagrion sjoestedti nigeriense GAMBLES, Entomologist's month. Mag., 110: 118. Type male: British Museum (Nat.Hist.), London; type-locality: Nigeria.

Distribution: Nigeria. sjoestedti pseudosjoestedti PINHEY 1984 Pseudagrion sjoestedti pseudosjoestedti PINHEY, 1964 - loc. cit. DAVIES & TOBIN, T h e D r a -

gonflies of the World, Utrecht, 1: 90.

Distribution: East Africa.

345 sjoestedti wittei

FRÄSER

1949 Pseudagrion wittei FRÄSER, Expl. Pare nat. Albert Miss, de Witte, fasc. 61: 7. Type male: Inst. Nat. Pares C.B.; type-locality: Belgian Congo. 1962 Pseudagrion wittei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129.

Distribution: Congo Republic. speneei

FRASER

1922 Pseudagrion spencei FRASER, Mem. Dept. Agr. India, 7: 47. Type female: originally in Pusa Collection, now unknown; type-locality: Bihar.

Distribution: India. spernatum spernatum

SELYS

1881 Pseudagrion spernatum SELYS, Annali Mus. civ. Stor. nat. Genova, 16: 223. Type male: R. Mus. Brussels; type-locality: Cap. 1936 Pseudagrion spernatum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 27. 1962 Pseudagrion spernatum - PINHEY, Publ. cult Comp. Diam. Angola, 59: 128 ("type-locality: Abyssinia") (Abyssinia, Belgian Congo, Uganda, Tanganyika, N. Rhodesia).

Distribution: East and South Africa. spernatum gerstaeckeri

KARSCH

1899 Pseudagrion gerstaeckeri KARSCH, Ent. Nachr., 25 (24): 379. Type male: Mus. Naturk. Berlin; type-locality: Zanzibar. 1936 Pseudagrion gerstaeckeri - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 29. 1958 Pseudagrion gerstaeckeri - PINHEY, Stuttg. Beitr. Naturk., 1958 (10): 1 (East and West Africa). 1962 Pseudagrion gerstaeckeri - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 124 (Tanganyika, Kenya, Uganda, Angola, Belgian Congo).

Distribution: tropical Africa. spernatum natalense Ris 1921 Pseudagrion natalense Ris, Annls S. Afr. Mus., 18: 307. Type male: ? S. Afr. Museum; typelocality: South Africa. 1936 Pseudagrion natalense - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 32. 1962 Pseudagrion natalense - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127 (Rhodesia, South Africa).

Distribution: South Africa, Rhodesia. spinithoracicum

LEGRAND

1984 Pseudagrion spinithoracicum LEGRAND, 1981 - loc. cit. DAVIES & TOBIN, The Dragonflies of

the World, Utrecht, 1: 91.

Distribution: West Africa. starreanum

LIEFTINCK

1949 Pseudagrion starreanum LIEFTINCK, Nova Guinea (N.S.), 5: 181. Type male: Lieftinck's Collection; type-locality: New Guinea.

Distribution: New Guinea.

346

stuckenbergi 1984

PINHEY

Pseudagrion stuckenbergi World, Utrecht, 1: 91.

PINHEY, 1 9 6 4

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Madagascar. symoensi

PINHEY

1984 Pseudagrion symoensi Utrecht, 1: 91.

PINHEY,

1967 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Rhodesia. sublacteum

(KARSCH)

1893 Coenagrion sublacteum KARSCH, Berl. ent. Z., 38:40. Type male: lost; type-locality: West Africa. 1936 Pseudagrion mortoni SCHMIDT, Abh. senckenb. naturf. Ges., 433: 55. Type male: Schmidt's Collection; type-locality: Palestine. 1938 Pseudagrion mortoni - SCHMIDT, SitzBer. Akad. Wiss. Wien (1) 147: 146. 1951 Pseudagrion pseudomassaicum PINHEY, Mem. Transvaal Mus., 5: 93. Type male: Transvaal Museum; type-locality: Natal. 1956 Pseudagrion pseudomassaicum f. rusingae PINHEY, Occ. Pap. Coryndon Mus., 4: 22. Male from Uganda. 1960 Pseudagrion pseudomassaicum doualae PINHEY, Entomologist's month. Mag. 96: 260. Type male: Nat. Mus. Bulawayo; type-locality: South Africa. 1962 Pseudagrion sublacteum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 128. 1962 Pseudagrion mortoni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: [proposed synonymy with sublacteum ( K A R S C H , 1 8 9 3 ) ] . 1962 Pseudagrion pseudomassaicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127. (South Africa, Mozambique, Rhodesia, Tanganyika, Kenya, Uganda, Nigeria, Congo). 1973 Pseudagrion pseudomassaicum - DUMONT, Israel J. Zool., 22 ( 2 - 4 ) : 176 [proposed synonymy with sublacteum (KARSCH, 1893)].

Distribution: Asia Minor to tropical and South Africa. sudanicum sudanicum LE ROI 1915 Pseudagrion sudanicum LE ROI, Ergebn. 2. Deutsch. Zentral-Afr., Esp., 1: 337. Type male: unknown; type-locality: Sudan. 1936 Pseudagrion sudanicum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 60. 1962 Pseudagrion sudanicum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 128. 1984 Pseudagrion sudanicum sudanicum - RAYAH & ZUBEIR, Entomologist's month. Mag., 120: 157.

Distribution: Sudan, sudanicum rubroviridis

PINHEY

1956 Pseudagrion rubroviridis PINHEY, OCC. Pap. Coryndon Mus., 4: 23. Type male: British Museum (Nat.Hist.), London; type-locality: Victoria Falls. 1962 Pseudagrion rubroviridis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 127 (Uganda, Zambezi). 1984 Pseudagrion sudanicum rubroviridis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 91.

Distribution: Uganda to Zambezi. syriacum

SELYS

1887 Pseudagrion praetextum race syriacum SELYS, Ann. Soc. Ent. Belg., 31: 47. Male from Beyrut. 1890 Pseudagrion syriacum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1 9 0 5 Pseudagrion

praetextum

syriacum

- JACOBSON & BIANKI, P r y a m . l o z h n o s . R o s s , i m p . : 8 3 0 .

347 1984 Pseudagrion kersteni syriacum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:91.

Distribution: Syria, Lebanon, Israel, Jordan, thenartum

FRASER 1955 Pseudagrion thenartum FRASER, Revue Zool.-Bot. Afr., 52: 17. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1962 Pseudagrion thenartum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 128.

Distribution: Congo Republic. tinctipennis

FRASER

1951 Pseudagrion tinctipennis FRASER, Mem. Inst. sei. Madagascar, 6A: 407. Type male: Inst. sei. Madagascar Rech.; type-locality: Madagascar. 1962 Pseudagrion tinctipennis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. torridum torridum

SELYS

1876 Agrion (Pseudagrion) torridum SELYS, Synopsis des Agrionines, Add. 5: 210. Type male: not given; type-locality: Senegal. 1890 Pseudagrion torridum - KIRBY, Syn. Cat. Neur.-Odon., London: 153. 1928 Pseudagrion torridum - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 23 (Egypt). 1936 Pseudagrion torridum - SCHMIDT, Abh. senckenb. naturf. Ges., 433: 58 (Abyssinia, Egypt, Sudan, Congo, Sansibar). 1949 Pseudagrion torridum f. orientis SCHMIDT, Arq. Mus. Bocage, 20: 155. Without locality. 1962 Pseudagrion torridum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 128 ("type female: R. Mus. Brussels") (Nigeria, Egypt, Sudan, Nubia, Kenya, Uganda, Abyssinia, Senegal, French Guinea, Zanzibar).

Distribution: tropical Africa, torridum hulae

DUMONT

1973 Pseudagrion torridum hulae DUMONT, Israel J. Zool., 22 (2-4): 189. Type male: Dumont's Collection; type-locality: Israel.

Distribution: Israel. tricornis 1984

PINHEY

Pseudagrion tricornis Utrecht, 1: 91.

PINHEY, 1 9 6 7

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Rhodesia. trigonale

SCHMIDT

1951 Pseudagrion trigonale SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 241. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Pseudagrion trigonale - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. umsingaziense

BALINSKY

1963 Pseudagrion umsingaziense Mus.; type-locality: Natal.

BALINSKY, J .

Distribution: South Africa.

ent. Soc. S. Afr.,

26:

237. Type male: South Afr.

348 ungulatum

FRÄSER

1951 Pseudagrion ungulatum FRÄSER, Mem. Inst. sei. Madagascar, 6A: 409. Type male: Inst. Rech. Sei. Madagascar; type-locality: Madagascar. 1962 Pseudagrion ungulatum - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 130.

Distribution: Madagascar. ustum SELYS 1876 Agrion (Pseudagrion) ustum SELYS, Synopsis des Agrionines, A d d . 5: 225. Type female: Selys's Collection; type-locality: Malaysia. 1890 Pseudagrion ustum - KIRBY, Syn. Cat. Neur.-Odon., London: 153.

Distribution: Malaysia. vaalense

CHUTTER

1963 Pseudagrion vaalense CHUTTER, Rev. Biol., 3: 173. Type male: unknown; type-locality: Transvaal.

Distribution: South Africa. vakoanae 1984

AGUESSE

Pseudagrion vakoanae Utrecht, 1: 91.

AGUESSE, 1 9 6 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Madagascar. vansomereni PINHEY 1961 Pseudagrion vansomereni PINHEY, Publ. Mus. Brit. (Nat.Hist.) Ent.: 38. Type male: British Museum (Nat.Hist.), London; type-locality: Uganda. 1962 Pseudagrion vansomereni - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 129. 1984 Pseudagrion sudanicum f. vansomereni - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 91.

Distribution: Uganda. vumbaense 1963

BALINSKY

Pseudagrion vumbaense BALINSKY, type-locality: Rhodesia.

J.

ent. Soc. S. Afr., 26:

228.

Type male: South Afr. Mus.;

Distribution: Rhodesia. williamsi 1964

PINHEY

Pseudagrion williamsi PINHEY, locality: N o r t h Rhodesia.

J.

ent. Soc. S. Afr., 27:

327.

Type male: South Afr. Mus.; type-

Distribution: North Rhodesia. williamsoni FRÄSER 1922 Pseudagrion williamsoni FRASER, Mem. Dept. Agr. India, 7: 46. Type male: British Museum (Nat.Hist.), London (ex Pusa Collection); type-locality: Burma.

Distribution: Burma. Genus Seychellibasis

KENNEDY

1920 Seychellibasis KENNEDY, Ohio J. Sei. Columbus, 21: 88. Type-species: Telebasis alluaudi MARTIN, 1 8 9 6 (original designation). - 1 9 5 1 Seychellibasis - SCHMIDT, Mem. Inst. sei. Madagascar,

349 (proposed synonymy with Teinobasis K I R B Y , 1 8 9 0 ) . - 1 9 6 2 Seychellibasis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 1 7 . (as syn. with Teinobasis K I R B Y , 1 8 9 0 ) . - 1 9 8 4 Seychellibasis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 2 (as good genus).

6A: 273

alluaudi alluaudi (MARTIN) 1896 Telebasis alluaudi MARTIN, Mem. Soc. zool. France, 9: 110. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Seychelles. 1920 Seychellibasis alluaudi - KENNEDY, Ohio J. Sei. Columbus, 21: 88 (as generotype of Seychellibasis K E N N E D Y , 1 9 2 0 ) . 1951 Teinobasis alluaudi alluaudi - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 274. 1962 Telebasis alluaudi alluaudi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 118.

Distribution: Seychelles. alluaudi berlandi (SCHMIDT) 1951 Teinobasis alluaudi berlandi SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 274. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Teinobasis alluaudi berlandi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 118. 1984 Seychellibasis alluaudi berlandi-DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 92.

Distribution: Madagascar. malawiensis PINHEY 1984

Sechellibasis malawiensis World, Utrecht, 1: 92.

PINHEY, 1 9 6 6 -

loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Malawi. Genus Stenagrion LAIDLAW 1915 Stenagrion LAIDLAW, Proc. zool. Soc. London, 1915: 39. Type-species: Pseudagrion dubium L A I D L A W , 1 9 1 2 (original designation). - 1 9 5 4 Stenagrion - LIEFTINCK, Treubia, 22, Suppl.: 6 2 .

dubium (LAIDLAW) 1912 Pseudagrion ? dubium LAIDLAW, J. Str. Br. R. Asiatic Soc., 63: 97. Type male: Laidlaw's Collection; type-locality: Borneo. 1915 Stenagrion dubium- LAIDLAW, Proc. zool. Soc. London, 1915: 39 (as generotype of Stenagrion LAIDLAW, 1915).

Stenagrion dubium - L A I D L A W , J . Fed. Mai. States Mus., 1954 Stenagrion dubium - LIEFTINCK, Treubia, 22, Suppl.: 62. 1934

17: 550.

Distribution: Borneo. Genus Teinobasis KIRBY 1877 Telebasis (Telebasis) SELYS, (nec SELYS, 1865), Bull. Acad. r. Belg., (2) 43: 97. Type-species: Telebasis superba SELYS, 1877 (monotypy) (with revision, key to subgenera Leptobasis, Stenobasis, Telebasis s. str.). - 1890 Teinobasis KIRBY, Syn. Cat. Neur.-Odon., London: 157 (new name for Telebasis SELYS, 1877, a junior primary homonymy of Telebasis SELYS, 1865). 1935 Teinobasis - LIEFTINCK, Nova Guinea, 17: 230 (revision, key to species). - 1939 Teinobasis - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 299 (key to species). - 1954 Teinobasis LIEFTINCK, Treubia, 22, Suppl.: 62. - 1962 Teinobasis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 1 7 . - 1962 Seychellibasis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 117 (incorrect synonymy with Teinobasis KIRBY, 1890).

350 aerides

LIEFTINCK

1962 Teinobasis aerides LIEFTINCK, Insecta of Micronesia, 5 (1): 34. Type male: US natn. Mus., Washington; type-locality: Ponape Island.

Distribution: East Caroline Islands. albula Ris 1916 Teinobasis albula Ris, Nova Guinea, 13 (2): 83. Type male: Mus. Naturk. Berlin; type-locality: New Guinea.

Distribution: New Guinea, alternans

LIEFTINCK

1935 Teinobasis alternans LIEFTINCK, Nova Guinea, Leiden, type-locality: New Guinea.

17:

260. Type male: Rijksmus. Natuurl. Hist.

Distribution: New Guinea, aluensis CAMPION 1 9 2 4 Teinobasis aluensis

C A M P I O N , Ann. Mag. nat. Hist., ( 9 ) (Nat.Hist.), London; type-locality: Solomon Islands. 1949 Teinobasis aluensis - LIEFTINCK, Treubia, 2 0 (2): 243.

14: 6 1 4 .

Type male: British Museum

Distribution: Solomon Islands. argiocnemis

LIEFTINCK

1949 Teinobasis argiocnemis LIEFTINCK, Nova Guinea (N.S.) 5: 165. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, ariel

LIEFTINCK

1962 Teinobasis ariel LIEFTINCK, Insecta of Micronesia, 5 (1): 31. Type male: US Natn. Mus. Washington; type-locality: Ponape Islands.

Distribution: East Caroline Islands. aurea

LIEFTINCK

1932 Teinobasis aurea LIEFTINCK, Nova Guinea, Leiden, type-locality: New Guinea.

15

(5): 101. Type male: Rijksmus. Natuurl. Hist.

Distribution: New Guinea, bradleyi

KIMMINS

1957 Teinobasis bradleyi KIMMINS, Bull. Brit. Mus. (Nat.Hist.), Ent.: 5: 315. Type male: British Museum (Nat.Hist.), London; type-locality: Guadalcanal.

Distribution: Guadalcanal. buwaldai

LIEFTINCK

1949 Teinobasis buwaldai LIEFTINCK, Nova Guinea (N.S.) 5: 171. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Aroe Island. D i s t r i b u t i o n : Aroe Island.

351 carolinensis

LIEFTINCK

1962 Teinobasis carolinensis LIEFTINCK, Insects of Micronesia, 5 (1): 23. Type male: US natn. Mus., Washington; type-locality: Truk.

Distribution: East Caroline Islands, combusta

(SELYS) 1877 Telebasis (Telebasis) combusta SELYS, Bull Acad. r. Belg. (2) 43: 120. Type female: Selys's Collection; type-locality: Sulu. 1 8 9 0 Teinobasis combusta - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 5 7 .

Distribution: Sulu. corolla 1939

NEEDHAM & GYGER

Teinobasis corolla N E E D H A M type-locality: Luzon.

& GYGER,

Philipp.

J.

Sei., 70

(3): 3 0 3 .

Type male: not given;

Distribution: Philippine Islands. debeauforti LIEFTINCK 1938 Teinobasis debeauforti LIEFTINCK, Nova Guinea (N.S.) 2: 107. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea. debeauxi

LIEFTINCK

1938 Teinobasis debeauxi New Guinea.

LIEFTINCK,

Nova Guinea (N.S.) 2:113. Type male: not given; type-locality:

Distribution: New Guinea. dolabrata 1938

LIEFTINCK

Teinobasis dolabrata LIEFTINCK, Nova Guinea (N.S.) type-locality: New Guinea.

2: 1 1 4 .

Type male: Ann Arbor Museum;

Distribution: New Guinea, dominula

LIEFTINCK 1937 Teinobasis dominula LIEFTINCK, Nova Guinea (N.S.) 1: 328. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, emarginata 1949

LIEFTINCK Teinobasis emarginata LIEFTINCK, Treubia, 20 Leiden, type-locality: Shortland Island.

(2): 3 4 4 .

Type female: Rijksmus. Natuurl. Hist.

Distribution: Shortland Island, euglena 1934

1934 1948 1954

LIEFTINCK Teinobasis euglena Java. Teinobasis euglena Teinobasis euglena Teinobasis euglena

LIEFTINCK,

Stylops, 3:

-

LIEFTINCK,

-

LIEFTINCK,

-

LIEFTINCK,

8.

Type male: Buitenzorg Museum; type-locality:

Treubia, 14 ( 4 ) : 3 9 8 (Java). Treubia, 19 ( 2 ) : 2 9 1 (Engano Island). Treubia, 22 Suppl.: 6 2 (Engano, Sumatra, Java).

Distribution: Engano, Sumatra, Java.

352 filamenta NEEDHAM & GYGER 1939 Teinobasis filamentum NEEDHAM type-locality: Mindanao.

& GYGER,

Philipp. J. Sei., 70 (3): 302. Type male: not given;

Distribution: Philippine Islands. filiformis (BRAUER) 1868 Amphicnemis filiformis BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 544. Type male: Semper's Collection; type-locality: Luzon. 1877 Telebasis (Telebasis) filiformis - SELYS, Bull Acad. r. Belg. (2) 43: 123 (Luzon) (type in Selys's Collection). 1890 Teinobasis filiformis - KIRBY, Syn. Cat. Neur.-Odon., London: 157.

Distribution: Philippine Islands. filum (BRAUER) 1868 Amphicnemis filum BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 545. Type male: Semper's Collection; type-locality: Mindanao. 1877 Telebasis (Telebasis) filum - SELYS, Bull Acad. r. Belg. (2) 43: 124 (type in Selys's Collection). 1890 Teinobasis filum - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1939 Teinobasis filum - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 301.

Distribution: Philippine Islands. fortis LIEFTINCK 1962 Teinobasis fortis LIEFTINCK, Insecta of Micronesia, 5 (1): 26. Type male: US natn. Mus., Washington; type-locality: Ponape.

Distribution: East Caroline Islands. fulgens LIEFTINCK 1949 Teinobasis fulgens LIEFTINCK, Nova Guinea (N.S.) 5: 172. Type male: Rijksmus. Natuurl. Hist., Leiden, type-locality: New Guinea.

Distribution: New Guinea. helvola LIEFTINCK 1930 Teinobasis helvola LIEFTINCK, Treubia, 12:155. Type male: Buitenzorg Museum; type-locality: Celebes.

Distribution: Celebes. kirbyi LAIDLAW 1902 Teinobasis kirbyi LAIDLAW, Proc. Zool. Soc. London, 1902: 386. Type male: Laidlaw's Collection; type-locality: Perak. 1 9 3 1 Teinobasis kirbyi - L A I D L A W , J . Fed. Mai. States Mus., 1 6 : 2 0 2 . 1954 Teinobasis kirbyi - LIEFTINCK, Treubia, 22, Suppl.: 62 (Malaysia).

Distribution: Malaysia. laglaizei (SELYS) 1878 Telebasis (Telebasis) laglaizei SELYS, Mitt. Zool. Mus. Dresden, 3: 322. Type male: not given; type-locality: Papua.

353 1890 Teinobasis laglaizei - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1935 Teinobasis laglaizei - LIEFTINCK, Nova Guinea, 17: 259 (redescription).

Distribution: New Guinea. laidlawi

KIMMINS

1936 Teinobasis laidlawi KIMMINS, J. Fed. Mal. States Mus., 18: 95. Type male: British Museum (Nat.Hist.), London; type-locality: Borneo. 1954 Teinobasis laidlawi - LIEFTINCK, Treubia, 22 Suppl.: 63 (Borneo).

Distribution: Borneo. lorquini

(SELYS)

1877 Telebasis (Telebasis) lorquini SELYS, Bull Acad. r. Belg. (2) 43: 118. Type female: Selys's Collection; type-locality: Moluccas. 1890 Teinobasis lorquini - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1935 Teinobasis lorquini - LIEFTINCK, Nova Guinea, 17: 238 (redescription).

Distribution: Malaysia. luciae

LIEFTINCK

1937 Teinobasis luciae LIEFTINCK, Nova Guinea (N.S.) 1: 326. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, metallica

(FÖRSTER) 1898 Telebasis metallica FÖRSTER (nec Ris, 1913), Termeszetr. Füz., Budapest, 21: 300. Type male: Termeszettud. Μύζ. Budapest; type-locality: New Guinea. 1935 Teinobasis metallica - LIEFTINCK, Nova Guinea, 17: 263.

Distribution: New Guinea. micans

LIEFTINCK

1949 Teinobasis micans LIEFTINCK, Nova Guinea (N.S.) 5: 174. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, nigra

CAMPION & LAIDLAW Teinobasis nigra C A M P I O N & L A I D L A W , Proc. Zool. Soc. London, 1928:136. Type male: British Museum (Nat.Hist.), London; type-locality: Luzon. 1939 Teinobasis nigra - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 302 (Luzon).

1928

Distribution: Philippine Islands. nigrolutea

LIEFTINCK

1962 Teinobasis nigrolutea LIEFTINCK, Insects of Micronesia, 5 (1): 29. Type male: US natn. Mus., Washington; type-locality: Ponape.

Distribution: East Caroline Islands. nitescens nitescens

LIEFTINCK

1913 Teinobasis metallica Ris, (nec FÖRSTER, 1898), Naturf. Ges., 34: 522. Type male: Inst. Tax. Zöol. Amsterdam; type-locality: New Guinea.

354 1935 Teinobasis nitescens LIEFTINCK, N o v a Guinea, 1 7 : 262 (a new name for Teinobasis Ris, 1913, a junior secondary homonymy of Telebasis metallica FÖRSTER, 1898).

metallica

Distribution: New Guinea. nitescens angusticlavia Ris 1913 Teinobasis metallica angusticlavia Ris, Abh. senckenb. Naturf. Ges., 34: 522. Type male: Inst. Tax. Zöol., Amsterdam; type-locality: Aru Island.

Distribution: Aru. olivacea Ris 1916 Teinobasis olivacea Ris, Nova Guinea, 13 (2): 102. Type male: Mus. N a t u r k . Berlin; typelocality: Philippines. 1918 Teinobasis olivacea - Ris, Denkschr. med.-naturw. Ges., Jena, 13: 102. 1939 Teinobasis olivacea - N E E D H A M & G Y G E R , Philipp. J. Sei., 7 0 (3): 306 (Mindoro, Luzon).

Distribution: Philippine Islands. olthofi

LIEFTINCK

1949 Teinobasis olthofi LIEFTINCK, N o v a Guinea, (N.S.) 5: 169. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea. palauensis 1962

LIEFTINCK

Teinobasis palauensis LIEFTINCK, Insects of Micronesia, Washington; type-locality: Palau.

5 (1): 21.

Type male:

US

natn. Mus.,

Distribution: West Caroline Islands. ponapensis

LIEFTINCK

1962 Teinobasis ponapensis LIEFTINCK, Insects of Micronesia, 5 (1): 24. Type male: U S natn. Mus., Washington; type-locality: Ponape.

Distribution: East Caroline Islands. pretiosa 1877

1890 1935

(SELYS)

Telebasis (Telebasis) pretiosa SELYS, Bull Acad. r. Belg. ( 2 ) 4 3 : 1 1 6 . Type male: sels; type-locality: New Guinea. Teinobasis praetiosa - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 5 7 (Papua). Teinobasis pretiosa - LIEFTINCK, N o v a Guinea, 1 7 : 2 6 5 (redescription).

R.

Mus., Brus-

Distribution: New Guinea. prothoracica

(SELYS)

1877 Telebasis (Telebasis) prothoracica SELYS, Bull. Acad. r. Belg., (2) 43: 117. Type male: R. Mus. Brussels; type-locality: Mysol. 1 8 9 0 Teinobasis prothoracica - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 5 7 . 1 9 3 5 Teinobasis prothoracica - LIEFTINCK, Nova Guinea, 1 7 : 2 3 9 (redescription). 1 9 3 8 Teinobasis prothoracica - LIEFTINCK, Nova Guinea (N.S.) 2 : 1 1 6 (description of allotype female).

Distribution: New Guinea.

355 perverulenta Ris 1916 Teinobasis perverulenta Ris, Nova Guinea, 13 (2): 104. Type male: Mus. Naturk. Berlin; typelocality: Philippines.

Distribution: Philippine Islands. rajah

LAIDLAW

1912 Teinobasis rajah LAIDLAW, J. Str. Br. R. Asiatic Soc., 63: 97. Type male: Laidlaw's Collection; type-locality: Borneo. 1937 Teinobasis leonorae LIEFTINCK, Treubia, 16 (1): 97. Type male: Univ. Mus. Michigan, A n n Arbor; type-locality: Malay Peninsula. 1954 Teinobasis rajah - LIEFTINCK, Treubia, 22 Suppl.: 63 (Penang, Borneo). 1954 Teinobasis leonorae - LIEFTINCK, Treubia, 2 2 Suppl.: 63 (proposed synonymy with rajah L A I D LAW, 1912).

Distribution: Borneo, Malaysia. ranee

NEEDHAM & GYGER

1941 Teinobasis ranee N E E D H A M locality: Mindanao.

& GYGER,

Philipp. J. Sei., 74: 149. Type male: not given; type-

Distribution: Philippine Islands. recurva

(SELYS)

1877 Telebasis (Telebasis) recurva SELYS, Bull. Acad. r. Belg. (2) 43: 114. Type male: Selys's Collection; type-locality: Mindanao. 1890 Teinobasis recura - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1939 Teinobasis recurva - N E E D H A M & G Y G E R , Philipp. J. Sei., 7 0 (3): 305 (Mindanao). 1957 Teinobasis recurva - KIMMINS, Bull. Brit. Mus. (Nat.Hist.) Ent., 5: 171 (redescription).

Distribution: Philippine Islands. rubicauda

LIEFTINCK

1974 Teinobasis rubicauda LIEFTINCK, Steenstrupia, 3 (12): 120. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Palawan.

Distribution: Palawan. ruficollis

(SELYS)

1877 Telebasis (Telebasis) ruficollis SELYS, Bull Acad. r. Belg. (2) 43: 119. Type male: Selys's Collection; type-locality: Singapore. 1890 Teinobasis ruficollis - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1927 Teinobasis ruficollis - Ris, Zool. Meded., Leiden, 10 (1): 25 (Sumatra). 1954 Teinobasis ruficollis - LIEFTINCK, Treubia, 22 Suppl.: 63 (Malaya, Sumatra).

Distribution: Malaysia, Sumatra. rufithorax

(SELYS)

1877 Telebasis (Telebasis) rufithorax SELYS, Bull Acad. r. Belg. (2) 43: 122. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Celebes. 1890 Teinobasis rufithorax - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1913 Telebasis rufithorax - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 473 (Celebes, New Guinea, Australia). 1949 Teinobasis rufithorax - LIEFTINCK, Treubia, 20 (2): 346 (Bismarck: New Ireland).

Distribution: Celebes, New Guinea, New Ireland, North Australia.

356 samaritis Ris 1916 Teinobasis samaritis Ris, Nova Guinea, 12 (2): 103. Type male: Mus. Naturk. Berlin; typelocality: Philippines. 1918 Teinobasis samaritis - Ris, Denkschr. med. naturw. Ges. Jena, 13: 103. 1939 Teinobasis samaritis - NEEDHAM & G Y G E R , Philipp. J. Sei., 7 0 (3): 306 (Luzon, Mindanao).

Distribution: Philippine Islands. scintillans LIEFTINCK 1932 Teinobasis scintillans LIEFTINCK, Nova Guinea, 15 (5): 96. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, serena serena LIEFTINCK 1932 Teinobasis serena LIEFTINCK, Nova Guinea, 15 (5): 98. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea. 1935 Teinobasis serena - LIEFTINCK, Nova Guinea, 17: 268. 1949 Teinobasis serena serena - LIEFTINCK, Nova Guinea, (N.S.) 5: 168.

Distribution: New Guinea. serena humeralis LIEFTINCK 1949 Teinobasis serena humeralis LIEFTINCK, Nova Guinea, (N.S.) 5: 168. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, stigmatizans LIEFTINCK 1938 Teinobasis stigmatizans LIEFTINCK, Nova Guinea, (N.S.) 2:109. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, strigosa NEEDHAM & GYGER 1939

Teinobasis strigoza N E E D H A M type-locality: Luzon.

& GYGER,

Philipp. J. Sei.,

7 0 (3): 304.

Type male: not given;

Distribution: Philippine Islands, suavis LIEFTINCK 1953 Teinobasis suavis LIEFTINCK, Treubia, 22: 244. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Borneo. 1954 Teinobasis suavis - LIEFTINCK, Treubia, 22 Suppl.: 63.

Distribution: Borneo. superba (SELYS) 1877 Telebasis (Telebasis) superba (HÄGEN MS) SELYS, Bull. Acad. r. Belg. (2) 43:114 (as generotype of Telebasis SELYS, 1877). Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Celebes. 1878 Telebasis (Telebasis) eximia SELYS, Mitt. Zool. Mus. Dresden, 3: 322. Type male: not given; type-locality: Ternate. 1890 Teinobasis superba - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1890 Teinobasis exima - KIRBY, Syn. Cat. Neur.-Odon., London: 157. 1 9 2 6 Teinobasis gracillima FRASER, Treubia, 8 ( 3 - 4 ) : 4 9 4 . Type male: Buitenzorg Museum; typelocality: Java.

357 1934 Telebasis

superba

- COWLEY, E n t o m o l o g i s t , 67: 253 (as g e n e r o t y p e of Telebasis

SELYS, 1865).

1935 Teinobasis superba - LIEFTINCK, Nova Guinea, 17: 236 (redescription).

Distribution: Celebes, Java, tenuis (MARTIN) 1898 Telebasis tenuis MARTIN, Ann. Soc. ent. France, 66: 584. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Celebes. 1984 Teinobasis tenuis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 94.

Distribution: Celebes. wallacei

CAMPION

1924 Teinobasis wallacei CAMPION, Ann. Mag. nat. Hist., (9) 14: 612. Type male: British Museum (Nat.Hist.), London; type-locality: New Guinea. 1935 Teinobasis wallacei - LIEFTINCK, Nova Guinea, 17: 267 (redescription).

Distribution: New Guinea.

Genus Telagrion

SELYS

1876 Agrion (Teleagrion) SELYS, Synopsis des Agrionines, Add. 5: 256. Type-species: Teleagrion longum SELYS, 1876 (original designation). - 1890 Teleagrion - KIRBY, Syn. Cat. Neur.-Odon., London: 155. - 1916 Teleagrion - Ris, Arch. Naturg. (A) 82: 136. - 1920 Aceratobasis KENNEDY, Ohio J. Sei. Columbus, 2: 88. Type-species: Metaleptobasis cornicauda CALVERT, 1909 (original designation). - 1960 Teleagrion - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 48 (key to species).

cornicauda (CALVERT) 1909 Metaleptobasis cornicauda CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 199. Type male: Carnegie Mus.; type-locality: Brazil. 1920 Aceratobasis cornicauda - KENNEDY, Ohio J. Sei. Columbus, 2: 88 (as generotype of Aceratobasis

KENNEDY, 1920).

1984 Telagrion cornicauda - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 94.

Distribution: Brazil. diceras (SELYS) 1877 Telebasis (Leptobasis) diceras SELYS, Bull. Acad. r. Belg. (2): 43: 102. Type male: Selys's Collection; type-locality: Para. 1890 Leptobasis diceras - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1984 Telagrion diceras - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 94.

Distribution: Brazil. fulvellum

SELYS

1876 Agrion (Telagrion) fulvellum (BATES MS) SELYS, Synopsis des Agrionines, Add. 5: 258. Type male: Termeszett. Muz., Budapest; type-locality: Amazons. 1890 Telagrion fulvellum - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. longum

SELYS

1876 Agrion (Telagrion) longum SELYS, Synopsis des Agrionines, Add. 5: 260 (as generotype of Telagrion SELYS, 1876). Type male: Selys's Collection; type-locality: Brazil. 1890 Telagrion longum - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

358 1957 Metaleptobasis sooretamae SANTOS, Rev. bras. Biol.: 17: 143. Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil. 1960 Leptobasis tenax ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 53. Type male: Naturhist. Mus., Wien; type-locality: Brazil.

Distribution: Brazil. macilentum

(RAMBUR)

1842 Agrion macilentum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 259. Type male: unknown; typelocality: Brazil. 1877 Telebasis (Leptobasis) macilenta - SELYS, Bull. Acad. r. Belg. (2): 43: 106 (Brazil). 1890 Leptobasis macilenta - KIRBY, Syn. Cat. Neur.-Odon., London: 156. 1984 Telagrion macilentum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 94.

Distribution: Brazil. maurei

SANTOS

1 9 8 4 T e l a g r i o n maurei SANTOS, 1 9 7 0 - l o c . cit. DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 94.

Distribution: Brazil. oreas Ris 1916 Telagrion oreas Ris, Arch. Naturg., (A) 82:138. Type male: not given; type-locality: Colombia.

Distribution: Colombia, quadricolor Ris 1916 Telagrion quadricolor Ris, Arch. Naturg. (A) 82:136. Type male: not given; type-locality: Peru. 1943 Telagrion quadricolor - SCHMIDT, Beitr. Fauna Perus, 2: 240.

Distribution: Peru. raineyi

WILLIAMSON

1915 Teleagrion raineyi WILLIAMSON, Proc. US natn. Mus., 48: 613. Type male: Williamson's Collection; type-locality: Trinidad. 1932 Telagrion raineyi -GEIJSKES, Zool. Meded., Leiden, 14: 248 (Trinidad) (description of female). 1946 Oligoclada raineyi - SANTOS, Livro hom. R. Ferreira d'Almeida: 303 (redescription of female).

Distribution: Trinidad. ribeiroi

SANTOS

1962 Teleagrion ribeiroi SANTOS, Bol. Mus. nac. Rio de Janeiro, (N.S.) (Zool.) 230: 1. Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. Genus Telebasis

SELYS

1865 Telebasis SELYS (nec SELYS, 1877), Bull. Acad. r. Belg., (2) 20: 378. Type-species: Agrion salvum HÄGEN, 1 8 6 1 ( d e s i g n a t e d b y KIRBY, 1 8 9 0 ) . - 1 8 7 6 A g r i o n ( E r y t h a g r i o n ) SELYS, B u l l . A c a d ,

r. Belg. (2): 41 1890 Telebasis - KIRBY, Syn. 1865). - 1916

( 2 - 3 ) : 245. Type-species: Agrion filiola PERTY, 1834 (original designation). - KIRBY, Syn. Cat. Neur.-Odon., London: 155. - 1890 Agrion (Erythagrion) Cat. Neur.-Odon., London: 155 (proposed synonymy with Telebasis SELYS, Telebasis - Ris, Arch. Naturg., (A) 82: 128. - 1919 Agrion (Erythagrion) -

CALVERT, T r a n s . A m . e n t . S o c . , 45: 3 5 4 . - 1 9 3 4 Telagrion

- COWLEY, E n t o m o l o g i s t , 67: 2 5 3 .

- 1960 Telagrion - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 49 (key to South American species).

359 aureipennis 1984

JURZITRA

Telebasis aureipennis Utrecht, 1: 94.

JURZITRA, 1 9 8 0

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Argentina. byersi

WESTFALL

1957 Telebasis byersi WESTFALL, Florida Ent., 40: 20. Type male: unknown; type-locality: Florida.

Distribution: carmesina

U.S.A.

CALVERT

1909 Telebasis carmesina CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 191. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil, carminita

CALVERT 1909 Telebasis carminita CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 194. Type male: Carnegie Museum; type-locality: Brazil. 1960 Telebasis carminita - ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 49 (description of female).

Distribution: Brazil. carota

KENNEDY

1936 Telebasis carota KENNEDY, Ann. Ent. Soc. Amer., 29 (4): 807. Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador. coccinata

CALVERT

1909 Telebasis coccinata CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 192. Type male: Carnegie Museum; type-locality: Brazil.

Distribution: Brazil, coccinea

(SELYS) 1876 Agrion (Erythagrion) coccineum SELYS, Synopsis des Agrionines, Add. 5: 255. Type male: Selys's Collection; type-locality: Minas Geraes. 1890 Telebasis coccinea - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. collopistes

CALVERT

1902 Telebasis collopistes CALVERT, Biol. Centr.-Amer. Neur., London: 116. Type male: US natn. Mus., Washington; type-locality: Mexico.

Distribution: Mexico, corallina

(SELYS)

1876 Agrion (Erythagrion) corallinum SELYS, Synopsis des Agrionines, Add. 5: 254. Type male: Selys's Collection; type-locality: Brazil. 1890 Telebasis corallina - KIRBY, Syn. Cat. Neur.-Odon., London: 155. 1 9 8 4 Telebasis corallina - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 4 .

Distribution: Brazil.

360 digiticollis CALVERT 1902 Telebasis digiticollis CALVERT, Biol. Centr.-Amer. Neur., London: 118. Type male: US natn. Mus., Washington; type-locality: Mexico. 1916 Telebasis digiticollis - Ris, Arch. Naturg., (A) 82: 128 (Mexico, Guatemala, Honduras, Colombia).

Distribution: Central America. dominicana (SELYS) 1857 Agrion dominicanum SELYS, Sagra, Hist. Cuba Ins.: 466. Type male: unknown; type-locality: Dominica. 1876 Agrion (Erythagrion) dominicanum - SELYS, Synopsis des Agrionines, Add. 5: 248 (Puerto Rico, Dominica, Guyana). 1890 Telebasis dominicana - KIRBY, Syn. Cat. Neur.-Odon., London: 155 (Haiti, Puerto Rico, Guyana). 1919 Agrion (Erythagrion) dominicanum - CALVERT, Trans. Am. ent. Soc., 45: 354 (Cuba).

Distribution: West Indies. erythrina (SELYS) 1876 Agrion (Erythagrion) erythrinum SELYS, Synopsis des Agrionines, Add. 5: 251. Type male: Selys's Collection; type-locality: Minas Geraes. 1890 Telebasis erythrina - KIRBY, Syn. Cat. Neur.-Odon., London: 155.

Distribution: Brazil. filiola (PERTY) 1834 Agrion filiola PERTY, Del. Anim. Art.: 125. Type male: unknown; type-locality: Brazil. 1876 Agrion (Erythagrion) filiola - SELYS, Synopsis des Agrionines, Add. 5: 246 (as generotype of Erythagrion

SELYS, 1 8 7 6 ) .

1890 Telebasis filiola - KIRBY, Syn. Cat. Neur.-Odon., London: 155. 1916 Telebasis filiola - Ris, Arch. Naturg. (A) 82: 128 (Guatemala, Mexico, Panama, Colombia, Brazil).

Distribution: from Mexico to Brazil. flammeola KENNEDY 1936 Telebasis flammeola KENNEDY, Ann. Ent. Soc. Amer., 29 (4): 804. Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador. fluviatilis ST. QUENTIN 1960 Telebasis fluviatilis ST. QUENTIN, Beitr. neotrop. Fauna, 2 (1): 50. Type male: Naturhist. Mus. Wien; type-locality: Brazil.

Distribution: Brazil. garleppi Ris 1916 Telebasis garleppi Ris, Arch. Naturg., (A) 82: 129. Type male: not given; type-locality: Costa Rica.

Distribution: Costa Rica.

361 griffinii

(MARTIN)

1895 Erithagrion griffinii MARTIN, Boll. Mus. Torino, No. 240: 2. Type male: Mus. Ist. Zool. sist. Univ., Torino; type-locality: Panama. 1902 Telebasis griffinii - CALVERT, Biol. Centr.-Amer. Neur., London: 116 (redescription). 1932 Telebasis griffinii - GEIJSKES, Zool. Meded., Leiden, 14: 257 (Trinidad).

Distribution: Central America. incolumis 1930

WILLIAMSON & WILLIAMSON

Telebasis incolumis WILLIAMSON & WILLIAMSON, OCC. Pap. Mus. Zool. Univ. Michigan, 1. Type male: Williamson's Collection; type-locality: Mexico.

216:

Distribution: Mexico. isthmica

CALVERT

1902 Telebasis isthmica CALVERT, Biol. Centr.-Amer. Neur., London: 118. Type male: US natn. Mus. Washington; type-locality: Panama.

Distribution: Panama. livida

KENNEDY

1936 Telebasis livida KENNEDY, Ann. Ent. Soc. Amer., 29 (4): 811. Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador. paraensis

MACHADO

1956 Telebasis paraensis MACHADO, Rev. bras. Biol., 16: 227. Type male: Nac. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. salva

(HÄGEN)

1861 Agrion salvum HAGEN, Syn. Neur. N. Amer., Washington: 85. Type male: Agassiz Mus., Cambridge; type-locality: Mexico. 1 8 6 5 Telebasis salva - SELYS, Bull. Acad. r. Belg. ( 2 ) : 2 0 : 3 7 8 (as generotype of Telebasis SELYS, 1 8 6 5 ) . 1869 Telebasis boucardi SELYS, C.R. Soc. Ent. Belg., 11: 120. Type male: Selys's Collection; typelocality: unknown. 1876 Agrion (Erythagrion) salvum - SELYS, Synopsis des Agrionines, Add. 5: 252 (Mexico, Texas, Guatemala). 1890 Telebasis salva - KIRBY, Syn. Cat. Neur.-Odon., London: 155. 1916 Telebasis salva - RlS, Arch. Naturg. (A) 82: 128 (Arizona, Mexico). 1989 Telebasis salva - BROOKS, Notul. odonatol., 3 (4): 50 (Costa Rica).

Distribution: Central America. sanguinalis

CALVERT

1909 Telebasis sanguinalis CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 192. Type male: Carnegie Mus.; type-locality: Brazil.

Distribution: Brazil. theodori 1984

NAVÄS

Telebasis theodori Utrecht, 1: 95.

NAVÄS, 1 9 3 4

Distribution: Brazil.

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

362 versicolor

FRÄSER

1946 Telebasis versicolor FRÄSER, Trans. R. ent. Soc. London, 96: 42. Type male: Leeds City Mus.; type-locality: Colombia. 1966 Telebasis versicolor - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 217 (holotype in British Museum (Nat.Hist.), London).

Distribution: Colombia. vulnerata

(HAGEN)

1861 Agrion vulneratum HAGEN, Syn. Neur. N. Amer., Washington: 86. Type female: Agassiz M u seum, Cambridge; type-locality: Puerto Rico. 1876 Agrion (Erythagrion) vulneratum - SELYS, Synopsis des Agrionines, Add. 5: 250. 1890 Telebasis vulnerata - KIRBY, Syn. Cat. Neur.-Odon., London: 155. 1919 Agrion (Pyrrhosoma) vulneratum - CALVERT, Trans. Am. ent. Soc., 45: 354 (Cuba, Puerto Rico).

Distribution: Central America. willinki

FRÄSER

1948 Telebasis willinki FRÄSER, Acta Zool. lilloana, 5: 54. Type male: Miguel Lille Inst., Tucuman; type-locality: Argentina.

Distribution: Argentina. Genus Xanthagrion

SELYS

1876 Agrion (Xanthagrion) SELYS, Synopsis des Agrionines, Add. 5: 230. Type-species: Agrion (Xanthagrion) erythroneurum SELYS, 1876 (original designation). - 1890 Xanthagrion - KIRBY, Syn. Cat. Neur.-Odon., London: 154.

erythroneurum

SELYS

1876 Agrion (Xanthagrion) erythroneurum SELYS, Synopsis des Agrionines, Add. 5: 231. Type male: Selys's Collection; type-locality: Melbourne (as generotype of Xanthagrion SELYS, 1876). 1890 Xanthagrion erythroneurum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1901 Xanthagrion erythroneurum - MARTIN, Mem. Soc. zool. France, 14: 247 (Australia). 1913 Xanthagrion erythroneurum - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 466 (Australia).

Distribution: Australia. Genus Xanthocnemis

TILL YARD

1913 Xanthocnemis TILL YARD, Proc. Linn. Soc. N.S. Wales, 37:465. Type-species: Telebasis zelandica - MCLACHLAN, 1873 (original designation).

zelandicum

(MCLACHLAN)

1873 Telebasis zelandica MCLACHLAN, Ann. Mag. nat. Hist., (4) 12: 35. Type male: McLachlan's Collection; type-locality: New Zealand. 1876 Agrion (Xanthagrion) zelandicum - SELYS, Synopsis des Agrionines, Add. 5: 232. 1876 Agrion (Xanthagrion) zelandicum race antipodum SELYS, Synopsis des Agrionines, Add. 5: 234. Female from New Zealand. 1890 Agrion (Xanthagrion) zelandicum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1890 Agrion (Xanthagrion) antipodum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1913 Xanthocnemis zelandicum - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 465 (as generotype of Xanthocnemis TILLYARD, 1913).

Distribution: New Zealand.

363

sobrinum

(MCLACHLAN)

1873 Telebasis sobrina MCLACHLAN, Ann. Mag. nat. Hist., (4) 12: 36. Type male: McLachlan's Collection; type-locality: New Zealand. 1876 Agrion (Xanthagrion) sobrinum - SELYS, Synopsis des Agrionines, Add. 5: 234. 1890 Xanthagrion sobrinum - KIRBY, Syn. Cat. Neur.-Odon., London: 154. 1984 Xanthocnemis sobrina - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 95.

Distribution: New Zealand. tuanuii 1984

ROWE

Xanthocnemis tuanuii Utrecht, 1: 95.

- loc. cit.

ROWE, 1981

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: New Zealand. Family Isostictidae Slim species with very narrow wings; CuP usually reduced to 1 - 2 cells; anal vein not visible. Larva with saccoid triquetral nodate gills. 1955 Isostictinae FRASER, Entomologist's month. Mag., 91: 227. - 1957 Isostictinae - FRÄSER, R. Zool. Soc. N.W. Wales, 1957: 44 (with genera Isosticta, Austrosticta, Neosticta, Oristicta, Amphisticta, Phasmosticta, Tanymecosticta, Selysioneura). - 1984 Isostictidae DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 95.

Genus Austrosticta

TILLYARD

1908 Austrosticta TILLYARD, Proc. Linn. Soc. N.S. Wales, 32: 764. Type-species: Austrosticta fieldi TILLYARD, 1908 (original designation).

fieldi

TILLYARD

1908 Austrosticta fieldi TILLYARD, Proc. Linn. Soc. N.S. Wales, 32:765 (as generotype of Austrosticta TILLYARD, 1908). Type male: not given; type-locality: Tennant's Creek. 1913 Austrosticta fieldi - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 434.

Distribution: North Australia. soror

SJÖSTEDT

1917 Austrosticta soror SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 29. Type male: Naturhist. Riksmus., Naturhist., Stockholm; type-locality: N.-W. Australia.

Distribution: Australia. Genus Eurysticta

WATSON

1984 Eurysticta WATSON, 1969 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 95. Type-species: Eurysticta coolawanyah WATSON, 1969 (original designation).

coolawanyah

WATSON

1984 Eurysticta coolawanyah World, Utrecht, 1: 95.

WATSON,

Distribution: West Australia.

1969 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

364 Genus Isosticta SELYS 1885 Isosticta SELYS, C.R. SOC. Ent. Belg., 29: 65. Type-species: Isosticta spinipes, SELYS, 1885 (monotypy). - 1975 Isosticta LIEFTINCK, Cah. Off. Rech. sei. techn. Hydrobiol., 9 (3): 149 (key to species).

banksi TILLYARD 1913

Isosticta banksi TILLYARD, Proc. Linn. Soc. N.S. Wales, lection; type-locality: Banks Island.

37: 433.

Type male: Tillyard's Col-

Distribution: Banks Island. gracilior 1975

LIEFTINCK

Isosticta gracilior LIEFTINCK, Cah. Off. Rech. sei. techn. Hydrobiol., Rijksmus. Natuurl. Hist., Leiden; type-locality: New Caledonia.

9 (3): 148.

Type male:

Distribution: New Caledonia. handschini

LIEFTINCK

Isosticta handschini LIEFTINCK, Revue suisse Zool., 40: 4 1 7 . Type male: Basel Museum; typelocality: Australia. 1955 Isosticta handschini - FRÄSER, Entomologist's month. Mag., 9 1 : 230. 1933

Distribution: Australia. humilior

LIEFTINCK

1975 Isosticta humilior LIEFTINCK, Cah. Off. Rech. sei. techn. Hydrobiol., 9 (3): 149. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Caledonia.

Distribution: New Caledonia. robustior Ris 1916 Isosticta robustior Ris, Nova Caledonia Zool., 2: 66. Type male: Mus. Naturk., Berlin; typelocality: New Caledonia.

Distribution: New Caledonia. simplex MARTIN 1901 Isosticta simplex M A R T I N , Mem Soc. zool. France, 14: 244. Type male: not given; type-locality: New Caledonia. 1913 Isosticta simplex - T I L L Y A R D , Proc. Linn. Soc. N.S. Wales, 3 7 : 432. 1917 Isosticta simplex - SJÖSTEDT, Ark. Zool. Uppsala, 1 1 (11): 29 (Queensland).

Distribution: New Caledonia, Australia. spinipes

SELYS

1885 Isosticta spinipes SELYS, C.R. Soc. Ent. Belg., 29: 65 (as generotype of Isosticta Type male: Selys's Colletion; type-locality: New Caledonia.

SELYS,

1885).

Distribution: New Caledonia. tillyardi

CAMPION

1921 Isosticta tillyardi CAMPION, Ann. Mag. nat. Hist., (6) (Nat.Hist.), London; type-locality: New Caledonia.

Distribution: New Caledonia.

8:

38. Type male: British Museum

365 Genus Neosticta TILLYARD 1913 Neosticta TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 435. Type-species: Neosticta canescens TILLYARD, 1913 (original designation). - 1917 Amphisticta SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 31. Type-species: Amphisticta silvarum SJÖSTEDT, 1917 (original designation).

canescens TILL YARD 1913 Neosticta canescens TILL γ ARD, Proc. Linn. Soc. N.S. Wales, 37:435 (as generotype of Neosticta TILL YARD, 1913). Type male: Tillyard's Collection; type-locality: New South Wales. 1913 Neosticta canescens f. dorrigoensis TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 435. Male from New South Wales.

Distribution: Australia. silvarum (SJÖSTEDT) 1917 Amphisticta silvarum SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 32 (as generotype of Amphisticta SJÖSTEDT, 1917). Type male: Naturhist. Riksmus., Stockholm; type-locality: Queensland. 1984 Neosticta silvarum - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 96.

Distribution: Australia. Genus Oristicta TILLYARD 1913 Oristicta TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 438. Type-species: Oristicta filicicola TILLYARD, 1913 (original designation). - 1951 Phasmosticta LIEFTINCK, Amer. Mus. Novit, No. 1488: 17. Type-species: Phasmosticta interposita LIEFTINCK, 1951 (original designation). - 1984 Phasmosticta - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 96 (as incl. of Oristicta TILLYARD, 1913).

filicicola TILL YARD 1913 Oristicta filicicola TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 438 (as generotype of Oristicta TILLYARD, 1913). Type male: Tillyard's Collection; type-locality: Cooktown. 1955 Oristicta filicicola - FRÄSER, Entomologist's month. Mag., 91: 229 (redescription).

Distribution: Australia. interposita (LIEFTINCK) 1951 Phasmosticta interposita LIEFTINCK, Amer. Mus. Novit., No. 1488: 22 (as generotype of Phasmosticta LIEFTINCK, 1951). Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: North Australia. 1984 Oristicta interposita - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 96.

Distribution: Australia. vallisi (FRÄSER) 1955 Phasmosticta wallisi FRÄSER, Entomologist's month. Mag., 91: 227. Type male: British Museum (Nat.Hist.), London; type-locality: Queensland. 1984 Oristicta vallisi - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 96.

Distribution: Australia. Genus Selysioneura FÖRSTER 1900 Selysioneura FÖRSTER, Termeszetr. Füz. Budapest, 23: 106. Type-species: Selysioneura cervicornu FÖRSTER, 1900 (original designation). - 1932 Selysioneura - LIEFTINCK, Nova Guinea, 15 (5): 57 (revision, key to New Guinean species). - 1953 Selysioneura - LIEFTINCK, Treubia, 21: 654 (key to species).

366 aglaia

LIEFTINCK

1953 Selysioneura aglaia LIEFTINCK, Treubia, 21: 646. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Morotai Island.

Distribution: Indonesia: Morotai Island. arboricola

LIEFTINCK

1958 Selysioneura arboricola LIEFTINCK, Nova Guinea, (N.S.) 10: 286. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Fergusson Island. 1 9 8 4 Selysioneura arboricola - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 6 (d'Etrecasteaux Island).

Distribution: Fergusson Island (near Papua New Guinea). bacillus Ris 1916 Selysioneura bacillus Ris, Nova Guinea, 13 (2): 93. Type male: Mus. Naturk. Berlin; typelocality: New Guinea.

Distribution: New Guinea. capreola

LIEFTINCK

1932 Selysioneura capreola LIEFTINCK, Nova Guinea, 15 (5): 61. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea. cervicornu

FÖRSTER

1900 Selysioneura cervicornu FÖRSTER, Termeszetr. Füz. Budapest, 23: 106 (as generotype of Selysioneura FÖRSTER, 1900). Type male: Termeszett. Muz. Budapest; type-locality: New Guinea.

Distribution: New Guinea. Cornelia

LIEFTINCK

1953 Selysioneura Cornelia LIEFTINCK, Treubia, 21: 647. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea. drymobia

LIEFTINCK

1958 Selysioneura drymobia LIEFTINCK, Nova Guinea, (N.S.), 10: 284. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Misima Island. 1984 Selysioneura drymobia - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 96.

Distribution: Luiseade Archipelago. phasma

LIEFTINCK

1932 Selysioneura phasma LIEFTINCK, Nova Guinea, 15 (5): 63. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea. ranatra

LIEFTINCK

1949 Selysioneura ranatra LIEFTINCK, Nova Guinea, (N.S.) 5: 78. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea.

367

rangifera

LIEFTINCK

1958 Selysioneura rangifera LIEFTINCK, Nova Guinea (N.S.) 10: 290. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: N o r m a n b y Island.

Distribution: Normanby Island (near Papua New Guinea), rhapia

LIEFTINCK

1958 Selysioneura rhaphia LIEFTINCK, N o v a Guinea (N.S.) 10: 288. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: N o r m a n b y Island.

Distribution: Normanby Island (near Papua New Guinea), stenomantis

LIEFTINCK

1932 Selysioneura stenomantis LIEFTINCK, N o v a Guinea 15 (5): 65. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, thalia

LIEFTINCK

1953 Selysioneura thalia LIEFTINCK, Treubia, 21: 642. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Halmahera Island.

Distribution: Indonesia: Halmahera Island. umbratilis L I E F T I N C K 1932 Selysioneura umbratilis LIEFTINCK, N o v a Guinea, 15 (5): 67. Type female: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea. 1949 Selysioneura umbratilis - LIEFTINCK, Nova Guinea (N.S.) 5: 81 (description of male).

Distribution: New Guinea. venilia

LIEFTINCK

1953 Selysioneura venilia LIEFTINCK, Treubia, 21:651. Type female: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, virgula

LIEFTINCK

1958 Selysioneura virgula LIEFTINCK, Nova Guinea, (N.S.) 10: 282. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality:Woodlark Island.

Distribution: Woodlark Island (near Papua New Guinea). Genus Tanymecosticta

LIEFTINCK

1932 Stenosticta LIEFTINCK, N o v a Guinea, 15 (5): 67. Type-species: Stenostictafissicollis LIEFTINCK, 1932 (original designation). - 1935 Tanymecosticta LIEFTINCK, N o v a Guinea, 17: 300 (a new name for Stenosticta LIEFTINCK, 1932, a junior primary homonymy of Stenosticta HAMPSON, 1912, in Lepidoptera).

capillaris

LIEFTINCK

1958 Tanymecosticta capillaris LIEFTINCK, N o v a Guinea, (N.S.) 10: 294. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Tanimbar Island.

Distribution: Indonesia: Tanimbar Island.

368 filiformis (Ris) 1898 Isosticta filiformis Ris, Ent. Nachr., 24: 325. Type male: Mus. Naturk. Berlin; type-locality: Bismarck Archipelago. 1932 Stenosticta filiformis - LIEFTINCK, Nova Guinea, 15 (5): 553. 1949 Tanymecosticta filiformis - LIEFTINCK, Treubia, 20 (2): 325 (New Britain).

Distribution: Bismarck Archipelago. fissicollis (LIEFTINCK) 1932 Stenosticta fissicollis LIEFTINCK, Nova Guinea, 15 (5): 71 (as generotype of Stenosticta LIEFTINCK, 1932). Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea. 1935 Tanymecosticta fissicollis - LIEFTINCK, Nova Guinea, 17: 300 (as generotype of Tanymecosticta LIEFTINCK, 1935).

Distribution: New Guinea. jejuna LIEFTINCK 1958 Tanymecosticta jejuna LIEFTINCK, Nova Guinea (N.S.) 10: 295. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: New Guinea.

Distribution: New Guinea, leptalea LIEFTINCK 1958 Tanymecosticta leptalea LIEFTINCK, Nova Guinea (N.S.) 10: 299. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Woodlark Island.

Distribution: Woodlark Island (near Papua New Guinea)., simonae LIEFTINCK 1984

Tanymecosticta simonae World, Utrecht, 1: 96.

LIEFTINCK, 1 9 6 9

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Key Island. Family Platycnemididae Small to medium sized Zygoptera. Anal vein and CuP present; Dc 2 - 3 times longer than wide, anterior & posterior sides subequal, MA and IRii run straight; pterostigma subtended by one cell. Old World only. 1899

Micronymphidae KARSCH, Ent. Nachr., 25: 1 6 6 (emandation). - 1 9 3 8 Platycnemididae TILLYARD& FRASER, Austr. Zool., 9:164 (with subfamiliy Platycnemidinae). -1951 Platycnemididae - S C H M I D T , Mitt, münch. ent. Ges., 41: 2 1 8 (key to genera). - 1 9 5 7 Platycnemididae - F R A S E R , R. Zool. N.S. Wales, 1957: 45 (with subfamilies Platycnemidinae, Calicneminae). - 1962 Platycnemididae- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 108 (key to genera Leptocnemis, Paracnemis, Paulianagrion, Allocnemis, Metacnemis, Stenocnemis, Platycnemis). - 1977 Platycnemididae - BELYSHEV & KHARITONOV, Nauka, Novosibirsk: 2 2 5 (key to subfamilies). 1986 Platycnemididae - KHARITONOV, Odonata, 1: 150 (key to Russian Far Eastern genera).

Subfamily Calicneminae Lower distal corner of Dc acute; anal vein reches 4 - 6 cells beyond origin of Riii. Tibia in males not dilated. 1957 Calicneminae FRASER, R . Zool. Soc. N.S. Wales, 1 9 5 7 : 4 6 (with genera Calicnemia, Coeliccia, Indocnemis, Risiocnemis, Paracnemis, Leptocnemis, Allocnemis, Arrhenocnemis, Asthenocnemis, Cyanocnemis,

Thaumatagrion,

Lochmanocnemis,

Paramecocrtemis).

369 Genus Allocnemis

SELYS

1863 Platycnemis (Allocnemis) SELYS, Synopsis des Agrionines, Add. 4: 29. Type-species: Allocnemis leucosticta SELYS, 1 8 8 6 (monotypy). - 1 8 9 0 Allocnemis- K I R B Y , Syn. Cat. Neur.-Odon., London: 1 3 1 . - 1 9 2 1 Allocnemis-Ris, Annals. S. Afr. Mus., 1 8 : 2 8 9 . - 1 9 6 2 Allocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 113.

leucosticta

SELYS

1863 Platycnemis (Allocnemis) leucosticta (in original description without author) SELYS, Synopsis des Agrionines, Add. 4: 30 (as generotype of Allocnemis SELYS, 1863). Type male: Selys's Collection; type-locality: Cape of Good Hope. 1890 Allocnemis leucosticta - KIRBY, Syn. Cat. Neur.-Odon., London: 131. 1921 Allocnemis leucosticta - Ris, Annals. S. Afr. Mus., 18: 289 (Transvaal, Cape Colony, Natal, Zuzuland). 1951 Allocnemis leucosticta - SCHMIDT, Mitt. münch. ent. Ges., 41: 227 (Natal, Transvaal, South Angola). 1962 Allocnemis leucosticta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 113.

Distribution: South Africa. mitwabae

PINHEY

I960 Allocnemis mitwabae PINHEY, Entomologist's month. Mag., 96: 258. Type male: Nat. Mus. Bulawayo; type-locality: South Africa.

Distribution: South Africa. Genus Arabicnemis

WATERSTON

1984 Arabicnemis WATERSTON, 1984 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 97. Type-species: Arabicnemis caerulea WATERSTON, 1984 (original designation).

caerulea

WATERSTON

1984 Arabicnemis caerulea WATERSTON, 1984 - loc. cit. World, Utrecht, 1: 97.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Arabia. Genus Asthenocnemis

LIEFTINCK

1949 Asthenocnemis LIEFTINCK, Nova Guinea (N.S.), 5: 88. Type-species: Asthenocnemis dera LIEFTINCK, 1949 (original designation).

stephanodera

stephano-

LIEFTINCK

1949 Asthenocnemis stephanodera LIEFTINCK, Nova Guinea, (N.S.), 5: 90 (as generotype of Asthenocnemis LIEFTINCK, 1949). Type male: Rijksmus. Natuurl, Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea.

Genus Calicnemia

STRAND

1863 Playtacnemis (Calicnemis) SELYS, Synopsis des Agrionines, Add. 4: 15. Type-species: Calicnemis eximia SELYS, 1 8 6 3 (monotypy). - 1 8 9 0 Calicnemis - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 3 1 . - 1 9 0 5 Calocnemis (misprint) - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 835 (key to species). - 1917 Calicnemis - LAIDLAW, Ree. Indian Mus., 13: 330 (key to species). - 1926 Calicnemia STRAND, Arch. Naturg., 92 (A): 46 (new name for Calicnemis

370 SELYS, 1863, a junior primary homonymy of Calicnemis LAPORTE, 1832, in Coleoptera). 1934 Calicnemia - COWLEY, Entomologist, 67: 201. - 1954 Calicnemia - LIEFTINCK, Treubia, 22 Suppl.: 45. - 1984 Calicnemia - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 97. (type-species: Calicnemis miniata SELYS, 1886).

chaseni

(CAMPION & LAIDLAW)

1928 Calicnemis chaseni CAMPION & LAIDLAW, Proc. Zool. Soc. London, 1928: 136. Type male: Raffles Mus., Singapore; type-locality: Pahang. 1931 Calicnemia chaseni - LAIDLAW, J. Fed. Mai. States Mus., 16: 193 (Perak). 1933 Calicnemia chaseni - LAIDLAW, Bull. Raffles Mus., 7: 553. 1954 Calicnemia chaseni - LIEFTINCK, Treubia, 22, Suppl.: 45 (Malaya).

Distribution: Malaya. doonensis doonensis

SANGAL & TYAGI

1985 Calicnemia miniata doonensis SANGAL & TYAGI, Proc. First Indian Symp. Odon., Madurei: 193. Type: unknown; type-locality: Nepal. 1989 Calicnemia doonensis doonensis - HÄMÄLÄINEN, Odonatologica, 18 (1): 17.

Distribution: Nepal. doonensis carminea

LIEFTINCK

1989 Calicnemia doonensis carminea LIEFTINCK, 1984 - loc. cit. HÄMÄLÄINEN, Odonatologica, 18 (1): 17.

Distribution: Nepal. erythromelas

(SELYS)

1892 Calicnemis erythromelas SELYS, Annali. Mus. civ. Stor. nat. Genova, (2) 10: 505. Type male: Mus. civ. Stor. nat., Genova, type-locality: Burma.

Distribution: Burma. eximia

(SELYS)

1863 Platycnemis (Calicnemis) eximia SELYS (in KOLLAR MS), Synopsis des Agrionines, Add. 4: 16 (as generotype of Calicnemis SELYS, 1863). Type male: Selys's Collection; type-locality: India. 1886 Calicnemis atkinsoni SELYS, (nec SELYS, 1891), Mem. Cour., 38: 131. Type female: Selys's Collection; type-locality: India. 1890 Calicnemis eximia - KIRBY, Syn. Cat. Neur.-Odon., London: 131. 1890 Calicnemis atkinsoni -KIRBY , Syn. Cat. Neur.-Odon., London: 131 [proposed synonymy with eximia

1891 1905 1917 1926 1927

(SELYS, 1 8 6 3 ) ] .

Calicnemis Calicnemis Calicnemis Calicnemia Calicnemis

eximia eximia eximia exima eximia

race atkinsoni - SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 72. - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 835 (Himalaya). - LAIDLAW, Ree. Indian Mus., 13: 327 (India). - STRAND, Arch. Naturg., 92 (A): 46. - LIEFTINCK, Tijdschr. Ent., 70: 87 (North India).

Distribution: India. imitans

LIEFTINCK

1948 Calicnemia imitans LIEFTINCK, Ark. Zool. Uppsala, 41A (10): 12. Type male: Naturhist. Riksmus., Stockholm; type-locality: Burma.

Distribution: Burma.

371 miles

(LAIDLAW)

Calicnemis atkinsoni SELYS, (nec SELYS, 1 8 8 6 ) . Annali civ. Stor. nat. Mus., Genova, ( 2 ) 10: 72. Type female: Mus. civ. Stor. nat., Genova; type-locality: Burma. 1917 Calicnemis miles LAIDLAW, Ree. Indian Mus., 13: 330 (new name for Calicnemis atkinsoni SELYS, 1891, a junior secondary homonymy of Calicnemis atkinsoni SELYS, 1886). 1891

Distribution: Burma. miniata

(SELYS)

1886 Calicnemis miniata SELYS, Mem. Cour. Acad. Belg., 38: 132. Type male: Selys's Collection; type-locality: Sikkim. 1 9 0 5 Calicnemis miniata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 5 . 1917 Calicnemis miniata - LAIDLAW, Ree. Indian Mus., 13: 328 (Darjeeling). 1926 Calicnemia miniata - STRAND, Arch. Naturg., 92 (A) 8: 46 (as generotype of Calicnemia STRAND, 1926).

Distribution: India. mortoni

(LAIDLAW)

1917 Calicnemis mortoni LAIDLAW, Ree. Indian Mus., 13: 326. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Darjeeling.

Distribution: India. mukherjeei 1984

LAHIRI

Calicnemia mukherjeei Utrecht, 1: 97.

LAHIRI, 1 9 7 6

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: India. nipalica

KIMMINS

1958 Calicnemia nipalica KIMMINS, Bull. Brit. Mus. (Nat.Hist.) Ent., 7: 350. Type male: British Museum (Nat.Hist.), London; type-locality: Nepal.

Distribution: Nepal. pulverulans

(SELYS)

1886 Calicnemis pulverulans SELYS, Mem. Cour., Acad. Belg., 38:133. Type male: Selys's Collection; type-locality: Sikkim. 1 9 0 5 Calicnemis pulverulans - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 5 . 1964 Calicnemia pulverulans - SCHMIDT, Ent. Tidschr., 85: 148 (Burma).

Distribution: India, Burma. rectangulata

LAIDLAW

1932 Calicnemia rectangulata LAIDLAW, Bull. Raffles Mus., 7: 97. Type male: Raffles Mus., Singapore; type-locality: Pahang. 1942 Calicnemia rectangulata - FRÄSER, Proc. R. ent. Soc. London; (B) 11: 96 (Malaysia). 1934 Calicnemia rectangulata - LAIDLAW, J. Fed. Mai. States Mus., 17: 552. 1954 Calicnemia rectangulata - LIEFTINCK, Treubia, 22, Suppl.: 45 (Malaya).

Distribution: Malaysia. Genus Coeiiccia

KIRBY

1857 Trichocnemis SELYS, in SAGRA, Hist. Cuba Ins.: 464. Type-species: Platycnemis membranipes RAMBUR, 1842. - 1863 Platycnemis (Trichnocnemis) - SELYS, Synopsis des Agrionines, Add.

372 4: 11. - 1890 Coeliccia KIRBY, Syn. Cat. Neur.-Odon., London: 128 (new name for Trichocnemis SELYS, 1 8 5 7 , a junior primary homonymy of Trichocnemis LECONTE, 1 8 5 2 , in Coleoptera). - 1 9 0 5 Trichocnemis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 5 (key to species). - 1912 Coeliccia - Ris, Suppl. ent. Berlin, 1: 60 (revision, key to species). - 1917 Coeliccia - L A I D L A W , Ree. Indian Mus., 13: 3 3 1 (key to species). - 1 9 3 2 Coeliccia - L A I D L A W , Ree. Indian Mus., 34: 7 (revision). - 1954 Coeliccia - LIEFTINCK, Treubia, 22 Suppl.: 45. - 1962 Trichocnemis PINHEY, Publ. cult. Comp. Diam. Angola, 59: 112 (as syn. with Leptocnemis SELYS, 1 8 8 6 ) .

albicauda

(FÖRSTER)

1907 Trichocnemis octogesima race albicauda FÖRSTER, in LAIDLAW, Fasc. Malaysenses Zool., 4: 5. Male from Perak. 1932 Coeliccia albicauda - LAIDLAW, Ree. Indian Mus., 34: 39 (redescription). 1954 Coeliccia albicauda - LIEFTINCK, Treubia, 22 Suppl.: 45 (Malaya). 1 9 8 4 Trichocnemis borneensis L A I D L A W , 1 9 0 7 , (nec SELYS, 1 8 8 6 ) - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 97, (Borneo).

Distribution: Malaysia. arcuata

LIEFTINCK

1940 Coeliccia arcuata LIEFTINCK, Treubia, 17: 353. Type male: Buitenzorg Mus.; type-locality: Borneo. 1954 Coeliccia arcuata - LIEFTINCK, Treubia, 22 Suppl.: 46 (Borneo).

Distribution: Borneo. axinocerca

LIEFTINCK

1974 Coeliccia axinocerca LIEFTINCK, Steenstrupia, 3 (12): 122. Type male: Rijksmus. Natuurl. Hist. Leiden, type-locality: Balabac Island.

Distribution: Balabac Island (near Philippines). bimaculata

LAIDLAW

1914 Coeliccia bimaculata LAIDLAW, Ree. Indian Mus., 8: 341. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Assam. 1917 Coeliccia bimaculata - LAIDLAW, Ree. Indian Mus., 13: 335. 1932 Coeliccia bimaculata - LAIDLAW, Ree. Indian Mus., 34: 12 (redescription) (India).

Distribution: India. boettcheri

SCHMIDT

1951 Coeliccia boettcheri SCHMIDT, Mitt, münch. ent. Ges., 41: 202. Type male: Mus. N a t u r k . Berlin; type-locality: Philippines.

Distribution: Philippine Islands. borneensis

(SELYS)

1886 Trichocnemis borneensis SELYS (nec L A I D L A W , 1907), Mem. Cour. Acad. Belg., 38: 116. Type female: McLachlan's Collection; type-locality: N o r t h Borneo. 1890 Coeliccia borneensis - KIRBY, Syn. Cat. Neur.-Odon., London: 128. 1932 Coeliccia borneensis - LAIDLAW, Ree. Indian Mus., 34: 41 (description of male). 1954 Coeliccia borneensis - LIEFTINCK, Treubia, 22 Suppl.:· 46 (Borneo).

Distribution: Borneo.

373

brachysticta Ris 1912 Coeliccia brachysticta Ris, Suppl. ent. Berlin, 1: 63. Type male: Ris's Collection; type-locality: Mindoro. 1932 Coeliccia brachysticta - LAIDLAW, Ree. Indian Mus., 34: 23 (redescription) (Mindoro). 1 9 3 9 Coeliccia brachysticta - N E E D H A M & G Y G E R , Philipp. J . Sei., 7 0 ( 3 ) : 2 6 9 .

Distribution: Philippine Islands. campioni

LAIDLAW

1917 Coeliccia campioni LAIDLAW, Proc. Zool. Soc. London, 1917: 224. Type male: British Museum (Nat.Hist.), London; type-locality: Borneo. 1932 Coeliccia campioni - LAIDLAW, Ree. Indian Mus., 34: 36 (redescription) (Borneo). 1 9 3 6 Coeliccia campioni - KIMMINS, J . Fed. Mai. States, Mus., 1 8 : 8 8 . 1954 Coeliccia campioni - LIEFTINCK, Treubia, 22, Suppl.: 46 (Borneo).

Distribution: Borneo. chromothorax

(SELYS)

1892 Calicnemis chromothorax SELYS, Annali Mus. civ. Stor. nat. Genova, (2) 10: 502. Type male: Mus. civ. Stor. nat., Genova; type-locality: Burma. 1917 Calicnemis chromothorax - LAIDLAW, Ree. Indian Mus., 13: 327 (Burma). 1932 Calicnemis chromothorax - LAIDLAW, Ree. Indian Mus., 34: 21 (redescription) (Shan States). 1984 Coeliccia chromothorax - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 97.

Distribution: Burma, coomansi

LIEFTINCK

1940 Coeliccia coomansi LIEFTINCK, Treubia, 17: 355. Type male: Buitenzorg Mus.; type-locality: Borneo. 1954 Coeliccia coomansi - LIEFTINCK, Treubia, 22 Suppl.: 46 (Borneo).

Distribution: Borneo. cyaneothorax

KIMMINS

1936 Coeliccia cyaneothorax KIMMINS, J. Fed. Mai. States Mus., 18: 89. Type male: British Museum (Nat.Hist.), London; type-locality: Borneo. 1954 Coeliccia cyaneothorax - LIEFTINCK, Treubia, 22 Suppl.: 46 (Borneo).

Distribution: Borneo, cyanomelas Ris 1912 Coeliccia cyanomelas Formosa. 1931 Coeliccia cyanomelas 1932 Coeliccia cyanomelas 1933 Coeliccia cyanomelas 1 9 5 6 Coeliccia cyanomelas

Ris, Suppl. ent. Berlin, 1: 66. Type male: Ris's Collection; type-locality: - SCHMIDT, Konowia, 10 (3): 183 (China). - LAIDLAW, Ree. Indian Mus., 34: 29 (South China). - SJÖSTEDT, Ark. Zool. Uppsala, 25 A (5): 18 (Szechwan). - A S A H I N A , Ent. Medd., 2 7 ( 4 - 5 ) : 2 1 8 (Central China).

Distribution: China, Taiwan.

dierli 1984

ST. QUENTIN

Coeliccia dierli Utrecht, 1: 98.

ST. QUENTIN, 1 9 7 0

Distribution: Nepal.

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

374

didyma

(SELYS)

1863 Platycnemis (Trichocnemis) didyma SELYS, Synopsis des Agrionines, Add. 4: 14. Type male: Selys Collection; type-locality: Tibet. 1890 Coeliccia didyma - KIRBY, Syn. Cat. Neur.-Odon., London: 128. 1905 Trichocnemis didyma - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 836 (Tibet, Sikkim). 1907 Trichocnemis octogesima L A I D L A W (nec SELYS, 1863), Fasc. Malaysenses Zool., 4: 2. Type male: Laidlaw's Collection; type-locality: Malaysia. 1932 Coeliccia didyma- LAIDLAW, Ree. Indian Mus., 34:16 (Burma, Simla, Assam) (redescription). 1954 Coeliccia didyma - LIEFTINCK, Treubia, 22 Suppl.: 47 (Malaya).

Distribution: East Asia. dinoceras

LAIDLAW

1925 Coeliccia dinoceras LAIDLAW, Phillip. J. Sei., 28: 562. Type male: unknown; type-locality: Mindanao. 1932 Coeliccia dinoceras - LAIDLAW, Ree. Indian Mus., 34: 33 (Mindanao). 1 9 3 9 Coeliccia dinoceras - N E E D H A M & G Y G E R , Philipp. J . Sei., 7 0 ( 3 ) : 2 6 8 (Mindanao). 1961 Coeliccia dinoceras - LIEFTINCK, Fieldiana (Zool.) 42: 138 (redescription).

Distribution: Philippine Islands. dorothea

FRASER

1933 Coeliccia dorothea FRÄSER, J. Bombay, nat. Hist. Soc., 36: 466. Type male: Fraser's Colletion; type-locality: Bengal.

Distribution: India. erici

LAIDLAW

1917 Coeliccia erici LAIDLAW, Ree. Indian Mus., 13: 334. Type male: Annandale's Collection; typelocality: Siam. 1931 Coeliccia erici - L A I D L A W , J . Fed. Mai. States, Mus., 1 6 : 195. 1932 Coeliccia erici - LAIDLAW, Ree. Indian Mus., 34: 20 (redescription). 1954 Coeliccia erici - LIEFTINCK, Treubia, 22 Suppl.: 47 (Siam, Malaya).

Distribution: Southeastern Asia. exoleta

LIEFTINCK

1961 Coeliccia exoleta LIEFTINCK, Fieldiana (Zool.), 42: 140. Type male: Lieftinck's Collection; type-locality: Mindanao.

Distribution: Philippine Islands. flavicauda flavicauda Ris 1912 Coeliccia flavicauda Ris, Suppl. ent. Berlin, 1: 64. Type male: Ris's Collection; type-locality: Formosa. 1932 Coeliccia flavicauda - LAIDLAW, Ree. Indian Mus., 34: 28 (Formosa). 1951 Coeliccia flavicauda flavicauda - ASAHINA, Kontyü, 19: 17.

Distribution: Taiwan. flavicauda masakii

ASAHINA

1951 Coeliccia flavicauda masakii ASAHINA, Kontyü, 19: 17. Type male: Asahina's Collection; typelocality: Ryukyu Island.

Distribution: Ryukyu Island.

375 flavostriata

LAIDLAW

1917 Coeliccia flavostriata LAIDLAW, Proc. Zool. Soc. London, 1917: 223. Type male: British Museum (Nat.Hist.), London; type-locality: Borneo. 1932 Coeliccia flavostriata - LAIDLAW, Ree. Indian Mus., 34: 34 (redescription) (Borneo). 1954 Coeliccia flavostriata - LIEFTINCK, Treubia, 22 Suppl.: 47 (Borneo).

Distribution: Borneo. fraseri

LAIDLAW

1932 Coeliccia fraseri LAIDLAW, Ree. Indian Mus., 34: 14. Type male: British Museum (Nat.Hist.), London; type-locality: Assam.

Distribution: India. furcata

HÄMÄLÄINEN

1986 Coeliccia furcata HÄMÄLÄINEN, Odonatologica, 15 (4): 445. Type male: Hämäläinen's Collection; type-locality: Burma.

Distribution: Burma. lieftincki

LAIDLAW

1932 Coeliccia lieftincki LAIDLAW, Ree. Indian Mus., 34: 32. Type male: Lieftinck's Collection; type-locality: Java. 1934 Coeliccia lieftincki - LIEFTINCK, Treubia, 14 (4): 392 (Java). 1954 Coeliccia lieftincki - LIEFTINCK, Treubia, 22 Suppl.: 47 (Java).

Distribution: Java. loogali

LAIDLAW

1932 Coeliccia loogali LAIDLAW (in FRASER MS), Ree. Indian Mus., 34: 26. Type male: Wall's Collection; type-locality: Maymyo. 1964 Coeliccia loogali - SCHMIDT, Ent. Tidschr., 85: 149 (Burma).

Distribution: Burma. loringae

LAIDLAW

1932 Coeliccia loringae LAIDLAW, Ree. Indian Mus., 34:17. Type male: Laidlaw's Collection; typelocality: Shan States.

Distribution: Burma. macrostigma 1917 Coeliccia Museum 1932 Coeliccia 1954 Coeliccia

LAIDLAW

macrostigma LAIDLAW, Proc. Zool. Soc. London, 1917: 225. Type male: British (Nat.Hist.), London; type-locality: Borneo. macrostigma - LAIDLAW, Ree. Indian Mus., 34: 35 (redescription) (Borneo). macrostigma - LIEFTINCK, Treubia, 22 Suppl.: 48 (Borneo).

Distribution: Borneo. membranipes membranipes

(RAMBUR)

1842 Platycnemis membranipes RAMBUR, Hist. nat. Ins. Neuropt., Paris: 240. Type female: Nation. Mus. nat. Hist., Paris; type-locality: Java. 1863 Platycnemis (Trichocnemis) membranipes - SELYS, Synopsis des Agrionines, Add. 4: 12 (as generotype of Trichocnemis SELYS, 1863). 1863 Platycnemis (Trichocnemis) silenta HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 13. Type male: Halle Museum; type-locality: Java.

376 1890 Coeliccia membranipes - KIRBY, Syn. Cat. Neur.-Odon., London: 128. 1890 Platycnemis (Trichocnemis) silenta- KIRBY, Syn. Cat. Neur.-Odon., London: [proposed synonymy with membranipes membranipes (RAMBUR, 1842)]. 1927 Coeliccia membranipes - Ris, Zool. Meded., Leiden, 10 (1): 16 (Sumatra, Java). 1932 Coeliccia membranipes - LAIDLAW, Ree. Indian Mus., 34: 30 (Borneo, Java). 1934 Coeliccia membranipes - SCHMIDT, Arch. Hydrobiol., Suppl.: 13 335 (Java). 1934 Coeliccia membranipes membranipes - LIEFTINCK, Treubia, 14 (4): 393 (Java). 1935 Coeliccia membranipes - LIEFTINCK, Misc. Zool. Sum., 92-93: 8 (Sumatra). 1951 Coeliccia membranipes - SCHMIDT, Mitt, miinch. ent. Ges., 41: 209 (Java). 1954 Coeliccia membranipes membranipes- LIEFTINCK, Treubia, 22 Suppl.:48.

Distribution: Sumatra, Java, Borneo, membranipes nemoricola

LAIDLAW

1912 Coeliccia (Trichocnemis) nemoricola LAIDLAW, J. Straits. Asiat. Soc., 63: 92. Type male: Laidlaw's Collection; type-locality: Borneo. 1915 Coeliccia nemoricola - LAIDLAW, Proc. Zool. Soc. London, 1915: 37. 1918 Coeliccia membranipes race nemoricola - LAIDLAW, Proc. Zool. Soc. London, 1918: 230. 1954 Coeliccia nemoricola - LIEFTINCK, Treubia, 22 Suppl.: 48 (Borneo).

Distribution: Borneo. membranipes nereis

LIEFTINCK

1948 Coeliccia membranipes nereis LIEFTINCK, Treubia, 19 (2): 287. Type male: Buitenzorg Museum; type-locality: Engano Island. 1954 Coeliccia membranipes nereis - LIEFTINCK, Treubia, 22 Suppl.: 48 (Engano).

Distribution: Engano Island. montana

FRASER

1933 Coeliccia montana FRÄSER, J. Siam Soc. nat. Hist. Suppl.: 9: 116. Type male: British Museum (Nat.Hist.), London; type-locality: Laos.

Distribution: Laos, nigrescens

LAIDLAW 1931 Coeliccia nigrescens LAIDLAW, J. Fed. Mai. States Mus., 16: 196. Type male: Raffles Museum, Singapore; type-locality: Siam. 1954 Coeliccia nigrescens - LIEFTINCK, Treubia, 22 Suppl.: 49 (Siam).

Distribution: Thailand. nigrohamata LAIDLAW 1917 Coeliccia nigrohamata LAIDLAW, Proc. Zool. Soc. London, 1917: 228. Type male: British Museum (Nat.Hist.), London; type-locality: Borneo. 1932 Coeliccia nigrohamata - LAIDLAW, Ree. Indian Mus., 34: 37 (redescription) (Borneo). 1936 Coeliccia nigrohamata - KIMMINS, J. Fed. Mai. States. Mus., 18: 88. 1954 Coeliccia nigrohamata - LIEFTINCK, Treubia, 22 Suppl.: 49 (Borneo).

Distribution: Borneo. octogesima

(SELYS)

1 8 6 3 P l a t y c n e m i s ( T r i c h o c n e m i s ) o c t o g e s i m a SELYS ( n e c LAIDLAW, 1 9 0 7 ) , S y n o p s i s d e s A g r i o n i n e s ,

Add. 4: 13. Type male: McLachlan's Collection; type-locality: Singapore. 1886 Trichocnemis octogesima - SELYS, Mem. Cour. Acad. Belg., 38: 117. 1890 Coeliccia octogesima - KIRBY, Syn. Cat. Neur.-Odon., London: 128. 1 9 3 2 Coeliccia

octogesima

- LAIDLAW, R e e . I n d i a n M u s . , 3 4 : 4 0 ( r e d e s c r i p t i o n ) .

377 1935 Coeliccia octogesima - LIEFTINCK, Misc. Zool. Sum., 92-93: 8 (Sumatra). 1940 Coeliccia octogesima - LIEFTINCK, Treubia, 17: 357 (redescription of holotype). 1954 Coeliccia octogesima - LIEFTINCK, Treubia, 22 Suppl.: 49 (Malaya, Sumatra, Borneo).

Distribution: Malaysia, Sumatra, Borneo. palawani

LIEFTINCK

1940 Coeliccia palawani LIEFTINCK, Treubia, 17: 368. Type male: Lieftinck's Collection; type-locality: Palawan. 1984 Coeliccia palawana - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 98.

Distribution: Palawan. poungyi

FRASER

1924 Coeliccia poungyi FRÄSER, J. Bombay nat. Hist. Soc., 30: 52. Type male: British Museum (Nat.Hist.), London; type-locality: India. 1932 Coeliccia poungyi - LAIDLAW, Ree. Indian Mus., 34: 24 (redescription) (Upper Burma).

Distribution: India, Burma. pyriformis

LAIDLAW

1932 Coeliccia pyriformis LAIDLAW, (in FRASER MS), Ree. Indian Mus., 34: 26. Type male: Steven's Collection; type-locality: Tonkin. 1984 Coeliccia pyriformis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 98 (China).

Distribution: Vietnam, China. renifera

(SELYS)

1886 Trichocnemis renifera SELYS, Mem. Cour. Acad. Belg., 38: 119. Type male: Selys's Collection; type-locality: Darjeeling. 1890 Coeliccia renifera - KIRBY, Syn. Cat. Neur.-Odon., London: 128. 1905 Trichocnemis renifera - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 836 (Sikkim). 1917 Coeliccia renifera - LAIDLAW, Ree. Indian Mus., 13: 333 (India). 1921 Coeliccia renifera - LAIDLAW, Ree. Indian Mus., 34: 18 (redescription) (India). 1954 Coeliccia kumaoensis SINGH & BAIJAL, Agra Univ. J. Res., 3 (2): 398. Type male: Nation. Coll. Zool. Surv. India, Calcutta; type-locality: Himalaya. 1964 Calicnemis mahesi SAHNI, Agra Univ. J. Res., 13: 82. Type male: Sahni's Collection; typelocality: Himalaya. 1989 Coeliccia kumaoensis - HÄMÄLÄINEN, Odonatologica, 18 (1): 17 [proposed synonymy with renifera

(SELYS, 1886)].

1989 Calicnemis mahesi - HÄMÄLÄINEN, Odonatologica, 18 (1): 17 [proposed synonymy with renifera (SELYS, 1886)].

Distribution: India. resecta

LIEFTINCK

1953 Coeliccia resecta LIEFTINCK, Treubia, 22: 236. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1954 Coeliccia resecta - LIEFTINCK, Treubia, 22 Suppl.: 49 (Borneo).

Distribution: Borneo.

378 ryukyuensis ryukyuensis ASAHINA 1951 Coeliccia ryukyuensis ASAHINA, Kontyü, 19:16. Type male: Asahina's Collection; type-locality: Ryukyu Island.

Distribution: Ryukyu Island. ryukyuensis amamii ASAHINA 1984

Coeliccia ryukyuensis amamii the World, Utrecht, 1: 97.

ASAHINA, 1 9 6 2

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of

Distribution: Ryukyu Island. scutellum scutellum

LAIDLAW

1932 Coeliccia scutellum scutellum LAIDLAW, Ree. Indian Mus., 34: 22. Type male: British Museum (Nat.Hist.), London; type-locality: Tonkin.

Distribution: Vietnam. scutellum hainanensis

LAIDLAW

1932 Coeliccia sacutellum hainanensis LAIDLAW, Ree. Indian Mus., 34: 23. Type male: British Museum (Nat.Hist.), London; type-locality: Hainan.

Distribution: China. simillima

LAIDLAW

1917 Coeliccia simillima LAIDLAW, Ree. Indian Mus., 13: 334. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Bukit Besar. 1954 Coeliccia simillima - LIEFTINCK, Treubia, 22 Suppl.: 47 [incorrent synonymy with didyma (SELYS, 1 8 6 3 ) ] .

Distribution: India. svihleri 1984

ASAHINA

Coeliccia svihleri Utrecht, 1: 99.

ASAHINA, 1 9 7 0

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Assam. vacca

LAIDLAW

1932 Coeliccia vacca LAIDLAW, Ree. Indian Mus., 34: 15. Type female: Nation. Coll. Zool. Surv. India, Calcutta; type-locality: Garo Hills.

Distribution: India. werneri LIEFTINCK 1961 Coeliccia werneri LIEFTINCK, Fieldiana (Zool.), 42: 137. Type male: Lieftinck's Collection; type-locality: Palawan.

Distribution: Palawan. Genus Cyanocnemis

LIEFTINCK

1949 Cyanocnemis LIEFTINCK, Nova Guinea (N.S.) 5: 107. Type-species: Cyanocnemis LIEFTINCK, 1 9 4 9 ( o r i g i n a l

designation).

aureofrons

379

aureofrons

LIEFTINCK

1949 Cyanocnemis aureofrons LIEFTINCK, N o v a Guinea (N.S.) 5:107 (as generotype of Cyanocnemis LIEFTINCK, 1949). Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Idiocnemis 1878

SELYS

Idiocnemis SELYS, Mitt. Zool. Mus. Dresden, 3 : 3 2 1 . Type-species: Idiocnemis inornata SELYS, 1 8 7 8 (designated by K I R B Y , Syn. Cat. Neur.-Odon., London: 1 2 7 ) . - 1 9 3 7 Idiocnemis - LIEFTINCK, Nova Guinea (N.S.) 1: 286.

bidentata

SELYS

Idiocnemis bidentata SELYS (nec FÖRSTER, 1 8 9 8 ) . Mitt. Zool. Mus. Dresden, 3: 3 2 1 . Type male: R. Mus. Brussels; type-locality: Papua. 1937 Idiocnemis bidentata - LIEFTINCK, Nova Guinea (N.S.) 1: 288 («description). 1878

Distribution: New Guinea. dagnyae

LIEFTINCK

1 9 8 4 I d i o c n e m i s d a g n y a e LIEFTINCK, 1 9 5 8 - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 99.

Distribution: New Guinea. fissidens 1984

LIEFTINCK

Idiocnemis fissidens Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: New Guinea. huonensis 1984

LIEFTINCK

Idiocnemis huonensis Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: New Guinea. inaequidens

LIEFTINCK

1932 Idiocnemis inaequidens LIEFTINCK, Nova Guinea 15 (5): 19. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea. 1937 Idiocnemis inaequidens - LIEFTINCK, N o v a Guinea, (N.S.) 1: 289 (description of female). 1 9 8 4 Idiocnemis bidentata FÖRSTER, 1 8 9 8 (nec SELYS, 1 8 7 8 ) - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 99.

Distribution: New Guinea. inornata

SELYS

1878 Idiocnemis inornata SELYS, Mitt. Zool. Mus. Dresden, 3: 321. Type male: R. Mus. Brussels; type-locality: Papua. 1890 Idiocnemis inornata - KIRBY, Syn. Cat. Neur.-Odon., London: 127 (as generotype of Idiocnemis SELYS, 1 8 7 8 ) .

1932 Idiocnemis inornata - LIEFTINCK, Nova Guinea, (N.S.) 1: 286 (redescription). 1932 Idiocnemis chloropleura LIEFTINCK, Nova Guinea, 15 (5): 18. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea. 1949 Idiocnemis inornata - LIEFTINCK, Treubia, 20 (2): 336 (Bismarck Archipelago).

Distribution: New Guinea, Bismarck Archipelago.

380 kimminsi 1984

LIEFTINCK

Idiocnemis kimminsi Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: New Guinea. leonardi 1984

LIEFTINCK

Idiocnemis leonardi Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

Distribution: New Guinea. leonorae

LIEFTINCK

1949 Idiocnemis leonorae LIEFTINCK Nova Guinea (N.S.) 5: 95. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. louisiadensis 1984

LIEFTINCK

Idiocnemis louisiadensis World, Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: New Guinea. mertoni Ris 1913 Idiocnemis mertoni Ris, Abh. Senckenb. Ges., 34: 507. Type male: Mus. Naturk., Berlin; typelocality: Aru.

Distribution: Aru. nigriventris

LIEFTINCK

1937 Idiocnemis nigriventris LIEFTINCK, N o v a Guinea, (N.S.) 1: 290. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. obliterate

LIEFTINCK

1932 Idiocnemis obliterata LIEFTINCK, Nova Guinea, 15 (5): 15. type-locality: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, pruinescens

LIEFTINCK

1949 Idiocnemis pruinescens LIEFTINCK, Nova Guinea (N.S.) 5: 96. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. strumidens 1984

LIEFTINCK

Idiocnemis strumidens LIEFTINCK, World, Utrecht, 1: 99.

Distribution: New Guinea.

1958

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

381 zebrina zebrina LIEFTINCK 1984

Idiocnemis zebrina zebrina the World, Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of

DAVIES & TOBIN,

The Dragonflies of

Distribution: New Guinea. zebrina sufficiens LIEFTINCK 1984

Idiocnemis zebrina sufficiens the World, Utrecht, 1: 99.

LIEFTINCK, 1 9 5 8

- loc. cit.

Distribution: New Guinea. Genus Indocnemis LAIDLAW 1917

Indocnemis L A I D L A W , Ree. Indian Mus., 13: 3 2 5 . Type-species: Indocnemis kempi L A I D L A W , 1917 (original designation). - 1954 Indocnemis - LIEFTINCK, Treubia, 22 Suppl.: 50.

kempi LAIDLAW Indocnemis kempi L A I D L A W , Ree. Indian Mus., 13:326 (as generotype of Indocnemis L A I D L A W , 1917). Type male: Nation. Coll. zool. Surv. India, Calcutta;, type-locality: India. 1937 Indocnemis kempni (misprint) - ST. QUENTIN, Konowia, 16 (1): 85 (redescription) (Assam). 1 9 8 4 Indocnemis kempi - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 9 9 [incorrect synonymy with orang ( F Ö R S T E R , 1 9 0 7 ) ] . 1917

Distribution: India. orang (FÖRSTER) 1907 Trichocnemis orang FÖRSTER, Fase. Malayensis Zool., 4. Odonata, 2: 2. Type male: Förster's Collection; type-locality: Perak. 1912 Coeliccia (?) orang - Ris, Suppl. ent. Berlin, 1. 67 (redescription). 1 9 3 1 Indocnemis orang - L A I D L A W , J . Fed. Mai. States Mus., 16: 1 9 3 (Malaya). 1935 Indocnemis orang - LIEFTINCK, Misc. Zool. Sum., 92-93: 8 (Sumatra). 1954 Indocnemis orang - LIEFTINCK, Treubia, 22 Suppl.: 50 (Siam, Malaya).

Distribution: Malaysia, Sumatra. Genus Leptocnemis SELYS 1882

Hemicnemis SELYS, Congr. Sei. Ass. franc., 10: 6 6 8 . Type-species: Trichocnemis cyanops SELYS, 1869 (monotypy). - 1886 Leptocnemis SELYS, Mem. Cour. Acad. Belg., 38: 223 (new name for Hemicnemis SELYS, 1 8 8 2 , a junior primary homonymy of Hemicnemis M Ü L L E R , 1 8 4 0 , in Echinodermata). - 1890 Leptocnemis - KIRBY, Syn. Cat. Neur.-Odon., London: 128. 1962 Leptocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 112.

cyanops(SELYS) 1869 Trichocnemis cyanops SELYS, Ann. Soc. Ent. Belg., 12: 28. Type male: Nation. Hist. Nat. Mus., Paris; type-locality: Seychelles. 1869 Trichocnemis bilineata SELYS, Ann. Soc. Ent. Belg., 12: 28. Type: not designated; type-locality: Seychelles. 1869 Trichocnemis bivittata SELYS, Zool. Ree., 1869: 453 (nec 456) (error for bilineata, since bivittata does not occur in Journal cited). 1882 Hemicnemis cyanops - SELYS, Congr. Ass. franc. Sei., 10: 668 (as generotype of Hemicnemis SELYS, 1882).

1886 Leptocnemis cyanops - SELYS, Mem. Cour. Acad. Belg., 38:112 (as generotype of Leptocnemis SELYS, 1886).

382 1886 1890 1951 1962

Leptocnemis Leptocnemis Leptocnemis Leptocnemis

bilineata cyanops cyanops cyanops

-

SELYS, Mem. Cour. Acad. Belg., 38: 113. KIRBY, Syn. Cat. Neur.-Odon., London: 128. SCHMIDT, Mitt, münch. ent. Ges., 41: 219. PINHEY, Publ. cult. Comp. Diam. Angola, 59: 112.

Distribution: Seychelles.

Genus Lieftinckia

KIMMINS

1957 Lieftinckia KIMMINS, Bull. Brit. Mus. (Nat.Hist.) Ent., 5: 312. Type species: Lieftinckia lomonensis KIMMINS, 1957 (original designation).

isabellae

sa-

LIEFTINCK

1987 Lieftinckia isabellae LIEFTINCK, Tijdschr. Ent., 129 (9): 274. Type male: Rijksmus. Natuurl.Hist., Leiden; type-locality: Solomon Island.

Distribution: Solomon Islands, kimminsi

LIEFTINCK 1963 Lieftinckia kimminsi LIEFTINCK, Nova Guinea (Zool.) 21: 537. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Solomon Island.

Distribution: Solomon Islands, lairdi

LIEFTINCK 1963 Lieftinckia lairdi LIEFTINCK, Nova Guinea (Zool.) 21: 534. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Solomon Island. 1987 Lieftinckia lairdi - LIEFTINCK, Tijdschr. Ent., 129 (9): 271 (redescription).

Distribution: Solomon Islands. malaitae

LIEFTINCK

1987 Lieftinckia malaitae LIEFTINCK, Tijdschr. Ent., 129 (9): 272. Type male: Rijksmus. Natuurl.Hist., Leiden; type-locality: Solomon Island.

Distribution: Solomon Islands, ramosa

LIEFTINCK 1987 Lieftinckia ramosa LIEFTINCK, Tijdschr. Ent., 129 (9): 275. Type male: Rijksmus. Natuurl.Hist., Leiden; type-locality: Solomon Island.

Distribution: Solomon Islands. salomonensis

KIMMINS

1957 Lieftinckia salomonensis KIMMINS, Bull. Brit. Mus. (Nat.Hist.), Ent., 5: 314 (as generotype of Lieftinckia KIMMINS, 1957). Type male: British Museum (Nat. Hist.), London; type-locality: Guadalcanal. 1963 Lieftinckia salomonensis - LIEFTINCK, Nova Guinea (Zool.), 21: 527. 1987 Lieftinckia salomonensis - LIEFTINCK, Tijdschr. Ent., 129 (9): 271 (redescription).

Distribution: Guadalcanal.

Genus Lochmaeocnemis

LIEFTINCK

1949 Lochmaeocnemis LIEFTINCK, Nova Guinea (N.S.) 5: 100. Type species: Lochmaeocnemis lacodora

LIEFTINCK, 1 9 4 9 ( o r i g i n a l d e s i g n a t i o n ) .

ma-

383

malacodora

LIEFTINCK

1949 Lochmaeocnemis malacodora LIEFTINCK, Nova Guinea, (N.S.), 5: 100 (as generotype of Lochmaeocnemis LIEFTINCK, 1949). Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Mesocnemis

KARSCH

1891 Mesocnemis KARSCH, Ent. Nachr., 17 (5): 66. Type species: Mesocnemis singularis KARSCH, 1891 (monotypy). - 1936 Mesocnemis - LONGFIELD, Trans. R. ent. Soc. London, 85: 471. 1962 Mesocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 114 (incorrect synonymy w i t h Metacnemis

SELYS, 1 8 6 3 ) . - 1 9 8 4 Mesocnemis

- DAVIES & TOBIN, T h e D r a g o n f l i e s o f

the World, Utrecht, 1: 100 (as valid taxon).

robusta

(SELYS)

1886 Metacnemis robusta Wien; type-locality: 1951 Metacnemis robusta 1962 Metacnemis robusta 1984 Mesocnemis robusta System).

SELYS, Mem. Cour. Acad. Belg., 38: 135. Type male: Naturhist. Mus., Nubia. - SCHMIDT, Mitt, miinch. ent. Ges., 41: 222. - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 114 (Egypt, Sudan). - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 100 (Nile

Distribution: Nubia, Sudan, Egypt, singularis

KARSCH 1891 Mesocnemis singularis KARSCH, Ent. Nachr., 17 (5): 67. Type male: Mus. Naturk., Berlin; type-locality: Cameroon (as generotype of Mesocnemis KARSCH, 1891). 1912 Mesocnemis irregularis KARSCH (nom. nud., without description) in MARTIN, Fenill. jeun. Nat., (5) 42: 98. 1928 Metacnemis pruinosa FRASER, Trans, ent. Soc. London, 76: 129. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1936 Mesocnemis singularis - LONGFIELD, Trans, ent. Soc. London, 85: 471. 1936 Metacnemis pruinosa - LONGFIELD, Trans, ent. Soc. London, 85: 471 (proposed synonymy w i t h singularis

KARSCH, 1 8 9 1 ) .

1951 Metacnemis singularis - SCHMIDT, Mitt, miinch. ent. Ges., 41: 221 (Cameroon). 1962 Metacnemis singularis — PINHEY, Publ. cult. Comp. Diam. Angola, 59:114 (Continental Ethiopian region).

Distribution: Central Africa. Genus Metacnemis

SELYS

1863 Platycnemis (Metacnemis) SELYS, Bull. Acad. r. Belg., (2) 16: 160. Type species: Platycnemis (Metacnemis) valida HÄGEN, 1863 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 131). - 1890 Metacnemis - KIRBY, Syn. Cat. Neur.-Odon., London: 131. - 1905 Metacnemis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 835. - 1921 Metacnemis - Ris, Annls S. Afr. Mus., 18: 296. - 1951 Metacnemis - SCHMIDT, Mitt, miinch. ent. Ges., 41: 221 (key to African species). - 1962 Metacnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 114.

angusta

SELYS

1863 Platycnemis (Metacnemis) angusta SELYS, Synopsis des Agrionines, Add. 4: 18. Type female: Latreille's Collection; type-locality: Cap of Good Hope. 1890 Metacnemis angusta - KIRBY, Syn. Cat. Neur.-Odon., London: 131.

384 1905 Metacnemis angusta - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 835 (Nubia). 1962 Metacnemis angusta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 115 (Cape Province).

Distribution: South Africa. dupuyi

LEGRAND

1982 Metacnemis dupuyi LEGRAND, Odonatologica, 11 (2): 153. Type male: Legrand's Collection; type-locality: Senegal.

Distribution: Senegal. secundaris 1984

AGUESSE

Metacnemis secundaris AGUESSE, World, Utrecht, 1: 100.

1968

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: Madagascar. valida

HÄGEN

1863 Platycnemis (Metacnemis) valida HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 17. Type male: Hagen's Collection; type-locality: Cape of Good Hope. 1890 Metacnemis valida - KIRBY, Syn. Cat. Neur.-Odon., London: 131 (as generotype of Metacnemis SELYS, 1863). 1921 Metacnemis valida - Ris, Annals S. Afr. Mus., 18: 296 (Cap). 1951 Metacnemis valida - SCHMIDT, Mitt, münch. ent. Ges., 41: 221.

Distribution: South Africa. Genus Oreocnemis

PINHEY

1971 Oreocnemis PINHEY, Arnoldia, Rhodesia, 5 (9): 1. Type species: Oreocnemis phoenix 1971 (original designation).

phoenix

PINHEY,

PINHEY

1971 Oreocnemis phoenix PINHEY, Arnoldia, Rhodesia, 5 (9): 2 (as generotype of Oreocnemis 1971). Type male: Pinhey's Collection; type-locality: Malawi.

PINHEY,

Distribution: Malawi. Genus Paracnemis

MARTIN

1903 Paracnemis M A R T I N , Bull. Mus. Paris, 8: 509. Type species: Paracnemis alluaudi M A R T I N , 1903 (original designation). - 1917 Ciliagrion SJÖSTEDT, Ark.Zool., Uppsala, 11 (13): 5. Type species: Ciliagrion madagascariense SJÖSTEDT, 1917 (original designation). - 1951 Ciliagrion - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 166 (proposed synonymy with Paracnemis MARTIN, 1903). - 1962 Paracnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

alluaudi

MARTIN

1903 Paracnemis alluaudi M A R T I N , Bull. Mus. Paris, 8: 509 (as generotype of Paracnemis M A R T I N , 1903). Type male: lost; type-locality: Madagascar. 1917 Ciliagrion madagascariense SJÖSTEDT, Ark. Zool., Uppsala, 11 (13): 6 (as generotype of Ciliagrion SJÖSTEDT, 1917). Type male: Naturhist. Riksmus., Stockholm; type-locality: Madagascar. 1951 Ciliagrion madagascariense - SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 168 (proposed synonymy with alluaudi M A R T I N , 1 9 0 3 ) . 1962 Paracnemis alluaudi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 112.

Distribution: Madagascar.

385

Genus Paramecocnemis

LIEFTINCK

1932 Paramecocnemis LIEFTINCK, Nova Guinea, 15 (5): 20. Type species: Paramecocnemis throstigma LIEFTINCK, 1932 (original designation).

erythrostigma

ery-

LIEFTINCK

1932 Paramecocnemis erythrostigma LIEFTINCK, Nova Guinea, 15 (5): 22 (as generotype of Paramecocnemis LIEFTINCK, 1 9 3 2 ) . Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. stillacruoris

LIEFTINCK

1956 Paramecocnemis stilla-cruoris LIEFTINCK, Nova Guinea (N.S.) 7: 254. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Rhyacocnemis

LIEFTINCK

1956 Rhyacocnemis LIEFTINCK (nv. Torrenticnemis), Nova Guinea (N.S.) 7: 250. Type species: Rhyacocnemis sufficiens LIEFTINCK, 1956 (original designation).

prothoracica

LIEFTINCK

1987 Rhyacocnemis prothoracica LIEFTINCK, Tijdschr. Ent., 129 (9): 265. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. sufficiens

LIEFTINCK

1956 Rhyacocnemis sufficiens LIEFTINCK, N o v a Guinea (N.S.) 7: 251 (as generotype of Rhyacocnemis LIEFTINCK, 1956). Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: d'Entrecasteaux Island.

Distribution: D'Entrecasteaux I. (Fergusson Island). Genus Risiocnemis

COWLEY

1863 Hypocnemis HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 9. Type species: Hypocnemis serrata H A G E N , 1 8 6 3 (monotypy). - 1 8 8 6 Prionocnemis SELYS, Mem. Cour. Acad. Belg., 38: 2 2 3 (new name for Hypocnemis H A G E N , 1 8 6 3 , a junior primary homonymy of Hypocnemis CABANIS, 1 8 4 7 in Aves). - 1 9 3 4 Risiocnemis C O W L E Y , Entomologist, 67: 2 0 4 (new name for Prionocnemis SELYS, 1 8 8 6 , a junior primary homonymy of Prionocnemis SCHIODTE, 1 8 3 7 in Hymenoptera). Type species: Hypocnemis serrata H A G E N , in SELYS, 1 8 6 3 (original designation). - 1954 Risiocnemis - LIEFTINCK, Treubia, 22: Suppl.: 50.

appendiculata

(BRAUER)

1868 Hypocnemis appendiculata BRAUER, Verh. Zool.-Bot. Ges., Wien, 18: 548. Type male: ? Naturhist. Mus. Wien; type-locality: Mindanao. 1890 Prionocnemis appendiculata - KIRBY, Syn. Cat. Neur.-Odon., London: 127. 1939 Risiocnemis appendiculata - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 276 (Mindanao).

Distribution: Philippine Islands.

386

atripes (NEEDHAM & GYGER) comb. nov. 1941 Prionocnemis atripes N E E D H A M locality: Philippines.

& GYGER,

Philipp. J. Sei., 74: 146. Type male: not given; type-

Distribution: Philippine Islands, atropurpurea (BRAUER) comb. nov. 1868 Hypocnemis atropurpurea BRAUER, Verh. Zool.-Bot. Ges., Wien, 18: 549. Type male: ? Naturhist. Mus. Wien; type-locality: Manilla. 1890 Prionocnemis atropurpurea - KIRBY, Syn. Cat. Neur.-Odon., London: 127. 1 9 3 9 Prionocnemis atropurpurea - N E E D H A M & G Y G E R , Philipp. J . Sei., 70 ( 3 ) : 2 7 3 (redescription of female) (Luzon).

Distribution: Philippine Islands. cornuta (BRAUER) comb. nov. 1868 Hypocnemis cornuta BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 548. Type male: ? Naturhist. Mus. Wien; type-locality: Philippines. 1890 Prionocnemis cornuta - KIRBY, Syn. Cat. Neur.-Odon., London: 127. 1891 Hypocnemis cornuta - SELYS, Ann. Soc. Esp. nat. Hist., 20: 217 (description of female). 1 9 3 9 Prionocnemis cornuta - N E E D H A M & G Y G E R , Philipp. J . Sei., 70 ( 3 ) : 2 7 3 .

Distribution: Philippine Islands. erythrura (BRAUER) comb. nov. 1868 Hypocnemis erythrura BRAUER, Verh. Zool.-Bot. Ges., Wien, 18: 550. Type male: ? Naturhist. Mus., Wien; type-locality: Philippines. 1890 Prionocnemis erythrura - KIRBY, Syn. Cat. Neur.-Odon., London: 127. 1 9 3 9 Prionocnemis erythrura - NEEDHAM & G Y G E R , Philipp. J . Sei., 70 ( 3 ) : 2 7 4 .

Distribution: Philippine Islands. flammea (SELYS) comb. nov. 1882 Hypocnemis flammea SELYS, Ann. Soc. Esp. nat. Hist., 11: 26. Type male: not given; typelocality: Luzon. 1890 Prionocnemis flammea - KIRBY, Syn. Cat. Neur.-Odon., London: 127.

Distribution: Philippine Islands. haematopus (SELYS) 1882 Hypocnemis haematopus SELYS, Ann. Soc. Esp. nat. Hist., 11: 25. Type male: not given; typelocality: Mindanao. 1890 Prionocnemis haematopus - KIRBY, Syn. Cat. Neur.-Odon., London: 127. 1936 Risiocnemis reflexa KIMMINS, J . Fed. Mai. States Mus., 18: 92. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1939 Priocnemis haematopus - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 274 (Luzon). 1954 Risiocnemis reflexa - LIEFTINCK, Treubia, 22 Suppl.: 50 (Borneo).

Distribution: Philippine Islands, Borneo. ignea (BRAUER) comb. nov. 1868 Hypocnemis ignea BRAUER, Verh. Zool.-Bot. Ges., Wien, 18: 547. Type male: ? Naturhist. Mus. Wien; type-locality: Philippines. 1890 Prionocnemis ignea - KIRBY, Syn. Cat. Neur.-Odon., London: 127. 1 9 3 9 Prionocnemis ignea - NEEDHAM & G Y G E R , Philipp. J . Sei., 70 ( 3 ) : 2 7 5 (Luzon). Distribution: P h i l i p p i n e I s l a n d s .

387 incisa KIMMINS 1936 Risiocnemis incisa KIMMINS, J. Fed. Mai. States Mus., 18: 91. Type female: British Museum (Nat. Hist.), London; type-locality: Borneo. 1954 Risiocnemis incisa - LIEFTINCK, Treubia, 22 Suppl.: 50 (Borneo).

Distribution: Borneo. rubripes (NEEDHAM & GYGER) comb. nov. 1939 Prionocnemis rubripes NEEDHAM type-locality: Mindanao.

& GYGER,

Philipp. J. Sei., 70 (3): 275. Type male: not given;

Distribution: Philippine Islands. serrata (HÄGEN) 1863 Hypocnemis serrata HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 10 (as generotype of Hypocnemis HÄGEN, 1863). Type male: Hagen's Collection; type-locality: Manila. 1886 Prionocnemis serrata - SELYS, Mem. Cour. Acad. Belg., 38: 223 (as generotype of Prionocnemis SELYS, 1886).

1934 Risiocnemis serrata - C O W L E Y , Entomologist, 67: 204 (as generotype of Risiocnemis 1934). 1939 Prionocnemis serrata - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 270 (Luzon).

COWLEY,

Distribution: Philippine Islands. tendipes (NEEDHAM & GYGER) comb. nov. 1941 Prionocnemis tendipes NEEDHAM type-locality: Philippines.

& GYGER,

Philipp. J. Sei., 74: 147. Type male: not given;

Distribution: Philippine Islands. Genus Solomocnemis LIEFTINCK 1987 Solomocnemis LIEFTINCK, Tijdschr. Ent., 129 (9): 268. Type species: Solomocnemisgerdae TINCK, 1987 (original designation).

LIEF-

gerdae LIEFTINCK 1987 Solomocnemis gerdae LIEFTINCK, Tijdschr. Ent., 129 (9): 269. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Solomon Island.

Distribution: Solomon Islands. Genus Stenocnemis KARSCH 1899 Stenocnemis K A R S C H , Ent. Nachr., 25 (11): 6. Type species: Allocnemis pachystigma SELYS, 1886 (monotypy). - 1919 Stenocnemis - MUNZ, Mem. Am. ent. Soc., 3 (1): 54. - 1962 Stenocnemis- PINHEY, Publ. cult. Comp. Diam. Angola, 59: 113.

pachystigma (SELYS) 1886 Allocnemis pachystigma SELYS, Mem. Cour. Acad. Belg., 38:138. Type male: lost; type-locality: Sierra Leone. 1890 Allocnemis pachystigma - KIRBY, Syn. Cat. Neur.-Odon., London: 131. 1899 Stenocnemis pachystigma - KARSCH, Ent. Nachr., 25 (11): 6 (as generotype of Stenocnemis KARSCH, 1899) ( C a m e r o o n ) .

1940 Allocnemis pachystigma - FRÄSER, Proc. R. ent. Soc. London, (B) 10: 35 (description of female) (Cameroon).

388 1951 Coeliccia pachystigma - SCHMIDT, Mitt, miinch. ent. Ges., 41: 205 (redescription). 1962 Stenocnemis pachystigma - PINHEY, Pubk. cult. Comp. Diam. Angola, 59: 114 (Cameroon, Nigeria, Gold Coast, Sierra Leone).

Distribution: West Africa. Genus Thaumatagrion

LIEFTINCK

1932 Thaumatagrion LIEFTINCK, Nova Guinea, 15 (5): 24. Type species: Thaumatagrion

funereum

LIEFTINCK, 1 9 3 2 ( o r i g i n a l d e s i g n a t i o n ) .

funereum

LIEFTINCK

1932 Thaumatagrion funereum LIEFTINCK, Nova Guinea, 15: (5): 26 (as generotype of Thaumatagrion LIEFTINCK, 1932). Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Torrenticnemis

LIEFTINCK

1949 Torrenticnemis LIEFTINCK, Nova Guinea (N.S.) 5: 104. Type species: Torrenticnemis LIEFTINCK, 1 9 4 9 ( o r i g i n a l

filicornis

filicornis

designation).

LIEFTINCK

1949 Torrenticnemis filicornis LIEFTINCK, Nova Guinea (N.S.) 5: 104 (as generotype of Torrenticnemis LIEFTINCK, 1949). Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Subfamily Platycnemidinae Lower distal corner of Dc not acute; anal vein reaches level of origin of Riii. Tibia of male dilated. 1905 Platycnemidinae JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 831 (with genera Psilocnemis, Platycnemis, Calocnemis, Metacnemis, Trichocnemis). - 1929 Platycnemidinae - FRÄSER, J. Bombay nat. Hist. Soc., 33: 835. Type-genus: Platycnemis BURMEISTER, 1839 (with g e n e r a Platycnemis,

Copera).

-

1 9 3 8 Platycnemidinae

- T I L L YARD & FRASER, A u s t r .

Zool.,

9: 165 (with genera Metacnemis, Mesocnemis, Calicnemis, Stenocnemis, Platycnemis, Copera, Coeliccia, Indocnemis, Idiocnemis, Paracnemis, Leptocnemis, Allocnemis, Tatocnemis, Risiocnemis).

Genus Copera

KIRBY

1863 Platycnemis (Psilocnemis) SELYS, Synopsis des Agrionines, Add. 4: 24. Type species: Platycnemis marginipes R A M B U R , 1 8 4 2 (original designation). - 1 8 9 0 Copera K I R B Y , Syn. Cat. Neur.Odon., London: 1 2 9 [new name for Psilocnemis SELYS, 1 8 6 3 , a junior primary homonymy of Psilocnemis BURMEISTER, 1 8 4 2 in Coleoptera, et Psilocnemis (n.n. pro Torticella G L O G E R , 1 8 4 2 ) G L O G E R , 1 8 4 2 in Aves], - 1 9 0 5 Psilocnemis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 831 (key to species). - 1917 Copera - LAIDLAW, Ree. Indian Mus., 13: 336. 1 9 3 5 Copera

- LIEFTINCK, M i s c . Z o o l . S u m . , 9 2 - 9 3 : 8 - 1 9 4 0 Copera

- LIEFTINCK, T r e u b i a ,

17: 281 (revision, key to species and subspecies). - 1954 Copera - LIEFTINCK, Treubia, 22 Suppl.: 50. - 1962 Psilocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 109 (proposed synonymy with Platycnemis CHARPENTIER, 1 8 4 0 ) . - 1 9 6 2 Copera - PINHEY, Publ. cult. Comp. D i a m . A n g o l a , 5 9 : 1 0 9 ( i n c o r r e c t s y n o n y m y w i t h Platycnemis

CHARPENTIER, 1 8 4 0 ) .

389

atomaria

(SELYS)

1886 Psilocnemis atomaria SELYS (nec L A I D L A W , 1902), Mem. Cour. Acad. Belg., 38: 122. Type male: Selys's Collection; type-locality: Labuan. 1890 Copera atomaria - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1 9 4 0 Copera atomaria - LIEFTINCK, Treubia, 1 7 : 2 8 4 [incorrect synonymy with vittata vittata (SELYS, 1863)].

1954 Copera atomaria-

LIEFTINCK, Treubia, 22 Suppl.: 52 [as syn. with vittata vittata /"SELYS, 1863)].

Distribution: Malaysia: Labuan. annulata annulata

(SELYS)

1863 Platycnemis (Psilocnemis) annulata SELYS, Synopsis des Agrionines, Add. 4: 28. Type male: Selys's Collection; type-locality: Shanghai. 1863 Platycnemis (Psilocnemis) ciliata SELYS, Synopsis des Agrionines, Add. 4: 29. Type female: Selys's Collecion; type-locality: Malacca. 1886 Psilocnemis subannulata SELYS, Mem. Cour. Acad. Belg., 38:125. Type male: Selys's Collection; type-locality: India. 1886 Psilocnemis annulata race ciliata - SELYS, Mem. Cour. Acad. Belg., 38: 125. 1890 Copera annulata - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1890 Copera subannulata - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1 9 0 5 Psilocnemis annulata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 2 . 1916 Copera annulata - Ris, Suppl. ent. Berlin, 5: 17 (Formosa, Japan, China). 1922 Pseudocopera arachnoides FRASER, Mem. Dept.-Agric. India, 7: 56. Type male: British Museum (Nat. Hist.), Lodon; type-locality: Assam (as generotype of Pseudocopera FRASER, 1922). 1927 Copera annulata - Ris, Zool. Meded., Leiden, 10 (1): 17 (Sumatra). 1931 Copera annulata - SCHMIDT, Konowia, 10 (3): 183 (China). 1934 Copera annulata - LIEFTINCK, Treubia, 14 (4): 393 (Java). 1966 Pseudocopera arachnoides - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 179 [proposed synonymy with annulata annulata (SELYS, 1 8 6 3 ) ] .

Distribution: East Asia.

annulata stevensi

LAIDLAW

1914 Copera annulata stevensi LAIDLAW, Ree. Indian Mus., 8: 341. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Assam.

Distribution: India. imbricata

(HÄGEN)

1863 Platycnemis (Psilocnemis) imbricata HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 27. Type male: Halle Museum; type-locality: Sumatra. 1890 Copera imbricata - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1898 Psilocnemis lobimargo KRÜGER, Stettiner ent. Ztg., 59: 103. Type male: Selys's Collection; type-locality: Sumatra. 1927 Copera lobimargo - Ris, Zool. Meded., Leiden, 10 (1): 17 (redescription). 1935 Copera imbricata - LIEFTINCK, Misc. Zool.Sum., 92-93: 8 (Sumatra). 1940 Psilocnemis lobimargo - LIEFTINCK, Treubia, 17: 284 [proposed synonymy with imbricata (HAGEN, 1863)]. 1954 Copera imbricata - LIEFTINCK, Treubia, 22 Suppl.: 59 (Sumatra).

Distribution: Sumatra.

390 marginipes (RAMBUR) Platycnemis marginipes RAMBUR, Hist. nat. Ins. Neuropt., Paris: 2 4 0 . Type: sex not given: Serville's Collection; type-locality: Java. 1863 Platycnemis (Psilocnemis) marginipes - SELYS, Synopsis des Agrionines, Add. 4 : 24. 1863 Platycnemis (Platycnemis) lacteola SELYS, Synopsis des Agrionines, Add. 4 : 23. Type: sex not given: Rijksmus. Natuurl. Hist., Leiden; type-locality: Japan. 1863 Platycnemis (Psilocnemis) striatipes SELYS, Synopsis des Agrionines, Add. 4 : 25. Type female: Latreille's Collection; type-locality: Java. 1 8 9 0 Copera marginipes - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 2 9 (as generotype of Copera 1842

KIRBY, 1890). 1890 1890 1898 1905

1916 1917 1924 1927 1934 1934

1936 1942 1954 1955 1964

1965 1987

Platycnemis (Platycnemis) lacteola - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 2 9 [proposed synonymy with marginipes ( R A M B U R , 1 8 4 2 ) ] . Platycnemis (Psilocnemis) striatipes - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 2 9 [proposed synonymy with marginipes ( R A M B U R , 1 8 4 2 ) ] . Psilocnemis marginipes - K R Ü G E R , Stettiner ent. Ztg, 1 8 9 8 : 1 0 2 . Psilocnemis marginipes - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 832. Copera marginipes - Ris, Suppl. ent. Berlin, 5: 18 (Formosa, India, China). Copera marginipes - L A I D L A W , Ree. Indian Mus., 1 3 : 337 (Conchin State). Copera marginipes - FRÄSER, Ree. Indian Mus., 2 6 : 498 (India). Copera marginipes - Ris, Zool. Meded., Leiden, 10 (1): 17 (Sumatra). Copera marginipes - SCHMIDT, Arch. Hydrob. Suppl., 13: 335 (Java). Copera marginipes - LIEFTINCK, Treubia, 1 4 ( 4 ) : 3 9 3 (Java). Copera marginipes - LIEFTINCK, Revue suisse Zool., 4 3 (5): 212 (Flores). Copera marginipes - FRÄSER, Proc. R. ent. Soc. London, (B) 11: 97. Copera marginipes-LIEFTINCK, Treubia, 22 Suppl.: 51 (Siam, Malaya, Sumatra, Java, Borneo). Copera marginipes - LIEFTINCK, Zool. Meded., Leiden, 3 4 (5): 73 (Ceylon). Copera marginipes - SCHMIDT, Ent. Tidschr., 8 5 : 1 4 9 (Burma). Disparoneura bhatnagri SAHNI, Indian J. Ent., 2 7 : 209. Type male: Sahni's Collection; typelocality: India. Disparoneura bhatnargi- HÄMÄLÄINEN, Odonatologica, 1 8 ( 1 ) : 1 7 [proposed synonymy with marginipes ( R A M B U R , 1 8 4 2 ) ] .

Distribution: Oriental region. rubripes (NAVÄS) 1934

1984

Platycnemis rubripes NAVÄS, Mus. Heude Notes ent. chinoise, 2 ( 1 ) : 5. Type female: not given; type-locality: China. Copera rubripes - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 0 1 .

Distribution: China. superplatypes

FRASER

1927 Copera superplatypes FRÄSER, Ree. Indian Mus., 29: 88. Type male: British Museum (Nat. Hist.), London; type-locality: Hasimara. 1966 Platycnemis superplatypes - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 215.

Distribution: India. tokyoensis

ASAHINA

1948 Copera tokyoensis ASAHINA, Mushi, 18: 103. Type male: Asahina's Collection; type-locality: Japan. Distribution: J a p a n .

391 trotteri

(FRÄSER)

1922 Pseudocopera trotteri FRÄSER, J. nat. Hist. Soc. Siam, 4: 237. Type male: originally in Fraser's Collection, now unknown; type-locality: Bangkok. 1966 Pseudocopera trotteri - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 216 [probably syn. of annulata

annulata

(SELYS, 1863)].

1984 Copera trotteri - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 102 (Siam).

Distribution: Thailand. vittata vittata

(SELYS)

1863 Platycnemis (Psilocnemis) vittata SELYS, Synopsis des Agrionines, Add. 4: 26. Type male: Selys's Collection; type-locality: Malacca. 1890 Copera vittata - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1914 Copera annulata vittata - LAIDLAW, Ree. Indian Mus., 8: 342 (Assam). 1917 Copera vittata vittata - LAIDLAW, Ree. Indian Mus., 13: 338. 1940 Copera vittata vittata - LIEFTINCK, Treubia, 17: 295. 1954 Copera vittata vittata - LIEFTINCK, Treubia, 22 Suppl.: 52 (Siam, Malaya, Sumatra, Borneo).

Distribution: Thailand, Malaysia, Sumatra, Borneo, India. vittata acutimargo

KRÜGER

1898 Copera acutimargo KRÜGER, Stettiner ent. Ztg, 59: 103. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1915 Copera acutimargo - Ris, Tijdschr. Ent., Leiden 58: 7. 1934 Copera acutimargo - SCHMIDT, Arch. Hydrob. Suppl., 13: 336 (Sumatra). 1935 Copera acutimargo - LIEFTINCK, Misc. Zool. Sum., 92-93: 8 (Sumatra). 1940 Copera vittata acutimargo - LIEFTINCK, Treubia, 17: 290. 1948 Copera vittata acutimargo - LIEFTINCK, Treubia, 19: 284. 1954 Copera vittata acutimargo - LIEFTINCK, Treubia, 22 Suppl.: 52. 1984 Copera atomaria LAIDLAW, 1902 (nec SELYS, 1886) - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s

of the World, Utrecht, 1984, 1: 101.

Distribution: Sumatra. vittata assamensis

LAIDLAW

1914 Copera (Psilocnemis) vittata assamensis LAIDLAW, Ree. Indian Mus., 8:342. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Assam.

Distribution: India. vittata deccanensis

LAIDLAW

1917 Copera vittata deccanensis LAIDLAW, Ree. Indian Mus., 13: 338. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Parambikulum. 1924 Copera vittata deccanensis - FRÄSER, Ree. Indian Mus., 26: 498.

Distribution: India. vittata javana

LIEFTINCK

1940 Copera vittata javana LIEFTINCK, Treubia, 17: 298. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Java. 1954 Copera vittata javana - LIEFTINCK, Treubia, 22 Suppl.: 52 (Java).

Distribution: Java.

392 vittata palawana

LIEFTINCK

1940 Copera vittata palawana LIEFTINCK, Treubia, 17: 303. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Palawan.

Distribution: Palawan, vittata serapica

(HÄGEN)

1863 Platycnemis (Psilocnemis) serapica HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 27. Type male: Univ. zool. Mus.; Kobenhavn; type-locality: Nicobar. 1890 Copera serapica - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1 9 8 4 Copera vittata serapica - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 0 1 .

Distribution: Nicobar. Genus Platycnemis

CHARPENTIER

1840 Agrion (Platycnemis) CHARPENTIER, Libell. europ. Lipsiae: 21. Type species: Agrion lacteum CHARPENTIER, 1 8 2 5 (original designation). - 1 8 3 9 Agrion (Platycnemis) - BURMEISTER, H a n d . Ent., 2: 8 2 2 . - 1 8 5 0 Platycnemis - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 6 4 (revision). - 1857 Platycnemis - SELYS, in Sagra, Hist. fis. pol. nat. Cuba, 7 (2): 197. - 1863 Platycnemis (Platycnemis) - SELYS, Synopsis des Agrionines, Add. 4: 11 (with subgenera Trichocnemis, Psilocnemis, Allocnemis) (revision). - 1890 Platycnemis -KIRBY, Syn. Cat. Neur.-Odon., London: 1 2 9 (type: Libellula pennipes PALLAS, 1 7 7 1 ) . - 1 8 9 4 Platyscelus nom.nov. for Platycnemis CHARPENTIER, 1 8 4 0 (nec N O R D M A N N , 1 8 3 7 ) - WALLENGREN, Ent.Tidschr., 25: 2 6 2 . - 1 9 0 5 Platycnemis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 2 (type: Libellula pennipes PALLAS, 1 7 7 1 ) (key to species). - 1 9 2 0 Proplatycnemis K E N N E D Y , Ohio J . Sei. Columbus, 21: 8 5 . Type species: Platycnemis hova (SELYS) M A R T I N , 1 9 0 8 (original designation). - 1 9 2 2 Pseudocopera FRASER, Mem. Dept. Agric. India, 7: 56. Type species: Pseudocopera arachnoides FRASER, 1 9 2 2 (original designation). - 1 9 5 0 Platycnemis - M O R E R A , Inst. Esp. Ent., Madrid: 79 (key to Spanish species). - 1951 Proplatycnemis - SCHMIDT, Mitt, münch. ent. Ges., 41: 1 7 1 (proposed synonymy with Platycnemis CHARPENTIER, 1 8 4 0 ) . - 1 9 5 1 - Platycnemis SCHMIDT, Mitt, münch. ent. Ges., 41: 171 (key to Malagasy species). - 1954 Platycnemis LONGFIELD, Entomologist's month. Mag., 90: 145. - 1962 Platycnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59:109. - 1962 Platyscelus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 0 9 (proposed synonymy with Platycnemis CHARPENTIER, 1 8 4 0 ) .

acutipennis

SELYS

1841 Platycnemis acutipennis SELYS, Rev. Zool., 1841: 246. Type male: Selys's Collection; typelocality: France. 1842 Platycnemis diversa RAMBUR, Hist. nat. Ins. Neuropt., Paris: 243. Type: sex not given: unknown; type-locality: France. 1850 Platycnemis acutipennis - SELYS & H Ä G E N , Revue des Odonates, Paris: 164 (France). 1863 Platycnemis (Platycnemis) acutipennis - SELYS, Synopsis des Agrionines, Add. 4: 19 (redescription). 1885 Platycnemis algira KOLBE, Berl. ent. Z.,29:154. Type male: Mus. Naturk., Berlin; type-locality: Algeria. 1887 Platycnemis algira - SELYS, Ann. Soc. ent. Belg., 31: 64 (proposed synonymy with acutipennis SELYS, 1841).

1890 Platycnemis 1905 Platycnemis onymy with 1950 Platycnemis

acutipennis - KIRBY, Syn. Cat. Neur.-Odon., London: 130. diversa - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 833 (proposed synacutipennis SELYS, 1841). acutipennis - MORERA, Inst. Esp. Ent., Madrid: 80 (Spain).

Distribution: France, Spain, Algeria.

393 agrioides Ris 1915 Platycnemis agrioides Ris, Ent. Mitt., 4:138. Type: not designated, lectotype male: Ann Arbor Museum; type-locality: Mayotte. 1920 Proplatycnemis agrioides - KENNEDY, Ohio J. Sei., 21: 85. 1962 Platycnemis agrioides - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111 (Comoro Island).

Distribution: Comoro Island. alatipes

(MCLACHLAN)

1872 Psilocnemis alatipes MCLACHLAN, Entomologist's month. Mag., 9: 1. Type male: British Museum (Nat. Hist.), London; type-locality: Madagascar. 1886 Psilocnemis alatipes - SELYS, Mem. Cour. Acad. Belg., 38: 126. 1890 Copera alatipes - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1949 Platycnemis alatipes - FRASER, Mem. Inst. sei. Madagascar, 3 (9): 10. 1962 Platycnemis alatipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

Distribution: Madagascar. aurantipes

LIEFTINCK

1984 Platycnemis aurantipes LIEFTINCK, 1965 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 102.

Distribution: Madagascar. bilineata

BARTENEF

1910 Platycnemis bilineata BARTENEF, Verz. Tomsk, 11:9. Type male: unknown; type-locality: Sikok.

Distribution: Russia. congolensis

MARTIN

1908 Platycnemis congolensis MARTIN, Ann. Mus. civ. Stor. nat. Genova, 43: 664. Type male: Mus. civ. Stor. nat., Genova; type-locality: Port. Guinea. 1951 Platycnemis congolensis - SCHMIDT, Mitt, münch. ent. Ges., 41: 223 (Belgian Congo). 1962 Platycnemis congolensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110 (Uganda, Belgian Congo, French Congo, French Guinea, Ivory Coast).

Distribution: West Africa. dealbata

KLUG

1863 Platycnemis (Platycnemis) latipes race dealbata KLUG, in SELYS, Synopsis des Agrionines, Add. 4: 23. Male from Egypt (and Syria). 1 8 5 0 Platycnemis syriaca H A G E N & SELYS, Revue des Odonates, Paris: 3 8 9 . Type: unknown; typelocality: Syria. 1887 Platycnemis latipes race dealbata - SELYS, Ann. Soc. ent. Belg., 31: 46 (Syria, Egypt, Transcaucasica). 1890 Platycnemis dealbata - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1905 Platycnemis dealbata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 834 (Egypt, Syria). 1 9 0 5 Platycnemis syriaca - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 4 (proposed synonymy with dealbata K L U G , 1 8 6 3 ) . 1938 Platycnemis latipes dealbata - SCHMIDT, SitzBer.Akad. wiss. Wien (1) 147: 141 (Lebanon, Israel). 1984 Platycnemis syriaca - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 103 (as good species). 1 9 8 9 Platycnemis dealbata - D U M O N T , DEMIRSOG & MERTENS, Notul. odonatol., 3 ( 2 ) : 2 4 (Turkey).

Distribution: Transcaucasica, Turkey, Lebanon, Syria, Jordan, Israel, Egypt.

394 echigoana ASAHINA 1955 Platycnemis echigoana ASAHINA, Akitu, 4: 101. Type male: Asahina's Collection; type-locality: Japan.

Distribution: Japan. escherichi SCHMIDT 1951 Platycnemis escherichi SCHMIDT, Mitt, münch. ent. Ges., 41: 224. Type female: Mus. Naturk., Berlin; type-locality: Cameroon. 1962 Platycnemis escherichi - PINHEY, Pub. cult. Comp. Diam. Angola, 59: 110.

Distribution: Cameroon. flavipes NAVÄS 1924 Platycnemis flavipes NAVÄS, Mem. R. Acad. Barcelona, 18 (13): 12. Type: sex not given: Barcelona Museum; type-locality: Kenya. 1962 Platycnemis flavipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110.

Distribution: Kenya. foliacea foliacea SELYS 1886 Platycnemis foliacea SELYS, C.R. Soc. Ent. Belg., 30: 138. Type male: Selys's Collection; typelocality: Peking. 1890 Platycnemis foliacea - KIRBY, Syn. Cat. Neur.-Odon., London: 130. 1905 Platycnemis foliacea - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 834 (Pekin). 1916 Platycnemis foliacea - Ris, Suppl. ent., 5: 18 (Japan). 1930 Platycnemis foliacea - Ris, Ark. Zool., Uppsala, 21A (31): 23 (Japan). 1949 Platycnemis foliacea foliacea - ASAHINA, Mushi, 20: 32 (redescription). 1955 Platycnemis foliacea foliacea - ASAHINA, Ent. Meded., 27 (3): 133 (North China).

Distribution: North China, Japan. foliacea sasakii ASAHINA 1949 Platycnemis foliacea saskii ASAHINA, Insectamatsum., 17: 33. Type male: Asahina's Collection; type-locality: Japan.

Distribution: Japan, f o l i o s a NAVÄS 1932 Platycnemis foliosa NAVÄS, Mus. Heude Notes ent. chinoise, 8: 9. Type male: unknown; typelocality: China.

Distribution: China. guttifera FRASER 1950 Platycnemis guttifera FRASER, Bull. Inst, franc. Afr. noire, 12: 615. Type male: Nation. Mus. hist, nat., Paris; type-locality: Liberia. 1962 Platycnemis guttifera - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110.

Distribution: Liberia. hova MARTIN 1908 Platycnemis hova (SELYS MS) MARTIN, Ann. Mus. civ. Stor. Genova, 43: 665. Type male: Mus. civ. Stor. nat., Genova; type-locality: Madagascar. 1920 Proplatycnemis hova - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Proplatycnemis

KENNEDY, 1920).

395 1962 Platycnemis hova - PINHEY, Publ. cult.Comp. Diam. Angola, 59: 111.

Distribution: Madagascar. kervillei

(MARTIN)

1909 Psilocnemis kervillei MARTIN, Bull. Soc. ent. France, 1909: 214. Type male: Martin's Collection; type-locality: Syria. 1924 Copera kervillei - MORTON, Trans, ent. Soc. London, 1924: 30 (redescription) (Palestine). 1984 Platycnemis kervillei - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 102. 1989 Platycnemis

kervillei

- DUMONT, DEMIRSOG & MERTENS, N o t u l . o d o n a t o l . , 3 (2): 24 ( T u r k e y ) .

Distribution: Near East. latipes

RAMBUR

1842 Platycnemis latipes RAMBUR, Hist. nat. Ins. Neuropt. Paris: 242. Type sex and locality not given (? Montpellier). 1850 Platycnemis latipes - SELYS & HÄGEN, Revue des Odonates, Paris: 166 (France). 1863 Platycnemis (Platycnemis) latipes - SELYS, Synopsis des Agrionines, Add. 4: 22 (Europe merid., and Syria). 1890 Platycnemis latipes - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1905 Platycnemis latipes - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 833 (Greece, France). 1923 Platycnemis latipes var. brachygastra NAVÄS, Arx. Inst. Cienc., 8 (1920): 9. Male from Catalonia. 1928 Platycnemis latipes - ANDERS, Mem. Soc. R. ent. Egypt, 3 (1): 21 (Egypt). 1950 Platycnemis latipes - MOREIRA, Inst. Esp. Ent. Madrid: 80 (Spain).

Distribution: Syria, Greece, France, Spain, Egypt, Lybia, Morocco. longiventris

SCHMIDT

1949 Platycnemis longiventris SCHMIDT, Mem. Inst. sei. Madagascar, 3A: 23. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Platycnemis longiventris - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

Distribution: Madagascar. malgassica

SCHMIDT

1951 Platycnemis malgassica SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 180. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Platycnemis malgassica - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

Distribution: Madagascar. mauriciana

SELYS

1863 Platycnemis (Platycnemis) latipes race mauriciana SELYS, Synopsis des Agrionines, Add. 4: 23. Type male: R. Mus. Brussels; type-locality: Mauritius. 1890 Platycnemis mauriciana - KIRBY, Syn. Cat. Neur.-Odon., London: 129. 1949 Platycnemis mauriciana - FRASER, Trans. R. ent. Soc. London, 100: 145 (Mauritius). 1962 Platycnemis mauriciana - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

Distribution: Mauritius. nyansana

FÖRSTER

1916 Platycnemis nyansana FÖRSTER, Neue Beitr. syst. Insektenk., 1: 25. Type: not located in Ann Arbor; type-locality: Uganda. 1962 Platycnemis nyasana - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110.

Distribution: Uganda.

396 oedipus (EICHWALD) 1829 Agrion oedipus EICHWALD, Zool. Soec., Moscow, 1: 223. Type male: unknown; type-locality: Transcaucasica. 1890 Platycnemis oedipus - KIRBY, Syn. Cat. Neur.-Odon., London: 130. 1948 Platycnemis oedipus - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 150 (Armenia).

Distribution: Armenia.

pennipes pennipes (PALLAS) 1771 Libellula pennipes PALLAS, Eur. Ins. Reise, 1: 469. Type: unknown. 1789 Libellula albidella VILLIERS, Linn. Ent.: 11. Type male: Mus. Nation. Hist, nat., Paris; typelocality: France. 1823 Agrion platypoda VAN DER LINDEN, Opusc. Sei., 4: 102. Type male: Mus. Naturk., Berlin; type-locality: unknown. 1825 Agrion lacteum CHARPENTIER, Hor. Ent.: 11. Type male: Mus. Nation. Hist, nat., Paris; typelocality: Europe. 1840 Agrion (Platycnemis) lacteum - CHARPENTIER, Libell. europ. Lipsiae: 21 (as generotype of Platycnemis CHARPENTIER, 1840). 1 8 5 0 Platycnemis pennipes - SELYS & H Ä G E N , Revue des Odonates, Paris: 1 6 8 . 1863 Platycnemis pecchioli SELYS, Bull. Acad. r. Belg., (2) 16: 166. Type female: R. Mus., Brussels; type-locality: unknown. 1863 Platycnemis (Platycnemis) pennipes - SELYS, Synopsis des Agrionines, Add. 4: 21 (redescription). 1890 Platycnemis pennipes - KIRBY, Syn. Cat. Neur.-Odon., London: 129 (Europe, West Asia). 1890 Libellula albidella - KIRBY, Syn. Cat. Neur.-Odon., London: 129 [proposed synonymy with pennipes

pennipes

(PALLAS, 1771)].

1890 Agrion platypoda - KIRBY, Syn. Cat. Neur.-Odon., London: 129 [proposed synonymy with pennipes

pennipes

(PALLAS, 1771)].

1890 Agrion lacteum - KIRBY, Syn. Cat. Neur.-Odon., London: 129 [proposed synonymy with pennipes

pennipes

(PALLAS, 1771)].

1890 Plytacnemis pecchioli - KIRBY, Syn. Cat. Neur.-Odon., London: 129 [proposed synonymy with pennipes pennipes (PALLAS, 1 7 7 1 ) ] . 1 9 0 5 Platycnemis pennipes - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 2 . 1948 Platycnemis pennipes - AKRAMOVSKI, Zool. Zborn., Moscow, 5: 150 (Armenia). 1948 Platycnemis pennipes ab. albioctopunctata GÖRTLER, Ent. Listy, Brno, 11: 39. Male from Czechoslovakia. 1948 Platycnemis pennipes ab. brunneioctopunctata GÖRTLER, Ent. Listy, Brno, 11: 39. Male from Czechoslovakia. 1956 Platycnemis pennipes - FRÄSER, Proc. R. Ent. Soc. London, 1 (10): 32 (England). 1960 Platycnemis pennipes - HOFFMANN, Archives, 27: 223 (Holland, Belgium, France, Germany, Luxembourg).

Distribution: Europe, West Asia, pennipes nitidula (BRULLE) 1832 Puella nitidula BRÜLLE, Expl. Moree, 2 (1): 105. Type: not given. 1832 Puella hyalinata BRÜLLE, Expl. Moree, 2 (1): 106. Type: unknown; type-locality: Greece. 1863 Platycnemis (Platycnemis) pennipes race insularis HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 22. Type male: Naturhist. Mus. Wien, type-locality: Corfu. 1890 Platycnemis hyalinata - KIRBY, Syn. Cat. Neur.-Odon., London: 130 (Greece). 1890 Platycnemis insularis - KIRBY, Syn. Cat. Neur.-Odon., London: 130 (Corfu). 1 9 0 5 Platycnemis insularis - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 3 2 (Corfu, Algeria, Smyrna).

397 1905

Puella hyalinata - JACOBSON & with insularis H Ä G E N , 1 8 6 3 ) .

BIANKI,

Pryam. lozhnos. Ross, imp.:

832

(proposed synonymy

Distribution: Greece, Turkey, Algeria. phyllopoda

DJAKONOV

1926 Platycnemis phyllopoda DJAKONOV, Rev. russ. Ent., 20:231. Type male: not given; type-locality: Ussuri. 1930 Platycnemis phyllopoda- Ris, Ark. Zool. Uppsala, 21A (31): 29 (description of male) (China). 1930 Platycnemis ulmifolia Ris, Ark. Zool. Uppsala, 21A (31): 26. Type male: Naturhist. Riksmus., Stockholm; type-locality: China. 1933 Platycnemis hummeli SJÖSTEDT, Ark. Zool. Uppsala, 25A (5): 19. Type male: Naturhist. Riksmus., Stockholm; type-locality: Szechwan.

Distribution: Siberia, China. pierrati

NAVAS

1935 Platycnemis pierrati N A V Ä S , type-locality: China.

MUS.

Heude Notes ent. chinoise, 2 (5): 91. Type male: not given;

Distribution: China. protostictoides

FRASER

1953 Platycnemis protostictoides FRASER, Nat. malgache, 5 (2): 165. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Madagascar. 1962 Platycnemis protostictoides - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

Distribution: Madagascar. pseudalatipes

SCHMIDT

1951 Platycnemis pseudalatipes SCHMIDT, Mem. Inst. sei. Madagascar, 6A: 186. Type male: Naturhist. Mus. Wien; type-locality: Madagascar. 1962 Platycnemis pseudalatipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59, 111. 1 9 8 4 Platycnemis pseudalatipes pallida AGUESSE, 1 9 6 8 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 103.

Distribution: Madagascar. rufipes

(SELYS)

1886 Metacnemis rufipes SELYS, Mem. Cour. Acad. r. Belg., 38: 139. Type male: British Museum (Nat. Hist.), London; type-locality: Cameroon. 1890 Allocnemis rufipes - KIRBY, Syn. Cat. Neur.-Odon., London: 131. 1962 Platycnemis rufipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110 (Cameroon, Nigeria).

Distribution: Central Africa. sanguinipes

SCHMIDT

1951 Platycnemis sanguinipes SCHMIDT, Mem. Inst. sei. Madagascar, 6Α: 190. Type male: Schmidt's Collection; type-locality: Madagascar. 1962 Platycnemis sanguinipes - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111.

Distribution: Madagascar. sikassoensis

(MARTIN)

1912 Psilocnemis sikassoensis MARTIN, Feuille Jeun. Nat., 42: 98. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: French Guinea.

398 1928 Copera subaequistyla FRÄSER, Trans, ent. Soc. London, 76: 127. Type male: British Museum (Nat. Hist.), London; type-locality: Lake Victoria. 1962 Platycnemis sikassoensis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110. 1962 Platycnemis subaequistyla - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 110.

Distribution: Central Africa. subdilatata

SELYS

1849 Platycnemis subdilatata SELYS, LUC. Expl. Alg., 3: 132. Type male: Mus. Nation. Hist. Nat.; Paris; type-locality: Algeria. 1 9 0 5 Platycnemis subdilatata - JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 8 2 3 .

Distribution: Algeria. xanthopus

NAVÄS

1924 Platycnemis xanthopus NAVÄS, Mem. R. Acad. Barcelona, 18 (3): 13. Type male: Naväs's Collection; type-locality: East Africa. 1962 Platycnemis xanthopus - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 111 (Kenya).

Distribution: Kenya. Family Platystictidae Small to medium sized Zygoptera. Anal vein absent or reduced; cross vein joins Cup with border of wing; Dc obtuse apically, sub-Dc, contiguous with posterior border of wing. Pterostigma wide with extended inner angle. 1938 Platystictidae T I L L Y A R D & FRASER, Austr. Zool., 9: 161 (with subfamilies Platystictinae, Palaemnematinae). - 1957 Platystictidae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 41 (with subfamily Platystictinae).

Subfamily Palaemnematinae IRii & Riii arising midway between subnodus and stigma; PuP extending over halfway along posterior border of wing. Pterostigma subtending one or slightly over one cell. 1938 Palaemnematinae T I L L Y A R D & FRASER, Austr. Zool., 9: 1962 (for genus Palaemnema). - 1957 Palaemnematinae - FRASER, R. Zool. Soc. N.S. Wales, 1957: 42. Type-genus: Palaemnema SELYS, 1860).

Genus Palaemnema

SELYS

I860 Platysticta (Palaemnema) SELYS, Bull. Acad. r. Belg., (2) 10:434. Type species: Libellulapaulina DRURY, 1773 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon.: 132). - 1903 Palaemnema - CALVERT, Biol. Centr.-Amer. Neur., London: 135 (key to Central American species). - 1916 Palaemnema- Ris, Arch. Naturg., (A) 82: 91 (key to species). - 1931 Palaemnema - CALVERT, Trans. Am. ent. Soc., 57: 1 (revision, key to species).

abbreviata

KENNEDY

1938 Palaemnema abbreviata KENNEDY, Ann. ent. Soc. Amer., 31: 256. Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador.

399 angelina

SELYS

I860 Platysticta (Palaemnema) angelina SELYS, Bull. Acad. r. Belg., (2) 10: 435. Type male: Mus. Nation.Hist, nat., Paris; type-locality: Guatemala. 1890 Palaemnema angelina - KIRBY, Syn. Cat. Neur.-Odon., London: 132. 1915 Palaemnema angelina - WILLIAMSON, Proc. U.S. natn. Mus., 48: 608. 1916 Palaemnema angelina - Ris, Arch. Naturg., (A) 82: 95 (Costa Rica). 1931 Palaemnema angelina - CALVERT, Trans. Am. ent.Soc., 57: 58 (redescription) (Guatemale, Honduras).

Distribution: Central America, apicalis

NAVÄS

1924 Palaemnema apicalis NAVÄS, Mem. R. Acad. Barcelona, 18 (13): 14. Type male: Naväs's, Collection; type-locality: Colombia. 1923 Palaemnema apicalis - NAVÄS, Bol. Soc. Colombiana, Cien. Nat., 12: 361. 1931 Palaemnema apicalis - CALVERT, Trans. Am. ent. Soc., 57: 81 (redescription).

Distribution: Colombia. azupizui

CALVERT

1931 Palaemnema azupizui CALVERT, Trans. Am. ent. Soc., 57: 78. Type male: Cornell Univ., Ithaca; type-locality: Peru. 1943 Palaemnema azupizui - SCHMIDT, Beitr. Fauna Perus, 2: 231.

Distribution: Peru. brucei

CALVERT

1931 Palaemnema brucei CALVERT, Trans. Am. ent. Soc., 57: 75. Type male: Williamson's Collection; type-locality: Panama.

Distribution: Panama. brucelli

KENNEDY

1938 Palaemnema brucelli KENNEDY, Ann. ent. Soc. Amer., 31: 260. Type male: unknown; typelocality: Ecuador.

Distribution: Ecuador, carmelita Ris 1916 Palaemnema carmelita Ris, Arch. Naturg., (A) 82: 100. Type male: not given; type-locality: Colombia. 1931 Palaemnema carmelita - CALVERT, Trans. Am. ent. Soc., 57: 83 (redescription).

Distribution: Colombia. chiriquita

CALVERT

1931 Palaemnema chiriquita CALVERT, Trans.Am. ent.Soc., 57: 100. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Costa Rica.

Distribution: Costa Rica, dementia

SELYS

1886 Palaemnema dementia SELYS, Mem. Cour. Acad. Belg., 38:148. Type male: Selys's Collection; type-locality: Venezuela. 1909 Palaemnema dementia - CALVERT, Ann. Carnegie Mus., Pittsburgh, 6: 211 (Colombia).

400 1931 Palaemnema 1943 Palaemnema

dementia - CALVERT, Trans. Am. ent. Soc., 57: 93 (redescription). clementia - SCHMIDT, Beitr. F a u n a Perus, 2: 231 (Peru).

Distribution: Colombia, Venezuela, Peru. croceicauda

CALVERT

1931 Palaemnema croceicauda CALVERT, Trans. Am. ent. So.c, 57: 96. Type male: Williamson's Collection; type-locality: Colombia.

Distribution: Colombia, desiderata

SELYS

1886 Palaemnema desiderata SELYS, Mem. Cour. Acad. Belg., 38:146. Type male: Selys's Collection; type-locality: Mexico. 1903 Palaemnema desiderata - CALVERT, Biol. Centr.-Amer. Neur., London: 134. 1931 Palaemnema desiderata - CALVERT, Trans. Am. ent. Soc. 57: 50 (redescription).

Distribution: Mexico. distadens

CALVERT

1931 Palaemnema distadens CALVERT, Trans. Am. ent. Soc., 57: 35. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Costa Rica.

Distribution: Costa Rica. domina

CALVERT

1903 Palaemnema domina CALVERT, Biol. Centr.-Amer. Neur., London: 137. Type male: Mus. Comp. Zool. Cambridge (Mass.); type-locality: Mexico. 1931 Palaemnema domina ~ CALVERT, Trans. Am. ent. Soc., 57: 61 (redescription).

Distribution: Mexico. edmondi

CALVERT

1931 Palaemnema edmondi CALVERT, Trans. Am. ent. Soc., 57: 99. Type male: Williamson's Collection; type-locality: Colombia.

Distribution: Colombia, gigantula

CALVERT

1931 Palaemnema gigantula CALVERT, Trans. Am. ent. Soc., 57: 72. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Costa Rica.

Distribution: Costa Rica. joanetta

KENNEDY

1940 Palaemnema joanetta KENNEDY, Ann. ent. Soc.Amer., 33: 626. Type male: unknown; typelocality: Panama.

Distribution: Panama. lorena

KENNEDY

1942 Palaemnema lorena KENNEDY, Ann. ent. Soc. Amer., 35: 97. Type male: unknown; type-locality: Ecuador. Distribution: E c u a d o r .

maruni ^uwLti 1934 Palaemnema martini COWLEY, Stylops, 3: 203. Type male: Cowley's Collection; type-locality: Peru. 1943 Palaemnema martini - SCHMIDT, Beitr. Fauna Perus, 2: 231.

Distribution: Peru, melanocauda KENNEDY 1942 Palaemnema melanocauda KENNEDY, Ann. ent. Soc. Amer., 35: 100. Type male: unknown; type-locality: Ecuador.

Distribution: Ecuador, melanostigma HÄGEN I860 Platysticta (Palaemnema) melanostigma HAGEN, in SELYS, Bull. Acad.r. Belg., (2) 10: 436. Type male: Hagen's Collection; type-locality: Puerto Cabello. 1890 Palaemnema melanostigma - KIRBY, Syn. Cat. Neur.-Odon., London: 132. 1931 Palaemnema melanostigma - CALVERT, Trans. Am. ent. Soc., 57: 68 (redescription).

Distribution: Venezuela. melanota Ris 1916 Palaemnema melanota Ris, Arch. Naturg., (A) 82: 99. Type male: not given; type-locality: Costa Rica. 1931 Palaemnema melanota - CALVERT, Trans. Am. ent. Soc., 57: 102 (redescription).

Distribution: Costa Rica. mutans CALVERT 1931 Palaemnema mutans CALVERT, Trans. Am. ent. Soc., 57: 63. Type male: Williamson's Collection; type-locality: Venezuela.

Distribution: Venezuela. nathalia

SELYS

1886 Palaemnema nathalia SELYS, Mem. Cour. Acad. Belg., 38: 147. Type male: R. Mus., Brussels; type-locality: Panama. 1915 Palaemnema nathalia - WILLIAMSON, Proc. U.S. natn. Mus., 48: 609. 1916 Palaemnema nathalia - Ris, Arch. Naturg., (A) 82: 96 (Costa Rica). 1931 Palaemnema nathaliä - CALVERT, Trans. Am. ent. Soc., 57: 86 (redescription) (Panama, Guatemala, Costa Rica).'

Distribution: Central America. paulicaxa

CALVERT

1931 Palaemnema paulicaxa CALVERT, Trans. Am. ent.Soc., 57: 54. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Mexico.

Distribution: Mexico. paulicoba

CALVERT

1931 Palaemnema paulicoba CALVERT, Trans. Am. ent. Soc., 57: 45. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Mexico. 1933 Palaemnema paulicoba - CALVERT, Trans. Am. ent. Soc., 59: 379 (redescription).

Distribution: Mexico.

402 paulina (DRURY) 1773 1837 1842 1860

Libellula paulina DRURY, 111. Ex. Ent., 2: 46. Type male: unknown; type-locality: Mexico. Lestes paulina - WESTWOOD, 111. Exot. Ent. (New Ed.), 2: 46. Euphaea paulina - RAMBUR, Hist. nat. Ins. Neuropt. Paris: 231. Platysticta (Palaemnema) paulina - SELYS, Bull. Acad. r. Belg., (2) 10:434 (Mexico, Honduras) (as generotype of Palaemnema SELYS, 1860). 1890 Palaemnema paulina - KIRBY, Syn. Cat. Neur.-Odon., London: 132. 1916 Palaemnema paulina - Ris, Arch. Naturg., (A) 82: 93 (Mexico, Guatemala). 1931 Palaemnema paulina - CALVERT, Trans. Am. ent. Soc., 57: 39 (redescription) (Honduras).

Distribution: Central America. paulirica CALVERT 1931 Palaemnema paulirica CALVERT, Trans. Am. ent. Soc., 57: 32. Type male: Acad. Nat. Sei. Philadelphia; type-locality: Costa Rica.

Distribution: Costa Rica, paulitaba CALVERT 1931 Palaemnema paulitaba CALVERT, Trans. Am. ent. Soc., 57: 56. Type male: Acad. Nat. Sei. Philadelphia; type-locality: Mexico.

Distribution: Mexico, p a u l i t o y a c a CALVERT 1931 Palaemnema paulitoyaca CALVERT, Trans. Am. ent. Soc., 57: 47. Type male: British Museum (Nat. Hist.), London; type-locality: Mexico. 1933 Palaemnema paulitoyaca - CALVERT, Trans. Am. ent. Soc., 59: 377 (redescription) (Mexico).

Distribution: Mexico. peruviana Ris 1916 Palaemnema peruviana Ris, Arch. Naturg., (A) 82:97. Type male: not given; type-locality: Peru. 1931 Palaemnema peruviana - CALVERT, Trans. Am. ent. Soc., 57: 66 (redescription) (Peru). 1943 Palaemnema peruviana - SCHMIDT, Beitr. Fauna Perus, 2: 231.

Distribution: Peru. p i c i c a u d a t a KENNEDY 1938 Palaemenma picicaudata KENNEDY, Ann. ent. Soc. Amer., 31: 253. Type male: Kennedy's Collection; type-locality: Ecuador.

Distribution: Ecuador. r e v e n t a z o n i CALVERT 1931 Palaemnema reventazori CALVERT, Trans. Am. ent. Soc., 57: 85. Type male: Acad. Nat. Sei. Philadelphia; type-locality: Costa Rica.

Distribution: Costa Rica. Subfamily Platystictinae IRii & Riii arising nearer subnodus than to stigma; CuP ending at or before halfway along posterior border of wing. Pterostigma subtending from one to five cells. 1924 Platystictinae LAIDLAW, Spolia Zeylanica, 12: 360. Type-genus: Platysticta

SELYS, 1860. -

403 1 9 3 8 Platystictinae - T I L L Y A R D & FRÄSER, Austr. Zool., 9: 1 6 2 (with genera Platysticta, Protosticta, Drepanosticta, Ceylonosticta). - 1957 Platysctictinae - FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 42 (with genera Platyscticta, Protosticta, Drepanosticta, Ceylonosticta).

Genus Drepanosticta LAIDLAW 1917 Drepanosticta L A I D L A W , Ree. Indian Mus., 13:341. Type species: Protosticta carmichaeli L A I D LAW, 1915 (original designation). - 1931 Ceylonosticta FRÄSER, J . Bombay nat. Hist. Soc., 35: 329. Type species: Platysticta (Platysticta) hilaris SELYS, 1860 (original designation). 1932 Drepanosticta - LIEFTINCK, Nova Guinea, 15 (5): 30 (revision, key to New Guinean species). - 1933 Ceylonosticta - FRÄSER, Fauna brit. India, Odon., 1: 207 (key to Indian species). - 1938 Drepanosticta - LIEFTINCK, New Guinea (N.S.) 2: 86 (key to New Guinean species). - 1939 Drepanosticta - N E E D H A M & G Y G E R , Philipp. J. Sei., 70 (3): 263 (key to Philippiean species). - 1954 Drepanosticta - LIEFTINCK, Treubia, 22 Suppl.: 32. - 1955 Ceylonosticta - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 72 (proposed synonymy with Drepanosticta

LAIDLAW, 1917). - 1984 Drepanosticta

- DA VIES & TOBIN, T h e D r a g o n f l i e s of the

World, Utrecht, 1: 104.

actaeon LAIDLAW 1934 Drepanosticta actaeon LAIDLAW, J. Fed. Mai. States, Mus., 17: 558. Laidlaw's Collection; type-locality: Borneo. 1954 Drepanosticta acteon - LIEFTINCK, Treubia, 22 Suppl.: 32 (Borneo).

Distribution: Borneo. adami (FRÄSER) comb. nov. 1933 Ceylonosticta adami FRASER, Spolia Zeylanica, 17: 211. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon.

Distribution: Sri Lanka. anascephala FRASER 1933 Drepanosticta anascephala FRASER, J. Siam Soc. nat. Hist., Suppl., 9: 114. Type male: British Museum (Nat. Hist.), London; type-locality: Laos.

Distribution: Laos. annandalei FRASER 1924 Drepanosticta annandalei FRASER, Ree. Indian Mus., 26: 412. Type male: British Museum (Nat. Hist.), London: type-locality: Andamans.

Distribution: Andaman Islands. annulata (SELYS) 1886 Platysticta annulata SELYS, Mem. Cour. Acad. Belg., 38: 156. Type female: Selys's Collection; type-locality: Luzon. 1939 Protosticta annulata - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 264. 1984 Drepanisticta annulata - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 105.

Distribution: Philippine Islands. arcuata LIEFTINCK 1927 Drepanosticta kruegeri Ris (nec LAIDLAW, 1926), Zool. Meded., Leiden, 10: 19. 1934 Drepanosticta arcuata LIEFTINCK, Treubia, 14 (4): 469. Type male: Buitenzorg Mus.; typelocality: Sumatra.

404 1954 Drepanosticta arcuata-

LIEFTINCK, Treubia, 22 Suppl.: 32.

Distribution: Sumatra. aries NEEDHAM & GYGER 1941 Drepanosticta aries NEEDHAM locality: Philippines.

& GYGER,

Philipp. J. Sei., 74: 144. Type male: not given; type-

Distribution: Mindanao. attala LIEFTINCK 1934 Drepanosticta attala LIEFTINCK, Treubia, 14 (4): 472. Type male: Buitenzorg Mus.; type-locality: Borneo. 1954 Drepanosticta attala - LIEFTINCK, Treubia, 22 Suppl.: 32.

Distribution: Borneo. auriculata (SELYS) Platysticta auriculata SELYS (nec Ris, 1 9 2 9 ) , Mitt. Zool. Mus. Dresden, 3: 3 2 2 . Type male: R. Mus. Brussels; type-locality: New Guinea. 1938 Drepanosticta auriculata - LIEFTINCK, Nova Guinea (N.S.) 2: 81 (redescription of holotype). 1878

Distribution: New Guinea, austeni LIEFTINCK 1940 Drepanosticta austeni LIEFTINCK, Spolia zeylan., 22: 89. Type male: Lieftinck's Collection; type-locality: Ceylon.

Distribution: Sri Lanka, barbatula LIEFTINCK 1940 Drepanosticta barbatula LIEFTINCK, Treubia, 17: 351. Type male: Buitenzorg Museum; typelocality: Borneo. 1954 Drepanosticta barbatula - LIEFTINCK, Treubia, 22 Suppl.: 33 (Borneo).

Distribution: Borneo. bartelsi LIEFTINCK 1937 Drepanosticta bartelsi LIEFTINCK, Treubia, 16 (1): 70. Type male: Buitenzorg Museum; typelocality: Java. 1954 Drepanosticta bartelsi - LIEFTINCK, Treubia, 22 Suppl.: 33 (Java).

Distribution: Java. berlandi LIEFTINCK 1939 Drepanosticta berlandi LIEFTINCK, Rev. franc. Ent., 6: 145. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Lombok. 1953 Drepanosticta berlandi - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 149.

Distribution: Lombok. bicornuta (SELYS) Platysticta bicornuta SELYS (nec Ris, 1 9 1 5 ) , Mitt. Zool. Mus. Dresden, 3: 3 2 2 . Type male: R. Mus., Brussels; type-locality: New Guinea. 1937 Drepanosticta bicornuta - LIEFTINCK, Nova Guinea (N.S.) 2: 84 (redescription). 1878

Distribution: New Guinea.

405 bispina

FRÄSER

1932 Drepanosticta bispina FRÄSER, Mem. Mus. Hist.nat. Belg., 4 (3): 5. Type female: Fraser's Collection; type-locality: Sumatra. 1954 Drepanosticta bispina - LIEFTINCK, Treubia, 22 Suppl.: 33 (Sumatra).

Distribution: Sumatra. brinki

LIEFTINCK

1970 Drepanosticta brinki LIEFTINCK, Entom. scand. (Suppl.) 1: 190. Type male: Lieftinck's Collection; type-locality: Sri Lanka.

Distribution: Sri Lanka, brownelli

(TINKHAM)

1938 Ceylonosticta brownelli TINKHAM, Lingnan. Sei., J., 17:17. Type: unknown; type-locality: South China. 1 9 8 4 Drepanosticta brownelli - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 0 5 .

Distribution: China. carmichaeli

(LAIDLAW)

1915 Protosticta carmichaeli LAIDLAW, Ree. Indian Mus., 11: 390. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: India. 1917 Drepanosticta carmichaeli - LAIDLAW, Ree. Indian Mus., 13: 341 (as generotype of Drepanosticta LAIDLAW, 1917).

Distribution: India. ceratophora

LIEFTINCK

1974 Drepanosticta ceratophora LIEFTINCK, Steenstrupia, 3 (12): 117. Type male: unknown; typelocality: Palawan.

Distribution: Pacific Ocean Islands. claaseni

LIEFTINCK

1938 Drepanosticta claaseni LIEFTINCK, Nova Guinea (N.S.) 2: 80. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, clavata

LIEFTINCK

1932 Drepanosticta clavata LIEFTINCK, Nova Guinea, 15 (5): 31. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, conica

(MARTIN)

1909 Platysticta conica MARTIN, Bull. Soc. ent. Ital., 60: 204. Type female: Mus. civ. Stor. Nat. Genova; type-locality: New Guinea. 1938 Drepanosticta conica - LIEFTINCK, Nova Guinea (N.S.), 2: 78 (redescription of holotype, and allotype, male). Distribution: New Guinea.

406 crenitis

LIEFTINCK

1932 Drepanosticta crenitis LIEFTINCK, Konowia, 11 (4): 288. Type female: Buitenzorg Museum; type-locality; Borneo. 1954 Drepanosticta crenitis - LIEFTINCK, Treubia, 22 Suppl.: 33 (Borneo).

Distribution: Borneo. dendrolagina

LIEFTINCK

1938 Drepanosticta dendrolagina LIEFTINCK, Nova Guinea (N.S.) 2: 83. Type male: A n n A r b o r Museum; type-locality: New Guinea.

Distribution: New Guinea, dentifera

KIMMINS

193 Drepanosticta dentifera KIMMINS, J. Fed. Mai. States Mus., 13: 101. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Drepanosticta dentifera - LIEFTINCK, Treubia, 22 Suppl.: 33 (Borneo).

Distribution: Borneo. digna

(HAGEN)

comb. nov.

I860 Platysticta (Platysticta) digna HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 10: 440. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Platysticta digna - KIRBY, Syn. Cat. Neur.-Odon., London: 132. 1931 Ceylonosticta digna - FRÄSER, J. Bombay nat. Hist. Soc., 35: 333.

Distribution: Sri Lanka. dorcadion

LIEFTINCK

1949 Drepanosticta dorcadion LIEFTINCK, N o v a Guinea (N.S.) 5: 59. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, drusilla

LIEFTINCK

1934 Drepanosticta drusilla LIEFTINCK, Treubia, 14 (4): 474. Type male: Buitenzorg Museum; typelocality: Borneo. 1954 Drepanosticta drusilla - LIEFTINCK, Treubia, 22 Suppl.: 33 (Borneo).

Distribution: Borneo. dulitensis KIMMINS 1936 Drepanosticta dulitensis KIMMINS, J. Fed. Mai. States Mus., 13: 98. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Drepanosticta dulitensis - LIEFTINCK, Treubia, 22 Suppl.: 33 (Borneo).

Distribution: Borneo. dupophila

LIEFTINCK

1932 Drepanosticta dupophila LIEFTINCK, Konowia, 11 (4): 286. Type male: Buitenzorg Museum; type-locality: Borneo. 1954 Drepanosticta dupophila - LIEFTINCK, Treubia, 22 Suppl.: 34 (Borneo).

Distribution: Borneo.

407 ephippiata 1937

LIEFTINCK

Drepanosticta ephippiata locality: Celebes.

LIEFTINCK,

Treubia,

1 6 (1): 7 2 .

Type male: Buitenzorg Museum; type-

Distribution: Celebes. eucera

LIEFTINCK

1949 Drepanosticta eucera LIEFTINCK, N o v a Guinea (N.S.) 5: 56. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. exoleta

LIEFTINCK

1932 Drepanosticta exoleta LIEFTINCK, Nova Guinea, 15 (5): 33. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, floresiana

LIEFTINCK

1939 Drepanosticta floresiana LIEFTINCK, Rev. franc. Ent., 6: 147. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Flores. 1 9 5 3 Drepanosticta floresiana - LIEFTINCK, Verh. naturf. Ges. Basel, 6 4 : 1 4 9 (redescription) (Flores).

Distribution: Flores. fontinalis

LIEFTINCK

Drepanosticta fontinalis LIEFTINCK, Treubia, 16 ( 1 ) : 6 4 . Type male: Univ. Mus. Michigan; type-locality: Malaya. 1942 Drepanosticta fontinalis race wheeleri FRÄSER, Proc. R. ent. Soc. London, (B) 11: 96. Male from Malaysia. 1 9 5 4 Drepanosticta fontinalis - LIEFTINCK, Treubia, 2 2 Suppl.: 3 4 (Malaysia). 1937

Distribution: Malaysia. forficula

KIMMINS

1936 Drepanosticta forficula KIMMINS, J. Fed. Mai. States Mus., 13:100. Type male: British Museum (Nat. Hist.), London; type-locality: Borneo. 1 9 5 4 Drepanosticta forficula - LIEFTINCK, Treubia, 2 2 Suppl.: 3 4 (Borneo).

Distribution: Borneo. fraseri

LIEFTINCK

1955 Drepanosticta fraseri LIEFTINCK, Zool. Meded., Leiden, 34 (5): 70. Type male: Naturhist. Mus., Basel; type-locality: Ceylon.

Distribution: Sri Lanka, gazella 1929

1934 1954

LIEFTINCK

Drepanosticta gazella LIEFTINCK, Tijdschr. Ent., 72: 1 1 0 . Type male: Buitenzorg Mus.; typelocality: Java. Drepanosticta gazella - LIEFTINCK, Treubia, 1 4 ( 4 ) : 3 8 9 (Java). Drepanosticta gazella - LIEFTINCK, Treubia, 2 2 Suppl.: 3 4 (Java).

Distribution: Java.

408 halterata

(BRAUER)

1868 Platysticta halterata BRAUER, Verh. Zool.-Bot. Ges. Wien., 18: 551. Type male: Naturhist. Mus. Wien; type-locality: Philippines. 1 9 8 4 Drepanosticta halterata - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 0 6 (Luzon).

Distribution: Philippine Islands, hamadryas

LAIDLAW 1931 Drepanosticta hamadryas LAIDLAW, J. Fed. Mai. States Mus., 16: 187. Type male: Laidlaw's Collection; type-locality: Pahang. 1954 Drepanosticta hamadryas - LIEFTINCK, Treubia 22 Suppl.: 34 (Malaya).

Distribution: Malaysia. hilaris

(HÄGEN)

I860 Platysticta (Platysticta) hilaris HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 10: 439. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Platysticta hilaris - KIRBY, Syn. Cat. Neur.-Odon., London: 132. 1915 Platysticta hilaris - LAIDLAW, Ree. Indian Mus., 11: 389 (Ceylon). 1931 Ceylonosticta hilaris - FRÄSER, J. Bombay nat. Hist. Soc., 35: 329 (as generotype of Ceylonosticta FRASER, 1931). 1 9 8 4 Drepanosticta hilaris - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 0 6 .

Distribution: Sri Lanka. inconspicua

LIEFTINCK

1938 Drepanosticta inconspicua LIEFTINCK, Nova Guinea, (N.S.), 2: 85. Type male: Senckenberg Mus.; type-locality: New Guinea.

Distribution: New Guinea, inversa

LIEFTINCK

1949 Drepanosticta inversa LIEFTINCK, Nova Guinea (N.S.) 5: 55. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. krugeri

LAIDLAW

1926 Drepanosticta krugeri L A I D L A W (nec Ris, 1927), J. Malay, Br. R. Asiat. Soc., 4: 218. Type male: Laidlaw's Collection; type-locality: Mentawi Is. 1927 Drepanosticta kruegeri- Ris, Zool. Meded., Leiden, 10 (1): 19 (redescription) (Sumatra). 1934 Drepanosticta krugeri - SCHMIDT, Arch. Hydrob. Suppl., 13: 334 (Sumatra). 1934 Drepanosticta krugeri - LIEFTINCK, Treubia, 14 (4): 468. 1954 Drepanosticta krugeri - LIEFTINCK, Treubia, 22 Suppl.: 34 (Mentawei Island).

Distribution: Mentawei Island, Sumatra. lankanensis

(FRASER)

1931 Ceylonosticta lankanensis FRASER, J. Bombay nat. Hist. Soc., 35: 335. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1966 Drepanosticta lankanensis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 200. Distribution: Sri L a n k a .

409 lepyricollis LIEFTINCK 1949 Drepanosticta lepyricollis LIEFTINCK, Nova Guinea (Ν.S.) 5:57. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality; New Guinea.

Distribution: New Guinea. l e s t o i d e s (BRAUER) 1868 Platysticta lestoides BRAUER, Verh. Zool.-Bot. Ges. Wien, 18: 552. Type male: Naturhist. Mus. Wien; type-locality: Philippines. 1984 Drepanosticta lestoides - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 106 (Mindanao).

Distribution: Philippine Islands. l y m e t t a COWLEY 1936 Drepanosticta lymetta COWLEY, Trans. R. ent. Soc. London, 85: 161. Type male: British Museum (Nat. Hist.), London; type-locality: Surigao. 1939 Drepanosticta lymetta - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 263 (Surigao).

Distribution: Philippine Islands. m a r s y a s LIEFTINCK 1984 Drepanosticta marsyas LIEFTINCK, 1965 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s o f t h e

World, Utrecht, 1: 106.

Distribution: Malaysia. megametta COWLEY 1936 Drepanosticta megametta COWLEY, Trans. R. ent. Soc. London, 85: 163. Type male: British Museum (Nat. Hist.), London; type-locality: Surigao. 1939 Drepanosticta megametta - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 264 (Mindanao).

Distribution: Philippine Islands. moluccana LIEFTINCK 1915 Platysticta bicornuta Ris (nec SELYS, 1878), Nova Guinea, 13 (2): 89. Type female: Rijksmus. Natuurl. Hist., Leiden; type-locality: Ceram. 1938 Drepanosticta moluccana LIEFTINCK, Nova Guinea, (N.S.), 2: 82 (new name for Platysticta bicornuta Ris, 1915, a junior secondary homonymy of Drepanosticta bicornuta SELYS, 1878). 1984 Drepanosticta auriculata R i s 1929 ( n e c SELYS, 1878) - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s

of the World, Utrecht, 1: 105.

Distribution: Ceram. m o n t a n a (HÄGEN) I860 Platysticta (Platysticta) montana HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 10: 438. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Platysticta montana - KIRBY, Syn. Cat. Neur.-Odon. London: 132. 1931 Ceylonosticta montana - FRÄSER, J. Bombay nat. Hist. Soc., 35: 331. 1984 Drepanosticta montana - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 106.

Distribution: Sri Lanka. m y l i t t a COWLEY 1936 Drepanosticta mylitta COWLEY, Trans. R. ent. Soc. London, 85: 160. Type male: British Museum (Nat. Hist.), London; type-locality: Borongan.

410 1939 Drepanosticta myletta (misprint) - NEEDHAM & GYGER, Philipp. J. Sei., 70 (3): 254 (Samer).

Distribution: Philippine Islands. nietneri (FRASER) 1931 Ceylonosticta nietneri FRÄSER, J. Bombay nat. Hist. Soc., 35: 334. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1966 Drepanosticta nietneri - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 206.

Distribution: Sri Lanka. palauensis LIEFTINCK 1962 Drepanosticta palauensis LIEFTINCK, Insecta of Micronesia, 5 (1): 14. Type male: US natn. Mus., Washington; type-locality: Palau.

Distribution: Western Caroline Island, p a n LAIDLAW 1931 Drepanosticta pan LAIDLAW, J. Fed. Mai. States Mus., 16: 185. Type male: Laidlaw's Collection; type-locality: Perak. 1954 Drepanosticta pan - LIEFTINCK, Treubia, 22 Suppl.: 35 (Malaya).

Distribution: Malaysia. philippa LIEFTINCK 1961 Drepanosticta philippa LIEFTINCK, Fieldiana (Zool.), 42: 132. Type male: unknown; type-locality: Luzon.

Distribution: Philippine Islands, polychromatica FRASER 1931 Drepanosticta polychromatica FRASER, J. Bombay nat. Hist. Soc., 35: 338. Type male: British Museum (Nat. Hist.), London; type-locality: India.

Distribution: India. pytho LIEFTINCK 1937 Drepanosticta pytho LIEFTINCK, Treubia, 16 (1): 68. Type male: Univ. Mus. Michigan; typelocality: Sumatra. 1954 Drepanosticta pytho - LIEFTINCK, Treubia, 22 Suppl.: 35 (Sumatra).

Distribution: Sumatra. quadrata (SELYS) I860 Platysticta (Platysticta) quadrata SELYS, Bull. Acad. r. Belg., (2) 10: 441. Type male: Selys's Collection; type-locality: Singapore. 1890 Platysticta quadrata - KIRBY, Syn. Cat. Neur.-Odon., London: 133. 1924 Platysticta quadrata - LAIDLAW, J. Mal. Br. R. Asiatic Soc., 2: 304. 1954 Drepanosticta quadrata - LIEFTINCK, Treubia, 22 Suppl.: 35 (Malaya).

Distribution: Malaysia, Singapore. robusta FRASER 1926 Drepanosticta robusta FRASER, Treubia, 8 (3-4): 491. Type female: Buitenzorg Mus.; typelocality: Key Is. Distribution: K e y I s l a n d .

411 r u f o s t i g m a (SELYS) 1886 Platysticta rufostigma SELYS, Mem. Cour. Acad. Belg., 38:155. Type male: Selys's Collection; type-locality: Lebuan. 1 9 2 4 Drepanosticta rufostigma - L A I D L A W , J . Mai. Br. R . Asiatic Soc., 2: 3 0 6 . 1932 Drepanosticta rufostigma - LIEFTINCK, Konowia, 11 (4): 292 (redescription). 1 9 3 6 Drepanosticta rufostigma - KIMMINS, J . Fed. Mai. States Mus., 18: 9 8 . 1954 Drepanosticta rufostigma - LIEFTINCK, Treubia, 22 Suppl.: 35 (Borneo).

Distribution: Borneo. septima

NEEDHAM &

GYGER

1939 Drepanosticta septima N E E D H A M given; type-locality: Samar.

& GYGER,

Philipp. J. Sei., 70 (3): 264. Type female: not

Distribution: Samar Island, sharpi (LAIDLAW) 1907 Platysticta sharpi LAIDLAW, Fasc. Malayensis Zool., 4. Odonata 2: 10. Type male: Univ. Mus., Michigan; type-locality: Malaysia. 1924 Drepanosticta sharpi - LAIDLAW, J. Mal. Br. R. Asiatic Soc., 2: 304. 1954 Drepanosticta sharpi - LIEFTINCK, Treubia, 22 Suppl.: 35 (Malaya).

Distribution: Malaysia. siebersi

FRASER

Drepanosticta cality: Java. 1929 Drepanosticta 1934 Drepanosticta 1954 Drepanosticta 1926

siebersi

FRASER,

Treubia,

8 ( 3 - 4 ) : 490.

Type male: Buitenzorg Mus.; type-lo-

siebersi - LIEFTINCK, Tijdschr. Ent., 57: 114. siebersi - LIEFTINCK, Treubia, 14 (4): 390 (Java). siebersi - LIEFTINCK, Treubia, 22 Suppl.: 36 (Java).

Distribution: Java. silenus

LAIDLAW

Drepanosticta silenus L A I D L A W , J . Fed. Mai. States Mus., 17: lection; type-locality: Malaysia. 1954 Drepanosticta silenus - LIEFTINCK, Treubia, 22 Suppl.: 36. 1934

557.

Type male: Laidlaw's Col-

Distribution: Malaysia. sinhalensis

LIEFTINCK

1970 Drepanosticta sinhalensis LIEFTINCK, Entom. scand. (Suppl.), 1: 191. Type male: not given; type-locality: Sri Lanka.

Distribution: Sri Lanka, spathulifera

LIEFTINCK

1929 Drepanosticta spathulifera LIEFTINCK, Tijdschr. Ent., 72:112. Type male: Buitenzorg Museum; type-locality: Java. 1934 Drepanosticta ? spathulifera - SCHMIDT, Arch. Hydrob. Suppl., 13: 334 (Java). 1954 Drepanosticta spathulifera - LIEFTINCK, Treubia, 22 Suppl.: 36 (Java).

Distribution: Java.

412 starmuehlneri ST. QUENTIN 1972 Drepanosticta starmuehlneri ST. QUENTIN, Bull. Fish. Res. Stn. Sri Lanka, 23 (1-2): 137. Type male: ? Naturhist. Mus. Wien; type-locality: Sri Lanka.

Distribution: Sri Lanka. submontana (FRASER) 1933 Ceylonosticta submontana FRASER, Spolia zeylan., 17: 214. Type male: British Museum (Nat. Hist.)? London; type-locality: Ceylon. 1966 Drepanosticta submontana - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 215.

Distribution: Sri Lanka. subtropica (FRASER) 1933 Ceylonosticta subtropica FRASER, Spolia zeylan., 17: 209. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1984 Drepanosticta subtropica - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 106.

Distribution: Sri Lanka, sundana (KRÜGER) 1898 Platysticta sundana KRÜGER, Stettiner ent. Ztg., 59:107. Type male: Stettin Mus.; type-locality: Java. 1929 Drepanosticta sundana - LIEFTINCK, Tijdschr. Ent., 72: 113. 1934 Drepanosticta sundana - LIEFTINCK, Treubia, 14 (4): 390, and 470 (redescription) (Java). 1954 Drepanosticta sundana - LIEFTINCK, Treubia, 22 Suppl.: 36 (Java).

Distribution: Java.

taurus NEEDHAM & GYGER 1941 Drepanosticta taurus NEEDHAM & GYGER, Philipp. J. Sei., 74: 145. Type male: not given; type-locality: Philippines.

Distribution: Philippine Islands, tenella LIEFTINCK 1935 Drepanosticta tenella LIEFTINCK, Misc. Zool. Sum., 92-93: z. Type male: Buitenzorg Mus.; type-locality: Sumatra. 1954 Drepanosticta tenella - LIEFTINCK, Treubia, 22 Suppl.: 37 (Sumatra).

Distribution: Sumatra. trimaculata LIEFTINCK 1939 Drepanosticta trimaculata LIEFTINCK, Rev. franc. Ent., 6: 149. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Philippines.

Distribution: Philippine Islands. tropica (HÄGEN) I860 Platysticta (PLatysticta) tropica HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 10: 438. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Platysticta tropica - KIRBY, Syn. Cat. Neur.-Odon., London: 132. 1931 Ceylonosticta tropica - FRASER, J. Bombay nat. Hist. Soc., 35: 332. 1984 Drepanosticta tropica - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 107. Distribution: Sri L a n k a .

413

viridis

FRÄSER

1922 Drepanosticta viridis FRÄSER, Mem. Dept. Agr., India, 7: 39. Type male: British Museum (Nat. Hist.), London (ex Pusa Collection); type-locality: Burma.

Distribution: Burma. walli

(FRASER)

1931 Ceylonosticta walli FRÄSER, J. Bombay nat. Hist. Soc., 35: 335. Type male: originally in Fraser's Collection, now unknown; type-locality: Ceylon. 1984 Drepanosticta walli - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 107.

Distribution: Sri Lanka. Genus Platysticta

SELYS

I860 Platysticta SELYS, Bull. Acad. r. Belg., (2) 10: 433. Type species: Platysticta maculata NIETNER, 1860 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 132).

apicalis

KIRBY

1894 Platysticta apicalis KIRBY, J. Linn. Soc., 24: 561. Type male: British Museum (Nat.Hist.), London; type-locality: Ceylon. 1955 Platysticta apicalis - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 69 (Ceylon).

Distribution: Sri Lanka. maculata maculata

NIETNER

I860 Platysticta (Platysticta) maculata NIETNER, in SELYS, Bull. Acad. r. Belg., (2) 10: 437. Type male: Hagen's Collection; type-locality: Ceylon. 1890 Platysticta maculata - KIRBY, Syn. Cat. Neur.-Odon., London: 132 (as generotype of Platysticta

SELYS, 1 8 9 0 ) .

1891 Platysticta greeni KIRBY, Proc. Zool. Soc. London, 1891: 204. Type male: British Museum (Nat.Hist.), London; type-locality: Ceylon.

Distribution: Sri Lanka. maculata deccanensis

LAIDLAW

1915 Platyscticta maculata deccanensis LAIDLAW, Ree. Indian Mus., 11: 388. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Kavalai. 1917 Platysticta deccanensis - LAIDLAW, Ree. Indian Mus., 13: 340 (as species). 1924 Platysticta deccanensis - FRÄSER, Ree. Indian Mus., 26: 498 (West India).

Distribution: East Asia. Genus Protosticta

SELYS

1885 Protosticta SELYS, C.R. Soc. Ent. Belg., 29: Species typica: Protosticta simplicinervis SELYS, 1885 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon. London: 133). - 1931 Protosticta - FRÄSER, Ree. Indian Mus., 33: 466. - 1954 Protosticta - LIEFTINCK, Treubia, 22 Suppl.: 31.

antelopoides

FRASER

1931 Protosticta antelopoides FRASER, Ree. Indian Mus., 33: 467. Type male: British Museum (Nat.Hist.), London; type-locality: West India.

Distribution: West India.

414 bivittata

LIEFTINCK

1939 Protosticta bivittata LIEFTINCK, Rev. franc. Ent., 6: 151. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Celebes.

Distribution: Celebes. curiosa

FRASER

1934 Protosticta curiosa FRASER, Stylops, 3: 134. Type male: British Museum (Nat. Hist.), London; type-locality: Lower Burma.

Distribution: Burma, davenporti

FRASER

1931 Protosticta davenporti FRASER, J. Bombay nat. Hist. Soc., 35: 70. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1931 Ceylonolestes davenporti - FRASER, Ree. Indian Mus., 33: 464. 1931 Protosticta davenporti - FRASER, Ree. Indian Mus., 33: 467.

Distribution: India. feronia

LIEFTINCK

1933 Protosticta feronia LIEFTINCK, Konowia, 11 (4): 281. Type male: Buitenzorg Mus.; type-locality: West Borneo. 1933 Protosticta feronia - LIEFTINCK, Konowia, 11: 281. 1954 Protosticta feronia - LIEFTINCK, Treubia, 22 Suppl.: 31 (Borneo).

Distribution: Borneo. foersteri

LAIDLAW

1902 Protosticta foersteri LAIDLAW, Proc. Zool. Soc. London, 2: 383. Type female: Laidlaw's Collection; type-locality: Perak. 1907 Protosticta försteri - LAIDLAW, Malay. Zool., 4: 2 (description of male). 1931 Protosticta foersteri - LAIDLAW, J. Fed. Mai. States Mus., 16: 190. 1954 Protosticta foersteri - LIEFTINCK, Treubia, 22 Suppl.: 31 (Malaya).

Distribution: Malaysia. fraseri

KENNEDY

1936 Protosticta fraseri KENNEDY, Proc. R. ent. Soc. London, (B) 5: 67. Type male: Ohio State Univ., Columbus; type-locality: Assam.

Distribution: Assam. gracilis

KIRBY

1889 Protosticta gracilis KIRBY, Proc. Zool. Soc. London, 1889: 302. Type male: British Museum (Nat. Hist.), London; type-locality: Menado.

Distribution: Menado. gravelyi

LAIDLAW

1915 Protosticta gravelyi LAIDLAW, Ree. Indian Mus., 11: 389. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Kavalai. 1917 Protosticta gravelyi - LAIDLAW, Ree. Indian Mus., 13: 242 (India). 1922 Protosticta stevensi FRASER, Ree. Indian Mus., 24: 7. Type male: British Museum (Nat. Hist.), London; type-locality: Nilgiris. 1924 Protosticta

gravelyi

- FRASER, Ree. I n d i a n M u s . , 26: 499 (West India).

415 1924 Protosticta stevensi - FRÄSER, Ree. Indian Mus., 26: 499 (West India). 1924 Protosticta mortoni FRÄSER, Ree. Indian Mus., 26: 500. Type male: British Museum (Nat. Hist.), London; type-locality: West India. 1931 Protosticta gravelyi - FRASER, J. Bombay nat. Hist. Soc., 35: 69. 1931 Protosticta stevensi - FRÄSER, J. Bombay nat. His. Soc., 35: 69 (proposed synonymy with gravelyi

LAIDLAW, 1 9 1 5 ) .

1931 Protosticta gravelyi

mortoni - FRÄSER, J. Bombay nat. Hist. Soc., 35: 69 (proposed synonymy with

LAIDLAW, 1 9 1 5 ) .

Distribution: India. hearseyi

FRASER

1922 Protosticta hearseyi FRÄSER, Ree. Indian Mus., 24: 5. Type male: British Museum (Nat. Hist.), London; type-locality: Nilgiris.

Distribution: India. himalaiaca

LAIDLAW

1917 Protosticta himalaiaca LAIDLAW, Ree. Indian Mus., 13: 342. Type male: Nation Coll. zool. Surv. India, Calcutta; type-locality: Darjeeling. 1920 Protosticta lindgreni FRÄSER, J. Bombay nat. Hist. Soc., 27: 150. Type male: British Museum (Nat. Hist.), London; type-locality: Darjeeling. 1931 Protosticta himalaiaca - FRÄSER, J. Bombay nat. Hist. Soc., 35: 74. 1 9 3 1 Protosticta lindgreni - FRÄSER, J . Bombay nat. Hist. Soc., 35: 7 4 (proposed synonymy with himalaiaca LAIDLAW, 1 9 1 7 ) . 1966 Protosticta himalaiaca - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 201.

Distribution: India. kiautai

ZHON

1986 Protosticta kiautai ZHON, Odonatologica, 15 (4): 465. Type male: Zhon's Collection; typelocality: China.

Distribution: China. kinabaluensis

LAIDLAW

1915 Protosticta Collection; 1 9 2 4 Protosticta 1933 Protosticta 1954 Protosticta

kinabaluensis type-locality: kinabaluensis kinabaluensis kinabaluensis

LAIDLAW, Proc. Zool. Soc. London, 1915: 37. Type male: Laidlaw's Borneo. - L A I D L A W , J . Mal. Br. R . Asiatic Soc., 2 : 3 0 6 . - LIEFTINCK, Konowia, 11: 285. - LIEFTINCK, Treubia, 22 Suppl.: 31 (Borneo).

Distribution: Borneo. medusa

FRASER

1934 Protosticta medusa FRASER, Stylops, 3: 135. Type male: British Museum (Nat. Hist.), London; type-locality: Lower Burma.

Distribution: Burma. robusta

FRASER

1933 Protosticta robusta FRASER, J. Siam Soc. nat. Hist. Suppl., 9:111. Type female: British Museum (Nat. Hist.), London; type-locality: Laos.

Distribution: Laos.

416

rufostigma

KIMMINS

1958 Protosticta rufostigma KIMMINS, Bull. Brit. Mus. Nat. Hist. (Ent.), 7: 349. Type male: British Museum (Nat. Hist.), London; type-locality: South India.

Distribution: South India. sanguinostigma

FRÄSER

1922 Protosticta sanguinostigma FRÄSER, Ree. Indian Mus., 24: 6. Type male: British Museum (Nat. Hist.), London; type-locality: Nilgiris. 1924 Protosticta sanguinostigma - FRÄSER, Ree. Indian Mus., 26: 499 (West India). 1924 Protosticta cerinostigma FRÄSER, Ree. Indian Mus., 26: 499. Type male: British Museum (Nat. Hist.), London; type-locality: West India. 1966 Protosticta cerinostigmaKIMMINS, Bull. Brit.Mus. nat. Hist. (Ent.), 18: 186 (as var. of sanguinostigma 1984

FRASER, 1 9 2 2 ) .

Protosticta sanguinostigma Utrecht, 1: 107.

f. cerinostigma

- DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: India. simplicinervis

SELYS

1885 Protosticta simplicinervis SELYS, C.R. Ent. Belg., 2 9 : 145 (as generotype of Protosticta SELYS, 1885). Type male: Selys's Collection; type-locality: Celebes. 1890 Protosticta simplicinervis - KIRBY, Syn. Cat. Neur.-Odon., London: 133. 1 9 2 6 Protosticta annulata FRASER, Treubia, 8 ( 3 - 4 ) : 4 9 2 . Type male: Buitenzorg Mus.; type-locality: Celebes. 1930 Protosticta simplicinervis - LIEFTINCK, Treubia, 12: 138 (Celebes). 1930 Protosticta annulata - LIEFTINCK, Treubia, 12: 138 (proposed synonymy with simplicinervis SELYS, 1 8 8 5 ) .

Distribution: Celebes. taipokauensis 1987

ASAHINA &

DUDGEON

Protosticta taipokauensis ASAHINA & D U D G E O N , Tombo, Kawaga Univ., Tokyo; type-locality: Hong-Kong.

3 0 ( 1 - 4 ) : 2.

Type male: Lab. Ent.

Distribution: Hong-Kong. trilobata

FRASER

1933 Protosticta trilobata FRASER, J. Siam. nat. Hist. Suppl., 9: 112. Type male: British Museum (Nat. Hist.), London; type-locality: Laos.

Distribution: Laos. uncata 1931

FRASER

Protosticta uncata FRASER, J . Bombay nat. Hist. Soc., (Nat. Hist.), London; type-locality: Upper Burma.

35: 75.

Type male: British Museum

Distribution: Burma. versicolor

LAIDLAW

1913 Protosticta Collection; 1924 Protosticta 1933 Protosticta 1954 Protosticta

versicolor LAIDLAW, Proc. Zool. Soc. London, 1913: 78. Type female: Laidlaw's type-locality: Borneo. versicolor - LAIDLAW, J. Mal. Br. R. Asiatic Soc., 2: 307. versicolor - LIEFTINCK, Konowia, 11: 285. versicolor - LIEFTINCK, Treubia, 22 Suppl.: 32 (Borneo).

Distribution: Borneo.

417 Family Protoneuridae Small to medium sized Zygoptera. Abdomen frequently very long. Anal vein absent or reduced; CuP present or reduced, no cross vein in cubital-anal space at base of wing, Dc a long regular rectangle, at least 3 times as long as wide, sub Dc contiguous with posterior border of wing. Pterostigma short, trapezoidal. 1926 Protoneuridae TILL YARD, Insects of Austr. and New Zeal.: 7 6 . - 1 9 3 8 Protoneuridae-TILLY ARD & FRASER, Austr. Zool., 9: 163 (with genera Peristicta, Neoneura, Idioneura, Microneura, Protoneura, Epipleoneura, Epipotoneura, Phasmoneura, Psaironeura, Chlorocnemis, Disparoneura, Caconeura, Esme, Phylloneura, Melanoneura, Elattoneura, Isomecocnemis, Nososticta, Austrosticta, Neosticta, Notoneura, Orosticta, Selysioneura, Isosticta, Prodasineura). - 1962 Protoneuridae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 102. - 1977 Protoneuridae BELYSHEV & KHARITONOV, N a u k a Novosibirsk: 225 (key to boreal genera). - 1984 Protoneuridae - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 108.

Subfamily Caconeurinae Relatively robust; CuP up to 10 or more cells distal to level of vein descending from subnodus. 1957 Caconeurinae FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 44. Type-genus: Caconeura KIRBY, 1890 (with genera Caconeura, Esme, Phylloneura, Melanoneura).

Genus Caconeura

KIRBY

I860 Alloneura (Alloneura) SELYS (nec SELYS, 1886), Bull. Acad. r. Belg., (2) 10: 446. Type species: Argia gomphoides RAMBUR, 1842 (with subgenera Peristicta, Disparoneura, Alloneura, Nososticta). - 1 8 9 0 Caconeura KIRBY, Syn. Cat. Neur.-Odon., London: 134 (new name for Alloneura SELYS, 1 8 6 0 ( n e c SELYS, 1 8 8 6 ) , a j u n i o r p r i m a r y h o m o n y m y o f Alloneura

RONDANI, 1 8 5 6 i n

Diptera). - 1917 Indoneura LAIDLAW, Ree. Indian Mus., 13: 344. Type species: Argia gomphoides RAMBUR, 1842 (original designation). - 1919 Hypostrophoneura MUNZ, Mem. Amer. ent. Soc., 3: 64. Type species: Argia gomphoides RAMBUR, 1842 (original designation). - 1931 Indoneura - FRÄSER, Ree. Indian Mus., 33: 469. - 1934 Indoneura - COWLEY, Entomologist, 67: 203 (proposed synonymy with Caconeura KIRBY, 1890). - 1934 Hypostrophoneura - COWLEY, Entomologist, 67: 203 (proposed synonymy with Caconeura KIRBY, 1890). - 1936 Caconeura - COWLEY, Ann. Mag. nat. Hist., (10) 17: 518. - 1962 Alloneura - PINHEY, Publ. cult. Comp. Diam. Angola, 59:104 (as syn. with Elattoneura COWLEY, 1935). - 1962 Indoneura - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104 (incorrect synonymy with Prodasineura COWLEY, 1934). - 1962 Caconeura - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104 (inc o r r e c t s y n o n y m y w i t h Prodasineura

COWLEY, 1 9 3 4 ) . - 1 9 8 4 Indoneura

- DAVIES & TOBIN,

The Dragonflies of the World, Utrecht, 1: 108 (as "incl." in Caconeura KIRBY, 1890).

gomphoides

(RAMBUR)

1842 Argia gomphoides RAMBUR, Hist. nat. Ins. Neuropt., Paris: 256. Type male: Guerin's Collection; type-locality: Nilgris. 1860 Alloneura (Alloneura) gomphoides - SELYS, Bull.Acad. r. Belg., (2) 10: 448 (as generotype of Alloneura

SELYS, 1 8 6 0 ) .

1890 Disparoneura gomphoides - KIRBY, Syn. Cat. Neur.-Odon., London. 134. 1917 Indoneura gomphoides - LAIDLAW, Ree. Indian Mus., 13: 347 (as generotype of

Indoneura

LAIDLAW, 1 9 1 7 ) .

1919 Hypostrophoneura postrophoneura

gomphoides - MUNZ, Mem. Amer. ent. Soc., 3: 64 (as generotype of HyMUNZ, 1919).

1924 Indoneura gomphoides - FRÄSER, Ree. Indian Mus., 26: 505 (West India).

418 1934 Caconeura gomphoides - COWLEY, Entomologist, 67: 203.

Distribution: India. longjingensis

ZHON

1984 Caconeura longjingensis ZHON, 1981 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 108.

Distribution: China. obscura

(FRASER)

1933 Indoneura obscura FRASER, Fauna brit. India, Odon., 1: 257. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1966 Caconeura obscura - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 207.

Distribution: India. ramburi

(FRASER)

1922 Indoneura ramburi FRASER, Mem. Dept. Agr. India, 7 (2): 2. Type male: British Museum (Nat. Hist.), London; type-locality: Madras. 1924 Indoneura ramburi - FRASER, Ree. Indian Mus., 26: 505 (West India). 1933 Indoneura t-coerulea FRASER, Fauna brit. India, Odon., 1: 256. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1966 Caconeura ramburi - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 211. 1966 Caconeura t-coerulea - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 216. 1984 Caconeura ramburi f. t-coerulea - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 108.

Distribution: India. risi

(FRASER)

1931 Indoneura risi FRASER, Ree. Indian Mus., 33: 469. Type male: British Museum (Nat. Hist.), London; type-locality: West India. 1966 Caconeura risi - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 211.

Distribution: India. Genus Esme

FRASER

1922 Esme FRASER, Mem. Dept. Agr. India, 7: 44. Type species: Esme cyaneovittata FRASER, 1922 (original designation). - 1936 Esme - COWLEY, Ann. Mag. nat. Hist., (10) 17: 516 (list of species).

cyaneovittata

FRASER

1922 Esme cyaneovittata FRASER, Mem. Dept. Agr. India, 7: 45 (as generotype of Esme FRASER, 1922). Type male: British Museum (Nat. Hist.), London; type-locality: Palni Hills. 1924 Esme cyaneovittata - FRASER, Ree. Indian Mus., 26: 506 (West India).

Distribution: India. longistyla

FRASER

1931 Esme longistyla FRASER, Ree. Indian Mus., 33: 471. Type male: British Museum (Nat. Hist.), London; type-locality: West India. 1966 Esme longistyla - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 202 (type lost?). 1984 Esme longistyla-DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:108 (South India).

Distribution: India.

419 mudiensis

FRÄSER

1931 Esme mudiensis FRÄSER, Ree. Indian Mus., 33: 472. Type male: British Museum (Nat. Hist.), London; type-locality: West India.

Distribution: India. Genus Melanoneura

FRASER

1922 Melanoneura FRASER, Mem. Dept. Agr. India, Ent., 7: 55. Type species: Melanoneura bilineata FRASER, 1 9 2 2 (original designation). - 1 9 3 6 Melanoneura - C O W L E Y , Ann. Mag. nat. Hist., ( 1 0 ) 17: 5 1 9 .

bilineata

FRASER

1922 Melanoneura bilineata FRASER, Mem. Dept. Agr.India, Ent., 7: 55 (as generotype of Melanoneura FRASER, 1922). Type female: British Museum (Nat. Hist.), London; type-locality: Coorg. 1924 Melanoneura bilineata - FRASER, Ree. Indian Mus., 26: 504 (redescription) (West India). 1966 Melanoneura bilineata - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 182.

Distribution: India. Genus Phylloneura 1922

FRASER

Phylloneura FRASER, Ree. Indian Mus., 24: 3 . Type species: Alloneura westermanni SELYS, 1860 (monotypy). - 1934 Phylloneura - COWLEY, Entomologist, 67: 203. - 1936 Phylloneura - COWLEY, A n n . M a g . n a t . Hist., (10) 17: 516.

westermanni

(SELYS)

Alloneura (Alloneura) westermanni SELYS, Bull. Acad. r. Belg., ( 2 ) 10: 4 4 7 . Type male: Hagen's Collection; type-locality: Nilgiris. 1890 Disparoneura westermanni - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1922 Phylloneura westermanni - FRASER, Ree. Indian Mus., 24: 3 (as generotype of Phylloneura I860

FRASER, 1922).

1924 Phylloneura westermanni - FRASER, Ree. Indian Mus., 26: 506 (West India).

Distribution: India. Subfamily Disparoneurinae CuP extends several cells beyond vein descending from subnodus; male anal appendages trigger shaped in profile, triangular from above; anal vein reduced. 1957 Disparoneurinae FRASER, R. Zool. Soc. N.S. Wales, 1957: 44. Type-genus: Disparoneura SELYS, 1860 (with genera Elattoneura, Isomecocnemis, Notoneura, Prodasineura, Nososticta). - 1962 Disparoneurinae - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 102 (key to genera Chlorocnemis, Prodasineura, Isomecocnemis, Elattoneura, Notoneura).

Genus Chlorocnemis

SELYS

1863 Chlorocnemis SELYS, Synopsis des Agrionines, Add. 4: 31. Type species: Chlorocnemis elongata HÄGEN, 1863 (designated by KIRBY, 1890; Syn. Cat. Neur.-Odon., London: 131). - 1921 Chlorocnemis - Ris, Annals S. Afr. Mus., 18: 291. - 1936 Chlorocnemis - COWLEY, Ann. Mag. nat. Hist., (10) 17: 512 (list of species). - 1951 Chlorocnemis - SCHMIDT, Mitt, miinch. ent. Ges., 41: 229 (key to African species). - 1962 Chlorocnemis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 103.

420 abbotti

(CALVERT)

1896 Disparoneura abbotti CALVERT, Proc. U.S. natn. Mus., 48: 140. Type male: U.S. natn. Mus., Washington; type-locality: Kilimandjaro. 1902 Chlorocnemis inepta GRÜNBERG, SitzBer. Ges. naturf. Fr. Berlin, 9: 233. Type male: Mus. Naturk., Berlin; type-locality: Kilimandjaro. 1951 Chlorocnemis abbotti - SCHMIDT, Mitt, münch. ent. Ges., 41: 236 (as Chlorocnemis inepta GRÜNBERG, 1902 type).

1962 Chlorocnemis abbotti - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 103. 1962 Chlorocnemis inepta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 103 [proposed synonymy with abbotti (CALVERT, 1896)]. 1 9 8 4 Chlorocnemis abbotti ( „ C A L V E R T , 1 8 9 2 , 1 9 , 1 6 4 " ) - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 108.

Distribution: East Africa. contraria

SCHMIDT

1951 Chlorocnemis contracta SCHMIDT, Mitt, münch. ent. Ges., 41: 236. Type male: Mus. Naturk., Berlin; type-locality: Cameroon. 1962 Chlorocnemis contraria - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 103.

Distribution: Cameroon. eisentrauti 1974

PINHEY

Chlorocnemis eisentrauti PINHEY, Bonner zool. Beitr., type-locality: Cameroon.

2 5 ( 1 - 3 ) : 188.

Type male: Bonn Mus.;

Distribution: Cameroon. elongata

HÄGEN

1863 Chlorocnemis elongata HAGEN, in SELYS, Synopsis des Agrionines, Add. 4: 32 (as generotype of Chlorocnemis SELYS, 1 8 6 3 ) . Type male: Univ. zool. Mus., Kebenhavn; type-locality: Guinea. 1893 Chlorocnemis elongata - KARSCH, Berl. ent. Z., 38: 36 (description of female). 1951 Chlorocnemis elongata - SCHMIDT, Mitt, münch. ent. Ges., 41: 236. 1962 Chlorocnemis elongata - PINHEY, Publ. cult. Comp. Diam. Angola, 59:103 [type male in Mus. Comp. Zool., Cambridge (Mass.) from Togoland].

Distribution: West Africa. flavipennis

SELYS

1863 Chlorocnemis flavipennis SELYS, Synopsis des Agrionines, Add. 4: 32. Type male: Naturhist. Mus., Wien; type-locality: Sierra Leone. 1890 Chlorocnemis flavipennis - KIRBY, Syn. Cat. Neur.-Odon., London: 131. 1962 Chlorocnemis flavipennis- PINHEY, Publ. cult. Comp. Diam. Angola, 59:103 (type male: Brussles Mus., and „type not designated").

Distribution: Sierra Leone. interrupta 1984

LEGRAND

Chlorocnemis interrupta World, Utrecht, 1: 109.

LEGRAND, 1 9 8 4

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Gabon. marshalli marshalli Ris 1921 Chlorocnemis marshalli Ris, Annals S. Afr. Mus., 18: 291. Type male: British Museum (Nat. Hist.), London; type-locality: Mashonaland.

421 1928 Chlorocnemis marshalli - FRÄSER, Trans, ent. Soc. London, 76: 128 (Nyasaland). 1962 Chlorocnemis marshalli marshalli - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104 (S. Rhodesia, Nyasaland). 1984 Chlorocnemis marshalli marshalli f. impedita PINHEY, 1969 from Vumba - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 109.

Distribution: South Africa, marshalli superba

SCHMIDT

1951 Chlorocnemis superba SCHMIDT, Mitt, miinch. ent. Ges., 41: 234. Type male: Mus. Naturk., Berlin; type-locality: Ruwenzori. 1962 Chlorocnemis marshalli race superba - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104 (Belgian Congo, Uganda). 1984 Chlorocnemis marshalli superba - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 109.

Distribution: Belgian Congo, Uganda, montana montana ST. QUENTIN 1941 Chlorocnemis montana ST. QUENTIN, Ann. N a t u r h . Mus. Wien, 52: 108. Type male: Naturhist. Mus., Wien; type-locality: Tanganyika. 1951 Chlorocnemis montana - SCHMIDT, Mitt, miinch. ent. Ges., 41: 237. 1962 Chlorocnemis montana - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104. 1969 Chlorocnemis montana montana - PINHEY, Occ. Pap. natn. Mus., Rhod., 4 (29B): 245.

Distribution: Tanzania. montana maccleeryi PINHEY 1969 Chlorocnemis montana maccleeryi PINHEY, Occ. Pap. natn. Mus., Rhod., 4 (29B): 245. Type male: Mus. Rhod.; type-locality: Malawi.

Distribution: Malawi. nigripes nigripes

SELYS

1886 Chlorocnemis nigripes SELYS, Mem. Cour. Acad. Belg., 38: 141. Type subadult: Liege Univ.; type-locality: West Africa. 1907 Disparoneura oculata MARTIN (nec KIRBY, 1894), Mem. Soc. esp., Madrid, 1: 428. Type male: Cienc. Mus. Madrid; type-locality: Guinea. 1908 Chlorocnemis gestroi MARTIN, Annali Mus. civ. Stor. nat. Genova, 43: 654. Type male: Mus. civ. Stor. nat. Genova; type-locality: West Africa. 1921 Disparoneura williamsoni MARTIN, Ann. Soc. ent. France, 90: 96 (new name for Disparoneura oculata MARTIN, 1907, a junior secondary homonymy of Disparoneura oculata KIRBY, 1894). 1936 Chlorocnemis williamsoni - COWLEY, Ann. Mag. nat. Hist., (10) 17: 512. 1946 Chlorocnemis nigripes - FRÄSER, Proc. R. ent. Soc. London, (B) 15: 41 (redescription of male) (Belgian Congo). 1951 Chlorocnemis nigripes - SCHMIDT, Mitt, miinch. ent. Ges., 41: 234. 1951 Chlorocnemis gestroi - SCHMIDT, Mitt, miinch. ent. Ges., 41: 234 (proposed synonymy with nigripes nigripes SELYS, 1886). 1951 Disparoneura williamsoni - SCHMIDT, Mitt, miinch. ent. Ges., 41: 234 (proposed synonymy with nigripes nigripes SELYS, 1886). 1962 Chlorocnemis nigripes - PINHEY, Publ. cul. Comp. Diam. Angola, 59: 104 (Uganda, Belgian Congo, Cameroon, Spanish Guinea). 1969 Chlorocnemis nigripes semlikiensis PINHEY, OCC. Pap. natn. Mus. Rhod., 4 (29B): 252. Type male: Pinhey's Collection; type-locality: Uganda.

422 1984 Chlorocnemis nigripes f. semlikiensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 109.

Distribution: West and Central Africa. nigripes coeruleocauda

PINHEY

1969 Chlorocnemis nigripes coeruleocauda PINHEY, OCC. Pap. natn. Mus. Rhod., 4 (29B): 252. Type male: Pinhey's Collection; type-locality: Central Africa.

Distribution: Central Africa. nubilipennis nubilipennis 1893 Chlorocnemis type-locality: 1951 Chlorocnemis 1962 Chlorocnemis roon).

KARSCH

nubilipennis KARSCH, Berl. ent. Z., 38: 37. Type male: Mus. Naturk., Berlin; Togo. nubilipennis - SCHMIDT, Mitt, münch. ent. Ges., 41: 237. nubilipennis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104 (Togo, Came-

Distribution: West Africa. nubilipennis rossii

PINHEY

1969 Chlorocnemis nubilipennis rossii PINHEY, OCC. Pap. natn. Mus. Rhod., 4 (29B): 257. Type male: Pinhey's Collection: Sierra Leone.

Distribution: Sierra Leone. pauli

LONGFIELD

1939 Chlorocnemis pauli LONGFIELD, Trans. R. ent. Soc. London, 85: 470. Type male: British Museum (Nat. Hist.), London; type-locality: Uganda. 1951 Chlorocnemis pauli - SCHMIDT, Mitt, münch. ent. Ges., 41: 237 (Uganda). 1962 Chlorocnemis pauli - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104.

Distribution: Uganda. wittei

FRASER

1955 Chlorocnemis wittei FRASER, Explor. Pare nat. Upemba, Miss, de Witte, 38: 7. Type male: Inst. Pare Nat. C.B.; type-locality: Belgian Congo. 1962 Chlorocnemis wittei - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 104. 1984 Chlorocnemis lascellesi PINHEY, 1961 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 109 (as syn. with wittei FRASER, 1955).

Distribution: Congo Republic. Genus Disparoneura

SELYS

1860 Alloneura (Disparoneura) SELYS (nec SELYS, 1886), Bull. Acad. r. Belg., (2) 10:443. Type species: Argia quadrimaculata RAMBUR, 1842 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 133). - 1890 Disparoneura - KIRBY, Syn. Cat. Neur.-Odon., London: 133. - 1917 Disparoneura - LAIDLAW, Ree. Indian Mus., 13: 344 (redescription, pseudotype: Disparoneura frenulata SELYS, 1860). - 1917 Chloroneura LAIDLAW, Ree. Indian Mus., 13: 344. Type species: Argia quadrimaculata RAMBUR, 1842 (original designation). - 1 9 2 1 Disparoneura - Ris, Annals S. Afr. Mus., 18: 293. - 1933 Disparoneura - FRASER, Fauna brit. India, Odon., 1: 228 (pseudotype: Disparoneura mutata SELYS, 1886). - 1934 Chloroneura - COWLEY, Entomologist, 67: 203 (syn with Disparoneura SELYS, 1860, a proposed name for Chloroneura LAIDLAW, 1917, a junior primary homonymy of Chloroneura WALSH, 1864 in Hemiptera). - 1936 Disparoneura - COWLEY, A n n . M a g . nat. Hist., (10) 17: 517 (list o f species).

423

apicalis

(FRÄSER)

1924 Chloroneura apicalis FRÄSER, Ree. Indian Mus., 26: 501. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1934 Disparoneura apicalis - COWLEY, Entomologist, 67: 203.

Distribution: India. quadrimaculata

(RAMBUR)

1842 Argia quadrimaculata RAMBUR, Hist. nat. Ins. Neuopt. Paris: 255. Type sex not given: Serville's Collection; type-locality: Bombay. 1860 Alloneura (Disparoneura) quadrimaculata - SELYS, Bull. Acad. r. Belg., (2) 10: 446 (as gener o t y p e of Disparoneura

SELYS, 1860).

1890 Disparoneura quadrimaculata - KIRBY, Syn. Cat. Neur.-Odon., London: 133. 1917 Chloroneura quadrimaculata - LAIDLAW, Ree. Indian Mus., 13: 344 (as generotype of Chloroneura L A I D L A W , 1 9 1 7 ) . 1924 Chloroneura quadrimaculata - FRÄSER, Ree. Indian Mus., 26: 501 (West India). 1934 Disparoneura quadrimaculata - COWLEY, Entomologist, 67: 203. 1936 Disparoneura quadrimaculata - COWLEY, Ann. Mag. nat. Hist., (10) 17: 577.

Distribution: India. ramajana 1984

LIEFTINCK

Disparoneura ramajana World, Utrecht, 1: 109.

LIEFTINCK, 1 9 7 1

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the

Distribution: Sri Lanka. Genus: Elattoneura

COWLEY

1935 Elattoneura COWLEY, Entomologist's month. Mag., 71:14. Type species: Disparoneura glauca SELYS, 1860 (original designation) (new name for Disparoneura SELYS, 1886, nec SELYS, 1860). - 1954 Elattoneura - LIEFTINCK, Treubia, 22 Suppl.: 37. - 1962 Elattoneura - PINHEY, Publ. cult. Comp. Diam. Angola, 59:104. - 1976 Elattoneura - LINDLEY, Odonatologica, 5 (2): 153.

acuta acuta

KIMMINS

1938 Elattoneura acuta KIMMINS, Ann. Mag. nat. Hist., (11) 1: 300. Type male: British Museum (Nat. Hist.), London; type-locality: Nigeria. 1954 Elattoneura henrardi FRASER, Rev. Zool. Bot. afr., 50: 271. Type male: Mus. r. Afr. centr., Tervuren; type-locality: Belgian Congo. 1961 Elattoneura henrardi - PINHEY, Entomologist's month. Mag., 97: 101 (Rhodesia). 1962 Elattoneura acuta - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105.

Distribution: Central Africa. acuta lindleyi 1984

LEGRAND

Elattoneura acuta lindleyi World, Utrecht, 1: 110.

LEGRAND, 1 9 8 0

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Central Africa. analis

(SELYS)

1860 Alloneura (Alloneura) analis SELYS, Bull. Acad. r. Belg., (2) 10: 451. Type male: Selys's Collection; type-locality: Malacca. 1890 Disparoneura analis - KIRBY, Syn. Cat. Neur.-Odon., London: 134.

424 1935 Caeconeura analis - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 8 (Sumatra). 1937 Elattoneura analis - LIEFTINCK, Treubia, 16 (1): 74 (redescription) (Borneo).

Distribution: Malaysia, Borneo, Sumatra. atkinsoni

(SELYS)

1886 Disparoneura atkinsoni SELYS, Mem. Cour. Acad. Belg., 38:168. Type male: Selys's Collection; type-locality: Khasia Hill. 1 9 0 5 Disparoneura atkinsoni - JACOBSON & B I A N K I , Pryam. lozhnos. Ross, imp.: 8 3 6 . 1917 Disparoneura atkinsoni - LAIDLAW, Ree. Indian Mus., 13: 347 (Burma). 1 9 1 9 Disparoneura fletcheri FRÄSER, J . Bombay nat. Hist. Soc., 26: 8 7 6 . Type male: British Museum (Nat. Hist.), London; type-locality: Shillong. 1933 Disparoneura atkinsoni - FRASER, F a u n a brit. India, Odon., 1: 230. 1933 Disparoneura fletcheri - FRASER, F a u n a brit. India, Odon., 1: 230 [proposed synonymy with atkinsoni (SELYS, 1 8 8 6 ) ] . 1966 Elattoneura atkinsoni - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 192.

Distribution: India, Burma. aurantiaca

(SELYS)

1886 Disparoneura aurantiaca SELYS, Mem. Cour. Acad. Belg., 38: 169. Type male: Selys's Collection; type-locality: Sarawak. 1937 Elattoneura aurantiaca - LIEFTINCK, Treubia, 16 (1): 80 (redescription) (Sumatra). 1954 Elattoneura aurantiaca - LIEFTINCK, Treubia, 22 Suppl.: 37 (Malaya, Sumatra, Borneo).

Distribution: Malaysia, Borneo, Sumatra. balli

KIMMINS

1938 Elattoneura (Nat. Hist.), 1961 Elattoneura 1962 Elattoneura

balli KIMMINS, Ann. Mag. nat. Hist., (11) 1: 296. Type male: British Museum London; type-locality: Sierra Leone. balli - PINHEY, Entomologist's month. Mag., 97: 101 (Rhodesia). balli - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105.

Distribution: Central and West Africa. bigemmata

LIEFTINCK

1970 Elattoneura bigemmata LIEFTINCK, Entom. scand. (Suppl.), 1:188. Type male: Lieftinck's Collection; type-locality: Sri Lanka.

Distribution: Sri Lanka. caesia

(HÄGEN)

I860 AUoneura (AUoneura) caesia HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 10: 450. Type male: Hagen's Collection; type-locality: Ceylon.

campioni campioni

(FRASER)

1922 Disparoneura campioni FRASER, Mem. Dept. Agr. India, 7: 43. Type male: British Museum (Nat. Hist.), London; type-locality: India. 1966 Elattoneura campioni - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 185.

Distribution: India. campioni cacharensis

(FRASER)

comb. nov.

1933 Disparoneura campioni cacharensis FRASER, F a u n a brit. India, Odon., 1: 244. Type male: British Museum (Nat. Hist.), London; type-locality: Assam.

Distribution: Assam.

425 1890 Disparoneura caesia - KIRBY, Syn. Cat. Neur.-Odon., London: 133. 1955 Elattoneura caesia - LIEFTINCK, Zool. Meded., Leiden, 3 4 (5): 73 (Ceylon). Distribution: Sri Lanka. centrafricana

LINDLEY

1976 Elattoneura centrafricana LINDLEY, Odonatologica, 5(2): 153. Type male: Lindley's Collection; type-locality: Central African Republic.

Distribution: Central African Republic. centralis I860

1890 1894

1919 1955 1962

1966

(HÄGEN)

Alloneura (Alloneura) centralis H A G E N , in SELYS, Bull. Acad. r. Belg., ( 2 ) 10: 4 4 9 . Type male: Hagen's Collection; type-locality: Ceylon. Disparoneura centralis - KIRBY, Syn. Cat. Neur.-Odon., London: 133. Disparoneura oculata K I R B Y (nec M A R T I N , 1 9 0 7 ) , J . Linn. Soc., 24: 5 6 3 . Type female: British Museum (Nat. Hist.), London; type-locality: Ceylon. Caconeura mackwoodi FRÄSER, J. Bombay nat. Hist. Soc., 26: 876. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. Elattoneura centralis - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 73 (Ceylon). Disparoneura oculata - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 0 4 [proposed synonymy with centralis ( H Ä G E N , I 8 6 0 ) ] . Caconeura mackwoodi - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 202 [proposed synonymy with centralis (HÄGEN, I860)].

Distribution: Sri Lanka. coomansi

LIEFTINCK

1937 Elattoneura coomansi LIEFTINCK, Treubia, 16 (1): 79. Type male: Buitenzorg Mus.; type-locality: Borneo. 1954 Elattoneura coomansi - LIEFTINCK, Treubia, 22 Suppl.: 38 (Borneo).

Distribution: Borneo. dorsalis dorsalis

KIMMINS

1938 Elattoneura dorsalis KIMMINS, Ann. Mag. nat. Hist., (11) 1: 299. Type male: British Museum (Nat. Hist.), London; type-locality: Sierra Leone. 1946 Caconeura dorsalis dorsalis - FRASER, Entomologist's month. Mag., 82: 201. 1962 Elattoneura dorsalis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105.

Distribution: Sierra Leone. dorsalis auricolor

(FRASER)

comb. nov.

1927 Caconeura auricolor FRASER, Ree. Indian Mus., 29:90. Type male: British Museum (Nat. Hist.), London; type-locality: Upper Burma. 1946 Caconeura dorsalis auricolor - FRASER, Entomologist's month. Mag., 82: 201. 1948 Prodasineura auricolor - LIEFTINCK, Treubia, 19 (2): 222 (Burma).

Distribution: Burma. erythromma

LIEFTINCK

1953 Elattoneura erythromma LIEFTINCK, Treubia, 22: 283. Type male: Lieftinck's Collection; typelocality: Borneo. 1954 Elattoneura erythromma - LIEFTINCK, Treubia, 22 Suppl.: 38 (Borneo).

Distribution: Borneo.

426 frenulata

(HAGEN)

Alloneura (Disparoneura) frenulata H A G E N , in SELYS, Bull. Acad. r. Belg., ( 2 ) 10: 4 4 4 . Type male: Mus. Comp. Zool., Cambridge (Mass.); type-locality: Cape Province. 1890 Disparoneura frenulata - KIRBY, Syn. Cat. Neur.-Odon., London: 133. 1936 Elattoneura frenulata - COWLEY, Ann. Mag. nat. Hist., (10) 17: 518. 1962 Elattoneura frenulata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105 (Cape Province, Victoria Falls, North Rhodesia, Belgian Congo, North Uganda). I860

Distribution: Central and South Africa. girardi 1984

LEGRAND

Elattoneura girardi Utrecht, 1: 110.

LEGRAND, 1 9 8 0

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: West Africa. glauca

(SELYS)

I860 Alloneura (Disparoneura) glauca SELYS, Bull. Acad. r. Belg., (2) 10: 443. Type male: Selys's Collection; type-locality: Cap of Good Hope, Natal. 1886 Disparoneura mutata SELYS, Mem. Cour. Acad. r. Belg., 38: 164. Type male: British Museum (Nat. Hist.), London; type-locality: East Africa. 1906 Disparoneura alba FÖRSTER, Jb. Nassau Ver. Natk., 59: 338. Type: unknown; type-locality: Somalia. 1906 Disparoneura simba MARTIN, Bull. Mus. Hist. nat. Paris, 1906: 509. Type male: Mus. Nation. Hist. Na., Paris; type-locality: Kenya. 1921 Disparoneura mutata - Ris, Annls S. Afr. Mus., 18: 293 (Transvaal, Natal, Zululand). 1935 Elattoneura glauca - COWLEY, Entomologist's month. Mag., 71: 14 (as generotype of Elattoneura COWLEY, 1935). 1947 Elattoneura mutata - FRASER, Trans. R. ent. Soc. London, 98: 24 (Ivory Coast). 1962 Elattoneura glauca - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105. 1962 Disparoneura mutata - PINHEY, Publ. cult. Comp. Diam. Angola, 59:105 [proposed synonymy with glauca (SELYS, I 8 6 0 ) ] . 1962 Disparoneura alba - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105 [proposed synonymy with glauca (SELYS, I 8 6 0 ) ] . 1962 Disparoneura simba - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105 [proposed synonymy with glauca (SELYS, I 8 6 0 ) ] .

Distribution: Central and South Africa. josemorai

COMPTE SART

1984 Elattoneura josemorai COMPTE World, Utrecht, 1: 110.

SART,

1964 - loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: West Africa. leucostigma

(FRASER)

1933 Disparoneura leucostigma FRASER, Spolia zeylan., 17: 225. Type male: Colombo Museum; type-locality: Ceylon. 1955 Elattoneura leucostigma - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 73 (Ceylon). 1966 Elattoneura leucostigma - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 200.

Distribution: Sri Lanka.

427 longispina

LIEFTINCK

1937 Elattoneura longispina LIEFTINCK, Treubia, 16 (1): 76. Type male: Buitenzorg Museum; typelocality: Borneo. 1954 Elattoneura longispina - LIEFTINCK, Treubia, 22 Suppl. 38 (Borneo).

Distribution: Borneo. nigerrima

(LAIDLAW)

1917 Disparoneura nigerrima LAIDLAW, Ree. Indian Mus., 13: 346. Type male: Indian Museum, Calcutta; type-locality: Nagpur. 1924 Elattoneura nigerrima - FRÄSER, Ree. Indian Mus., 26: 503 (West India).

Distribution: West India. nigra

KIMMINS

1938 Elattoneura nigra KIMMINS, Ann. Mag. nat. Hist., (11) 1: 297. Type male: British Museum (Nat. Hist.), London; type-locality: Nigeria. 1949 Disparoneura nigra - SCHMIDT, Arq. Mus. Bocage, 20: 129. 1962 Elattoneura nigra - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 106 (Nigeria, Portuguese West Africa).

Distribution: West Africa. pasquinii 1984

CONSIGLIO

Elattoneura pasquinii CONSIGLIO, World, Utrecht, 1: 110.

1978

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Ethiopia. pluotae 1984

LEGRAND

Elattoneura pluotae Utrecht, 1: 110.

LEGRAND, 1 9 8 2

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: West Africa. pruinosa

(SELYS)

1886 Disparoneura pruinosa SELYS, Mem. Cour. Acad. Belg., 38:166. Type male: Brussels Museum; type-locality: Sierra Leone. 1936 Elattoneura pruinosa - COWLEY, Ann. Mag. nat. Hist., (10) 17: 518. 1938 Elattoneura pruinosa - KIMMINS, Ann. Mag. nat. Hist., (11) 1: 295. 1941 Elattoneura aethiopia FRÄSER, Proc. R. ent. Soc. London, (B) 10: 36. Type male: Mus. Nation. Hist. Nat. Paris; type-locality: Cameroon. 1962 Elattoneura pruinosa - PINHEY, Publ. cult. Comp.Diam. Angola, 59: 106 (Belgian Congo, Cameroon, Sierra Leone). 1 9 7 1 Elattoneura aethiopia - PINHEY, J . ent. Soc. sth. Afr., 3 4 : 2 1 6 [proposed synonymy with pruinosa (SELYS, 1 8 8 6 ) ] .

Distribution: West Africa. souteri

(FRASER)

1924 Disparoneura souteri FRÄSER, Ree. Indian Mus., 26: 521. Type male: British Museum (Nat. Hist.), London; type-locality: West India. 1931 Disparoneura souteri - FRÄSER, Ree. Indian Mus., 33: 468. 1966 Elattoneura souteri - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 214.

Distribution: West India.

428 tenax

(HAGEN)

Alloneura (Alloneura) tenax H A G E N , in SELYS, Bull. Acad. r. Belg., ( 2 ) 10: Hagen's Collection; type-locality: Ceylon. 1890 Disparoneura tenax - KIRBY, Syn. Cat. Neur.-Odon., London: 133. 1955 Elattoneura tenax - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 73 (Ceylon). I860

449.

Type male:

Distribution: Sri Lanka. tetrica

(LAIDLAW)

1917 Disparoneura tetrica LAIDLAW, Ree. Indian Mus., 13: 345. Type male: Nation. Coll. zool. Surv. India, Calcutta; type-locality: Talewadi. 1924 Disparoneura tetrica - FRÄSER, Ree. Indian Mus., 26: 503 (India). 1984 Elattoneura tetrica - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: India. tropicalis

PINHEY

1974 Elattoneura tropicalis PINHEY, Arnoldia, Rhod., 7 (2): 1. Type male: Pinhey's Collection; typelocality: Victoria Falls.

Distribution: Rhodesia. vridaghi

FRASER

1954 Elattoneura vridaghi FRASER, Rev. Zool. Bot. afr., 50: 269. Type male: Mus. r. Afr. Centr., Tervuren; type-locality: Belgian Congo. 1962 Elattoneura vridaghi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 106. 1984 Elattoneura tsiamae AGUESSE, 1966 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the W o r l d ,

Utrecht, 1: 111 (proposed synonymy with vridaghi FRASER, 1954).

Distribution: Congo Republic. Genus: Isomecocnemis

COWLEY

1936 Isomecocnemis COWLEY, Ann. Mag. nat. Hist., (10) 17: 514. Type species: Chlorocnemis cyanura FÖRSTER, 1 9 0 9 (originaldesignation). - 1 9 6 2 Isomecocnemis-PINHEY, Publ. cult. Comp. Diam. Angola, 59: 106.

cellularis

(GRÜNBERG)

1902 Disparoneura cellularis GRÜNBERG, SitzBer. Naturf. Frde., Berlin, 9: 232. Type male: Mus. Naturk., Berlin; type-locality: Tanganyika. 1962 Elattoneura cellularis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 105. 1984 Isomecocnemis cellularis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: Tanzania. cyanura

(FÖRSTER)

1909 Chlorocnemis cyanura FÖRSTER, Jb. Wiesbaden Ver. Natk. 62: 234. Type male: Förster's Collection; type-locality: Cameroon. 1936 Isomecocnemis cyanura - COWLEY, Ann. Mag. nat. Hist., (10) 17: 514 (as generotype of Isomecocnemis COWLEY, 1936). 1954 Isomecocnemis cyanura - FRASER, Rev. Zool. Bot. afr., 50: 273 (Belgian Congo). 1962 Isomecocnemis cyanura - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 107 (Cameroon, Belgian Congo). Distribution: W e s t A f r i c a .

429 subnodalis

(SELYS)

1886 Disparoneura cotype male: 1890 Disparoneura 1893 Disparoneura 1936 Isomecocnemis 1947 Isomecocnemis 1951 Isomecocnemis 1962 Isomecocnemis

subnodalis SELYS, Mem. Cour. Acad. r. Belg., 38: 162. Type: not designated, British Museum (Nat. Hist.), London; type-locality: Old Calabar. subnodalis - KIRBY, Syn. Cat. Neur.-Odon., London: 133. subnodalis - KARSCH, Berl. ent. Z., 38: 36. subnodalis - COWLEY, Ann. Mag. nat. Hist., (10) 17: 515. subnodalis - FRASER, Trans. R. ent. Soc. London, 96: 23 (Ivory Coast). subnodalis - FRASER, Bull. Inst, franc. Afr. noire, 13: 1088 (redescription). subnodalis - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 107.

Distribution: Central Africa. Genus: Nososticta

SELYS

I860 Alloneura (Nososticta) SELYS, Bull. Acad. r. Belg., (2) 10: 456. Type species: Nososticta solida SELYS, 1860 (monotypy). - 1913 Notoneura TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 430. Type species: Alloneura solitaria TILLYARD, 1906 (original designation). - 1919 Risioneura MUNZ, Mem. Amer. Ent. Soc., 3: 66. Type species: Alloneura plagiata SELYS, 1886 (original designation). - 1932 Notoneura - LIEFTINCK, Nova Guinea, 15 (5): 35 (revision, key to New Guinean species). - 1 9 3 3 Notoneura - LIEFTINCK, Revue suisse Zool., 40:413 (key to Australian species). - 1934 Notoneura - COWLEY, Entomologist, 67: 203. - 1934 Risioneura - COWLEY, Entomologist, 67: 203 (proposed synonymy with Notoneura TILLYARD, 1913). - 1936 Notoneura - COWLEY, Ann. Mag. nat. Hist., (10) 17: 521 (list of species). - 1937 Notoneura LIEFTINCK, Nova Guinea (N.S.) 1: 292 (revision, key to New Guinean species). - 1954 Notoneura - LIEFTINCK, Treubia, 22 Suppl.: 44. - 1 9 8 4 Notoneura - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111 (as "incl." with Nososticta SELYS, 1860).

africana

(SCHMIDT)

1944 Notoneura africana SCHMIDT, Mitt, dtsch. ent. Ges., 12: 43. Type male: Mus. Naturk., Berlin; type-locality: Erithrea. 1962 Notoneura africana - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 108. 1984 Nososticta africana - D AVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:111 (Solomon Islands?)

Distribution: Ethiopia. astrolabica

(FÖRSTER)

comb. nov.

1898 Caconeura finisterrae subrasse astrolabica FÖRSTER, Termeszetr. Fiiz., Budapest, 21: 299. Type male: unknown; type-locality: New Guinea. 1903 Caconeura torrenticola FÖRSTER, Ann. Mus. nat. hung., Budapest, 1: 552. Type male: Termeszett. Μύζ.; Budapest; type-locality: New Guinea. 1932 Notoneura torrenticola - LIEFTINCK, Nova Guinea, 15 (5): 48. 1937 Notoneura astrolabica - LIEFTINCK, N o v a Guinea (N.S.) 1: 552.

Distribution: New Guinea. atrocyana

LIEFTINCK

1984 Notosticta atrocyana LIEFTINCK, 1960 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the

World, Utrecht, 1: 111.

Distribution: New Guinea. aurantiaca

(LIEFTINCK)

comb. nov.

1938 Notoneura aurantiaca LIEFTINCK, Nova Guinea (N.S.) 2: 94. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea.

430 1984 Caconeura plagiata Ris, 1915 (nec SELYS, 1886) - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: New Guinea. baroalba

W A T S O N & THEISCHINGER

1984 Nososticta baroalba WATSON & THEISCHINGER, 1984 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: N o r t h Australia. beatrix

(LIEFTINCK)

comb. nov.

1949 Notoneura beatrix LIEFTINCK, Nova Guinea (N.S.) 5: 63. Type male: Lieftinck's Collection; type-locality: New Guinea.

Distribution: New Guinea. callisphaena

(LIEFTINCK)

comb. nov.

1937 Notoneura callisphaena LIEFTINCK, Nova Guinea (N.S.) 1: 298. Type male: Rijksmus. Natuurl. hist.; Leiden; type-locality: New Guinea.

Distribution: New Guinea. chalybeostoma

(LIEFTINCK)

comb. nov.

1932 Notoneura chalybeostoma LIEFTINCK, Nova Guinea, 15 (5): 44. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. circumscripta

(SELYS)

1886 Alloneura circumscripta SELYS, Mem. Cour. Acad. Belg., 38:190. Type male: R. Mus. Brussels; type-locality: Moluccas. 1890 Caconeura circumscripta - KIRBY, Syn. Cat. Neur.-Odon., London: 135. 1937 Notoneura circumscripta - LIEFTINCK, N o v a Guinea (N.S.) 1: 301 (redescription) (New Guinea). 1 9 8 4 Nososticta circumscripta - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 1 1 .

Distribution: New Guinea. coelestina

(TILLYARD)

1906 Alloneura coelestina TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 184. Type male: not given; type-locality: Queensland. 1913 Notoneura coelestina - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 432. 1984 Nososticta coelestina - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: Australia. commutata

(LIEFTINCK)

1900 Caconeura tain). 1913 Caconeura 1938 Notoneura Ris, 1900, 1949 Notoneura

comb. nov.

exul Ris (nec SELVS, 1886), Arch. f. Naturg., 66: 194 (misidentification) (New Briexul - Ris, Abh. Senckenb. Naturf. Ges., 34: 509. commutata LIEFTINCK, Nova Guinea (N.S.) 2: 90 (new name for Caconeura a junior secondary homonymy of Caconeura exul SELYS, 1 8 8 6 ) . commutata - LIEFTINCK, Treubia, 20 (2): 335 (Bismarck Archipelago).

Distribution: Bismarck Archipelago.

exul

431

cyanura (LIEFTINCK) 1932 Notoneura cyanura LIEFTINCK, Nova Guinea, 15 (5): 45. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea. 1937 Notoneura cyanura - LIEFTINCK, Nova Guinea (N.S.) 1: 293. 1984 Nososticta cyanura - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: New Guinea. diadesma (LIEFTINCK) comb. nov. 1936 Notoneura diadesma LIEFTINCK, Revue suisse Zool., 43 (5): 120. Type male: Buitenzorg Mus.; type-locality: Soemba. 1953 Notoneura diadesma - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 152 (redescription).

Distribution: Sumba. dorsonigra (MARTIN) 1903 Alloneura dorsonigra MARTIN, Bull. Mus. nat. hist., Paris, 8: 508. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: New Guinea. 1984 Nososticta dorsonigra - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 111.

Distribution: New Guinea. egregia (LIEFTINCK) comb. nov. Caconeura eburnea Ris (nec FÖRSTER, 1 8 9 7 ) , Treubia, 7: Suppl.: 1 4 4 . Type and locality: not given. 1937 Notoneura egregia LIEFTINCK, Nova Guinea (N.S.) 1: 306. Type female: Rijksmus. Natuurl. Hist. Leiden; type-locality: New Guinea (new name for Caconeura eburnea Ris, 1929, a junior secondary homonymy of Caconeura eburnea FÖRSTER, 1 8 9 7 ) . 1929

Distribution: New Guinea. emphyla (LIEFTINCK) comb. nov. 1936 Notoneura emphyla LIEFTINCK, Revue suisse Zool., 43 (5): 118. Type male: Buitenzorg Mus.; type-locality: Flores. 1953 Notoneura emphyla - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 151 (redescription) (Flores).

Distribution: Flores. erythroprocta (SELYS) comb. nov. 1886 Alloneura erythroprocta SELYS, Mem. Cour. Acad. Belg., 38:182. Type male: R. Mus. Brussels; type-locality: Waigiou. 1890 Caconeura erythroprocta - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1937 Notoneura erythroprocta - LIEFTINCK, Nova Guinea (N.S.) 1: 292 (redescription).

Distribution: Waigeu. erythrura erythrura (LIEFTINCK) comb. nov. 1932 Notoneura erythrura LIEFTINCK, Nova Guinea, 15 (5): 46. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea, erythrura efasciata (LIEFTINCK) comb. nov. 1949 Notoneura erythrura efasciata LIEFTINCK, Nova Guinea (N.S.) 5: 73. Type male: not given; type-locality: New Guinea.

Distribution: New Guinea.

432 evelynae

(LIEFTINCK)

comb. nov.

1960 Notoneura evelynae LIEFTINCK, N o v a Guinea (N.S.) 1: 120. Type male: Buitenzorg Mus.; type-locality: New Guinea.

Distribution: New Guinea, exul

(SELYS)

comb. nov.

1886 Alloneura exul SELYS (nec Ris, 1900), Mem. Cour. Acad. Belg., 38: 185. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Moluccas. 1890 Caconeura exul - KIRBY, Syn. Cat. Neur.-Odon., London: 135. 1938 Notoneura exul - LIEFTINCK, N o v a Guinea (N.S.) 2: 89 (redescription).

Distribution: Moluccas. finisterroe

(FÖRSTER)

1897 Caconeura finisterroe FÖRSTER, Ann. Soc. ent. Belg., 51: 41. Type male: Brussels Mus.; typelocality: New Guinea. 1 9 8 4 Nososticta finisterroe - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 1 2 .

Distribution: New Guinea. flavipennis

(SELYS)

1886 Alloneura flavipennis SELYS, Mem. Cour. Acad. Belg., 38: 188. Type male: Selys's Collection; type-locality: Celebes. 1890 Caconeura flavipennis - KIRBY, Syn. Cat. Neur.-Odon., London: 135. 1896 Caconeura flavipennis - FÖRSTER, Ent. Nachr., 22: 324 (description of female). 1 9 8 4 Nososticta flavipennis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 1 2 .

Distribution: Celebes, fonticola

(LIEFTINCK)

comb. nov.

1932 Notoneura fonticola LIEFTINCK, N o v a Guinea, 15 (5): 49. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea. 1953 Notoneura fonticola - LIEFTINCK, Treubia, 21: 674.

Distribution: New Guinea. fraterna

(LIEFTINCK)

comb. nov.

1933 Notoneura fraterna LIEFTINCK, Revue suisse Zool., 40: 410. Type male: not given; type-locality: Australia.

Distribution: Australia. insignis

(SELYS)

comb. nov.

1886 Alloneura insignis SELYS, Mem. Cour. Acad. Belg., 38: 181. Type male: Brussels Mus.; typelocality: Sumatra. 1890 Caconeura insignis - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1898 Alloneura fruhstorferi KRÜGER, Stettiner ent. Ztg., 59: 138. Type male: Mus. Naturk., Berlin; type-locality: Java. 1926 Caconeura fruhstorferi - FRASER, Treubia, 8: 493. 1930 Risioneura fruhstorferi - LIEFTINCK, Treubia, 12: 147 (Java, Sumatra). 1934 Notoneura insignis - LIEFTINCK, Treubia, 14 (4): 392 (Java). 1953 Notoneura insignis - LIEFTINCK, Verh. naturf. Ges. Basel, 64: 150 (Sumatra, Java, Bali). 1954 Notoneura insignis - LIEFTINCK, Treubia, 22 Suppl.: 44 (Java, Bali, Sumatra).

433 Alloneura fruhstorferi

1954

-

LIEFTINCK,

Treubia,

22

Suppl.:

44

[proposed synonymy with insignis

(SELYS, 1 8 8 6 ) ] .

Distribution: Sumatra, Java, Bali. irene

(LIEFTINCK)

comb. nov.

1949 Notoneura irene LIEFTINCK, N o v a Guinea (N.S.) 5: 64. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. kalumburu

WATSON & THEISCHINGER - loc. cit.

D A V I E S & TOBIN,

The Dra-

1984 Nososticta koolpinyah WATSON & THEISCHINGER, 1984 - loc. cit. gonflies of the World, Utrecht, 1: 112.

D A V I E S & TOBIN,

The Dra-

D A V I E S & TOBIN,

The Dra-

1984

Nososticta kalumburu WATSON & THEISCHINGER, gonflies of the World, Utrecht, 1: 112.

1984

Distribution: North Australia. koolpinyah

WATSON & THEISCHINGER

Distribution: North Australia. koongarra

WATSON & THEISCHINGER

1984 Nososticta koongarra WATSON & THEISCHINGER, 1984 - loc. cit. gonflies of the World, Utrecht, 1: 112.

Distribution: North Australia. liveringa

WATSON & THEISCHINGER

1984 Nososticta liveringa WATSON & THEISCHINGER, 1984 - loc. cit. of the World, Utrecht, 1: 112.

D A V I E S & TOBIN,

The Dragonflies

Distribution: North-West Australia. lorentzi

(LIEFTINCK)

comb. nov.

1938 Notoneura lorentzi LIEFTINCK, Nova Guinea (N.S.) Leiden; type-locality: New Guinea.

2:

91. Type male: Rijksmus. Natuurl. hist.,

Distribution: New Guinea. marina (Ris) comb. nov. 1913 Caconeura marina Ris, Abh. Senckenb. Ges., 34: 513. Type male: Mus. Naturk., Berlin; typelocality: Aru.

Distribution: Aru. melanoxantha 1949

(LIEFTINCK)

Notoneura melanoxantha locality: New Guinea.

comb. nov.

LIEFTINCK,

Nova Guinea (N.S.)

5: 6 8 .

Type male: unknown; type-

Distribution: New Guinea. moluccensis

(SELYS)

1886 Alloneura moluccensis SELYS, Mem. Cour. Acad. Belg., 38: 186. Type male: Selys's Collection; type-locality: Amboina.

434 1890 Caconeura moluccensis - KIRBY, Syn. Cat. Neur.-Odon., London: 135. 1953 Notoneura moluccensis - LIEFTINCK, Treubia, 21: 672. 1984 Nososticta moluccensis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 112.

Distribution: Malaysia. nigrifrons (Ris) comb. nov. 1913 Caconeura nigrifrons Ris, Abh. Senckenb. Ges., 34: 513. Type male: Mus. Naturk., Berlin; type-locality: New Guinea.

Distribution: New Guinea, nigrofasciata (LIEFTINCK) comb. nov.

1932 Notoneura nigrofasciata LIEFTINCK, Nova Guinea, 15 (5): 53. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea. 1949 Notoneura nigrofasciata - LIEFTINCK, Treubia, 20 (2): 336.

Distribution: New Guinea. phoenissa (Ris) comb. nov. 1929 Caconeura phoenissa Ris, Treubia, Suppl. 7: 143. Type male: Mus. Naturk., Berlin; type-locality: Buru.

Distribution: Buru. pilbara WATSON 1984 Nososticta pilbara WATSON, 1969 - loc. cit. DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 112.

Distribution: West Australia. plagiata (SELYS) comb. nov. 1886 Alloneura plagiata SELYS (nec Ris, 1915), Mem. Cour. Acad. Belg., 38:184. Type male: Brussels Mus.; type-locality: Mysol. 1890 Caconeura plagiata - KIRBY, Syn. Cat. Neur.-Odon., London: 135. 1916 Caconeura xanthomelaena Ris, Nova Guinea, 13 (2): 91. Type male: Mus. Naturk., Berlin; type-locality: New Guinea. 1919 Risioneura plagiata - MUNZ, Mem. Am. ent. Soc., 3: 66 (as generotype of Risioneura MUNZ, 1919).

1937 1938 1949 1949

Notoneura plagiata - LIEFTINCK, Nova Guinea (N.S.) 1: 296 (redescription) (New Guinea). Notoneura plagiata - LIEFTINCK, Nova Guinea (N.S.) 2: 95. Notoneura plagiata - LIEFTINCK, Nova Guinea (N.S.) 5: 68. Caconeura xanthomelaena - LIEFTINCK, Nova Guinea (N.S.) 5: 68 [proposed synonymy with plagiata

(SELYS, 1886)].

1953 Notoneura plagiata - LIEFTINCK, Treubia, 21: 676.

Distribution: New Guinea. plagioxantha (LIEFTINCK) comb. nov. 1932 Notoneura plagioxantha LIEFTINCK, Nova Guinea, 15 (5): 54. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea. Distribution: N e w G u i n e a .

435 pseudexul (Ris) comb. nov. 1913 Caconeura pseudexul Ris, Abh. Senckenb. Ges., 34: 513. Type male: Mus. Naturk., Berlin; type-locality: New Guinea.

Distribution: New Guinea. pyroprocta

(LIEFTINCK)

comb. nov.

I960 Notoneura pyroprocta LIEFTINCK, Nova Guinea (N.S.) 10: 122. Type male: Lieftinck's Collection; type-locality: New Guinea.

Distribution: New Guinea. rangifera

(LIEFTINCK)

comb. nov.

1949 Notoneura rangifera LIEFTINCK, Nova Guinea (N.S.) 5: 70. Type male: Rijksmus. Natuurl. hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. rosea rosea (Ris) comb. nov. 1913 Caconeura rosea Ris, Abh. Senckenb. Ges., 34: 513. Type male: Ris's Collection; type-locality: Aru. 1915 Caconeura rosea - Ris, Nova Guinea, 13 (2): 90. 1937 Notoneura rosea rosea - LIEFTINCK, Nova Guinea (N.S.) 1: 300.

Distribution: Aru. rosea cruentata

(LIEFTINCK)

comb. nov.

1932 Notoneura cruentata LIEFTINCK, Nova Guinea, 15 (5): 56. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea. 1937 Notoneura rosea cruentata - LIEFTINCK, Nova Guinea (N.S.) 1: 300.

Distribution: New Guinea. salomonis salomonis (SELYS) comb. nov. 1886 Alloneura salomonis SELYS, Mem. Cour. Acad. Belg., 38: 189. Type male: Selys's Collection; type-locality: Solomon Islands. 1890 Caconeura salomonis - KIRBY, Syn. Cat. Neur.-Odon., London: 135. 1932 Notoneura salomonis - LIEFTINCK, Nova Guinea, 15 (5): 51 (New Britain, New Guinea, Solomon Islands: Bougainville). 1949 Notoneura salomonis - LIEFTINCK, Treubia, 20 (2): 336 (Guadalcanal, Shortland Island).

Distribution: Oceania. salomonis eburnea

(FÖRSTER)

comb. nov.

1897 Caconeura eburnea FÖRSTER (nec Ris, 1929), Ent. Nachr. 23 (22): 336. Type male: Ann Arbor Mus.; type-locality: New Guinea. 1919 Risioneura eburnea - MUNZ, Mem. Amer. ent. Soc., 3: 20. 1926 Caconeura salomonis geminata FRASER, Treubia, 8 (3-4): 492. Type male: Sieber's Collection; type-locality: Key Island. 1937 Notoneura eburnea - LIEFTINCK, Nova Guinea (N.S.) 1: 304.

Distribution: New Guinea, Key Island.

436 selysi

(FÖRSTER)

1896 Caconeura selysi FÖRSTER, Ann. Soc. ent. Belg., 40: 423. Type male: Mus. Naturk., Berlin; type-locality: Sumba. 1930 Risioneura selysi - LIEFTINCK, Treubia, 12: 143 (Sumba, Flores). 1936 Notoneura selysi - LIEFTINCK, Revue suisse Zool., 43 (5): 117. 1984 Nososticta selysi - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:113 (Solomon Islands).

Distribution: Sumba, Solomon Islands. silvicola

(LIEFTINCK)

comb. nov.

1949 Notoneura silvicola LIEFTINCK, N o v a Guinea (N.S.) 5: 72. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. solida

(HÄGEN)

I860 Alloneura (Nososticta) solida HAGEN, in SELYS, Bull. Acad. r. Belg., (2) 10: 457. Type male: Mus. Naturk., Berlin; type-locality: Australia. 1890 Nososticta solida - KIRBY, Syn. Cat. Neur.-Odon., London: 135 (as generotype of Nososticta SELYS, 1 8 6 0 ) .

1901 Nososticta 1913 Nososticta

solida - MARTIN, Mem. Soc. zool. France, 14: 244 (South Australia, Victoria). solida - TILL YARD, Proc. Linn. Soc. N.S. Wales, 37: 429 (redescription, auctor:

SELYS).

1917 Nososticta

solida - SJÖSTEDT, Ark. Zool. Uppsala, 11 (11): 29 (Queensland).

Distribution: Australia. solitaria

(TILLYARD)

comb. nov.

1906 Alloneura solitaria TILLYARD, Proc. Linn. Soc. N.S. Wales, 31: 182. Type male: Tillyard's Collection; type-locality: Queensland. 1913 Notoneura solitaria - TILLYARD, Proc. Linn. Soc. N.S. Wales, 37: 431 (as generotype of Notoneura TILLYARD 1 9 1 3 ) . 1934 Notoneura solitaria - COWLEY, Entomologist, 67: 203.

Distribution: Australia. taracumbi

WATSON & THEISCHINGER

1984 Nososticta taracumbi WATSON & THEISCHINGER, 1984 - loc. cit. D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 113.

Distribution: N o r t h Australia. thalassina

(LIEFTINCK)

comb. nov.

1949 Notoneura thalassina LIEFTINCK, N o v a Guinea (N.S.) 5: 72. Type male: unknown; type-locality: New Guinea.

Distribution: New Guinea. wallacei

(SELYS)

comb. nov.

1886 Alloneura wallacei type-locality: New 1890 Caconeura wallacei 1932 Notoneura wallacii

SELYS, Mem. Cour. Acad. Belg., 38: 183. Type male: R. Mus. Brussels; Guinea. - KIRBY, Syn. Cat. Neur.-Odon., London: 135. - LIEFTINCK, Nova Guinea (N.S.) 1: 294 (redescription).

Distribution: New Guinea.

437 xanthe

(LIEFTINCK)

comb. nov.

1938 Notoneura xanthe LIEFTINCK, Nova Guinea (N.S.) 2: 93. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: New Guinea.

Distribution: New Guinea. Genus Prodasineura 1886

COWLEY

Alloneura SELYS, (nec SELYS, 1 8 6 0 ) , Mem. Cour. Acad. Belg., 3 8 : 1 7 9 . Type species: Alloneura gracillima SELYS, 1 8 8 6 (monotypy). - 1 8 7 1 Brachyneura SELYS, Trans, ent. Soc. London, 1871: 415 (nom.nud.). - 1 9 3 4 Prodasineura COWLEY, Entomologist, 67:202 (new name for Alloneura SELYS, 1 8 8 6 , a junior primary homonymy of Alloneura SELYS, 1 8 6 0 ) . Type species: Alloneura (Alloneura) dorsalis SELYS, 1 8 6 0 (original designation). - 1 9 3 5 Prodasineura - C O W L E Y , Entomologist's month. Mag., 71: 15. - 1936 Prodasineura - COWLEY, Ann. Mag. nat. Hist., (10) 17: 519 (list of species). - 1933 Cycloneura FRASER, F r a u n a brit. India, Odon., 1: 212. (nom.nud.). - 1948 Prodasineura - LIEFTINCK, Treubia, 19 (2): 229 (key to species). - 1954 Prodasineura - LIEFTINCK, Treubia, 22 Suppl.: 38. - 1962 Prodasineura - PINHEY, Publ. cult. Comp. Diam. Angola, 59:107. - 1 9 6 2 Brachyneura - PINHEY, Publ. cult. Comp. Diam. Angola, 5 9 : 1 0 7 (proposed synonymy with Prodasineura C O W L E Y , 1 9 3 4 ) . - 1 9 6 2 Cycloneura - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 1 0 7 (proposed synonymy with Prodasineura C O W L E Y , 1 9 3 4 ) . - 1 9 8 4 Alloneura SELYS, 1 8 8 6 - D A VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 1 3 (as „incl." with Prodasineura C O W L E Y , 1 9 3 4 ) .

abbreviata

LIEFTINCK

1951 Prodasineura abbreviata LIEFTINCK, Idea, 8: 76. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1954 Prodasineura

abbreviata - LIEFTINCK, Treubia, 22 Suppl.: 38.

Distribution: Borneo. autumnalis

(FRASER)

1922 Caconeura autumnalis FRASER, Mem. Dept. Agr. India, 7: 43. Type male: British Museum (Nat. Hist.), London; type-locality: Shillong. 1930 Caconeura corrina LIEFTINCK, Treubia, 12: 138. Type male: Buitenzorg Mus.; type-locality: Java. 1930 Indoneura dolorosa NEEDHAM, Zool. Sinica, (A) 11: 284. Type male: unknown; type-locality: China. 1934 Caconeura corrina - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 336. 1934 Caconeura autumnalis - LIEFTINCK, Treubia, 14 (4): 390. 1934 Caconeura corrina - LIEFTINCK, Treubia, 14 (4): 390 [proposed synonymy with autumnalis (FRASER, 1 9 2 2 ) ] .

1946 Caconeura autumnalis gandawricus SAHNI, Agra Univ. J. Res. (Sei.), 13: 84. Type male: Univ. Agra; type-locality: India. 1954 Prodasineura autumnalis - LIEFTINCK, Treubia, 22 Suppl.: 38 (Sumatra, Java). 1965 Prodasineura dolorosa - ASAHINA, Kontyü, 33 (4): 497 (Hong-Kong). 1966 Prodasineura autumnalis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent., 18: 181. 1989 Caconeura autumnalis gandawricus - HÄMÄLÄINEN, Odonatologica, 18 (1): 17 [proposed synonymy with autumnalis (FRASER, 1 9 2 2 ) ] .

Distribution: Oriental region, collaris

(SELYS)

I860 Alloneura (Alloneura) collaris SELYS, Bull. Acad. r. Belg., (2) 10: 455. Type male: Selys's Collection; type-locality: Malacca.

438 1890 1907 1915 1922 1926 1927 1935 1936 1954 1966

Disparoneura collaris - KIRBY, Syn. Cat. Neur.-Odon., London: 134 (Malacca, Borneo). Disparoneura notostigma collaris - FÖRSTER, in LAIDLAW, Malayenses Zool., 4: 13. Disparoneura dohrni Ris, Tijdsch. Ent., 58: 9. Type female: unknown; type-locality: Simalur. Caconeurabotti FRASER, Mem. Dept. Agr. India, 7:41. Type male: British Museum (Nat. Hist.), London; type-locality: Burma. Caconeura collaris dohrni - LAIDLAW, J. Mal. Br. R. Asiatic Soc., 4: 230. Disparoneura dohrni - Ris, Zool. Meded., Leiden, 10 (1): 21 (Sumatra). Caconeura collaris - LIEFTINCK, Misc. Zool. Sum., 92-93: 8. Prodasineura dohrni - COWLEY, Ann. Mag. nat. Hist., (10) 17: 521. Prodasineura collaris - LIEFTINCK, Treubia, 22 Suppl.: 39 (Siam, Sumatra, Borneo). Caconeura botti - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 183 [as syn. with collaris (SELYS, I860)].

1984 Prodasineura collaris f. dohrni - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1:114.

Distribution: East Asia, coerulescens

(FRASER)

1932 Caconeura coerulescens FRASER, J. Siam Soc. nat. Hist., Suppl., 8: 289. Type male: British Museum (Nat. Hist.), London; type-locality: Siam. 1966 Prodasineura coerulescens - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 186.

Distribution: Thailand. croconota (Ris) comb. nov. 1916 Disparoneura croconota Ris, Suppl. ent. Berlin, 5:18. Type male: Ris's Collection; type-locality: Formosa.

Distribution: Taiwan. delicatula

(LIEFTINCK)

1930 Caconeura delicatula LIEFTINCK, Treubia, 12: 141. Type male: Buitenzorg Mus.; type-locality: Java. 1934 Caconeura delicatula - LIEFTINCK, Treubia, 14 (4): 391 (Java). 1946 Caconeura dorsalis delicatula - FRASER, Entomologist's month. Mag., 82: 201. 1948 Prodasineura delicatula - LIEFTINCK, Treubia, 19 (2): 223 (Java). 1954 Prodasineura delicatula - LIEFTINCK, Treubia, 22 Suppl.: 39 (Java).

Distribution: Java. dorsalis

(SELYS)

I860 Alloneura (Alloneura) dorsalis SELYS, Bull. Acad. r. Belg., (2) 10: 456. Type male: R. Mus. Brussels; type-locality: Borneo. 1890 Caconeura dorsalis - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1934 Prodasineura dorsalis - COWLEY, Entomologist, 67: 202 (as generotype of Prosasineura COWLEY, 1934).

1946 Caconeura dorsalis dorsalis - FRASER, Entomologist's month. Mag., 82: 201. 1948 Prodasineura dorsalis - LIEFTINCK, Treubia, 19 (2): 224 (redescription) (Borneo). 1954 Prodasineura dorsalis - LIEFTINCK, Treubia, 22 Suppl.: 40 (Borneo).

Distribution: Borneo. flammula

LIEFTINCK

1948 Prodasineura flammula LIEFTINCK, Treubia, 19 (2): 227. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: B o r n e o .

439 1954 Prodasineura flammula - LIEFTINCK, Treubia, 22 Suppl.: 40 (Borneo).

Distribution: Borneo. flavifacies

PINHEY

1984 Prodasineura flavifacies PINHEY, 1981 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the W o r l d ,

Utrecht, 1: 114.

Distribution: Zambia. gracillima (SELYS) comb. nov. 1886 Alloneura gracillima SELYS, Mem. Cour. Acad. Belg., 38: 179. Type male: Selys's Collection; type-locality: Celebes (as generotype of Alloneura SELYS, 1 8 8 6 ) . 1890 Caconeura gracillima - KIRBY, Syn. Cat. Neur.-Odon., London: 134.

Distribution: Celebes. haematosoma LIEFTINCK 1937 Prodasineura haematosoma LIEFTINCK, Treubia, 16 (1): 84. Type male: Buitenzorg Mus.; typelocality: Borneo. 1954 Prodasineura haematosoma - LIEFTINCK, Treubia, 22 Suppl.: 40 (Borneo).

Distribution: Borneo. hosei (LAIDLAW) 1913 Disparoneura hosei LAIDLAW, Proc. zool. Soc. London, 1913: 78. Type male: Laidlaw's Collection; type-locality: Borneo. 1936 Disparoneura hosei - KIMMINS, J. Fed. Mai. States Mus., 18: 93. 1948 Prodasineura hosei - LIEFTINCK, Treubia, 19 (2): 231 (redescription) (Borneo). 1954 Prodasineura hosei - LIEFTINCK, Treubia, 22 Suppl.: 40 (Borneo).

Distribution: Borneo. hyperythra (SELYS) 1886 Alloneura hyperythra SELYS, Mem. Cour. Acad. Belg., 38: 180. Type male: Selys's Collection; type-locality: Labuan. 1890 Caconeura hyperythra - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1912 Disparoneura moultoni LAIDLAW, J. Str. Br. R.Asiatic Soc., 63: 98. Type male: Laidlaw's Collection; type-locality: Borneo. 1913 Disparoneura hyperythra - LAIDLAW, Proc. zool. Soc. London, 1913: 76. 1920 Caconeura moultoni - LAIDLAW, Proc. zool. Soc. London, 1920: 340. 1 9 3 6 Prodasineura hyperythra - KIMMINS, J . Fed. Mai. States Mus., 18: 9 4 . 1954 Prodasineura hyperythra - LIEFTINCK, Treubia, 22 Suppl.: 41 (Borneo).

Distribution: Borneo, incerta PINHEY 1962 Prodasineura incerta PINHEY, J. ent. Soc. S.Afr., 25: 25. Type male: unknown; type-locality: Congo.

Distribution: Congo Republic, integra (SELYS) comb. nov. 1882 Alloneura integra SELYS, Ann. Soc. Esp. nat.Hist., 11: 30. Type male: Semper's Collection; type-locality: Mindanao. 1890 Disparoneura integra - KIRBY, Syn. Cat. Neur.-Odon., London: 134.

Distribution: Philippine Islands.

440 interrupta

(SELYS)

I860 Alloneura (Alloneura) interrupta SELYS, Bull. Acad. r. Belg., (2) 10: 453. Type male: Selys's Collection; type-locality: Singapore. 1890 Disparoneura interrupta - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1920 Caconeura interrupta - LAIDLAW, Proc. zool. Soc. London, 1920: 339. 1936 Prodasineura interrupta - COWLEY, Ann. Mag. nat. Hist., (10) 17: 521. 1954 Prodasineura interrupta - LIEFTINCK, Treubia, 22 Suppl.: 41 (Malaya, Sumatra, Borneo).

Distribution: Malaysia, Singapore, Sumatra, Borneo. laidlawi

(FÖRSTER)

1907 Disparoneura notostigma laidlawi FÖRSTER, in LAIDLAW, Fasc. Malayensis Zool., 2: 12. Type male: unknown; type-locality: Malaya. 1931 Caconeura laidlawi - L A I D L A W , J . Fed. Mai. States Mus., 1 6 : 192. 1936 Prodasineura laidlawi - COWLEY, Ann. Mag. nat. Hist., (10) 17: 521. 1942 Prodasineura laidlawi - FRASER, Proc. R. ent. Soc. London, (B) 11: 96 (Malaysia). 1954 Prodasineura laidlawi - LIEFTINCK, Treubia, 22 Suppl.: 41 (Malaya).

Distribution: Malaysia. lansbergei

(SELYS)

1886 Alloneura lansbergei SELYS, Mem. Cour. Acad. Belg., 38: 178. Type male: Selys's Collection; type-locality: Borneo. 1890 Caconeura lansbergei - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1951 Prodasineura lansbergei - LIEFTINCK, Idea, 8: 74. 1954 Prodasineura lansbergei - LIEFTINCK, Treubia, 22 Suppl.: 42 (Borneo).

Distribution: Borneo. longjingensis

ZHON

1984 P r o d a s i n e u r a longjingensis ZHON, 1981 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of t h e

World, Utrecht, 1: 114.

Distribution: China. nigra

(FRASER)

1922 Caconeura nigra FRASER, Mem. Dept. Agr. India, 7: 42. Type male: British Museum (Nat. Hist.), London; type-locality: Burma. 1966 Prodasineura nigra - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 206.

Distribution: Burma. notostigma

(SELYS)

I860 Alloneura (Alloneura) notostigma SELYS, Bull. Acad. r. Belg., (2) 10: 452. Type male: Selys's Collection; type-locality: Singapore. 1890 Disparoneura notostigma - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1907 Disparoneura notostigma notostigma - FÖRSTER, in LAIDLAW, Fasc. Malayensis Zool., 4: 12 (Malaya). 1935 Caconeura notostigma - LIEFTINCK, Misc. Zool.Sum., 9 2 - 9 3 : 8 (Sumatra). 1936 Prodasineura notostigma - COWLEY, Ann. Mag. nat. Hist., (10) 17: 521. 1942 Prodasineura notostigma - FRASER, Proc. R. ent. Soc. London, (B) 11: 96 (Malaysia). 1954 Prodasineura notostigma - LIEFTINCK, Treubia, 22 Suppl.: 42 (Malaya, Sumatra, Borneo).

Distribution: Malaysia, Singapore, Sumatra, Borneo.

441 obsoleta (SELYS) comb. nov. 1882 Alloneura obsoleta SELYS, Ann. Soc. Esp. nat.Hist., 11: 31. Type male: Semper's Collection; type-locality: Philippines. 1890 Dispaeroneura obsoleta - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1939 Caconeura obsoleta - NEEDHAM & G Y G E R , Philipp. J. Sei., 70 (3): 265.

Distribution: Philippine Islands. odoneli (FRASER) 1924 Caconeura o'doneli FRÄSER, J. Bombay nat. Hist. Soc., 30: 51. Type male: British Museum (Nat. Hist.) London; type-locality: Bengal. 1966 Prodasineura odoneli - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 207.

Distribution: India, odzalae AGUESSE 1984

Prodasineura odzalae Utrecht, 1: 114.

AGUESSE, 1 9 6 6

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the World,

Distribution: Congo Republic. palawana LIEFTINCK 1948 Prodasineura palawana LIEFTINCK, Treubia, 19 (2): 232. Type male: R. Mus. Brussels; typelocality: Palawan.

Distribution: Palawan, peramoena (LAIDLAW) 1913 Disparoneura peramoena LAIDLAW, Proc. zool. Soc. London, 1913: 74. Type female: Laidlaw's Collection; type-locality: Borneo. 1936 Prodasineura peramoena - COWLEY, Ann. Mag. nat. Hist., (10) 17: 521. 1954 Prodasineura peramoena - LIEFTINCK, Treubia, 22 Suppl.: 42 (Borneo).

Distribution: Borneo. perisi SART 1964 Prodasineura perisi SART, Bol. Soc. esp. Hist.nat., (B( 62: 19. Type male: Dept. Zool. Suel. Ent.Apl., Madrid; type-locality: West Africa.

Distribution: West Africa. quadristigma LIEFTINCK 1951 Prodasineura quadristigma LIEFTINCK, Idea, 8:80. Type male: Rijksmus. Natuurl. Hist., Leiden; type-locality: Borneo. 1954 Prodasineura quadristigma - LIEFTINCK, Treubia, 22 Suppl.: 42 (Borneo).

Distribution: Borneo. sita (KIRBY) 1894 Disparoneura sita KIRBY, J. Linn. Soc., London, 24: 563. Type male: British Museum (Nat. Hist.), London; type-locality: Ceylon. 1917 Caconeura sita - LAIDLAW, Ree. Indian Mus., 13: 348. 1919 Caconeura canningi FRÄSER, J. Bombay Soc. nat.Hist., 26: 877. Type male: British Museum (Nat. Hist.), London; type-locality: Nilgiris. 1924 Caconeura canningi - FRÄSER, Ree. Indian Mus., 26: 504 (West India). 1955 Prodasineura sita - LIEFTINCK, Zool. Meded., Leiden, 34 (5): 73 (Ceylon).

442 1966 Caconeura canningi - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 185 [proposed synonymy with sita ( K I R B Y , 1 8 9 4 ) ] .

Distribution: Sri Lanka, India. tenebricosa

LIEFTINCK

1937 Prodasineura tenebricosa LIEFTINCK, Treubia, 16 (1): 82. Type male: Buitenzorg Mus.; typelocality: Borneo. 1954 Prodasineura tenebricosa - LIEFTINCK, Treubia, 22 Suppl.: 43 (Borneo).

Distribution: Borneo. theebawi

(FRÄSER)

comb. nov.

1922 Caconeura theebawi FRASER, Mem. Dept. Agr. India, 7: 40. Type male: British Museum (Nat. Hist.), London; type-locality: Burma.

Distribution: Burma. verticalis verticalis

(SELYS)

1860 AUoneura (Alloneura) verticalis SELYS, Bull. Acad. r. Belg., (2) 10: 453. Type male: Selys's Collection; type-locality: Borneo. 1890 Disparoneura verticalis - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1920 Caconeura verticalis - LAIDLAW, Proc. zool. Soc. London, 1920: 339. 1933 Caconeura verticalis - FRASER, F r a u n a brit. India, Odon., 1: 212. 1935 Caconeura verticalis - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 9. 1936 Prodasineura verticalis - COWLEY, Ann. Mag. nat. Hist., (10) 17: 520. 1954 Prodasineura verticalis - LIEFTINCK, Treubia, 22 Suppl.: 43 (Siam, Malaya, Sumatra, Java, Borneo). 1964 Caconeura verticalis - SCHMIDT, Ent. Tidschr., 85: 147 (Burma).

Distribution: Oriental region. verticalis andamanensis

(FRASER)

comb. nov.

1924 Caconeura verticalis andamanensis FRASER, Ree. Indian Mus., 26: 142. Type: not given; typelocality: A n d a m a n s (emandation without description).

Distribution: Andaman Island. verticalis annadalei

(FRASER)

comb. nov.

Caconeura annandalei FRASER, J . Bombay nat. Hist.Soc., 2 7 : 4 5 3 . Type male: British Museum (Nat. Hist.), London; type-locality: Mahableshwar. 1924 Caconeura verticalis annandalei - FRASER, Ree. Indian Mus., 26: 503 (West India). 1921

Distribution: India. verticalis burmanensis

(FRASER)

1933 Caconeura verticalis burmanensis FRASER, F a u n a brit. India, Odon., 1: 216. Type male: British Museum (Nat. Hist.), London; type-locality: Burma. 1966 Prodasineura verticalis burmanensis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.) 18: 184.

Distribution: Burma. verticalis delia

(KARSCH)

1891 Dispaeroneura delia KARSCH, Ent. Nachr., 17 (16): 242. Type female: Mus. Naturk., Berlin; type-locality: Sumatra.

443 1898 Disparoneura arba KRÜGER, Stettiner ent. Ztg, 59: 112. Type male: Mus. Naturk., Berlin; type-locality: Sumatra. 1926 Caconeura karnyi LAIDLAW, J. Mal. Br. R. Asiatic Soc., 4: 219. Type male: Laidlaw's Collection; type-locality: Mentawei Islands. 1927 Disparoneura delia arba - Ris, Zool. Meded., Leiden, 10 (1): 20 (sumatra). 1931 Caconeura delia delia - LIEFTINCK, Misc. Zool. Sum., 59: 4 (Sumatra). 1934 Disparoneura delia delia - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 337. 1934 Disparoneura arba - SCHMIDT, Arch. Hydrobiol. Suppl., 13: 338 [proposed synonymy with verticalis

delia

(KARSCH, 1891)].

1934 Disparoneura risi SCHMIDT, Arch. Hydrobiol. Suppl., 13: 338. Type male: Senckenberg Mus.; type-locality: Sumatra. 1935 Caconeura verticalis delia - LIEFTINCK, Misc. Zool. Sum., 9 2 - 9 3 : 9. 1948 Prodasineura verticalis delia - LIEFTINCK, Treubia, 19: 284. 1954 Prodasineura verticalis delia- LIEFTINCK, Treubia, 22 Suppl.: 43 [as syn. with verticalis verticalis (SELYS, I 8 6 0 ) ] .

Distribution: Sumatra. verticalis humeralis

(SELYS)

comb. nov.

I860 Alloneura (AUoneura) humeralis SELYS, Bull. Acad. r. Belg., (2) 10: 454. Type male: Selys's Collection; type-locality: Malacca. 1890 Disparoneura humeralis - KIRBY, Syn. Cat. Neur.-Odon., London: 134. 1930 Caconeura humeralis - LIEFTINCK, Treubia, 12: 151. 1934 Caconeura humeralis - LIEFTINCK, Treubia, 14 (4): 391 (Java). 1954 Caconeura humeralis - LIEFTINCK, Treubia, 22 Suppl.: 43 [as syn. with verticalis verticalis (SELYS, I 8 6 0 ) ] .

Distribution: Malaysia, Java. villiersi

FRASER

1948 Prodasineura villiersi FRÄSER, Proc. R. ent. Soc. L o n d o n , (B) 17: 7. Type male: Inst. fr. d'Afr., Dakar; type-locality: Ivory Coast. 1962 Prodasineura

villiersi - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 107.

Distribution: Ivory Coast. vittata

(SELYS)

1886 Disparoneura vittata SELYS, Mem. Cour. Acad. r. Belg., 38: 174. Type: not designated, cotype male: British Museum (Nat. Hist.), London; type-locality: West Africa. 1907 Disparoneura agrioniformis MARTIN, Mem. Soc. Espan. Madrid, 1: 430. Type male: R. Mus. Brussels; type-locality: Guinea. 1934 Prodasineura vittata - COWLEY, Entomologist's month. Mag., 70: 202. 1962 Prodasineura vittata - PINHEY, Publ. cult. Comp. Diam. Angola, 59: 107.

Distribution: West Africa. Subfamily Protoneurinae CuP extends to level of vein descending from subnodus (or slightly distal to this). 1905 Protoneurinae JACOBSON & BIANKI, Pryam. lozhnos. Ross, imp.: 836. Type-genus: not designated. - 1919 Protoneurinae - MUNZ, Mem. Amer. ent. Soc., 3: 63 (key to genera: Chlorocnemis, Proneura, Disparoneura, Peristicta, Neoneura, Idioneura, Palaemnema, Platysticta, Hypostrophoneura, Protosticta, Microneura, Protoneura, Epipleoneura, Epipotoneura, Phasmoneura, Psaironeura, Neosticta, Austrosticta, Notoneura, Oristicta, Nososticta, Isosticta, Risioneura, Selysioneura). - 1957 Protoneurinae - FRÄSER, R. zool. Soc. N.S. Wales, 1957: 43. Type-genus: Protoneura SELYS, 1857 (with genera: Peristicta, Idioneura, Microneura, Epipleo-

444 neura, Epipotoneura, Phasmoneura, Psaironeura). - 1 9 8 4 Protoneurinae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1:115 (with genera: Epipleoneura, Epipotoneura, Idioneura, Junix, Microneura, Neoneura, Peristicta, Phasmoneura, Proneura, Protoneura, Psaironeura, Roppaneura).

Genus Epipleoneura

WILLIAMSON

1915 Epipleoneura WILLIAMSON, Proc. U.S. natn. Mus., 48: 618. Type species: Epipleoneura WILLIAMSON, 1915 (original designation).

albuquerquei 1984

lamina

MACHADO

Epipleoneura albuquerquei M A C H A D O , the World, Utrecht, 1: 115.

1964

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of

Distribution: Brazil. ephippigera

(SELYS)

1886 Protoneura ephippigera SELYS, Mem. Cour. Acad. Belg., 38: 208. Type male: Selys's Collection; type-locality: Pebas. 1890 Protoneura ephippigera - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1943 Epipleoneura ephippigera - SCHMIDT, Beitr. F a u n a Perus, 2: 232 (Peru).

Distribution: Peru. fernandezi

RÄCENIS

1960 Epipleoneura fernandezi RACENIS, Acta biol. venezuel., 3: 28. Type male: unknown; type-locality: Venezuela.

Distribution: Venezuela. fuscaenea

WILLIAMSON

1915 Epipleoneura fuscaenea WILLIAMSON, Proc. U.S. natn. Mus., 48: 628. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. haroldoi

SANTOS

1964 Epipleoneura haroldoi SANTOS, Atas Soc. Biol., Rio de Janeiro, 8: 42. Type male: Rio de Janeiro Mus.; type-locality: Brazil.

Distribution: Brazil. humeralis

(SELYS)

1886 Protoeura humeralis SELYS, Mem. Cour. Acad. Belg., 38: 209. Type female: Selys's collection; type-locality: Coary. 1917 Protoneura humeralis - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 26 (description of male from Amazonas). 1 9 8 4 Epipleoneura humeralis - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 1 1 5 (Brazil).

Distribution: Brazil. incusa

WILLIAMSON

1915 Epipleoneura incusa WILLIAMSON, Proc. U.S. natn. Mus., 48: 267. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana.

445 lamina

WILLIAMSON

1915 Epipleoneura lamina WILLIAMSON, Proc. U.S. natn. Mus., 48: 625 (as generotype of Epipleoneura WILLIAMSON, 1915). Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. machadoi

RÄCENIS

I960 Epipleoneura machadoi RÄCENIS, Acta biol. venezuel., 3: 37. Type male: unknown; type-locality: Brazil.

Distribution: Brazil. manauensis

SANTOS

1964 Epipleoneura manauensis SANTOS, Atas Soc. Biol., Rio de Janeiro, 8: 40. Type male: Cienc. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. metallica

RÄCENIS

1955 Epipleoneura metallica RÄCENIS, Mem. Soc. Cienc. nat. La Dalle, 15: 56. Type male: unknown; type-locality: Venezuela.

Distribution: Venezuela. pallida

RÄCENIS

I960 Epipleoneura pallida RÄCENIS, Acta biol. venezuel., 3: 32. Type male: unknown; type-locality: Venezuela.

Distribution: Venezuela. pereirai 1984

MACHADO

Epipleoneura pereirai M A C H A D O , World, Utrecht, 1: 115.

1964

- loc. cit.

DAVIES & TOBIN,

The Dragonflies of the

Distribution: Brazil. spatulata

RÄCENIS

I960 Epipleoneura spatulata RÄCENIS, Acta biol. venezuel., 3: 34. Type male: unknown; type-locality: Venezuela.

Distribution: Venezuela. venezuelensis

RÄCENIS

1955 Epipleoneura venezuelensis RÄCENIS, Mem. Soc. Cienc. nat. La Salle, 15: 59. Type male: unknown; type-locality: Venezuela. 1957 Epipleoneura venezuelensis - SANTOS, Rev. bras. Biol., 17: 188 (description of female).

Distribution: Venezuela. williamsoni

SANTOS

1957 Epipleoneura williamsoni SANTOS, Bol. Mus. nac. Rio de Janeiro, Zool., 168: 1. Type male: Cienc. nat. Mus., Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil.

446 Genus Epipotoneura

WILLIAMSON

1915 Epipotoneura WILLIAMSON, Proc. U.S. natn. Mus., 48: 618. Type species: Epipotoneura lennia WILLIAMSON, 1915 (original designation).

nehalennia

neha-

WILLIAMSON

1915 Epipotoneura nehalennia WILLIAMSON, Proc. U.S. natn. Mus., 48: 630 (as generotype of Epipotoneura WILLIAMSON, 1915). Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. Genus Idioneura

SELYS

1860 Protoneura (Idioneura) SELYS, Bull. Acad. r. Belg., (2) 10: 458. Type species: Idioneura

ancilla

SELYS, 1860 ( m o n o t y p y ) .

ancilla

SELYS

I860 Protoneura (Idioneura) ancilla (HOFFMANNSEGG MS) Selys, Bull. Acad. r. Belg., (2) 10: 458 (as generotype of Idioneura SELYS, 1 8 6 0 ) . Type male: Mus. Naturk., Berlin; type-locality: Bahia. 1890 Idioneura ancilla - KIRBY, Syn. Cat. Neur.-Odon., London: 135.

Distribution: Brazil. Genus Junix

RACENIS

1968 Junix RACENIS, Publ. ocas. Mus. Ci. nat. Caracas (Zool.), 1 1 : 2 . Type species: Junix RACENIS, 1 9 6 8 (original designation).

elumbis

elumbis

RACENIS

1968 Junix elumbis RACENIS, Publ. ocas. Mus. Ci. nat. Caracas (Zool.), 11: 2 (as generotype of Junix RACENIS, 1968). Type male: Caracas Mus.; type-locality: Venezuela.

Distribution: Venezuela. Genus Microneura 1886

SELYS

Microneura SELYS, Mem. Cour. Acad. Belg., 38: 2 0 6 . Type species: Microneura caligata SELYS, 1886 (monotypy). - 1910 Microneura - MUTTKOWSKI, Bull. Publ. Mus. Millwaukee, 1 (1): 73. - 1919 Protoneura (Microneura) - CALVERT, Trans. Am. ent. Soc., 45: 349.

caligata

SELYS

1886 Microneura caligata SELYS, Mem. Cour. Acad. Belg., 38: 206 (as generotype of Microneura SELYS, 1886). Type male: Poey's Collection; type-locality: Cuba. 1919 Protoneura (Microneura) caligata - CALVERT, Trans. Am. ent. Soc., 45: 349 (Cuba).

Distribution: Cuba. Genus Neoneura I860

SELYS

Neoneura SELYS, Bull. Acad. r. Belg., (2) 10: 4 5 9 . Type species: Protoneura (Neoneura) rubriventris SELYS, 1 8 6 0 (original designation). - 1 8 9 0 Caenoneura K I R B Y , Syn. Cat. Neur.-Odon., London: 1 3 6 . Type species: Neoneura bilinearis SELYS, 1 8 6 0 (original designation) (new name for Neoneura SELYS, 1 8 6 0 ) . - 1 9 1 0 Neoneura - M U T T K O W S K I , Bull. Publ. Mus. Milwaukee, 1 (1): 72. - 1910 Caenoneura - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 72 (proposed synonymy with Neoneura S E L Y S , 1 8 6 0 ) . - 1 9 1 7 Neoneura - WILLIAMSON, Trans. Am. ent. Soc.,

447 43: 211 (revision, key to species). - 1919 Protoneura Soc., 4 5 : 350.

aaroni

(Neoneura)

- CALVERT, Trans. Am. ent.

CALVERT

1903 Neoneura aaroni CALVERT, Biol. Centr.-Amer., London: 139. Type male: Acad. Nat. Sei., Philadelphia; type-locality: Texas. 1917 Neoneura aaroni - WILLIAMSON, Trans. Am. ent. Soc., 43: 241.

Distribution: U.S.A. amelia

CALVERT

1903 Neoneura amelia CALVERT, Biol. Centr.-Amer., London: 138. Type male: Champion's Collection; type-locality: Mexico. 1916 Neoneura amelia - Ris, Arch. Naturg., (A) 82: 101 (Mexico, Guatemala). 1917 Neoneura amelia - WILLIAMSON, Trans. A m . ent. Soc., 43: 239 (redescription) (Guatemala, Costa Rica).

Distribution: Central America. bilinearis

SELYS

I860 Protoneura (Neoneura) bilinearis SELYS, Bull. Acad. r. Belg., (2) 10: 460. Type male: Selys's Collection; type-locality: Para. 1890 Caenoneura bilinearis - KIRBY, Syn. Cat. Neur.-Odon., London: 136 (as generotype of Caenoneura K I R B Y , 1 8 9 0 ) . 1917 Neoneura bilinearis - WILLIAMSON, Trans. Am. ent. Soc., 43: 219 (redescription) (Brazil, British Guiana).

Distribution: Brazil, British Guyana. carnatica

SELYS

1886 Protoneura (Neoneura) carnatica KIRBY, Mem. Cour. Acad. Belg., 38: 200. Type male: Hagen's Collection; type-locality: Cuba. 1890 Caenoneura carnatica - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1910 Neoneura carnatica - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 72. 1917 Neoneura carnatica - WILLIAMSON, Trans. Am. ent. Soc., 43: 236. 1919 Protoneura (Neoneura) carnatica - CALVERT, Trans. Am. Ent. Soc., 45: 350 (Cuba).

Distribution: Cuba. cristina

RÄCENIS

1955 Neoneura cristina RACENIS, Mem. Soc. Cienc. nat. La Salle, 15: 49. Type male: unknown; type-locality: Venezuela.

Distribution: Venezuela. denticulata

WILLIAMSON

1917 Neoneura denticulata WILLIAMSON, Trans. Am. ent. Soc., 43: 227. Type male: Mus. Comp. Zool., Cambridge; type-locality: Peru. 1917 Neoneura pardalina SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 24. Type male: Naturhist. Riksmus., Stockholm; type-locality: Amazonas. 1943 Neoneura denticulata - SCHMIDT, Beitr. F a u n a Perus, 2: 231. 1946 Neoneura denticulata - FRASER, Trans. R. ent. Soc. London, 96: 43 (Peru). 1946 Neoneura pardalina - FRASER, Trans. R. ent. Soc. London, 96: 43 (proposed synonymy with denticulata

WILLIAMSON, 1 9 1 7 ) .

448 1961 Neoneura denticulata - CALVERT, Proc. Acad. nat. Sei., Philadelphia, 113: 10 (redescription) (Peru).

Distribution: Peru. esthera

WILLIAMSON

1917 Neoneura esthera WILLIAMSON, Trans. Am. ent. Soc., 43: 232. Type male: Acad. Sei., Philadelphia; type-locality: Costa Rica. 1932 Neoneura esthera - GEIJSKES, Zool. Meded., Leiden, 14: 238 (Trinidad).

Distribution: Central America. ethela

WILLIAMSON

1917 Neoneura ethela WILLIAMSON, Trans. Am. ent. Soc. 43: 222. Type male: Acad. Sei., Philadelphia; type-locality: Brazil.

Distribution: Brazil. fulvicollis

SELYS

1886 Protoneura (Neoneura) fulvicollis SELYS, Mem. Cour. Acad. Belg., 38: 202. Type male: Selys's Collection; type-locality: Para. 1890 Caenoneura fulvicollis - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1917 Neoneura fulvicollis - WILLIAMSON, Trans. Am. ent. Soc., 43: 241.

Distribution: Brazil. gaida

RÄCENIS

1953 Neoneura gaida RÄCENIS, Acta biol. venezuel., 1:155. Type male: Caracas Mus.; type-locality: Venezuela.

Distribution: Venezuela. joana

WILLIAMSON

1917 Neoneura joana WILLIAMSON, Trans. Am. ent. Soc., 43: 215. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. maria

(SCUDDER)

1866 Agrion maria SCUDDER, Proc. Boston Soc. nat. Hist., 10: 188. Type female: Scudder's Collection; type-locality: Isle de Pino (Cuba). 1886 Protoneura (Neoneura) maria - SELYS, Mem. Cour. Acad. Belg., 38: 199. 1867 Agrion palustris HAGEN, Proc. Boston Soc. nat.Hist., 11: 190. Type: unknown; type-locality: Cuba. 1890 Caenoneura maria - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1910 Neoneura maria - MUTTKOWSKI, Publ. Mus. Milwaukee, 1 (1): 72. 1910 Agrion palustris - MUTTKOWSKI, Publ. Mus. Milwaukee, 1 (1): 72 [proposed synonymy with maria

(SCUDDER, 1866)].

1917 Neoneura maria - WILLIAMSON, Trans. Am. ent. Soc., 43: 236 (redescription) (Cuba). 1919 Protoneura (Neoneura) maria - CALVERT, Trans. Am. ent. Soc., 45: 350 (Cuba).

Distribution: Cuba. mariana

WILLIAMSON

1917 Neoneura mariana WILLIAMSON, Trans. Am. ent. Soc., 43: 224. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana.

449 myrthea

WILLIAMSON

1917 Neoneura myrthea WILLIAMSON, Trans. Am. ent. Soc., 43: 228. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. paya

CALVERT /

1907 Neoneura paya CALVERT, Biol. Centr.-Amer., London: 393. Type male: Calvert's Collection; type-locality: Guatemala. 1917 Neoneura paya - WILLIAMSON, Trans. Am. ent. Soc., 43: 241 (Guatemala).

Distribution: Guatemala. rubriventris

SELYS

1860 Protoneura (Neoneura) rubriventris (in original description: rurriventris) SELYS, Bull. Acad. r. Belg., (2) 10: 459 (as generotype of Neoneura SELYS, 1860). Type male: Selys's Collection; type-locality: Para. 1890 Caenoneura rubriventris - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1917 Neuneura rubriventris - WILLIAMSON, Trans. Am. ent. Soc., 43: 230 (redescription) (Brazil, British Guiana). 1955 Neoneura rubriventris - RACENIS, Mem. Soc. Cienc. nat. La Salle, 15: 53 (as sp.nov., perhaps new for Venezuela).

Distribution: Brazil, British Guyana, Venezuela. rufithorax

SELYS

1886 Protoneura (Neoneura) rufithorax SELYS, Mem. Cour. Acad. Belg., 38: 205. Type male: Selys's Collection; type-locality: Säo Paulo. 1890 Caenoneura rufithorax - KIRBY, Syn. Cat. Neur.-Odon., London: 136.

Distribution: Brazil. schreiberi

MACHADO

1975 Neoneura schreiberi MACHADO, Ciencia Cult., S. Paulo, 27 (7): 764. Type male: Cien. Mus. nat. Hist., Säo Paulo; type-locality: Brazil.

Distribution: Brazil. sylvatica

SELYS

1886 Protoneura (Neoneura) sylvatica SELYS, Mem. Cour. Acad. Belg., 38: 204. Type male: Selys's Collection; type-locality: Rio de Janeiro. 1890 Caenoneura sylvatica - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1917 Neoneura sylvatica - WILLIAMSON, Trans. Am. ent. Soc., 43: 222. 1955 Neoneura sylvatica - RACENIS, Mem. Soc. Cienc. nat. La Salle, 15: 52 (as sp.nov., perhaps new for Venezuela).

Distribution: Brazil, Venezuela. waltheri

SELYS

1886 Protoneura (Neoneura) waltheri SELYS, Mem. Cour. Acad. Belg., 38: 198. Type male: Selys's Collection; type-locality: Entre Rios. 1890 Caenoneura waltheri - KIRBY, Syn. Cat. Neur.-Odon., London: 136.

Distribution: Brazil.

450 Genus Peristicta

SELYS

I860 Alloneura (Peristicta) SELYS, Bull. Acad. r. Belg., (2) 10: 442. Type species: Alloneura (Peristicta) forceps SELYS, 1860 (monotypy). - 1890 Peristicta - KIRBY, Syn. Cat. Neur.-Odon., L o n d o n : 135.

aeneoviridis

CALVERT

1909 Peristicta aeneoviridis CALVERT, A n n . Carnegie Mus., Pittsburgh, 6: 124. Type male: Carnegie Mus.; type-locality: Paraguay.

Distribution: Paraguay. forceps

SELYS

I860 Alloneura (Peristicta) forceps (HAGEN in SELYS), Bull. Acad. r. Belg., (2) 10: 442 (as generotype of Peristicta SELYS, 1860). Type male: H a g e n ' s Collection; type-locality: Brazil. 1890 Peristicta forceps - KIRBY, Syn. Cat. Neur.-Odon., L o n d o n : 135.

Distribution: Brazil. gauchae

SANTOS

1968 Peristicta gauchae SANTOS, A t a s Soc. Biol. R i o de Janeiro, 12: 79. Type male: Cienc. Mus. Rio de Janeiro; type-locality: Brazil.

Distribution: Brazil. lizeria

NAVÄS

1920 Peristicta lizeria NAVÄS, An. Soc. Cienc. Argent., 90: 45. Type male: u n k n o w n .

Distribution: Argentina. missionera

JURZITZA

1 9 8 4 Peristicta missionera JURZITZA, 1981 - l o c . cit. D A VIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 116.

Distribution: Argentina. Genus Phasmoneura

WILLIAMSON

1916 Phasmoneura WILLIAMSON, Ent. News, 27: 30. Type species: Phasmoneura 1916 (original designation).

ciganae

o/tfryra WILLIAMSON,

SANTOS

1 9 8 4 P h a s m o n e u r a c i g a n a e SANTOS, 1968 - l o c . cit. DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 116.

Distribution: Argentina. itatiaiae

SANTOS

1 9 8 4 P h a s m o n e u r a itatiaiae SANTOS, 1970 - loc. cit. DAVIES & TOBIN, T h e D r a g o n f l i e s o f t h e W o r l d ,

Utrecht, 1: 116.

Distribution: Brazil. olmyra

WILLIAMSON

1916 Phasmoneura olmyra WILLIAMSON, Ent. News, 27: 31 (as generotype of Phasmoneura LIAMSON, 1916). Type male: Williamson's Collection; type-locality: G u i a n a .

Distribution: Guyana.

WIL-

451 Genus Proneura

SELYS

1889 Proneura SELYS, C.R. ent. Soc. Belg., 33: 172. Type species: Proneura prolongata (monotypy).

prolongata

SELYS, 1889

SELYS

1889 Proneura prolongata SELYS, C.R. ent. Soc. Belg., 33: clxxiv (as generotype of Proneura SELYS, 1889). Type male: Selys's Collection; type-locality: Pebas (Amazonas). 1890 Proneura prolongata - KIRBY, Syn. Cat. Neur.-Odon., London: 186. 1943 Proneura prolongata - SCHMIDT, F a u n a Perus, 2: 232 (Ecuador, Peru).

Distribution: Brazil, Ecuador, Peru. Genus Protoneura

SELYS

1857 Protoneura SELYS, Sagra, Hist. Ins. Cuba: 470. Type species: Agrion capillaris RAMBUR, 1842 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 136). - 1860 Protoneura (Protoneura) - SELYS, Bull. Acad. r. Belg., (2) 10: 457 (with subgenera Idioneura, Neoneura, Protoneura). - 1915 Protoneura - WILLIAMSON, Proc. U.S. natn. Mus., 48: 616 (revision, key to species). - 1919 Protoneura - CALVERT, Trans. Am. ent. Soc., 91: 169 (with subgenera Protoneura s.str., Microneura, Neoneura). - 1941 Protoneura - COWLEY, Trans. R. ent. Soc. London, 45: 349 (revision, key to species). - 1988 Protoneura - MICHALSKI, Univ. West Indies Dep. Occ. Pap., 6: 51 (key to Trinidad species).

ailsa

DONNELLY

1961 Protoneura ailsa DONNELLY, Florida Ent., 44: 119. Type male: Donnelly's Collection; typelocality: St. Lucia.

Distribution: St. Lucia. amatoria

CALVERT

1907 Protoneura amatoria CALVERT, Biol. Centr.-Amer. Neur., London: 395. Type male: British Museum (Nat. Hist.), London; type-locality: Guatemala. 1915 Protoneura amatoria - WILLIAMSON, Proc. U.S. natn. Mus., 48: 623 (Guatemala, Honduras). 1932 Protoneura amatoria - GEIJSKES, Zool. Meded., Leiden 14: 238 (Trinidad).

Distribution: Central America. aurantiaca

SELYS

1886 Protoneura aurantiaca SELYS, Mem. Cour. Acad. Belg., 38: 213. Type male: R. Mus. Brussels; type-locality: Panama. 1915 Protoneura aurantiaca - WILLIAMSON, Proc. U.S. natn. Mus., 48: 624 (Guatemala).

Distribution: Central America. bifurcata

SJÖSTEDT

1917 Protoneura bifurcata SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 28. Type male: Rijksmus. Naturhist. Stockholm; type-locality: Amazonas. 1 9 8 4 Protoneura

bifurcata

"SJÖSTEDT, 1 9 1 8 2: 2 8 " - l o c . c i t . D A VIES & TOBIN, T h e D r a g o n f l i e s o f

the World, Utrecht, 1: 116.

Distribution: Brazil.

452 calverti

WILLIAMSON

1915 Protoneura calverti WILLIAMSON, Proc. U.S. natn. Mus., 48: 620. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana, capillaris

(RAMBUR)

1842 Agrion capillare RAMBUR, Hist. nat. Ins. Neuropt., Paris: 280. Type male: unknown; typelocality: not given. 1857 Protoneura capillaris - SELYS, Sagra, Hist. Ins. Cuba: 471 (as generotype of Protoneura SELYS, 1857). 1860 Protoneura (Protoneura) capillaris - SELYS, Bull. Acad. r. Belg., (2) 10: 461 (Cuba). 1890 Protoneura capillaris - KIRBY, Syn. Cat. Neur.-Odon., London: 136. 1910 Protoneura capillaris - MUTTKOWSKI, Publ. Mus. Milwaukee, 1 (1): 73 (type in Coll. Selys). 1915 Protoneura capillaris - WILLIAMSON, Proc. U.S. natn. Mus., 48: 625 (Cuba). 1919 Protoneura (Protoneura) capillaris - CALVERT, Trans. Am. ent. Soc., 45: 349 (Cuba).

Distribution: Cuba. capilliformis

SELYS

1886 Protoneura capilliformis SELYS, Mem. Cour. Acad. Belg., 38:211. Type male: Selys's Collection; type-locality: Santarem.

Distribution: Santarem. cara

CALVERT

1903 Protoneura cara CALVERT, Biol. Centr.-Amer., Neur., London: 142. Type male: British Museum (Nat. Hist.), London; type-locality: Central America. 1915 Protoneura cara - WILLIAMSON, Proc. U.S. natn. Mus., 48: 624 (Guatemala).

Distribution: Central America. corculum

CALVERT

1907 Protoneura corculum CALVERT, Biol. Centr.-Amer., Neur., London: 396. Type male: British Museum (Nat. Hist.), London; type-locality: Guatemala. 1915 Protoneura corculum - WILLIAMSON, Proc. U.S. natn. Mus., 48: 623 (Guatemala).

Distribution: Guatemala. cupida

CALVERT

1903 Protoneura cupida CALVERT, Biol. Centr.-Amer., Neur., London: 142. Type male: unknown; type-locality: Central America. 1915 Protoneura cupida - WILLIAMSON, Proc. U.S. natn. Mus., 48: 624 (Guatemala).

Distribution: Guatemala. exigua

SELYS

1886 Protoneura exigua SELYS, Mem. Cour. Acad. Belg., 38: 216. Type male: R. Mus. Brussels; type-locality: Amazonas. 1915 Protoneura exigua - WILLIAMSON, Proc. U.S. natn. Mus., 48: 619. 1943 Protoneura exigua - SCHMIDT, F a u n a Perus, 2: 232 (Peru). Distribution: Brazil, P e r u .

453 klugi

COWLEY

1941 Protoneura klugi COWLEY, Trans. R. ent. Soc. London, 91: 145. Type male: British Museum (Nat. Hist.), London; type-locality: Peru.

Distribution: Peru. macintyrei

KENNEDY

1939 Protoneura macintyrei KENNEDY, Pap. Occ. Mus. Zool. Univ. Mich., 410: 2. Type male: Ann Arbor Mus.; type-locality: Ecuador.

Distribution: Ecuador. paucinervis

SELYS

1886 Protoneura paucinervis SELYS, Mem. Cour. Acad. Belg., 38: 214. Type male: Selys's Collection; type-locality: Pebas. 1917 Protoneura paucinervis - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 29 (Rio Antaz). 1943 Protoneura paucinervis - SCHMIDT, Beitr. Fauna Perus, 2: 232 (Peru).

Distribution: Brazil, Peru. peramans 1984

CALVERT

Protoneura peramans CALVERT, Utrecht, 1: 117.

1902

- loc. cit.

D A VIES & TOBIN,

The Dragonflies of the World,

Distribution: Guatemala. peruviensis

FRASER

1946 Protoneura peruviensis FRASER, Trans. R. ent. Soc. London, 97: 459. Type male: Leeds Mus.; type-locality: Peru. 1966 Protoneura peruviensis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 209 (holotype in British Museum (Nat. Hist.), London).

Distribution: Peru. protostictoides

FRASER

1946 Protoneura protostictoides FRASER, Trans. R. ent. Soc. London, 97: 458. Type male: Leeds Mus.; type-locality: Peru. 1966 Protoneura proctostictoides - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 210 (holotype in British Museum (Nat. Hist.), London).

Distribution: Peru. scintilla

GLOYD

1939 Protoneura scintilla GLOYD, Occ. Pap. Mus. Zool. Univ. Mich., 408:1. Type male: Ann Arbor Mus.; type-locality: Brazil.

Distribution: Brazil. tenuis

SELYS

I860 Protoneura (Protoneura) tenuis (DALE MS) SELYS, Bull. Acad. r. Belg., (2) 10: 461. Type male: Dale's Collection; type-locality: Para. 1890 Protoneura tenuis - KIRBY, Syn. Cat. Neur.-Odon., London: 136.

Distribution: Brazil.

454 viridis

WESTFALL

1984 Protoneura viridis WESTFALL, 1964 - loc. cit. DA VIES & TOBIN, T h e D r a g o n f l i e s of the W o r l d ,

Utrecht, 1: 117.

Distribution: Virgin Island. woytkowskii 1939

GLOYD

Protoneura woytkowskii G L O Y D , OCC. Pap. Mus. Zool. Univ. Mich., 408: Arbor Mus.; type-locality: Peru.

8.

Type male: Ann

Distribution: Peru. Genus Psaironeura

WILLIAMSON

1915 Psaironeura WILLIAMSON, Proc. U.S. natn. Mus., 48: 618. Type species: Protoneura remissa CALVERT, 1 9 0 3 (original designation).

cerasina

WILLIAMSON

1915 Psaironeura cerasina WILLIAMSON, Proc. U.S. natn. Mus., 48: 629. Type male: Williamson's Collection; type-locality: British Guiana.

Distribution: British Guyana. remissa

(CALVERT)

1903 Protoneura remissa CALVERT, Biol. Centr.-Amer., Neur., London: 144. Type male: British Museum (Nat. Hist.), London; type-locality: Central America. 1915 Psaironeura remissa - WILLIAMSON, Proc. U.S. natn. Mus., 48: 629 (as generotype of Psaironeura WILLIAMSON, 1 9 1 5 ) (Guatemala, Costa Rica).

Distribution: Central America. sancta (HAGEN) comb. nov. Protoneura (Protoneura) sancta H A G E N , in SELYS, Bull. Acad. r. Belg., Hagen's Collection; type-locality: Venezuela. 1890 Protoneura sancta - KIRBY, Syn. Cat. Neur.-Odon., London: 137. I860

(2) 10:462.

Type female:

Distribution: Venezuela. sanguinipes

WESTFALL

1987 Psaironeura sanguinipes WESTFALL, Odonatologica, 16 (1): 94. Type male: Westfall's Collection; type-locality: Dominica.

Distribution: Dominican Republic. tenuissima

(SELYS)

1886 Protoneura tenuissima SELYS, Mem. Cour. Acad. Belg., 38: 217. Type male: R. Mus., Brussels; type-locality: Pebas. 1890 Protoneura tenuissima - KIRBY, Syn. Cat. Neur.-Odon., London: 137. 1915 Psaironeura tenuissima - WILLIAMSON, Proc. U.S. natn. Mus., 48: 620. 1943 Psaironeura tenuissima - SCHMIDT, Beitr. Fauna Perus, 2: 232 (Peru). 1946 Psaironeura tenuissima - FRASER, Trans. R. ent. Soc. London, 97: 460 (Peru).

Distribution: Peru.

455

Genus Roppaneura

SANTOS

1984 Roppaneura SANTOS, 1966 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 117. Type species: Roppaneura beckeri SANTOS, 1966 (original designation).

beckeri

SANTOS

1984 Roppaneura beckeri SANTOS, 1966 - loc. cit. DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 117.

Distribution: Brazil. Family Pseudostigmatidae Very large and slender forms. Pterostigma absent or replaced by pseudostigma which is traversed by crossveins and differs widely between fore and hind wings. Nodus close to wing base; Dc nearly quadrilateral but distal angle slightly acute; postnodals very numerous, intercalated veins present. 1938 Pseudostigmatidae TILLYARD & FRASER, Austr. Zool., 9: 158 (with subfamily Pseudostigmatinae). - 1957 Pseudostigmatidae- FRÄSER, R. Zool. Soc. N.S. Wales, 1957: 50 (with subfamily Pseudostigmatinae). - 1977 Pseudostigmatidae - BELYSHEV & KHARITONOV, Nauka, Novosibirsk: 231 (key to boreal genera). - 1984 Pseudostigmatidae - DAVIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 117.

Subfamily Pseudostigmatinae 1886 Pseudostigmatina SELYS, Mem. Cour. Acad. Belg., 38: 4 (as legion). - 1890 Pseudostigmatina - KIRBY, Syn. Cat. Neur.-Odon., London: 119 (as divisio). - 1903 Anormostigmatini NEEDHAM, Proc. U.S. natn. Mus., 26: 748 (as tribe). - 1910 Pseudostigmatinae - MUTTKOWSKI, Publ. Mus. Bull. Milwaukee, 1 (1): 41 (with genera Pseudostigma, Mecistogaster) (not defined). 1919 Pseudostigmatinae MUNZ, Mem. Amer. Ent. Soc., 3: 20 (first defined). Type-genus: Pseudostigma SELYS, 1860 (with genera Anonisma, Pseudostigma, Micrastigma, Mecistogaster, Megaloprepus). -1938 Pseudostigmatinae - TILL YARD & FRASER, Austr. Zool., 9:158 (with genera Mecistogaster, Pseudostigma, Microstigma, Anonisma, Megaloprepus).

Genus Anonisma

MCLACHLAN

1877 Anonisma MCLACHLAN, Ent. Mag., 14: 87. Type species: Microstigma terminata MCLACHLAN, 1877 (designated by KIRBY, 1890, Syn. Cat. Neur.-Odon., London: 120).

terminatum

(MCLACHLAN)

1877 Microstigma terminata MCLACHLAN, Ent. Mag., 14: 87. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador. 1877 Anonisma abnorma MCLACHLAN, Ent. Mag., 14: 87. Type male: British Museum (Nat. Hist.), London; type-locality: Peru. 1890 Anonisma terminatum - KIRBY, Syn. Cat. Neur.-Odon., London: 120 (as generotype of Anonisma

MCLACHLAN, 1877).

1943 Anonisma terminatum - SCHMIDT, Beitr. Fauna Perus, 2: 230. 1943 Anonisma abnorma - SCHMIDT, Beitr. Fauna Perus, 2: 230 [proposed synonymy with terminatum

(MCLACHLAN, 1877)].

1946 Anonisma terminatum - FRASER, Trans. R. ent. Soc. London, 97: 463 (Colombia, Peru).

Distribution: Ecuador, Peru, Colombia.

456 Genus Mecistogaster RAMBUR 1842 Mecistogaster RAMBUR, Hist. nat. Ins. Neuropt., Paris: 281. Type species: Mecistogaster linearis RAMBUR, 1842 (original designation). - 1860 Mecistogaster (Mecistogaster) - SELYS, Synopsis des Agrionines: 1: 11 (with subgenera Pseudostigma, Mecistogaster). - 1910 Mecistogaster MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 41 (type-species: Libellula lucretia DRURY, 1773). - 1920 Goniostigma KENNEDY, Ohio J. Sei. Columbus, 21: 85. Type species: Agrion amalia BURMEISTER, 1839 (original designation). - 1920 Haplostigma KENNEDY, Ohio J. Sei. Columbus, 21: 84. Type species: Mecistogaster (Mecistogaster) modestus SELYS, 1860 (original designation). - 1920 Platystigma KENNEDY, Ohio J. Sei. Columbus, 21: 84. Type species: Mecistogaster jocaste HAGEN, 1869 (original designation). - 1920 Xanthostigma KENNEDY, Ohio J. Sei. Columbus, 21: 84. Type species: Mecistogaster ornatus RAMBUR, 1842 (original designation).

amalia (BURMEISTER) 1839 Agrion amalia BURMEISTER, H a n d b . Ent., 2: 818. Type male: Mus. Comp. Zool., Cambridge (Mass.); type-locality: Brazil. 1842 Mecistogaster virgatus RAMBUR, Hist. nat. Ins. Neuropt., Paris: 284. Type sex not given, male in Serville's Collection; type-locality: Brazil. 1842 Mecistogaster filiformis RAMBUR, Hist. nat. Ins. Neuropt., Paris: 285. Type female: Serville's Collection; type-locality: Brazil. 1842 Mecistogaster leueostigma RAMBUR, Hist. nat. Ins. Neuropt., Paris: 286. Type female: Marchal's Collection; type-locality: Brazil. 1890 Agrion amalia-ΚΛΚΒΎ, Syn. Cat. Neur.-Odon., London: 121 [incorrect synonymy with lucretia (DRURY, 1773)].

1920 Goniostigma amalia - KENNEDY, Ohio J. Sei. Columbus, 21: 85 (as generotype of Goniostigma KENNEDY, 1 9 2 0 ) .

Distribution: Brazil. amazonica SJÖSTEDT 1917 Mecistogaster amazonicus SJÖSTEDT, Ark. Zool. Uppsala, 11 (15): 29. Type female: Naturhist. Riksmus., Stockholm; type-locality: Amazonas.

Distribution: Brazil. astica SELYS I860 Mecistogaster (Mecistogaster) asticus (BURMEISTER MS) SELYS, Synopsis des Agrionines, 1: 15. Type female: Mus. Naturk., Berlin; type-locality: Brazil. 1890 Mecistogaster asticus - KIRBY, Syn. Cat. Neur.-Odon., London: 120.

Distribution: Brazil. buckleyi MCLACHLAN 1881 Mecistogaster buckleyi MCLACHLAN, Trans, ent. Soc. London, 1881: 32. Type male: British Museum (Nat. Hist.), London; type-locality: Ecuador.

Distribution: Ecuador, garleppi FÖRSTER 1903 Mecistogaster garleppi FÖRSTER, Insekten-Börse, 20: 355. Type male: unknown; type-locality: Ecuador. 1943 Mecistogaster garleppi - SCHMIDT, Beitr. Fauna Perus, 2: 230 (Peru).

Distribution: Ecuador, Peru.

457 jocaste jocaste

HAGEN

1869 Mecistogaster jocaste HAGEN, Stettiner ent. Ztg, 30: 260. Type male: originally in Stettin Mus.; type-locality: Colombia. 1916 Mecistogaster jocaste jocaste - Ris, Arch. Naturg., (A) 82: 72 (Colombia). 1920 Platystigma jocaste - KENNEDY, Ohio J. Sei. Columbus, 21: 84 (as generotype of Platystigma KENNEDY, 1920).

Distribution: Colombia. jocaste sincerus

MCLACHLAN

1877 Mecistogaster sincerus MCLACHLAN, Entomologist's month. Mag., 14: 88. Type male: British Museum (Nat. Hist.), London; type-locality: Peru. 1886 Mecistogaster jocaste race sincerus - SELYS, Bull. Acad. r. Belg., (3) 38: 19. 1890 Mecistogaster sincerus - KIRBY, Syn. Cat. Neur.-Odon. London: 120 (incorrect synonymy with jocaste H Ä G E N , 1 8 6 9 ) . 1920 Mecistogaster syncerus (misprint) - NAVAS, Bol. Soc. Ent. Esp., 3: 92. 1943 Mecistogaster iocaste (misprint) sincerus - SCHMIDT, Beitr. Fauna Perus, 2: 230 (Peru, Colombia).

Distribution: Peru, Colombia, jocaste vicentius Ris 1916 Mecistogaster jocaste vicentius Ris, Arch. Naturg., (A) 82: 72. Type male: not given; typelocality: Colombia. 1946 Mecistogaster

jocaste vicentius - FRASER, Trans. R. ent. Soc. London, 97: 462 (Colombia).

Distribution: Colombia. linearis

(FABRICIUS)

1776 Aegrion linearis FABRICIUS, Gen. Ins.: 249. Type sex not given; lost; type-locality: not given. 1842 Mecistogaster filigerus RAMBUR, Hist. nat. Ins. Neuropt., Paris: 282. Type male: Serville's Collection; type-locality: Surinam. 1842 Mecistogaster linearis -RAMBUR, Hist. nat. Ins. Neuropt., Paris: 282 (Amer.merid., description of male). 1842 Mecistogaster flavistigma RAMBUR, Hist. nat. Ins. Neuropt. Paris: 287. Type female: Mus. Nation. Hist. Nat. Paris; type-locality: not given. 1860 Mecistogaster (Mecistogaster) linearis - SELYS, Synopsis des Agrionines, 1: 16. 1860 Mecistogaster filigerus - SELYS, Synopsis des Agrionines, 1: 16 [proposed synonymy with linearis

(FABRICIUS, 1 7 7 6 ) ] .

1860 Mecistogaster linearis

flavistigma

- SELYS, Synopsis des Agrionines, 1 : 1 6 [proposed synonymy with

(FABRICIUS, 1 7 7 6 ) ] .

1890 Mecistogaster linearis - KIRBY, Syn. Cat. Neur.-Odon., London: 121. 1916 Mecistogaster linearis - Ris, Arch. Naturg., (A) 82: 73 (Colombia, Peru, Guyana, Surinam, Brazil). 1917 Mecistogaster linearis - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 53 (Amazonas). 1920 Mecistogaster linearis - NAVAS, An. Soc. Cienc. Argentina, 90: 92 (Argentina). 1943 Mecistogaster linearis - SCHMIDT, Beitr. F a u n a Perus, 2: 229 (Peru, Colombia, Amazonas, Costa Rica, Bolivia). 1946 Mecistogaster linearis - FRASER, Trans. R. ent. Soc. London, 97: 462 (Colombia, Peru). 1946 Mecistogaster linearis var. infumata FRASER, Trans. R. ent. Soc. London, 97: 462. Male from Colombia. 1 9 4 8 Mecistogaster marchali var. selysia NAVÄS, 1 9 2 3 - CALVERT, Zoologica, 33: 5 5 [incorrect synonymy with linearis (FABRICIUS, 1 7 7 6 ) ] .

Distribution: South America.

458

lucretia

(DRURY)

1773 Libellula lucretia D R U R Y , 111. Ex. Ent., 2: 48. Type female: unknown; type-locality: Brazil. 1 8 4 2 Mecistogaster marchali R A M B U R , Hist. nat. Ins. Neuropt., Paris: 2 8 3 . Type male: Marchal's Collection; type-locality: unknown. 1842 Mecistogaster filum R A M B U R , Hist. nat. Ins. Neuropt., Paris: 284. Type male: Serville's Collection; type-locality: Amer. merid. 1 8 4 2 Mecistogaster pedicillatus R A M B U R , Hist. nat. Ins. Neuropt., Paris: 2 8 4 . Type male: Serville's Collection; type-locality: unknown. 1860 Mecistogaster (Mecistogaster) lucretia - SELYS, Synopsis des Agrionines, 1: 18. 1 8 6 0 Mecistogaster marchali - SELYS, Synopsis des Agrionines, 1: 1 8 [proposed synonymy with lucretia ( D R U R Y , 1 7 7 3 ) ] . 1 8 6 0 Mecistogaster filum - SELYS, Synopsis des Agrionines, 1 : 1 8 [proposed synonymy with lucretia (DRURY,

1773)].

Mecistogaster pedicillatus - SELYS, Synopsis des Agrionines, 1: 1 8 [proposed synonymy with lucretia ( D R U R Y , 1 7 7 3 ) ] . 1886 Mecistogaster hauxwelli SELYS, Mem. Cour. Acad. Belg., 38: 27. (originally as race of marchali from Amazonas). 1890 Mecistogaster lucretia - K I R B Y , Syn. Cat. Neur.-Odon., London: 121. 1943 Goniostigma marchali hauxwelli - S C H M I D T , Beitr. F a u n a Perus, 2: 230 (Peru).

1860

Distribution: South America. modesta

SELYS

I860 Mecistogaster (Mecistogaster) modestus SELYS, Synopsis des Agrionines, 1: 17. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: Mexico. 1886 Mecistogaster iphigenia SELYS, Mem. Cour. Acad.Belg., 38: 22. Type male: R. Mus., Brussels: type-locality: Colombia. 1 8 9 0 Mecistogaster modestus - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 2 0 . 1916 Mecistogaster modestus - Ris, Arch. Naturg., (A) 82: 73 (Mexico, Panama, Colombia, Costa Rica). 1920 Haplostigma modesta - K E N N E D Y , Ohio J. Sei. Columbus, 21: 84 (as generotype of Haplostigma K E N N E D Y , 1920). 1984 Mecistogaster modesta f. iphigenia - D A V I E S & TOBIN, The Dragonflies of the World, Utrecht, 1: 118.

Distribution: Central America, ornata

RAMBUR

1842 Mecistogaster ornatus R A M B U R , Hist. nat. Ins. Neuropt., Paris: 288. Type broken, sex not given; type-locality: not given. 1860 Mecistogaster (Mecistogaster) ornatus - SELYS, Synopsis des Agrionines, 1: 14 (Mexico, Venezuela, Lima, Caracas, Surinam). 1860 Mecistogaster (Mecistogaster) luctuosus SELYS, Synopsis des Agrionines, 1: 14. Type male: Agassiz Mus; type-locality: Puerto-Capello. 1 8 9 0 Mecistogaster ornatus - K I R B Y , Syn. Cat. Neur.-Odon., London: 1 2 0 (Mexico, South America). 1886 Mecistogaster acutipennis SELYS, Mem. Cour. Acad. Belg., 38:18. Type male: R. Mus., Brussels; type-locality: Trinidad. 1916 Mecistogaster ornatus - Ris, Arch. Naturg., (A) 82: 70 (Mexico). 1917 Mecistogaster ornatus - SJÖSTEDT, Ark. Zool. Uppsala, 11 (15): 32 (Amazonas). 1920 Xanthostigma ornatus - K E N N E D Y , Ohio J. Sei. Columbus, 21: 84 (as generotype of Xanthostigma K E N N E D Y , 1920). 1920 Mecistogaster ornatus - N A V Ä S , An. Soc. Cient. Argentina, 90: 35 (Argentina). 1 9 4 3 Mecistogaster ornatus - SCHMIDT, Beitr. Fauna Perus, 2: 2 2 9 (Peru).

459 1946 Mecistogaster ornatus - FRÄSER, Trans. R. ent. Soc. London, 97: 461 (Mexico, Guatemala, Honduras, Nicaragua, Panama, Colombia, Venezuela, Guiana, Brazil, Costa Rica, Peru). 1984 Mecistogaster ornata f. acutipennis - DA VIES & TOBIN, The Dragonflies of the World, Utrecht, 1: 117.

Distribution: South and Central America, pronoti

SJÖSTEDT

1917 Mecistogaster pronoti SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 31. Type female: not given; type-locality: Espirito Santo.

Distribution: Brazil. Genus Megaloprepus

RAMBUR

1842 Megaloprepus RAMBUR, Hist. nat. Ins. Neuropt., Paris: 290. Type species: Libellula caerulatus caerulatus DRURY, 1782 (monotypy).

caerulatus caerulatus

(DRURY)

Libellula caerulata D R U R Y , 111. Ex. Ent., 3 : 5 0 . Type male: unknown; type-locality: Honduras. 1842 Megaloprepus caerulatus - RAMBUR, Hist. nat. Ins. Neuropt., Paris: 290 (as generotype of Megaloprepus RAMBUR, 1842) (Colombia, Honduras). 1860 Megaloprepus caerulatus - SELYS, Synopsis des Agrionines, 1: 7 (Guatemala). 1860 Megaloprepus caerulatus race latipennis SELYS, Synopsis des Agrionines, 1: 7. Male from New Granada and Mexico. 1916 Megaloprepus caerulatus caerulatus - Ris, Arch. Naturg., (A) 82: 65 (Mexico, Costa Rica, Panama). 1782

Distribution: Central America, caerulatus brevistigma

SELYS

I860 Megaloprepus caerulatus race brevistigma SELYS, Synopsis des Agrionines, 1: 7. Male from Bogota. 1916 Megaloprepus caerulatus brevistigma - Ris, Arch. Naturg., (A) 82: 65 (Colombia, Bolivia, Peru, Mexico). 1943 Megaloprepus caerulatus brevistigma - SCHMIDT, Beitr. Fauna Perus, 2: 229 (from Colombia to Bolivia).

Distribution: from Mexico to Bolivia.

Genus Microstigma

RAMBUR

1842 Microstigma RAMBUR, Hist. nat. Ins. Neuropt., Paris: 288. Type species: Microstigma anomalum anomalum R A M B U R , 1 8 4 2 (designated by K I R B Y , 1 8 9 0 , Cyn. Cat. Neur.-Odon., London: 119). - 1860 Microstigma - SELYS, Synopsis des Agrionines, 1: 8 (revision). - 1 9 5 8 Microstigma - SCHMIDT, Mem. Inst. Sei. nat. Belg., (2) 57: 29 (revision, key to Neotropical species).

anomalum anomalum

RAMBUR

1842 Microstigma anomalum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 289. Type male: Mus. Nation. Hist. Nat., Paris; type-locality: not given (as generotype of Microstigma R A M B U R , 1 8 4 2 ) . 1842 Microstigma proximum RAMBUR, Hist. nat. Ins. Neuropt., Paris: 289. Type female: Mus. Nation. Hist. Nat., Paris; type-locality: Cayenne. 1860 Microstigma anomalum - SELYS, Synopsis des Agrionines, 1: 10 (Para, Santarem, Cayenne).

460 I860

Microstigma anomalum

proximum

- SELYS, Synopsis des Agrionines, 1: 1 0 (proposed synonymy with

RAMBUR, 1 8 4 2 ) .

1886 Microstigma minor SELYS, Mem. type-locality: Para. 1890 Microstigma anomalum - KIRBY, 1919 Microstigma anomalum - MÜNZ, 1961 Microstigma anomalum anomalum

Cour. Acad. Belg., 38: 11. Type male: Selys's Collection; Syn. Cat. Neur.-Odon., London: 120. Mem. Amer. Ent. Soc., 3: 49. - CALVERT, Proc. Acad. nat. Sei. Philad., 113: 4 (Peru).

Distribution: South America. anomalum sjoestedti

SCHMIDT

1958 Microstigma anomalum sjöstedti SCHMIDT, Mem. Inst. Sei. nat. Belg., (2) 57: 37. Type male: R. Mus., Brussels; type-locality: Brazil.

Distribution: Brazil. anomalum vicinum

SCHMIDT

1958 Microstigma anomalum vicinum SCHMIDT, Mem. Inst. Sei. nat. Belg., (2) 57: 36. Type male: R. Mus., Brussels type-locality: Brazil.

Distribution: Brazil. calcipennis

FRASER

1946 Microstigma calcipennis FRASER, Trans, ent. R. Soc., London, 97: 463. Type female: Leeds Mus.; type-locality: Bolivia. 1966 Microstigma calcipennis - KIMMINS, Bull. Brit. Mus. nat. Hist. (Ent.), 18: 185 [type female: British Museum (Nat. Hist.), London]. 1948 Microstigma calcipennis - FRASER, Acta zool. lilloana, 5: 65 (Bolivia).

Distribution: Bolivia. maculatum maculatum

HÄGEN

1860 Microstigma maculatum HAGEN, in SELYS, Synopsis des Agrionines, 1:11. Type male: Guerin's Collection; type-locality: Cayenna.

Distribution: Brazil. maculatum albopictum

SCHMIDT

1958 Microstigma maculatum albopictum SCHMIDT, Mem. Inst. Sei. nat. Belg., (2) 57: 33. Type male: R. Mus., Brussels type-locality: Brazil.

Distribution: Brazil. maculatum fassli

SCHMIDT

1958 Microstigma maculatum fassli SCHMIDT, Mem. Inst. Sei. nat. Belg., (2) 57: 33. Type male: R. Mus.; Brussels type-locality: Brazil.

Distribution: Brazil. rotundatum rotundatum

SELYS

I860 Microstigma rotundatum SELYS, Synopsis des Agrionines, 1: 9. Type male: Selys's Collection; type-locality: Amazonas. 1860 Microstigma lunatum SELYS, Synopsis des Agrionines, 1: 10. Type male: Bates's Collection; type-locality: Para. 1886 Microstigma rotundatum race lunatum - SELYS, Mem. Cour. Acad. Belg., 38: 10. 1890 Microstigma rotundatum - KIRBY, Syn. Cat. Neur.-Odon., London: 119 (Peru, Ecuador). 1890 Microstigma

lunatum

- KIRBY, Syn. C a t . N e u r . - O d o n . , L o n d o n : 119 (Brazil).

461 1916 1917 1943 1946

Microstigma Microstigma Microstigma Microstigma

rotundatum rotundatum rotundatum rotundatum

- Ris, Arch. Naturg., (A) 82: 69 (Colombia, Peru). - SJÖSTEDT, Ark. Zool., Uppsala, 11 (15): 33 (Amazonas). rotundatum - SCHMIDT, Beitr. F a u n a Perus, 2: 229. - FRÄSER, Trans. R. ent. Soc. London, 97: 462.

Distribution: Peru, Ecuador, Brazil. rotundatum exustum

SELYS

1860 Microstigma exustum SELYS, Synopsis des Agrionines, 1: 9. Type male: Selys's Collection; type-locality: Peru. 1886 Microstigma rotundatum race exustum - SELYS, Mem. Cour. Acad. Belg., 38: 10. 1890 Microstigma exustum - KIRBY, Syn. Cat. Neur.-Odon., London: 119 (Colombia). 1943 Microstigma rotundatum exustum - SCHMIDT, Beitr. F a u n a Perus, 2: 229 (Peru, Colombia).

Distribution: Peru, Colombia. Genus Pseudostigma

SELYS

I860 Mecistogaster (Pseudostigma) SELYS, Synopsis des Agrionines, 1: 12. Type species: Mecistogaster (Pseudostigma) aberrans SELYS, 1860 (designated by KIRBY, 1890, Syn. Cat. Neur.Odon., London: 121). - 1890 Pseudostigma - KIRBY, Syn. Cat. Neur.-odon., London: 121.

aberrans

SELYS

I860 Mecistogaster (Pseudostigma) aberrans SELYS, Synopsis des Agrionines, 1: 12 (as generotype of Pseudostigma SELYS, 1860). Type female: Selys's Collection; type-locality: Colombia. 1890 Pseudostigma aberrans - KIRBY, Syn. Cat. Neur.-Odon., London: 121. 1910 Pseudostigma aberrans - MUTTKOWSKI, Bull. Publ. Mus. Milwaukee, 1 (1): 41 (Panama, Mexico).

Distribution: Mexico, Panama, Colombia, accendens

SELYS

1860 Mecistogaster (Pseudostigma) accendens SELYS, Synopsis des Agrionines, 1: 15. Type male: Saussure's Collection; type-locality: Mexico. 1861 Pseudostigma accendens - HAGEN, Syn. Neur.N. Amer., Washington: 63.

Distribution: Mexico.

463

Alphabetical Register aaroni, Neoneura 447 abbotti, Chlorocnemis 420 abbotti, Ortholestes, 166 abbreviata, Palaemnema 398 Prodasineura 437 abbreviata, subsp. of biforata, Heliocypha 55 abbreviatum, Pyrrhosoma 243 abercornensis, subs, of steeleae, Aciagrion 242 aberrans, Pseudostigma 461 abessinicum, Enallagma 260 ablutum, Acanthagrion 232 abnorma, Anonisma 455 aborense, Argiocnemis 186 abrogata, Paraphlebia 156 abunae, Acanthagrion 232 abyssinica, Ischnura 264 acaciae, Pseudagrion 323 Acanthagrion 232 Acanthallagma 237 accendens, Pseudostigma 461 Aciagrion 238 aciculare, Aciagrion 238 acicularis, subsp. of posita, Ischnura 274 acostai, Polythore 97 actaeon, Drepanosticta 403 aculeatum, Leptagrion 308 aculeatus, Perissolestes 173 aculeatus aculeatus, Rhipidolestes 175 - yakusimensis, Rhipidolestes 175 acuminata, Mesoleptobasis 288 acuta, Cyanallagma 247 acuta acuta, Elattoneura 423 - lindleyi, Elattoneura 423 acuta, Leptagrion 308 acuticauda, Ischnura 264 acutimargo, subsp. of vittata, Copera 391 acutipennis, Mecistogaster 458 acutipennis, Platycnemis 392 adamantina, Rhinocypha 45 adami, Drepanosticta 403 - , Libellago 59 adamsi, Argia 191

adelais, Coenagrion 219 aderces, Agriocnemis 178 adjuncta, subsp. of derivata, Polythore 98 aduncum, Agrion 256 adusta, subsp. of tincta, Rhinocypha 71 adustum, Acanthagrion 232 adytum, Megalagrion 311 adytum var. tillyardi, Agrion 311 aenea, Argia 192 - , Mnesarete 42 aeneoviridis, Peristicta 450 Aeolagrion 290 aequabilis aequabilis, Calopteryx 9 - californica, Calopteryx 9 - hudsonica, Calopteryx 10 - yakima, Calopteryx 10 aequatoriale, Heteragrion 148 aequatorialis, race of picta, Thore 100 aerides, Teinobasis 350 aeruginosum, Ceriagrion 299 aethiopia, Elattoneura 427 affinis, Micromerus 61 Africalestes 109 Africallagma 248 africana, Nososticta 429 africanum, Aciagrion 238 Africocypha 45 aglae, Agrion 268 aglaia, Selysioneura 366 Agnophilogenia 138 Agriidae 1 Agriinae 8 Agriocnemidinae 178 Agriocnemis 178 Agrioidea 3 agrioides, Amphipteryx 4 Argia 192 Platycnemis 393 Agriomorpha 138 Agrion 9 and 219 Agrionidae 6 and 177 agrioniformis, Disparoneura 443

464 Agrioninae 8 and 214 aguessei, Pseudagrion 323 ailsa, Protoneura 451 alacer, Lestes 110 alata, Libellago 53 alatipes, Platycnemis 393 alba, Disparoneura 426 alberta, Argia 192 albescens, race of griseus, Argiolestes 142 albicauda, Coeliccia 372 - , Episynlestes 103 Indolestes 131 Nesolestes 153 albicolor, Nesolestes 153 albidella, Libellula 396 albifrons, subsp. of ruberrima, Agriocnemis 184 - , Heteragrion 149 albioctopunctata, ab. of pennipes, Platycnemis 396 albistigma, Argia 192 Hetaerina 34 Ischnura 264 albistigma Matsumura, Rhinocypha 57 albistigma Selys, Rhinocypha 66 albofasciatus, Lestes 110 albopictum, subsp. of maculatum, Microstigma 460 alboterminatus, Nesolestes 153 albovittata, Thore 100 albuda, Teinobasis 350 albuquerquei, Epipleoneura 444 alcedo, Palaiargia 211 alcicorne alcicorne, Pseudagrion 324 - digitatum, Pseudagrion 324 alcyone, Cora 94 - , Mortonagrion 189 aldabrensis, subsp. of ochraceus, Lestes 177 aleison, Austrolestes 129 alfurus, Argiolestes 140 Indolestes 131 algira, Ischnura 271 - , Platycnemis 392 alienum, Heteragrion 149 aligulae, Agriocnemis 178 alleni, Indolestes 131 Allocnemis 369 Allolestes 139 Alloneura Selys, 1860 417 Alloneura Selys, 1886 437 Allophaea 77 Allopodagrion 167 allopterum, Anisagrion 296 allopterum race rubicaudum, Anisagrion 296 alluaudi, Paracnemis 384 alluaudi alluaudi, Seychellibasis 349

- berlandi, Seychellibasis 349 alma, Epallage 83 alpinus, Argiolestes 140 alternans, Teinobasis 350 aluensis, Teinobasis 350 amabilis, Amphicnemis 292 Argiolestes 140 - , Vestalis 30 amalia, Mecistogaster 456 amamii, subsp. of ryukyuensis, Coeliccia 378 amanda, subsp. of tincta, Rhinocypha 71 Amanipodagrion 139 amaryllis, Vestalis 31 amasina, subsp. of splendens, Calopteryx 15 amasine, subsp. of fatime, Epallage 83 amata, Calopteryx 10 amata, Lestes 112 amatoria, Protoneura 451 amaurodytum amaurodytum, Megalagrion 311 - fallax, Megalagrion 311 - peles, Megalagrion 311 - wainaeanum, Megalagrion 311 amazon, Agrion 231 amazona amazona, Heliocharis 75 - libera, Heliocharis 75 amazonicum, Acanthagrion 233 amazonica, Hetaerina 34 Mecistogaster 456 Metaleptobasis 288 amazonicum, Heteragrion 149 ambatoroae, Pseudagrion 324 ambigua, Argia 192 ambigua, Libellago 64 ambigua, subsp. of derivata, Polythore 98 ambiguum, Argentagrion 246 Enallagma 248 amboiniensis, Platycypha 63 amelia, Agriocnemis 265 amelia, Neoneura 447 americana, Hetaerina 34 amethystina, Vestalis 31 amicus, Lestes 110 amnicola, Vestalis 31 amoena, Vestalis 31 amoena, subsp. of apicalis, Vestalis 32 amoena, subsp. of gracilis, Vestalis 32 amoenum, Mortonagrion 189 Amorphostigma 243 Amphiagrion 243 Amphiallagma 244 Amphicneminae 290 Amphicnemis 292 amphicyana, Euphaea 84 amphidactylis, Arrhenocnemis 166

465 Amphilestes 163 amphion, Amphiagrion 243 Amphipterygidae 3 Amphipteryginae 3 Amphipteryx 4 Amphisticta 265 amphistylus, Argiolestes 140 ampolomitae, Pseudagrion 324 Anaciagrion 9 anacolosa, Vestalis 31 analis Agrion 227 analis, Austrolestes 129 Elattoneura 423 anambae, subsp. of biseriata, Heliocypha 56 Anapetes, subgen. of Agrion 109 anascephala, Drepanosticta 403 anatolica, subsp. of fatime, Epallage 83 ancilla, race of splendens, Calopteryx 15 ancilla, Calopteryx 10 - , Idioneura 446 andamanensis, subsp. of praemorsus, Lestes 118 andamanensis, Libellago 59 - , subsp. of verticalis, Prodasineura 442 andamanicum, Pseudagrion 324 andersoni, Mnais 20 anderssoni, Neurobasis 23 Andinagrion 244 andinum, Amphiagrion 283 andromache, Leptagrion 308 angelicum, Pseudagrion 324 angelina, Palaemnema 399 angolense, subsp. of caligata, Platycypha 64 - , Pseudagrion 324 angolense rubridorsum, Pseudagrion 333 angolensis angolensis, Agriocnemis 178 - spatulae, Agriocnemis 178 angolicum, Enallagma 257 angularis, Lestes 110 angulatum, Coenagrion 219 angulicollis, Nesobasis 318 angusta, Heliocypha 55 - , Metacnemis 383 angusta oceanis, Rhinocypha 55 angusti, Philogenia 157 angusticlavia, subsp. of nitescens, Teinobasis 354 angustipenne, Heteragrion 149 angustipennis, Calopteryx 10 angustirami, Agriocnemis 179 angydna, Nesolestes 154 Anisagrion 295 Anisoneura 6 Anisophaea 78 Anisopleura 79

anisoptera, Rhinocypha 66 anna, Enallagma 249 annae, Amphicnemis 292 annandalei, Drepanosticta 403 subsp. of rufostigma, Ischnura 276 annandalei, Micromerus 59 annandalei, subsp. of verticalis, Prodasineura 442 annexum, Agrion 252 annulare, Agrion 225 annulata annulata, Copera 389 - stevensi, Copera 389 annulata, Drepanosticta 403 Melanesobasis 316 annulata, Protosticta 416 annulata annulata, Sympecma 136 - braueri, Sympecma 136 - gobica, Sympecma 136 annulatum, Ceroagrion 299 annulipes, Argiolestes 140 annulosum, Ceriagrion 299 annulosus, Austrolestes 129 Anomalagrion 245 anomalum, Agrion 245 anomalum anomalum, Microstigma 459 - sjoestedti, Microstigma 460 - vicinum, Microstigma 460 Anomalura 263 anomalus, Indolestes 131 Anonisma 455 Anormostigmatini 455 antelopoides, Protosticta 413 antennatum, Enallagma 249 Antiagrion 285 antigone, Antiagrion 285 antipodium, race of zelandicum, Xanthagrion 362 Apanisagrion 296 aphrodite, Chlorocypha 48 apicale, Acanthagrion 233 apicale descendens, Acanthagrion 233 apicale, subsp. of dimidiata, Calopteryx 12 Pseudagrion 325 apicalis, Argia 192 Disparoneura 423 - , Neurobasis 23 - , Palaemnema 399 - , Platysticta 413 apicalis, Rhinocypha 55 apicalis, Rhipidolestes 176 apicalis apicalis, Vestalis 31 - nigrescens, Vestalis 32 - submontana, Vestalis 32 apicatus, Rhipidolestes 175 apollinaris, Lestes 110

466 appendiculata, Risiocnemis 385 appendiculatum, Mortonagrion 189 approximans, Aciagrion 239 approximatum, Pseudagrion 325 apricans, Chlorolestes 102 aquisextanum, Agrion 220 Arabicnemis 369 arabicum, subsp. of inconspicuum, Pseudagrion 335 arachne, Megapodagrion 167 arachnoides, Pseudocopera 389 aragonienses, Sympecma 136 aralensis, Ischnura 264 arawakai, Nehalennia 229 arba, Disparoneura 443 arboricola, Selysioneura 366 arcana, Rimanella 101 Archaeallagma 245 Archeopodagrion 139 Archibasis 296 Archboldargia 191 Archiargiolestes 140 Archilestes 108 Archineura 8 arcuata, Coeliccia 372 arcuata, Drepanostigma 403 Argentagrion 245 argentea, Agriocnemis 179 argenteolineatum, Hylaeagrion 291 Argia 191 Argiagrion 298 Argiallagma 229 Argiinae 191 Argiocnemis 185 argiocnemis, Teinobasis 350 Argiolestes 139 Argiolestinae 138 Argyrocnemis 178 argyroides argyroides, Devadatta 4 - tiomanensis, Devadatta 4 aridus, Austrolestes 129 ariel, Ischnura 264 - , Teinobasis 350 aries, Drepanosticta 404 Aristocypha 45 armageddoni, Chlorocypha 65 armatum, Coenagrion 219 armeniacum, Oreagrion 279 armeniacus, Argiolestes 140 armstrongi, Amorphostigma 243 Arrhenocnemis 166 arses, Palaiargia 211 arthuri, Mortonagrion 189 aruanus, Indolestes 132 ascendens, Acanthagrion 233

asclepiades, Libellago 59 asiatica, Ischnura 264 asiatica, Libellago 54 aspasia, Euphaea 84 aspersum, Enallagma 249 assamensis, subsp. of vittata, Copera 391 assamica, Enallagma 242 assamicus, Indolestes 132 assegaii, Pseudagrion 325 asteliae, Agrion 313 Asthenocnemis 369 asticta, var. of macropus, Hetaerina 38 astica, Mecistogaster 456 astrolabica, Nososticta 429 atkinsoni Selys, 1886, Calicnemia 370 atkinsoni Selys, 1891, Calicnemis 371 atkinsoni, Elattoneura 424 atomaria Laidlaw, Copera 391 atomaria Selys, Copera 389 atomarius, Podolestes 160 atra, subsp. of fumipennis, Argia 197 atrata, Calopteryx 10 atrinuchalis, Nehalennia 229 atripes, Risiocnemis 386 atrocyana, Calopteryx 11 - , Nososticta 429 - , Onychargia 211 atrodorsum, subsp. of vacillans, Leptobasis 287 atropha, Vestalis 32 atropurpurea, Risiocnemis 386 atrosanguinea, Dicterias 74 attala, Drepanosticta 404 attenuatum, subsp. of gracile, Aciagrion 240 attenuatus, Perilestes 172 augustana, Argia 192 aulicus, Argiolestes 140 aurantiaca, Agrion 274 aurantiaca, Elattoneura 424 - , Libellago 59 - , Nesobasis 318 - , Nososticta 429 - , Protoneura 451 aurantiacum, subsp. of olivaceum, Ceriagrion 306 - , Heteragrion 149 - , Ceriagrion 299 aurantinigrum, Tigriagrion 283 aurantipes, Platycnemis 393 aurea, Kalocora 92 - , Pachycypha 63 - , Teinobasis 350 aureipennis, Telebasis 359 aurentiacus, Argiolestes 141 aureofrons, Cyanocnemis 379 - , Pseudagrion 325

467 aureus, Argiolestes 141 auriceps, Leptagrion 308 auricolor, Amorphostigma 243 - , subsp. of dorsalis, Elattoneura 425 auriculata Ris, Drepanosticta 409 auriculata Selys, Drepanosticta 404 auriculatum, Papuagrion 320 auripennis, Hetaerina 35 - , Lestes 110 - , Mnais 21 auripes, Platycypha 64 auritum, Ceriagrion 300 auritus, Lestes 110 aurofulgens, Rhinocypha 66 aurora, Hetaerina 35 aurora aurora, Ischnura 265 - viduata, Ischnura 265 aurora, Polythore 97 aurulenta, Rhinocypha 66 austeni, Drepanosticta 404 australasiae, Pseudagrion 325 australis, Agriocnemis 179 - , Argiolestes 141 - , subsp. of disjuntus, Lestes 113 australis australis, Neurobasis 23 - misoolensis, Neurobasis 23 Austroagrion 215 Auslroallagma 246 Austroargiolestes 140 Austrocnemis 187 Austrocoenagrion 216 Austrolestes 129 Austrosticta 363 autazensis, Leptagrion 309 autumnalis, Prodasineura 437 autumnalis gandawricus, Caconeura 437 axinocerca, Coeliccia 372 azteca, subsp. of posita, Ischnura 274 azupizui, Palaemnema 399 azulum, Heteragrion 149 azureum, Aciagrion 239 - , Ceriagrion 300 - , Pseudagrion 326 bacillus, Selysioneura 366 baibarana, Rhynocypha 46 baicalensis, f. of najas, Erythromma 228 bakeri, Ceriagrion 300 balachowskyi, Aciagrion 239 balcanica, subsp. of splendens, Calopteryx 15 balica, subsp. of lara, Allophaea 77 balli, Elattoneura 424 bamptoni, subsp. of crocea, Chlorocypha 49 banksi, Isosticta 364 barbarae, Lestoidea 137

barbarata, Heterophaea 88 barbarus, Lestes 111 barbatula, Drepanosticta 404 barbatum, Coenagrion 218 barberi, Ischnura 265 baroalba, Nososticta 430 barretti, Argia 193 bartelsi, Drepanosticta 404 basale, Oxyagrion 279 basalis, Euphaea 84 basalis, Hetaerina 34 basalis, Lestes 34 basicornu Nielsen, Pseudagrion 326 basicornu Schmidt, Pseudagrion 326 basidens, Enallagma 249 basilactea, Archineura 9 basilanensis, subsp. of podolestoides, Devadatta 5 basilaris, Matrona 19 basilewskyi basilewskyi, Chlorocypha 48 - jacksoni, Chlorocypha 48 basitincta, Dysphaea 82 batesi Navas, Thore 97 batesi Selys, Polythore 97 baueri, Skiallagma 283 Bayadera 80 beadlei, Pseudagrion 326 beata, Polythore 97 beatifica, subsp. of perforata, Heliocypha 57 beatrix, Nososticta 430 beckeri, Roppaneura 455 Bedfordia 263 beebeanum, Leptagrion 309 beesoni, Rhinocypha 55 bellax, Indolestes 132 bellona bellona, Ceriagrion 300 - hoogerwerfi, Ceriagrion 300 belyshevi, Enallagma 249 bengalense, Pseudagrion 325 berenice, Philogenia 157 berlandi, Drepanosticta 404 - , subsp. of alluaudi, Seychellibasis 349 berlandi, race of cyanomelas, Xiphiagrion 284 beryllae, Vestalis 32 beschki, Heteragrion 149 bhatnagari, Disparoneura 390 bhintalensis, Ischnura 265 bicellulare, Melanesobasis 316 bicellulata, Argia 193 bicoerulans, Pseudagrion 326 bicolor, Amphicnemis 292 bicolor, race of curta, Chlorocypha 49 bicolor, Sapho 27 bicorne, Archeopodagrion 139 bicornis, Metaleptobasis 288

468 bicornuta, Drepanosticta 404 bicornuta, Platysticta 409 bidens, Nesobasis 318 bidens, Rhipidolestes 175 bidentata, Bayadera 80 bidentata Förster, Idiocnemis 379 bidentata Selys, Idiocnemis 379 bidentatum, Ceriagrion 301 bidentatum, Pseudagrion 326 biedermanni, Disparocypha 55 bifenestrata, Rhinocypha 47 biforata, Heliocypha 55 biforata abbreviata, Rhinocypha 55 -, delimbata, Heliocypha 55 bigemmata, Elattoneura 424 bilenearis, Neoneura 447 bilineata, Melanoneura 419 - , Platycnemis 393 bilineata, Trichocnemis 381 bilineatum, ab. of pulchellum, Agrion 226 bilineatus, Indolestes 132 billinghursti, Caliagrion 298 billitonis, Amphicnemis 292 bilobata, Archeopodagrion 139 bilobata, Micronympha 260 bimaculata, Coeliccia 372 binocellata, Agriocnemis 179 binotatum, Arion 208 bipartita, Hetaerina 41 bipunctatum, f. of najas, Erythromma 228 bipunctulata, Argia 193 bipupillatus, Lestes 111 birmanus, subsp. of gracilis, Indolestes 134 biseriata, Heliocypha 56 biseriata anambae, Rhinocypha 56 bisignata, Heliocypha 56 Libellago 59 bispina, Drepanosticta 405 bispinus, Perilestes 172 bivittata, Protosticta 414 bivittata, Trichocnemis 381 bizonata, Ischnura 265 blackburni, Megalagrion 312 blanchardi, Antiagrion 285 blandus, Micromerus 60 blumi, Ischnura 265 bocainense, Leptagrion 309 bocki, Euphaea 84 bogotensis, Cora 94 bogotensis, race of terminalis, Cora 95 boliviana, Philogenia 157 - , Polythore 97 bonariensis, Cyanallagma 247 Boninagrion 263 boninensis, Austrolestes 129

- , Lestes 111 bonita, Amphicnemis 292 borchgravii, Hetaerina 35 boreale, Enallagma 249 Bornargiolestes 146 borneense, Aciagrion 239 borneensis, Coeliccia 372 - , Rhinagrion 163 botacudo, Argia 193 botjanum, Ceriagrion 300 botti, Caconeura 438 boucardi, Telebasis 361 bovilla, Metaleptobasis 289 boettcheri, Coeliccia 372 Brachybasis 298 brachycerca, Nesobasis 318 brachygastra, var. of latipes, Platycnemis 395 Brachyneura 437 brachysticta, Coeliccia 373 bradleyi, Teinobasis 350 brasiliensis, Cora 94 - , Heliocharis 75 brasiliensis, Hetaerina 38 braueri, subsp. of annulata, Sympecma 136 braueri, Rhionocypha 71 bremii, Agrion 228 bravicauda brevicauda, Bayadera 80 - ishigakiana, Bayadera 80 brevispina, Enallagma 250 brevistigma, subsp. of caerulatus, Megaloprepus 459 - , Oxyagrion 279 brevistyla, Lais 40 bringhtwelli, Hetaerina 35 brinki, Drepanosticta 405 brisbanense, Coenagrion 219 britannica, subsp. of virgo, Calopteryx 17 brosetti, Aciagrion 239 brownelli, Drepanosticta 405 brucei, Palaemnema 399 brucelli, Palaemnema 399 bruchi, Oxyagrion 280 brunnea, subsp. of ochracea, Allophaea 78 - , Euphaea 84 brunneiopunctata, ab. of pennipes, Platycnemis 396 brysonima, Metaleptobasis 289 buchholzi, Chrysobasis 285 - , Enallagma 250 buchi, Philosina 171 buckleyi, Mecistogaster 456 buddha, Indolestes 134 buenavista, Philogenia 157 bukharensis, Ischnura 270

469 burmanensis, subsp. of verticalis, Prodasineura 442 Burmargiolestes 146 buwaldai, Podolestes 160 Teinobasis 350 buxtoni, Ischnura 266 bwamba, subsp. of straeleni, Chlorocypha 54 byersi, Telebasis 359 Caconeura 417 Caconeurinae 417 cacherensis, subsp. of campioni, Elattoneura 424 Caenoneura 446 caerulatus caerulatus, Megaloprepus 459 - brevistigma, Megaloprepus 459 caerulatus race latipennis, Megaloprepus 459 caerulea, Arabicnemis 369 - , Rhinoneura 73 caerulescens caerulescens, Coenagrion 220 - caesarum, Coenagrion 220 caerulescens var. pygmaea, Coenagrion 220 caerulescens theryi, Coenagrion 220 caeruleum, Acanthallagma 237 caesarum, subsp. of caerulescens, Coenagrion 220 caesia, Elattoneura 424 caffrum, Pseudagrion 326 caja, Hetaerina 35 calamineum, Ceriagrion 300 calamorum, Cercion 216 calcaris, Argiolestes 141 calcipennis, Microstigma 460 caldense, Minagrion 317 Caledargiolestes 147 Caledopteryx 147 calendulum, Heteragrion 149 Calepteryx 9 Caliagrion 298 Calicnemia 369 Calicneminae 368 Calicnemis, subgen. of Platycnemis 369 calidum, Agrion 196 californica, subsp. of aequabilis, Calopteryx 9 californica, Hetaerina 34 californicus, Archilestes 108 caligata, Microneura 446 caligata caligata, Platycypha 64 - angolense, Platycypha 64 caligata f. lacus, Platycypha 64 Calilestes 147 Caliphaea 7 Caliphaeinae 7 calliphya calliphya, Megalagrion 312 - micromelas, Megalagrion 312

callisphaena, Notoneura 430 Calocypha 48 Calopterygidae 6 Calopteryginae 8 Calopterygoidea 3 Calopteryx 9 calosomum, Pseudagrion 327 Calvertagrion 247 calverti, Agrion 312 calverti, Enallagma 249 - , Protoneura 452 camerunense, Enallagma 250 camerunensis, Libellago 53 camerunensis, Phaon 25 Camerunolestes 172 campioni, Caoeliccia 373 campioni campioni, Elattoneura 424 - cacharensis, Elattoneura 424 campioni, Nesobasis 318 canaanensis, Minagrion 317 canadense, Agrion 250 cancellata, Chlorocypha 49 cancellata insulana, Chlorocypha 49 candelaria, Leptobasis 287 Candibranchiata 2 candicans, Leucobasis 288 canescens, Neosticta 365 canescens f. dorrigoensis, Neosticta 365 canningi, Caconeura 441 cantralli, Mesoleptobasis 288 cantuga, Amphicnemis 292 capillaris, Protoneura 452 - , Tanymecosticta 367 capilliformis, Protoneura 452 capitalis, Hetaerina 36 capitalis var. colombiana, Hetaerina 36 capixabae, Leptagrion 309 caprai, subsp. of splendens, Calopteryx 15 capreola, Ischnura 266 - , Selysioneura 366 cara, Protoneura 452 cardenium, Enallagma 250 cardinale, Oxyagrion 280 cardinalis, Indophaea 88 - , Ischnura 266 caribbea, Enacantha 248 carissima, Pseudophaea 87 carlotta, Nehalennia 229 carmelita, Agriocnemis 179 - , Palaemnema 399 carmesina, Telebasis 359 carmichaeli, Drepanosticta 405 carminea, subsp. of doonensis, Calicnemia 370 carminita, Telebasis 359 carnatica, Neoneura 447

470 carnifex, Hetaerina 36 carnifex race fulgens, Hetaerina 36 carnifex carnifex, Palaiargia 211 - praeclave, Palaiargia 212 carolinensis, Teinobasis 351 carota, Telebasis 359 carpentieri, Ischnura 276 carrillica, Philogenia 158 cartvelica, subsp. of splendens, Calopteryx 15 carunculatum, Enallagma 250 cassandra, Philogenia 158 castellani, Coenagrion 220 castor, Perissolestes 173 casuarina, Podopteryx 161 cecilia, subsp. of intermedia, Calopteryx 13 Celaeneura 263 Celebargiolestes 148 celebense, Pseudagrion 327 cellularis, Isomecocnemis 428 centrafricana, Elattoneura 425 centralis, Elattoneura 425 centripunctata, Chlorocypha 49 cerasina, Psaironeura 454 ceratophora, Drepanosticta 405 Cercion 216 Ceriagrion 298 cerinomelas, subsp. of fallax, Ceriagrion 302 cerinorubellum, Ceriagrion 301 cerinostigma, Protosticta 416 cerinum, Agrion 302 ceromandelianum race erubescens, Ceriagrion 299 cervicornu, Selysioneura 366 cervula, Ischnura 266 Ceylanicolestes 131 ceylonicum, Mortonagrion 189 - , subsp. of rubiceps, Pseudagrion 342 Ceylonolestes 129 Ceylonosticta 403 ceyx ceyx, Palaiargia 212 - flammula, Palaiargia 212 chacoense, Acanthagrion 233 Chalcargia, subgen of Argia 191 Chalcolestes 109 Chalcopteryx 93 chalybeostoma, Nososticta 430 championi, Philogenia 158 chaoi, Ceriagrion 301 chapadae, Argia 193 chapadense, Oxyagrion 280 charalana, Argia 193 chararum, Acanthagrion 233 charca, Hetaerina 36 Charitopteryx 74 charmosyna charmosyna, Palaiargia 212

- cylopica, Palaiargia 212 - miniata, Palaiargia 212 charpentieri, Agrion 252 chaseni, Calicnemia 370 cheesmanae, Austrolestes 130 cheesmani, Ischnura 278 chelata, Argia 193 cheliferum, Argentagrion 246 - , Pseudagrion 327 chengi, Megalestes 106 chinensis chinensis, Neurobasis 23 chinensis var. florida, Neurobasis 24 chinensis longipes, Neurobasis 24 chingolae, subsp. of lacustris, Platycypha 65 chirihuanum, Helveciagrion 262 chiriquita, Palaemnema 399 chirripa, Cora 94 chloridion, Agrion 227 chloridum, Erythromma 231 chloroceps, Pseudagrion 327 Chlorocnemis 419 Chlorocypha 48 Chlorocyphidae 44 Chlorolestes 101 Chlorolestidae 101 Chlorolestinae 101 Chloroneura 422 chloropleura, Idiocnemis 379 chomae, Pseudagrion 337 chongwe, f. of salisburyense, Pseudagrion 343 Chorismagrion 105 Chorismagrioniae 105 Chromagrion 218 chromatus, Lestes 111 chromostigma, Ischnura 266 chromothorax, Coeliccia 373 Chrysobasis 285 chrysoides, Argiolestes 141 chrysops, Heteragrion 150 chrysopus, Podolestes 161 chusanicum, Coenagrion 220 ciganae, Phasmoneura 450 Ciliagrion 384 ciliata, Plagulibasis 323 ciliata, Psilocnemis 389 ciliata, Sapho 27 ciliata ab. fumipenne, Echo 27 cincta, Umma 28 cinctus, Celebargiolestes 148 cineraceus, Lestes 111 cingillum, Pseudagrion 327 cingulatus, Austrolestes 130 cinnamomeum, Heteragrion 149 circularis, Amphicnemis 292 circulatum, Enallagma 250

471 circumscripta, Nososticta 430 citricola, Pseudagrion 327 citrinum, Ceriagrion 301 civicum, Pseudagrion328 civile, Enallagma 250 civile race simile, Enallagma 250 claaseni, Drepanosticta 405 clara, Hypolestes 165 clauseni, Agriocnemis 179 clausseni, Argia 193 clausum, Enallagma 251 clavata, Drepanosticta 405 Cleis 28 dementia, Palaemnema 399 Climacobasis 18 coarctatum, Pseudagrion 328 coartans, Argiolestes 141 coccinata, Telebasis 359 coccinea, Telebasis 359 coecum coecum, Enallagma 251 - novaehispaniae, Enallagma 251 coeleste coeleste, Pseudagrion 328 - samfyae, Pseudagrion 328 colestina, Nososticta 430 Coeliccia 371 Coenagriinae 215 Coenagriocnemis 188 Coenagrion 219 Coenagrionidae 177 Coenagrionoidea 177 Coenagrioninae 214 Coenargiocnemis 188 coeruleiceps, Pseudagrion 328 coeruleipunctum, Pseudagrion 328 coeruleocauda, subsp. of nigripes, Chlorocnemis 422 coerulescens, Diphlebia 76 Prodasineura 438 coeruleum, Ceriagrion 301 coeruleum, Pseudagrion 215 coeruleus, Indolestes 132 cogmani, var. of massaicum, Pseudagrion 337 cognata, Calopteryx 11 cognata, Rhinocypha 66 collaris, Prodasineura 437 collarti, Libellago 59 collata, Argia 194 collopistes, Telebasis 359 colombiana, var. of capitalis, Hetaerina 36 colorata, Rhinocypha 66 Columbiana, Argia 194 colensonis, Austrolestes 130 combusta, Teinobasis 351 comes, Anisopleura 79 commoniae commoniae, Pseudagrion 328

- nigerrimum, Pseudagrion 328 commutata, Nososticta 430 comosa, Nesobasis 318 compar, Euphaea 85 concinna, Argia 194 - , Polythore 97 concinnum, Coenagrion 220 concinnus, Lestes 112 conditum, Chromagrion 218 confluens, subsp. of rutilans, Chalcopteryx 93 confusa, Caliphaea 7 Cora 94 confusus, Lestes 122 congener, Lestes 112 congoense, Aciagrion 239 congolensis, Platycnemis 393 conica, Drepanosticta 405 conjuncta, Lestoidea 137 connectens, Argiolestes 141 consimilis, Agriocnemis 181 Caliphaea 7 consisum, Enallagma 251 consors, Heteragrion 150 conspicuum, Pseudagrion 329 conspicuus, Chlorolestes 102 consueta, Libellago 52 continentale, subsp. of deserti, Enallagma 253 continentalis, Bayadera 80 contortum, Megapodagrion 167 contraria, Chlorocnemis 420 convalescens, Coenagrion 221 convergens, Argiolestes 142 coolawanyah, Eurysticta 363 coomani, Calopteryx 11 coomansi, Coeliccia 373 - , Elattoneura 425 - , Podolestes 161 - , Pseudagrion 329 Copera 388 Cora 94 cora, Euphaea 85 corallina, Telebasis 359 corallinum, Ceriagrion 301 corbeti, Agriocnemis 179 corculum, Protoneura 452 cordosa, race of dispar, Chlorocypha 50 coriaceum, Pseudagrion 329 Cornelia, Calopteryx 11 Selysioneura 366 cornelii, subsp. of fenestrata, Heliocypha 56 cornicauda, Telagrion 357 corniculata corniculata, Melanesobasis 316 - marginata, Melanesobasis 316 corniculatum, subsp. of pesechem, Papuagrion 322

472 cornuta, Risiocnemis 386 cornutus, Perissolestes 173 corolla, Teinobasis 351 coromandelianum, Ceriagrion 302 corrina, Caconeura 437 corruptum, Papuagrion 320 Coryphagrion 165 Coryphagrioninae 165 costalimai, Leptobasis 287 costalis, subsp. of pruinosa, Mnais 22 cothurnata, Dicterias 74 crassidens, Hetaerina 36 credula, subsp. of ramburi, Ischnura 275 crenitis, Drepanosticta 406 crenulatipennis, Tatocnemis 169 cretensis, var. of splendens, Calopteryx 15 cristalina, Philogenia 158 cristatus, Episynlestes 104 cristina, Neoneura 447 crocea crocea, Chlorocypha 49 - bamptoni, Chlorocypha 49 crocea, Leptobasis 287 croceicauda, Palaemnema 400 croceipennis, Argia 194 croceum, Leptagrion 309 croconota, Prodasineura 438 crocops, Pseudagrion 329 cruentata, Calopteryx 38 cruentata, subsp. of rosea, Nesosticta 435 crucigera, Archibasis 297 cruzi, Ischnura 266 cucullata, Rhinocypha 67 culicinovum, Enallagma 249 cultellatum, Neoerythromma 230 cuneata, Aristocypha 45 cuneatum, var. of gracile, Acanthagrion 233 cuneistigma, Enallagma 251 cupida, Protoneura 452 cupraurea, Argia 194 cuprea, Argia 194 cupraea, Argia 207 cupraea, Mnesarete 42 curiosa, Protosticta 414 curta, Chlorocypha 49 curta race bicolor, Chlorocypha 49 curtum, Megapodagrion 167 cuyabae, Acanthagrion 235 Cyanallagma 247 Cyanargia, subgen. of Argia 191 cyane, Austroagrion 215 cyane, Cora 95 cyaneipennis, Neurobasis 24 cyaneothorax, Coeliccia 373 cyaneovittata, Esme 418 cyaneus, Indolestes 132

cyanifrons, Chlorocypha 50 - , Cyclophaea 81 Cyanocharis 74 Cyanocnemis 378 cyanofrons, Oxystigma 156 cyanomelas, Coeliccia 373 cyanomelas race berlandi, Xiphiagrion 284 cyanomelas, Xiphiagrion 284 cyanops, Leptocnemis 381 cyanura, Isomecocnemis 428 - , Nososticta 431 cyathiforme, Pseudagrion 329 cyathigera, Argia 194 cyathigerum cyathigerum, Enallagma 251 - mongolicum, Enallagma 252 cyathigerum f. nigriprothorax, Enallagma 252 cyathigerum var. rotundatum, Enallagma 252 - , possompsei, Enallagma 252 cyclanthorum, Sciotropis 164 Cycloneura 437 Cyclophaea 81 cyclopica, subsp. of charmosyna, Palaiargia 212 Cyptolestes 108 Cyrano 54 dabreui, Agriocnemis 180 dactylostyla, Amphicnemis 293 daeckii, Enallagma 252 dagnina, Argia 194 dagnyae, Idiocnemis 379 dahli, Chlorocypha 50 dajakanus, Lestes 112 damula, Ischnura 266 danielli, Moroagrion 210 dardanoi, Leptagrion 309 davenporti, Ceylonolestes 134 davenporti, Protosticta 414 davisi, Enallagma 253 dealbata, Platycnemis 393 deami, Argia 195 debeauforti, Teinobasis 351 debeauxi, Teinobasis 351 deccanensis, subsp. of vittata, Copera 391 subsp. of maculata, Drepanosticta 413 deceptor, Agrion 312 deceptum, Acanthagrion 236 decipiens, subsp. of praemorsus, Lestes 119 deckeni, Agrion 335 declaratum, subsp. of pilidorsum, Pseudagrion 340 declivia, Umma 28 decolorata, Mnais 21 decoloratum, Libyagrion 260 deconcentrans, Pseudagrion 329 decorata, Anisophaea 78

473 decorata, Libellago 49 decorum, Pseudagrion 329 defixa, Ischnura 267 deflexum, subsp. of pilidorsum, Pseudagrion 340 degeneratum, Papuagrion 320 delia, subsp. of verticalis, Prodasineura 442 delicata, Ischnura 267 delicatula, Prodasineura 438 delicatulum, Agrion 265 delimbata, subsp. of biforata, Heliocypha 55 demerarum, Aeolagrion 291 demorsa, Ischnura 267 demorsum, Pseudagrion 330 dendrolagina, Drepanosticta 406 deningi, Pseudagrion 330 denticollis, Ischnura 267 denticularis, Tatocnemis 169 denticulata, Neoneura 447 dentifer, Amphicnemis 293 dentifera, Drepanosticta 406 dentiferum, Agrion 259 dentiplaga, subsp. of tincta, Rhinocypha 71 derivata derivata, Polythore 98 - adjuncta, Polythore 98 - ambigua, Polythore 98 derivata race originata, Polythore 98 derivata race terminata, Polythore 98 descendens, subsp. of apicale, Acanthagrion 233 deserti deserti, Enallagma 253 - continentale, Enallagma 253 - yezoense, Enallagma 253 desiderata, Palaemnema 400 Devadatta 4 devilli, Mnesarete 42 diadesma, Nososticta 431 Diargia 191 dicellulare, Calvertagrion 247 diceras, Telagrion 357 Diceratobasis 286 dichrostigma, Lestes 112 Dicterias 74 Dicteratidae 74 didyma, Coeliccia 374 dierli, Coeliccia 373 difficilis, Argia 195 digitatum, Pseudagrion 330 - , subsp. of alcicorne, Pseudagrion 324 digiticollis, Telebasis 360 digitiferum, Papuagrion 320 digna, Drepanosticta 406 Demeragrion 148 dimidiata dimidiata, Calopteryx 11 - apicale, Calopteryx 12 dimidiata, Dysphaea 82 dimissa, Argia 195

dinesiotes, Megalagrion 314 Dineura 76 dinoceras, Coeliccia 374 diotima, Nubiolestes 172 Diphlebia 76 Diphlebiidae 75 disarmatus, Lestes 112 discolor, Agrion 243 disjunctus disjunctus, Lestes 113 - australis, Lestes 113 dispar dispar, Chlorocypha 50 - ovulosa, Chlorocypha 50 - pyriformosa, Chlorocypha 50 dispar race cordosa, Chlorocypha 50 dispar, Indophaea 89 - , Leptagrion 309 - , Pseudagrion 330 disparilis, Calopteryx 24 Disparocypha 55 Disparoneura 422 Disparoneurinae 419 dissidens, Protallagma 282 dissimulans, Lestes 113 distadens, Palaemnema 400 distans, Megalestes 106 distigma, Agrion 271 distincta, Umma 29 distinctum, Agrion 226 divagans Selys, Enallagma 253 divagans Kellicott, Enallagma 255 divaricatum, Oxyagrion 280 - , Pseudagrion 330 diversa, Platycnemis 392 dives, Argia 195 divisus, Indolestes 133 dobsoni, Agriocnemis 180 dohrni, Disparoneura 438 dolabrata, Teinobasis 351 dolorosa, Indoneura 437 dolorosa, Pacificagrion 282 domina, Palaemnema 400 dominicana, Telebasis 360 dominula, Hetaerina 36 - , Teinobasis 351 donna, Hetaerina 36 doonensis doonensis, Calicnemia 370 - carminea, Calicnemia 370 dorcadion, Drepanosticta 406 dorsale, Aeolagrion 290 Heteragrion 150 dorsalis dorsalis, Elattonerura 425 - auricolor, Elattoneura 425 dorsalis, Prodasineura 438 dorsocyana, Libellago 60 dorsonigra, Nososticta 431

474 dorsosanguinea, Rhinocypha 67 dorothea, Coeliccia 374 - , Coenagrion 221 - , Ischnura 267 - , Lestes 113 - , subsp. of mandarinus, Phaon 26 dorrigoensis, f. of canescens, Neosticta 365 doualae, subsp. of pseudomassaicum, Pseudagrion 346 doubledayi, Enallagma 253 draconicus, Chlorolestes 102 Drepanosticta 403 drocus, Nesolestes 154 drusilla, Drepanosticta 406 - , Rhinocypha 67 dryas, Lestes 113 drymobila, Selysioneura 366 dualis, Cora 95 dubium, Enallagma 253 - , Stenagrion 349 ducatrix, Devadatta 4 dulitensis, Drepanosticta 406 dundoense, Pseudagrion 330 duodecima, Paraphlebia 156 dupex, Hetaerina 36 dupophila, Drepanosticta 406 dupuyi, Metacnemis 384 durum, Enallagma 253 dyeri, Coenagrion 221 Dysphaea 81 earnshawi earnshawi, Mnais 21 - thoracicus, Mnais 21 ebneri, Agriocnemis 185 ebneri, subsp. of elegans, Ischnura 268 ebona, Philogenia 158 eboracus, race of griseus, Argiolestes 142 eboratum, Heteragrion 150 ebrium, Enallagma 254 eburnea, Caconeura 431 eburnea, subsp. of salomonis, Nososticta 435 Ecchlorolestes 103 echigoana, Platycnemis 394 Echo 18 eclecta, Palaiargia 212 ecornuta, Amphicnemis 293 ecornutum, Coenagrion 221 ecuadorica, race of vacillans, Leptobasis 287 edai, Mnais 21 editus, Sinolestes 104 edmondi, Palaemnema 400 efasciata, subsp. of erythrura, Nososticta 431 egleri, Acanthagrion 233 egregia, Nososticta 431 eiseni, Enallagma 254

eisentrauti, Cholorcnemis 420 ekari, Papuagrion 320 Elattoneura 423 elatus, Lestes 114 electa, Umma 29 elegans, Calopteryx 10 elegans, Chlorolestes 102 elegans elegans, Ischnura 268 - ebneri, Ischnura 268 elegans var. infuscans, Ischnura 268 elegans var infuscens, Ischnura 268 elegans marquardti, Ischnura 268 - mortoni, Ischnura 269 elegans var. obsoleta, Ischnura 269 elegans pontica, Ischnura 269 - saharensis, Ischnura 269 eliasi, Inpabasis 286 eliptica, Argia 195 elisabeta, Philogenia 158 elisabethae, Nesolestes 154 - , subsp. of nymphula, Pyrrhosoma 231 elongata, Chlorocnemis 420 elongatum, Enallagma 254 - , Leptagrion 309 - , Pseudagrion 330 elopurae, Rhinagrion 163 elumbis, Junix 446 elwesi, Notholestes 7 emarginata, Teinobasis 351 emarginatipes, Tatocnemis 169 emarginatum, Pseudagrion 337 emma, Argia 195 emphyla, Nososticta 431 Enacantha 248 Enallagma 248 enganoense, subsp. of pilidorsum, Pseudagrion 340 ensifera, Argiocnemis 186 Eolestes 172 eos, Palaiargia 212 Epallage 83 Epallagidae 11 ephippiana, Drepanosticta 407 ephippiatus, Argiolestes 142 ephippigera, Epipleoneura 444 epiphonematicum, Pseudagrion 330 Epipleoneura 444 Epipotoneura 446 Episynlestes 103 erevanensis, subsp. of splendens, Calopteryx 15 erici, Coeliccia 374 erinys, Megapodagrion 167 erminea, Amphicnemis 293 ernstmayri, Palaiargia 212 erratica, Ischnura 269

475 erubescens, race of ceromandelianum, Ceriagrion 299 Erythagrion, subgen. of Agrion 358 erythraceum, Pseudagrion 335 erythrina, Telebasis 360 erythrogastrum, Heteragrion 150 erythromelas, Calicnemia 370 Erythromma 227 erythromma, Elattoneura 425 erythroneurum, Xanthagrion 362 erythroprocta, Nososticta 431 erythrops, Nesobasis 318 erythrostigma, Paramecocnemis 385 erythrura erythrura, Nososticta 431 - efasciata, Nososticta 431 Risiocnemis 386 esakii, Mnais 21 Esaulestes 131 escherichi, Platycnemis 394 Esme 418 espinalensis, Agia 195 estesi, Pseudagrion 331 esthera, Neoneura 448 esuriensis, Argiolestes 142 ethalae, Phylolestes 104 ethela, Dysphaea 82 - , Neoneura 448 eucera, Drepanosticta 407 Euchlorolestes 101 eudytum, Megalagrion 312 euglena, Teinobasis 351 Euphaea 83 Euphaeidae 77 euphoeoides, Diphlebia 76 euphorbia, Argia 195 eurinus, Lestes 114 Eurysticta 363 Euthore 90 Euthorinae 90 evanescens, Oxyagrion 280 evansi, Ischnura 269 evelynae, Nososticta 432 evidanum, Pseudagrion 331 excelsa, Agrion 268 excelsus, Orolestes 127 exclamationis, Austroagrion 215 - , Himalagrion 228 - , Zoniagrion 284 exigua, Agrion 268 exigua, Protoneura 452 exilis, Agriocnemis 180 eximia, Calicnemia 370 eximia, Libellago 68 Telebasis 356 exlis, var. of lindeni, Cercion 217

exoleta, Coeliccia 374 - , Drepanosticta 407 exoletus, Archilestes 108 expansa, Philogenia 158 exstriata, Ischnura 267 exsudans, Agriocnemis 180 extranea, Argia 196 extraneus, Indolestes 133 exul, Caconeura 430 exul, Calopteryx 12 Nososticta 432 exulans, Enallagma 254 exustum, subsp. of rotundatum, Microstigma 461 ezoin, Ischnura 269 ezonatum, Agrion 268 fabamacula, Chlorocypha 54 faivrei, var. of splendens, Calopteryx 15 falcatum, Mortonagrion 189 falcifer, Lestes 114 falcifera falcifera, Agriocnemis 180 - transvaalica, Agriocnemis 180 fallax fallax, Ceriagrion 302 - cerinomelas, Ceriagrion 302 - pendleburyi, Ceriagrion 302 fallax, subsp. of amaurodytum, Megalagrion 311 farinicolle, Pseudagrion 331 fasciata, Bayadera 80 - , Euthore 90 fasciatus, Chlorolestes 102 fasciatus nigerrimus, Chlorolestes 102 fasciculare, Aciagrion 239 fassli, Euthore 90 - , subsp. of maculatum, Microstigma 460 fastigiata, Euthore 90 fatime fatime, Epallage 83 - amasine, Epallage 83 - anatolica, Epallage 83 femina femina, Agriocnemis 181 - oryzae, Agriocnemis 181 femina, Umma 29 fenestrata fenestrata, Heliocypha 56 - cornelii, Heliocypha 56 fenestrella, Aristocypha 45 fernandezi, Epipleoneura 444 - , Metaleptobasis 289 fernandezianum, Leptagrion 309 feronia, Protosticta 414 ferox, Philogenia 158 ferrugineum, Agrion 302 festiva, subsp. of virgo, Calopteryx 17 feuerborni, Aciagrion 240 fickei, Protolestes 162

476 fieldi, Austrosticta 363 filamenta, Teinobasis 352 filicicola, Oristicta 365 filicornis, Torrenticnemis 388 filiforme, Pseudagrion 325 filiformis, Mecistogaster 456 filiformis, Tanymecosticta 368 Teinobasis 352 filigerus, Mecistogaster 457 filiola, Telebasis 360 filippina, Devadatta 5 filosa, Ischnura 269 filum, Mecistogaster 458 filum, Teinobasis 352 fimense, Acanthagrion 235 finalis, Libellago 60 finisterroe, Nososticta 432 fischeri, Enallagma 249 fisheri, Pseudagrion 331 fissa, Argia 196 fissicollis, Tanymecosticta 368 fissidens, Idiocnemis 379 fitzsimonsi fitzsimonsi, Platycypha 64 - inyangae, Platycypha 65 flammea, Risiocnemis 386 flammeola, Telebasis 360 flammeum, Aeolagrion 291 flammula, subsp. of ceyx, Palaiargia 212 Prodasineura 438 flavescens, var. of heterodoxum, Hesperagrion 263 flavicauda flavicauda, Coeliccia 374 - masakii, Coeliccia 374 flaviceps, Burmargiolestes 147 flavicornis, Amphicnemis 293 flavidorsum, Heteragrion 150 flavifacies, Prodasineura 439 flavifrons, Pseudagrion 331 flavilabris, Agriocnemis 181 - , Melanesobasis 316 flavipedum, Papuagrion 320 flavipennis, Chlorocnemis 420 Nososticta 432 flavipes, Platycnemis 394 - flavipes, Pseudagrion 331 - leonense, Pseudagrion 331 flavistigma, Mecistogaster 457 flavistyla, Ischnura 271 flavithorax, Papuagrion 320 flavostigma, Lestomima 174 flavostriata, Coeliccia 373 flavovittata, Onychargia 214 flavovittatum, Heteragrion 150 fletcheri, Disparoneura 424 floresiana, Drepanosticta 407

floresianus, Indolestes 133 florida, var. of chinensis, Neurobasis 24 floridana, subsp. of maculata, Calopteryx 14 floridense, subsp. of gracile, Acanthagrion 234 fluviatilis, Ischnura 269 - , Telebasis 360 foersteri, Protosticta 414 foliacea foliacea, Platycnemis 394 - saskii, Platycnemis 394 foliaceum, Aeolagrion 291 foliosa, Platycnemis 394 fonscolombii, Agrion 224 fontanus, Argiolestes 142 fonticola, Nososticta 432 fontinalis, Argiolestes 142 Drepanosticta 407 fontinalis race wheeleri, Drepanosticta 407 fontium, Agrion 208 forceps, Lestes 118 forceps, Peristicta 450 forcipata Le Roi, Agriocnemis 181 forcipata Sjöstedt, Agriocnemis 181 forcipata, Bayadera 80 - , Ischnura270 forcipata Hägen, Lestes 113 forcipata Rambur, Lestes 114 forcipula, Agrion 121 foreli, Metaleptobasis 289 forficula, Argia 196 - , Drepanosticta 407 - , Lestes 114 forficulatum, Mortonagrion 189 forficuloides, Nesolestes 154 formosa, Calopteryx 33 formosa, Euphaea 85 formosana, Ischnura 264 fortis, Teinobasis 352 fosteri, Argia 196 fountainei, Ischnura 270 fractum, Enallagma 254 fragile, Aciagrion 240 fragilis, Perilestes 172 fraseri, Chlorocypha 50 fraseri, Coeliccia 375 Drepanosticta 407 - , Indophaea 89 fraseri, Phaon 25 fraseri, Protosticta 414 fraseri, subsp. of pruinosum, Pseudagrion 341 fraterculum, Papuagrion 320 fraterna, Nososticta 432 fraudatricula, Argia 196 freiense, Acanthagrion 235 frenulata, Elattoneura 426

477 frequentula, Argia 197 freyi, Agrion 222 frigida, Chlorocypha 51 frontalis, Rhinocypha 67 fruhstorferi, Alloneura 432 fugax, f. of glaucum, Enallagma 255 - , f. of sinuatum, Enallagma 259 fulgens, race of carnifex, Hetaerina 36 fulgens, Teinobasis 352 fulgida, Argia 197 fulgida,Lais 43 fulgidipennis, Aristocypha 46 fuliginosa, subsp. of iridipennis, Phaon 26 fulvellum, Telagrion 357 fulvicollis, Neoneura 448 fulvipes, Agrion 231 fulvum, Aeolagrion 291 fumigata, Argia 197 fumipenne, ab. of ciliata, Echo 27 fumipennis fumipennis, Argia 197 - atra, Argia 197 fumosa, Sapho 27 funcki, Argia 197 funebris, Argia 198 funereum, Thaumatagrion 388 furcata, Amphicnemis 293 Anisopleura 79 - , Coeliccia 375 furcatum, Agrion 225 furcatus, Protolestes 162 furcifer, Podolestes 161 furcigerum, Pseudagrion 332 fusca, Agriomorpha 138 fusca, ab. of virgo, Calopteryx 17 fusca, Sympecma 136 fuscaenea, Epipleoneura 444 fuscibasis, Hetaerina 37 fuscipleuris, Umma 29 fuscoguttata, Hetaerina 37 fuscomarginis, Umma 29 gabonense, Enallagma 255 gaida, Neoneura 448 garbei, Leptagrion 310 garleppi, Telebasis 360 gamblesi, Pentaphlebia 5 Pseudagrion 332 gandawricus, subsp. of autumnalis, Caconeura 437 gangetica, Ischnura 270 garleppi, Mecistogaster 456 gauchae, Peristicta 450 gaumeri, Argia 198 gautama, Mortonagrion 190 gayi, Antiagrion 285

gazella, Drepanosticta 407 gemina, Ischnura 270 geminata, Lestes 115 - , subsp. of salomonis, Nososticta 435 geminatum, Enallagma 254 genei, Ischnura 270 georgfreyi, Ceriagrion 303 gerdae, Solomocnemis 387 gerhardi gerhardi, Argia 198 - nigrior, Argia 198 gerstaeckeri, subsp. of spernatum, Pseudagrion 345 gestroi, Chlorocnemis 421 gigantea, Polythore 98 giganteum, Pseudagrion 332 gigantula, Palaemnema 400 gigas, Pseudagrion 332 gilliesi, Amanipodagrion 139 girardi, Elattoneura 426 ghesquierei, Chlorocypha 51 glabrum glabrum, Ceriagrion 302 - longispinum, Ceriagrion 303 glaciale glaciale, Coenagrion 221 - Orientale, Coenagrion 222 gladiolatum, Neoerythromma 231 glauca, Amphicnemis 293 glauca glauca, Chlorocypha 51 - radix, Chlorocypha 51 Elattoneura 426 glaucescens glaucescens, Pseudagrion 332 - mozambicensis, Pseudagrion 332 - zambesiensis, Pseudagrion 333 glaucifrons, Libellago 50 glaucoideum, Pseudagrion 333 glaucum, Coenagrion 339 glaucum, Enallagma 255 glaucum f. fugax, Enallagma 255 globifer, Mnesarete 42 gloriosa, Archboldargia 191 - , Dysphaea 82 Sapho 28 gobica, subsp. of annulata, Sympecma 136 gomphoides, Caconeura 417 goniocerus, Indolestes 133 Goniostigma 456 gracile gracile, Acanthagrion 233 gracile var. cuneatum, Acanthagrion 233 gracile floridense, Acanthagrion 234 - minarum, Acanthagrion 234 - gracile, Aciagrion 240 - attenuatum, Aciagrion 240 gracilior, Isosticta 364 gracilis, Amphicnemis 294 gracilis gracilis, Indolestes 133 - birmanus, Indolestes 134

478 - peregrinus, Indolestes 134 gracilis, Libellago 51 gracilis, Nehalennia 229 - , Protosticta 414 - gracilis, Vestalis 32 - montana, Vestalis 32 gracilis amoena, Vestalis 32 gracillima, Prodasineura 439 gracillima, Teinobasis 356 gracillimus, Perilestes 172 graellsi, Ischnura 271 grahami, Matrona 19 grandaeva, Calopteryx 12 grande, Pseudagrion 326 grandis, Archilestes 108 - , Chlorocypha 51 - , Coryphagrion 165 granti, Enallagma 255 gratiosa, Agriocnemis 181 graueri, Libellago 52 gravelyi, Argiocnemis 221 gravelyi, Protosticta 414 greeni, Libellago 60 greeni, Platysticta 413 greeni, Pseudagrion 333 greenwayi, Platycypha 64 gregory, Mnais 21 greigi, Pseudagrion 344 greyi, Africocypha 45 griffinii, Telebasis 361 grilloti, Pseudagrion 333 grinbergi, Antiagrion 285 grisea, Mnesarete 42 griseus griseus, Argiolestes 142 - race albescens, Argiolestes 142 - race eboracus, Argiolestes 142 - race intermedius, Argiolestes 142 - subgriseus, Argiolestes 142 - race tenius, Argiolestes 142 guayaraca, Archilestes 108 guerini guerini, Euphaea 85 - inonei, Euphaea 85 guinae, race of vacillans, Leptobasis 287 guianensis, Perissolestes 173 guichardi, Pseudagrion 333 gurneyi, Papuagrion 320 guttifera, Mnesarete 42 - , Platycnemis 394 haemastigma, Ischnura 271 haematinum, Oxyagrion 280 haematopus, Risiocnemis 386 haematosoma, Prodasineura 439 haemorrhoidalis haemorrhoidalis, Calopteryx 12

- occasi, Calopteryx 12 hageni, Enallagma 255 hageni hageni, Pseudagrion 333 - tropicanum, Pseudagrion 333 hageni, Rhinocypha 67 hainanensis, subsp. of scitellum, Coeliccia 378 halecarpenteri, Ischnura 271 halcyon, Palaiargia 213 halterata, Drepanosticta 408 hamadryas, Drepanosticta 408 hamata Selys, Lestes 113 - Hägen, Lestes 114 hamoni, Aciagrion 240 - , Ceriagrion 303 Pseudagrion 334 hamulata, Argia 198 hamulum, Pseudagrion 334 handschini, Isosticta 364 Haplostigma 456 harknessi, Argia 198 haroldoi, Epipleoneura 444 harrissoni, Podolestes 161 hartmanni, Libellago 64 hasemani, Argia 198 hasta, Chlorocypha 52 hastatum, Anomalagrion 245 hastulatum, Burmeister Agrion 268 - Stephens, Agrion 251 hastulatum, Coenagrion 222 hauxwelli, Mecistogaster 458 hauxwelli, Mnesarete 43 hawaiense, Megalagrion 312 Hawaiiagrion 311 hearseyi, Protosticta 415 hebdomatica, Argia 198 hebe, Hetaerina 37 helena, Indolestes 132 helena, Philogenia 158 helenae, Chlorocypha 51 helenae helenae, Pseudagrion 334 - planivarum, Pseudagrion 334 Heliargia, subgen. of Argia 191 Heliocharis 75 Heliocharitidae 74 Heliocypha 55 helix, Lestes 114 Helveciagrion 262 helvola, Teinobasis 352 hemicolon, Pseudagrion 331 Hemicnemis 381 hemihylina, subsp. of quadrimaculata, Aristocypha 47 Hemiphlebia 176 Hemiphlebiidae 176 Hemiphlebioidea 176

479 hempeli, Oxyagrion 280 henrardi, Elattoneura 423 henshawi, Lestes 115 herberti, Argia 199 hermeticum, subsp. of mercuriale, Coenagrion 224 hermosae, Acanthagrion 234 heros, Megalestes 106 Hesperagrion 263 Hetaerina 34 Hetaerinae 34 hetaerinoides, Archineura 8 Heteragrion 148 Heterocypha 58 heterodoxum, Hesperagrion 263 heterodoxum var. flavescens, Agrion 263 heterogamias, Megalagrion 313 heteroneura, Nesobasis 318 Heterophaea 88 Heteropodagrion 153 Heteropteroides 177 heterosticta, Ischnura 271 heterosticta var. tasmanica, Ischnura 271 heterosticta, var.of macropus, Hetaerina 38 heterostictum heterostictum, Aciagrion 241 - karamoja, Aciagrion 241 heterostigma, Rhinocypha 67 heterostylus, Platylestes 128 hieroglyphicum, Cercion 216 hilaris, Drepanosticta 408 hilaryae, Aristocypha 46 hildegarda, Acanthagrion 234 hilli, Ischnura 272 Himalagrion 228 himalaiaca, Protosticta 415 hincksi, Mnesarete 43 hinei, Argia 199 hintzi, Libellago 49 hirosei, Mortonagrion 190 hisopa, Aciagrion 240 hisopa race krishna, Aciagrion 241 holdereri, Coenagrion 222 holosericea, Calopteryx 13 - , Euphaea 87 Homeoura 245 hoogerwerfi, subsp. of bellona, Ceriagrion 300 hosei, Prodasineura 439 hova, Platycnemis 394 huallaga, Argia 199 huanacina, Argia 199 hubelli, Inpabasis 286 hudsonica, subsp. of aequabilis, Calopteryx 10 hulae, subsp. of torridum, Pseudagrion 347 humerale, Erythromma 227 humeralis, Epipleoneura 444

- , subsp. of verticalis, Prodasineura 443 Rhinocypha 68 - , subsp. of serena, Teinobasis 356 humida, Palaiargia 213 humilior, Isosticta 364 hummeli, Platycnemis 397 humphriesi, Lestes 129 huonensis, Idiocnemis 379 hyacinthus, Austrocnemis 187 hyalina, Bayadera 80 Calopteryx 13 Euthore 90 Libellago 60 Mnesarete 43 - , Paraphlebia 157 hyalinata, Puella 396 hybridoides, Diphlebia 76 Hylaeagrion, 290 Hylaeargia 210 Hylaenonympha 308 hylas, Coenagrion 222 hylas ussurense, Coenagrion 222 hypermelas, Pseudagrion 334 hyperythra, Prodasineura 439 Hypocnemis 385 Hypolestes 165 Hypolestinae 165 Hyponeura, 191 Hypostrophoneura 417 ianthinipennis, Neurobasis 24 icterica, Argia 199 ictericum, Heteragrion 150 ictericus, Lestes 115 Icterolestes, subgen. of Lestes 109 icteromelas, Argiolestes 142 icteromelas, f. nobilis, Argiolestes 143 icterops, Heteragrion 151 icteroptera, Mnais 21 Idiocnemis 379 Idioneura 446 ifanicum, Enallagma 256 ignea, Risiocnemis 386 igniceps Fraser, Pseudagrion 327 igniceps Fraser, Pseudagrion 334 ignifer, Pseudagrion 334 ignitum, Ceriagrion 303 ignipennis, Rhinocypha 68 imbricata, Copera 389 imitans, Calicnemia 370 immaculata, Aristocypha 46 immaculata, var. of pumilio, Ischnura 275 immaculatum, f. of najas, Erythromma 228 immsi, Ischnura 244 immunda, Argia 199

480 impar, Coenagrion 222 - , Euphaea 85 impedita, f. of marshalli, Chlorocnemis 421 imperatrix, Mnesarete 43 impunctatum, Oxyagrion 280 impura, Argia 199 inaequale, Ceriagrion 303 inaequalis, Lestes 115 inaequalis, Thore 97 inaequidens, Idiocnemis 379 inaequipar, Euphaea 85 inaequistigma, Millotagrion 279 inalatum, Aeolagrion 291 inarmata, Ischnura 272 inas, Rhinocypha 57 inca, Aeolagrion 291 - , Cora 95 - , Heteragrion 151 Mesoleptobasis 288 incallida, Amphicnemis 294 incana, Cora 95 incarnata, Archineura 9 incerta, Prodasineura 439 incisa, Agrioncnemis 181 incisa race pulverulans, Agriocnemis 181 incisa, Risiocnemis 387 incisura, Archibasis 297 incisurum, Pseudagrion 335 incolor, Mnais 21 incolumnis, Telebasis 361 inconspicua, Drepanosticta 408 inconspicuum inconspicuum, Pseudagrion 335 - arabicum, Pseudagrion 335 inculta, Argia 199 incusa, Epipleoneura 444 Indagrion 188 indefensum, Acanthagrion 234 indefinita, Ischnura 272 indica, Bayadera 80 - , subsp.of lineata, Libellago 61 - , Onychargia 211 indicatrix, Argia 200 indicum. Pseudagrion 335 indicus, Indolestes 134 indivisa, Ischnura 272 indochinese, Ceriagrion 303 indocilis, Argia 200 Indocnemis 381 Indocypha 58 Indolestes 131 Indoneura 417 Indophaea 88 inepta, Chlorocnemis 420 iners, Agrion 257

inexpectum, Acanthagrion 234 infecta, Hetaerina 37 inflatus, Indolestes 134 infracavum, Pseudagrion 339 infrequentula, Argia 200 infumata, Argia 200 infumata, var. of linearis, Mecistogaster 457 infumata, Rhinocypha 56 infumosa Fraser, Umma 29 infuscans, var. of elegans, Ischnura 268 infuscens, f. of elegans, Ischnura 268 inlactea, Euthore 91 inonei, subsp. of guerini, Euphaea 85 inopinata, Thaumatoneura 171 inopinatum, Pseudagrion 335 inornata, Idiocnemis 379 inornatum, Anisagrion 296 Inpabasis 286 insignis, Nososticta 432 insipida, Argia 200 insula, Enallagma 256 insulana , subsp. of cancellata, Chlorocypha 49 insulare, Coenagriocnemis 188 Papuagrion 321 insularis, Austrolestes 130 insularis, race of pennipes, Platycnemis 396 integra, Prodasineura 439 integricollis, Nehalennia 229 intermedia, subsp. of rubescens, Argiocnemis 186 intermedia intermedia, Calopteryx 13 - cecilia, Calopteryx 13 - persica, Calopteryx 13 - , subsp. of variegata, Euphaea 88 - , Ischnura 272 intermedius, race of griseus, Argiolestes 142 interposita, Oristicta 365 - , subsp.of nymphula, Pyrrhosoma, 231 interrogatum, Coenagrion 223 interrupta, Agriocnemis 182 Prodasineura 440 - , Chlorocnemis, 420 interruptum, Agrion 226 - , Coenagrion 271 interruptum, Cyanallagma 247 intruda, Argia 203 inversa, Agriocnemis 182 - , Drepanosticta 408 inversum, Leptobasis 287 inyangae, subsp. of fitzsimonsi, Platycypha 65 io, Austrolestes 130 io, Rhinocypha 67 iphigeania, Mecistogaster 458 iralai, Argia 200 irene, Cora 95

481 Nehalennia 229 - , Nososticta 433 iricolor, Echo 19 iridea, Aristocypha 46 Iridictyon 19 iridipennis iridipennis, Phaon 26 - fuliginosus, Phaon 26 iridipennis occidentalis, Sapho 26 irina, Puella 225 irma, Megalestes 106 irregularis, Mesocnemis 383 isabellae, Lieftinckia 382 Ischnallagma 248 Ischnosoma 263 Ischnura 263 Ischnuragrion 238 Ischnuridia, subgen. of Ischnura 263 Ischnurinae 232 ishigakiana, subsp. of brevicauda, Bayadera 80 isidromorai, Pseudagrion 333 isoetes, Ischnura 272 Isomecocnemis 428 Isosticta 364 Isostictidae 363 isthmica, Telebasis 361 itatiaiae, Phasmoneura 450 jacksoni, subsp. of basilewskyi, Chlorocypha 48 - , subsp. of sjoestedti, Pseudagrion 344 jacobi, Lestes 115 janicea, Caledarargiolestes 147 javana, subsp. of vittata, Copera 391 - , subsp. of snellemanni, Melanocypha 63 japonica, Calopteryx 13 Ischnura 272 - , Matrona 20 japonicus, Lestes 115 jejuna, Libellago 52 jejuna, Tanymecocsticta 368 jessei, Acanthagrion 234 joana, Neoneura 448 joanetta, Palaemnema 400 jocaste jacaste, Mecistogaster 457 - sincerus, Mecistogaster 457 - vicentinus, Mecistogaster 457 jocosa, Argia 200 - , Josocora 92 joergenseni, Argia 200 johanella, Argia 201 johanseni, subsp. of splendens, Calopteryx 15 johanssoni, Agrion 220 josemorai, Elattoneura 426 Josocora 91 jucundus, Rhipidolestes 175 jugorum, Megalagrion 313

jujuya, Argia 201 Junix 446 kaffinum, Pseudagrion 335 kagiensis, Coenagrion 223 - , Enallagma 256 kahli, Perilestes 172 kali, Bayadera 81 Kalocora 92 kalumburu, Nososticta 433 karafutonis, Ischnura 273 karamoja, subsp. of heterostictum, Aciagrion 241 karnyi, Argiolestes 148 - , Caconeura 443 karschi, Rhinocypha 73 - , Xiphiagrion 284 kashmirensis, f. of paedisca, Sympecma 137 kashmirum, Coenagrion 223 katambovae, Ceriagrion 304 katharina, Rhinocypha 68 kauaiense, Megalagrion 313 kauderni, subsp. of nigridorsum, Enallagma 257 kaupi kaupi, Neurobasis 25 - pavo, Neurobasis 25 kellicotti, Ischnura 273 kempi, Indocnemis 381 kennedyi, Acanthagrion 235 - , Anisagrion 296 kerckhoffae, Protolestes 162 kerri, Aristocypha 47 kersteni, Pseudagrion 335 kervillei, Platycnemis 395 kevalensis, Agriocnemis 182 kiautai, Argia 201 Protosticta 415 kibalensis, Pseudagrion 336 Kilauagrion 311 kilimandjaricum, Pseudagrion 336 kimminsi, Idiocnemis 380 - , Lieftinckia 382 Neurobasis 25 inabaluensis, Protosticta 415 kirbyi, Argiolestes 143 - , Teinobasis 352 klenei, Josocora 92 klugi, Hetaerina 37 - , Perissolestes 173 - , Protoneura 453 kocheri, subsp. of puella, Coenagrion 225 koelanse, Megalagrion 313 kokama, Argia 201 koolpinyah, Nososticta 433 koongarra, Nososticta 433 kordofanicum, Ceriagrion 304

482 koxingai, Orolestes 127 kricheldorffi, Matrona 20 krishna, race of hisopa, Aciagrion 241 kruegeri Laidlaw, Drepanosticta 408 kruegeri Ris, Drepanosticta 403 krugi, Enallagma 251 kuiperi, Amphicnemis 294 kumaoensis, Coeliccia 377 kunjina, Agriocnemis 182 kurilis, Argia 210 kuroiwae, subsp. of mandarinus, Phaon 27 kusumi, Anisopleura 79 lachrymosa, Pacificagrion 282 lacrimans, Argia 201 lacus, f. of caligata, Platycypha 64 lacteola, Agriocnemis 182 lacteola, Platycnemis 390 lacteum, Agrion 396 lacus-elephantum, Africocypha 45 lacustris lacustris, Platycypha 65 - schingolae, Platycypha 65 laesa, Hetaerina 37 laglaizei, Teinobasis 352 laidlawi, Burmargiolestes 146 - , Calocypha 48 - , Euphaea 86 Prodasineura 440 Pseudagrion 336 - , Teinobasis 353 lairdi, Lieftinckia 382 lais, Apanisagrion 296 Lais, subgen. of Hetaerina 42 lamellata, Ischnura 273 lamina, Epipleoneura 445 laminatum, Papuagrion 321 lamprostomus, Argiolestes 143 lanceolata, Libellago 53 lanceolatum, Coenagrion 223 lanceum, Acanthagrion 235 lankanensis, Drepanosticta 408 lansbergei, Prodasineura 440 laosica, Calopteryx 13 lara lara, Allophaea 77 - balica, Allophaea 77 - lombokensis, Allophaea 78 lascellesi, Chlorocnemis 422 latapistylum, Acanthagrion 235 laterale, Cyanallagma 247 - , Enallagma 256 latericium latericium, Ceriagrion 304 - ryukyuanum, Ceriagrion 304 latialatus, Archilestes 108 latimacula, Rhinocypha 68 latipennis, race of caerulatus, Megaloprepus 459

latipes, Platycnemis 395 latipes var, brachygastra, Platycnemis 395 latistrigii, f. of viridulum, Erythromma 228 laurenti, Enallagma 262 leda, Austrolestes 130 lehmanni, Coenagrion 223 leonardi, Acanthagrion 235 Idiocnemis 380 leonense, subsp. of flavipes, Pseudagrion 331 leoninum, Argiagrion 298 leonora, Acanthagrion 236 leonora, Philogenia 159 leonarae, Idiocnemis 380 - , Protolestes 162 leonorae, Teinobasis 355 leopoldi, Neurobasis 23 lepidum, Megapodagrion 167 Leptagrion 308 leptalae, Tanymecosticta 368 Leptargia, subgen. of Argia 191 Leptocnemis 381 Leptobasinae 284 Leptobasis 286 leptodemas, Megalagrion 313 lepyricollis, Drepanosticta 409 leroii, Euthore 91 Lestes 109 Lestidae 107 Lestinae 107 Lestinoidea 101 Lestoidea 137 Lestoideidae 137 Lestoideinae 137 lestoides, Amphicnemis 294 - , Anisopleura 79 lestoides lestoides, Diphlebia 76 - tillyardi, Diphlebia 77 lestoides, Drepanosticta 409 Lestomima 174 Leucobasis 288 leucopsalis, Agrion 126 Leucopteryx 8 leucorhinum, Mesagrion 153 leucosticta, Allocnemis 369 leucostigma, Elattoneura 426 leucostigma, Mecistogaster 456 leveri, Nesobasis 319 Libellagidae 44 Libellago 58 libera, subsp. of amazona, Heliocharis 75 libera, Rhinocypha 68 Libyagrion 248 lieftincki,Ceriagrion 304 - , Coeliccia 375 - , Megalestes 106

483 Lieftinckia 382 lilacina, Argia 201 limbata, Dysphaea 82 - , var. of perforata, Heliocypha 57 - , Hetaerina 41 limitata, Argia 201 lincolniense,Agrion 227 lindeni lindeni, Cercion 216 lindeni var. exlis, Cercion 217 - , var. nigriceps, Cercion 217 lindeni zernyi, Cercion 217 lindgreni, Protosticta 415 lindicum, Pseudagrion 336 lindleyi, subsp. of acuta, Elattoneura 423 lindneri, Argentagrion 246 linearis, Mecistogaster 457 linearis var. infumata, Mecistogaster 457 lineata, Hetaerina 38 lineata lineata. Libellago 60 - indica, Libellago 61 lineolatum, Agrion 216 linsleyi, Indolestes 134 liveringa, Nososticta 433 livida, Telebasis 361 lizeria, Peristicta 450 lobata, Argia 201 lobata, Ischnura 264 lobimargo, Psilocnemis 389 Lochmaeocnemis 382 lombockensis, subsp. of lara, Allophaea 78 lombockensis, Libellago 69 longfieldi, subsp. of vaginale, Enallagma 261 longicauda, Agrion 103 longicauda, Bayadera 81 longipes, Hetaerina 37 longipes, subsp. of chinensis, Neurobasis, 24 longispina, Elattoneura 427 longispinosum, Acanthagrion 235 longispinum, subsp. of glabrum, Ceriagrion 303 longistigma, Umma 29 longistyla, Esme 418 - , Nesobasis 319 longiventris, Platycnemis 395 longjingensis, Caconeura 418 Prodasineura 440 longum, Telagrion 357 loogali. Coeliccia 375 lorena, Palaemnema 400 lorentzi, Nososticta 433 Oreagrion 279 loringae, Coeliccia 375 - , Philoganga 6 lorquini, Teinobasis 353 louisiadensis, Idiocnemis 380 lousiae, Amphicnemis 295

lucalaensis, subsp. of rufitibia, Platycypha 65 luciae, Teinobasis 353 lucidum, Pseudagrion 336 lucifer, Chrysobasis 285 lucretia, Mecistogaster 458 luctuosa, Vestalis 33 luctuosus, Mecistogaster 458 ludoviciana, Calopteryx 14 lugens, Argia 202 lugens, Climacobasis 18 lugens, Dysphaea 82 - , Vestalis 33 lugubris, Cora 95 luminosa, Chlorocypha 52 luna, Acanthagrion 236 lunatum, Microstigma 460 lundquisti, Indolestes 134 lunifera, Enallagma 219 lunulata, subsp. of rubescens, Argiocnemis 186 lunulatum, Agrion 227 luteipes, Argiolestes 143 luteola, Agriocnemis 182 - , Hetaerina 37 luteum, Acanthagrion 235 Acanthallagma 238 luxatus, Indolestes 134 luzonicum, Cercion 217 luzoniensis, Neurobasis 25 lyelli, Austrocoenagrion 216 lygisticerus, Indolestes 135 lymetta, Drepanosticta 409 maai, Megalestes 107 maccleeryi, subsp. of montana, Chlorocnemis 421 machadina, Argia 202 machadoi, Epipleoneura 445 - , Inpabasis 286 Oxyagrion 281 macilentum, Heteragron 151 - , Telagrion 358 macintyrei, Protoneura 453 mackwoodi, Caconeura 425 maclachlani, Agriocnemis 182 - , Allolestes 139 - , Mnais 22 macrocephalum, Rhinagrion 163 macrogaster, Diceratobasis 286 macrogaster, Nehalennia 286 macrolucidum, Pseudagrion 336 macrootithenae, Aciagrion 241 macropus, Hetaerina 38 macropus var. asticta, Hetaerina 38 macropus var. heterosticta, Hetaerina 38 macropus, Megapodagrion 167

484 macrostigma, Coeliccia 375 Lestes 115 macrostylis, Argiolestes 143 macrurum, Leptagrion 310 maculae, Acanthagrion 234 maculata, Caledopteryx 147 maculata maculata, Calopteryx 13 - floridana, Calopteryx 14 - maculata, Platysticta 413 - deccanensis, Platysticta 413 maculata, Rhinocypha 69 maculatum maculatum, Microstigma 460 - albopictum, Microstigma 460 - fassli, Microstigma 460 maculosa, Melanesobasis 316 macuma, Philogenia 159 madagascariense, Ciliagrion 384 madagazureum, Ceriagrion 304 madelenae, Amphicnemis 294 magdalenae, Perissolestes 173 magna, Agrion 268 magnanimum Lieftinck,Papuagrion 320 magnanimum Selys, Papuagrion 321 magoi, Hylaenonympha 308 mahesi, Calicnemis 377 major, Megalestes 107 major, f. of stellatum, Pseudagrion 326 majus, Hetaragrion 151 majuscula, Hetaeragrion 38 makabusiense, Pseudagrion 336 makoka, Argia 202 malabarica, Lestes 116 malabaricum, Pseudagrion 337 malacodora, Lochmaeocnemis 383 malagasoides, Pseudagrion 337 malaisei, Ceriagrion 304 Lestes 116 Rhipidolestes 175 malaitae, Lieftinckia 382 malawiensis, Seychellibasis 349 malayana, Libellago 60 malayanum, Cercion 217 - , Coenagrion 223 maldiviense, Enallagma 256 malgassica, Platycnemis 395 - , Tatocnemis 17P malgassicum, Pseudagrion 337 Malgassolestes, subgen. of Lestes 109 mammilaris, Leptobasis 287 manaliensis, Lestes 132 manauensis, Epipleoneura 445 magosisa, Philogenia 159 manicaria, Metaleptobasis 289 mandarinus mandarinus, Psolodesmus 26 - dorothea, Psolodesmus 26

- kuroiwae, Psolodesmus 27 marchali, Mecistogaster 458 margarita margarita, Echo 18 - tripartita, Echo 18 margarita, Philogenia 159 - , subsp. of corniculata, Melanesobasis 316 Mnesarete 43 marginipes, Copera 390 maria, Neoneura 448 mariae, Amphiocnemis 294 - , Heliocypha 57 - , Nesolestes 154 mariana, Neoneura 448 mariensis, Argia 202 marikovskii, subsp. of virens, Lestes 125 marina, Cora 95 - , Nesosticta 423 maroccana, Ischnura 271 marquardti, subsp. of elegans, Ischnura 268 marshalli marshalli, Chlorocnemis 420 marshalli f. impedita, Chlorocnemis 421 marshalli superba, Chlorocnemis 421 marsyas, Drepanosticta 409 martinae, Micromerus 61 martini, Amphicnemis 294 - , Mortonagrion 190 - , Nesolestes 154 - , Palaemnema 401 masakii, subsp. of flavicauda, Coeliccia 374 masoni, Euphaea 86 massaicum massaicum, Pseudagrion 337 massaicum var. cogmani, Pseudagrion 337 materna, Agriocnemis 181 - , Calopteryx 13 Matrona 19 Matronides, subgen. of Neurobasis 23 maurei, Telagrion 358 mauriciana, Platycnemis 395 mauritia, Metaleptobasis 289 mauritianum, Agrion 188 maxima, Echo 18 - , Hetaerina 38 mcgregori, Amphicnemis 295 mcleani, Melanesobasis 316 meccullochi, Austrocnemis 187 Mecistogaster 456 mecistogastrum, Minagrion 317 medinai, Hetaerina 39 medinensis, Argia 202 mediorufus, Lestes 116 medullaris, Argia 202 medusa, Protosticta 415 Megalagrion 311 Megalestes 106 Megalestinae 106

485 Megaloprepus 459 megalopus, Megapodagrion 168 megametta, Drepanosticta 409 Megapodagrion 167 Megapodagriidae 138 Megapodagriinae 166 Megapodagrionidae 138 Melanesobasis 316 melania, Bayadera 81 Vestalis 33 melanicterum, Pseudagrion 337 melanocauda, Palaemnema 401 melanocyana Selys, Archibasis 297 melanocyana Ris, Archibasis 298 Melanocypha 62 melanogaster, Diceratobasis 286 Melanoneura 419 melanoproctum, Coenagrion 223 melanopteryx, Bayadera 81 melanostigma, Palaemnema 401 melanota, Palaemnema 401 melanothorax, Argiolestes 146 melanothorax, Burmargiolestes 146 melanotum, Enallagma 256 melanoxantha, Nososticta 433 melanurum, Ceriagrion 304 Heteragrion 151 melidora, Palaiargia 213 melli, Calopteryx 14 mellisi, Pseudagrion 338 Tatocnemis 170 membranipes membranipes, Coeliccia 375 - , nemoricola, Coeliccia 376 - , nereis, Coeliccia 376 mendezi, Hetaerina 39 mercenarium, Megapodagrion 168 mercuriale mercuriale, Coenagrion 224 - hermeticum, Coenagrion 224 meridiana, Euthore 91 meridionale, Anisagrion 296 meridionalis, subsp. of virgo, Calopteryx 17 merina, Agriocnemis 183 Pseudagrion 338 mertoni, Idiocnemis 380 Mesagrion 153 Mesamphiagrion 278 Mesocnemis 383 Mesoleptobasis 288 Mesophaea 78 Mesopodagrion 169 mesostigma, Umma 29 Metacnemis 383 metagrion 140 Metaleptobasis 288 metallica, Epipleoneura 445

Mnesarete 43 metallica Förster, Teinobasis 353 metallica Ris, Teinobasis 354 metallicus, Argiagrion 143 miae, Libellago 59 micans, Megalestes 107 Teinobasis 353 Micrargia, subgen. of Argia 191 microcephalum, Pseudagrion 338 micromalgassica, Tatocnemis 170 micromelas, subsp. of calliphya, Megalagrion 312 Micromerus 58 Microneura 446 Micronympha 263 Micronymphidae 368 micropsitta, Palaiargia 213 Microstigma 459 microstigma, Argiolestes 143 Oxyagrion 281 migratum, Aciagrion 241 miles, Calicnemia 371 Millotagrion 278 milloti, Protolestes 162 mima, Libellago 61 mima, Rhinagrion 164 mimetes, Archibasis 297 Minagrion 317 minarum, subsp. of gracile, Acanthagrion 234 mindredae, subsp. of rufostigma, Ischnura 276 mingrelica, subsp. of splendens, Calopteryx 16 miniata, Calicnemia 371 - , Hetaerina 39 subsp. of charmosyna, Palaiargia 212 minima, Agriocnemis 183 minimum, Libellula 231 minimus, Argiolestes 143 miniopsis, Oxyagrion 281 minjerriba, Austrolestes 131 minor, Microstigma 460 minor, Perilestes 173 minteri, Philogenia 159 minusculum, Enallagma 256 minuta, Nehalennia 230 minutissima, Coenagrion 224 minutum, Acanthagrion 235 - , Anallagma 256 minutus, Lestes 116 Miocora 92 Miocorinae 91 mirifica, Archboldargia 191 mirabilis, Euthore 91 - , Hemiphlebia 176 mirabilis, Megalestes 127 mirabilis, Pseudolestes 175

486 mishuyaca, Argia 202 misoolensis, subsp. of australis, Neurobasis 23 missionensis, Argia 202 missionera, Peristicta 450 mista, Argia 203 mitratum, Heteragrion 151 mitwabae, Allocnemis 369 Mnais 20 mneme, Mnais 22 Mnesarete 42 modesta, Argia 203 Cora 95 - E c h o 18 - , Mecistogaster 458 modigliani, Euphaea 86 moesta, Argia 203 mohelli, Pseudagrion 338 molekulana, Nesobasis 319 molindica, Chlorocypha 52 mollis, Argia 203 mollusca, Argia 203 molakaiense, Megalagrion 313 moluccana, Drepanosticta 409 moluccensis, Nososticta 433 Mombagrion 238 monardi, Pseudagrion 324 mongolicum, subsp. of cyathigerum, Enallagma 252 monochroa, Rhinocypha 69 monostigma, Umma 29 monotis, Agnophilogenia 138 montana montana, Chlorocnemis 421 - maccleeryi, Chlorocnemis 421 montana, Coeliccia 376 Drepanosticta 409 - , Philoganga 6 - , Polythore 98 - , subsp. of gracilis, Vestalis 32 monteili, Lestes 133 montgomeryi, Euthore 91 montivagans, Argiolestes 144 moorei, Ceriagrion 305 moremi, Enallagma 257 moribunda, Hetaerina 39 Moroagrion 210 Mortonagrion 188 mortoni, Calicnemia 371 subsp. of elegans, Ischnura 269 mortoni, Protosticta 415 - , Pseudagrion 346 mortua, Hetaerina 39 motis, Orolestes 127 moultoni, Disparoneura 439 moultoni, Rhinocypha 69 mourae, Ceriagrion 305

mozambicensis, subsp. of glaucescens, Pseudagrion 332 mudiensis, Esme 419 mukherjeei, Calicnemia 371 multinervosa, Devadatta 5 munda, Argia 203 - , Cora 96 musa, Ischnura 270 mutans, Palaemnema 401 mutata, Disparoneura 426 mutata, Polythore 98 muzana, Argia 204 Myagrion 232 myersi, Iridictyon 19 mylitta, Drepanosticta 409 myrthea, Neoneura 449 mysticum, Neuragrion 155 myzomela, Palaiargia 213 nahuana, Argia 204 naia, Agriocnemis 183 naias, Libellago 61 nainitalensis, Agriocnemis 270 najas, Erythromma 227 najas f. baicalensis, Erythromma 228 - f. bipunctatum, Erythromma 228 - f. immaculatum, Erythromma 228 - f. puellare, Erythromma 228 nana, Agriocnemis 183 Nanosura 263 nasiterna, Palaiargia 213 natalense, subsp. of spernatus, Pseudagrion 345 natali, Africocypha 45 nathalia, Palaemnema 401 nawai, subsp. of pruinosa, Mnais 22 neblina, Archilestes 109 nebulosum, Megapodagrion 168 nectans, Rhipidolestes 175 needhami, Coenagrion 270 neglectus, Indolestes 135 Nahalennia 229 nehalennia, Epipotoneura 446 nemoricola, subsp. of membranipes, Coeliccia 376 Neocharis 74 Neocypha 54 Neoerythromma 230 Neoneura 446 Neosticta 365 Nepocnemis 299 nepos, Argentagrion 246 neptuna, Chlorocypha 52 nereis, subsp. of membranipes, Coeliccia 376 nasiotes, Megalagrion 314 Nesocnemis, subgen. of Prionocnemis 169

487 Nesobasis 318 Nesolestes 153 Neuragrion 155 Neurobasis 23 Neurolestes 155 newtoni, Pseudagrion 338 nexans, Diplebia 77 nielseni, subsp. of tenellum, Ceriagrion 307 nietneri, Drepanosticta 410 niger, Lestes 123 nigeriense, subsp. of sjoestedti, Pseudagrion 344 nigeriensis, Chlorocypha 52 - , Nesolestes 154 nigerrima, Elattoneura 427 nigerrimum, subsp. of commoniae, Pseudagrion 328 nigerrimus, subsp. of fasciatus, Chlorolestes 102 nigra, Allolestes 139 nigra, Bornargiolestes 146 Elattoneura 427 - , Prodasineura 440 - , Teinobasis 353 nigrescens, Coeliccia 376 Synlestes 105 - , subsp. of apicalis, Vestalis 32 nigricans, subsp. of rubescens, Argiocnemis 186 nigriceps, var. of lindeni, Cercion 217 nigriceps, Ischnura 273 - , Lestes 116 nigridorsum nigridorsum, Enallagma 257 - kauderni, Enallagma 257 nigrifrons, Nososticta 434 nigrinuhale, Cyanallagma 247 nigrior, subsp. of gerhardi, Argia 198 nigripectus, Matrona 20 nigripes nigripes, Chlorocnemis 421 - coeruleocauda, Chlorocnemis 422 nigripes semlikiensis, Chlorocnemis 421 nigripes, Pseudagrion 339 nigriprothorax, f. of cyathigerum, Enallagma 252 nigriventris, Idiocnemis 380 nigrofasciata, Nososticta 434 nigrofasciatum, Pseudagrion 339 nigrofasciatus, Lestes 119 nigroflavum, Ceriagrion 305 nigrohamata, Coeliccia 376 nigrohamatum nigrohamatum, Megalagrion 314 - nigrolineatum, Megalagrion 314 nigrolineatum, Ceriagrion 305 - , Enallagma 257 - , subsp. of nigrohamatum, Megalagrion 314 nigrolutea, Teinobasis 353 nigrostigma, Nesobasis 319

niloticum, Pseudagrion 339 nipalica, Calicnemia 371 nipponicum, Ceriagrion 305 nitens, Caliphaea 81 nitescens nitescens, Teinobasis 353 - angusticlavia, Teinobasis 354 nitidula, subsp. of pennipes, Platycnemis 396 nobilis, f. of icteromelas, Argiolestes 143 nodalis, Lestes 116 nodosum, Aciagrion 241 Noguchiphaea 8 Normopteroides 3 Nososticta 429 notabilitata, Agrion 24 Notholestes 1 Notoneura 429 notostigma, Prodasineura 440 notoxantha, Cora 96 novaehispaniae, subsp. of coecum, Enallagma 251 nubecula, Heliocypha 57 nubicum, Pseudagrion 339 nubilipennis nubilipennis, Chlorocnemis 422 - rossii, Chlorocnemis 422 Nubiolestes 172 nursei, Rhodischnura 283 nyansana, Platycnemis 395 nylephtha, Ecchlorolestes 103 nympha, Agrion 111 - , Lestes 113 nymphoides, Diphlebia 77 nymphula nymphula, Pyrrhosoma 231 - elisabethae, Pyrrhosoma 231 - interposita, Pyrrhosoma 231 Oahuagrion 311 oahuense, Megalagrion 314 oberthuri, Matrona 20 obiensis, Argiolestes 144 - , Palaiargia 213 obiri, Indolestes 135 obliterata, Idiocnemis 380 obliteratum, Enallagma 255 oblungulum, Ceriagrion 305 obscura, Argia 197 obscura, Argiolestes 144 obscura, subsp. of rubescens, Argiocnemis 187 - , Caconeura 418 Cora 96 obscurum, subsp. of pilidorsum, Pseudagrion 340 obscurus, Lestes 118 - , Micromerus 60 obsoleta, var. of elegans, Ischnura 268 obsoleta, Prodasineura 441

488 obsoletum, Acanthagrion 235 Heteragrion 151 Leptagrion 310 occasi, subsp. of haemorrhoidalis, Calopteryx 12 occidentale, Aciagrion 241 occidentalis, subsp. of iridipennis, Sapho 26 occipitale,Papuagrion 321 occisa, Hetaerina 38 occisa var. sublimata, Hetaerina 38 occultum, Mesamphiagrion 278 oceanicum, Megalagrion 314 oceanis, subsp. of angusta, Rhinocypha 55 ochracea ochracea, Allophaea 78 - brunnea, Allophaea 78 ochreacum, Heteragrion 151 ochraceus, Argiolestes 144 - ochraceus, Lestes 116 - aldabrensis, Lestes 117 - unicolor, Lestes 117 ochrostomus, Argiolestes 144 octogesima, Coeliccia 376 octogesima, Trichocnemis 374 octomaculatus, Orolestes 127 oculata, Argia 204 oculata Martin, Disparoneura 421 oculata Kirby, Disparoneura 425 odoneli, Prodasineura 441 odzalae, Prodasineura 441 Odonata 1 oedipus, Platycnemis 396 oenea, Argia 204 ogasawarensis, Rhinocypha 69 ogumi, f. of costalis, Mnais 22 okinawana, Calopteryx 14 okinawanus, Rhipidolestes 176 olivacea, Teinobasis 354 olivaceum olivaceum, Ceriagrion 305 - aurantiacum, Ceriagrion 306 olmyra, Phasmoneura 450 olsufiefTi, Pseudagrion 339 - , Tatocnemis 170 olthofi, Teinobasis 354 olympicum, Aciagrion 242 Onychargia 210 opaca, Calopteryx 13 opaca, Euphaea 86 oppositum, Papuagrion 321 optata, Palaiargia 213 orang, Indocnemis 381 ordosi, Ischnura 273 oreadum, Oreagrion 279 Oreagrion 279 oreas, Telagrion 358 Oreocnemis 384

oresitophum, Megalagrion 315 orichalcea, Argia 204 - , Sapho 28 Orientale, subsp. of glaciale, Coenagrion 222 Orientale, subsp. of viridulum, Erythromma 228 Orientale, Pseudagrion 339 orientalis orientalis, Calopteryx 14 - risi, Calopteryx 14 orientalis, Ischnura 264 orientalis, Platylestes 128 - , Podolestes 161 - , Sinhalestes 128 orientis, f. of torridum, Pseudagrion 347 originata, race of derivata, Polythore 98 Oristicta 365 ornata, Anisophaea 78 ornata, var. of pumilio, Ischnura 275 ornata, Mecistogaster 458 ornatum, Coenagrion 224 ornatus ornatus, Argiolestes 144 - subornatus, Argiolestes 144 ornatus, Sinolestes 104 orobates, Megalagrion 315 Orolestes 127 Or tholes tes 165 oryzae, subsp. of femina, Agriocnemis 181 oscillans, Archibasis 297 - , Megapodagrion 168 ovatum, Heteragrion 152 ovigerum, Archaellagma 245 ovulosa, subsp. of dispar, Chlorocypha 50 Oxyagrion 279 Oxyallagma 282 Oxystigma 156 Pachycypha 63 pachystigma, Stenocnemis 387 Pacificagrion 282 pacificum, Megalagrion 315 Pseudagrion 339 paedisca paedisca, Sympecma 137 paedisca f. kashmirensis, Sympecma 137 paedisca striata, Sympecma 137 pagenstecheri pagenstecheri, Rhinocypha 69 - pusilla, Rhinocypha 69 - timorana, Rhinocypha 69 palaeforme, Agriocnemis 183 Palaemnematinae 398 Palaemnema 398 Palaeobasis 298 Palaiargia 211 palauense, Pseudagrion 340 palauensis, Drepanosticta 410 - , Teinobasis 354 palawana, subsp. of vittata, Copera 392

489 - , Prodasineura 441 palawani, Coeliccia 377 pallens, Argia 204 palliatum, Agrion 326 pallida, Epipleoneura 445 pallida, subsp. of pseudalatipes, Platycnemis 397 pallidistigma, Calilestes 148 pallidistylus, Argiolestes 144 pallidifrons, Rhinocypha 70 pallidula, Nehalennia 230 pallidum, Aciagrion 242 Ceriagrion 306 - , Enallagma 257 pallidus Navas, Lestes 122 pallidus Rambur, Lestes 117 paludensis, Aciagrion 241 paludicola, Megalagrion 315 paludosus, Lestes 112 palustris, Agrion 448 pan, Drepanosticta 410 pandanicola, Amphicnemis 295 pandanicolum, Papuagrion 321 papaverina, Hetaerina 39 papilionacea, Calopteryx 13 paprzyckii, Perissolestes 174 Papuagrion 319 Papuargia 214 papuense, Pseudagrion 327 Paracnemis 384 Paracypha 63 paradisearum, Neurobasis 23 paraensis, Telebasis 361 Paralestes 109 Paramecocnemis 385 parameles, Papuagrion 321 paramoena, Prodasineura 441 Paraneuroptera 1 Paraphaea 88 Paraphlebia 156 pardalina, Neoneura 447 parthenias, Calopteryx 14 parvidens, subsp. of viridis, Lestes 126 parvulus, Argiolestes 144 parvum, Amphiallagma 244 pasquinii, Elattoneura 427 patricia, Ischnura 273 patricius, Lestes 117 paulicaxa, Palaemnema 401 paucinervis, Protoneura 453 pauli, Chlorocnemis 422 pauliani, Nesolestes 154 paulicoba, Palamnema 401 paulina, Palaemnema 402 Paulinagrion 323 paulirica, Palaemnema 402

paulistus, Lestes 117 paulitaba, Palaemnema 402 paulitoyaca, Palaemnema 402 pavidum, Oxyagrion 281 pavo, subsp. of kaupi, Neurobasis 25 pavonis, Chlorocypha 51 paya, Neoneura 449 pecchioli, Platycnemis 396 pectitus, Argiolestes 145 pedicillatus, Mecistogaster 458 pelecotomum, Pseudagrion 340 peles, subsp. of amaurodytum, Megalagrion 311 pellucida, Miocora 92 - , Thaumatoneura 171 pelops, Rhinocypha 70 pemon, Heteragrion 152 pendleburyi, subsp. of fallax, Ceriagrion 302 pendulum, Coenagrion, 225 pennipes pennipes, Platycnemis 396 pennipes ab. albioctopunctata, Platycnemis 396 pennipes ab. brunneiopunctata, Platycnemis 396 pennipes ab. insularis, Platycnemis 396 pennipes nitidula, Platycnemis 396 Pentaphlebia 5 peraltica, Miocora 92 paramens, Protoneura 453 percellulata, Argia 205 percornuta, Stenocora 93 percostale, Trineuragrion 165 percubitale, Dimeragrion 148 peregrinum, Heteragrion 152 peregrinus, subsp. of gracilis, Indolestes 134 pereirai, Epipleoneura 445 perengueyi, Ecchlorolestes 103 perforata, Heliocypha 57 perforata beatifica, Rhinocypha 57 perforata var. limbata, Rhinocypha 57 perfuscatum, Pseudagrion 340 Pericnemis 322 Perilestes 172 Perilestidae 171 perimecosoma, Palaiargia 214 perisi, Prodasineura 441 Perissolestes 173 Peristicta 450 perlongum, Leptagrion 310 perparva, Ischnura 273 perplex, Hetaerina 41 persica, subsp. of intermedia, Calopteryx 13 pertinax, Lestes 117 peruanum, Acanthagrion 236 peruviana, Dicterias 75 peruviana, Palaemnema 402 -Philogenia 159 perivianum, Acanthagrion 236

490 peruviensis, Protoneura 453 perverulenta, Teinobasis 355 pesechem pesechem, Papuagrion 321 - corniculatum, Papuagrion 322 peterseni, Andinagrion 244 petiolata, Sundacypha 73 petiolatum, Oxystigma 156 pfeifferi, subsp. of splendens, Calopteryx 16 phaethon, Libellago 61 phallatum, Agrion 136 phallicornis, Acanthagrion 236 Phantasma, Rhinocypha 70 Phaon 25 phasma, Selysioneura 366 Phasmoneura 450 Phasmosticta 365 philippa, Drepanosticta 410 philippinum, Rhinagrion 164 Philoganga 6 Philoganginae 5 Philogenia 157 Philosina 171 Philosininae 170 phoenissa, Nososticta 434 phoenix, Oreocnemis 384 Phylloneura 419 phyllopoda, Platycnemis 397 Phylolestes 104 picicaudata, Palaemnema 402 picta, Platycypha 65 - Polythore 99 picta race aequatorialis, Thore 100 picteti, Agrion 121 picteti, Lestes 115 pictum, Enallagma 257 picturata, Thore 99 pictus, Lestes 117 pieli, Agriocnemis 183 - Mnais 22 pieris, Agriocnemis 183 pierrati, Platycnemis 397 pilbara, Nososticta 434 pilidorsum pilidorsum, Pseudagrion 340 - declaratum, Pseudagrion 340 - deflexum, Pseudagrion 340 - enganoense, Pseudagrion 340 - obscurum, Pseudagrion 340 - simalurum, Pseudagrion 341 pilula, Hetaerina 39 pindrina, Austroagrion 216 pinheyi, Agriocnemis 180 pinheyi, Lestes 117 - Platycypha 65 pipila, Argia 205 piscinarium, Enallagma 255

pithoragarhicum, Himalagrion 229 plagiata, Euthore 90 plagiata, Nososticta 434 plagiatus, Lestes 118 plagiosum, Cercion 217 plagioxantha, Nososticta 434 Plagulibasis 323 plana, Argia 205 planivarum, subsp. of helenae, Pseudagrion 334 Platycnemididae 368 Platycnemidinae 388 Platycnemis 392 Platycypha 63 Platylestes 128 platypoda, Agrion 396 Platyscelus 392 Platysticta 413 Platystictidae 398 Platystictinae 402 platystictoides, Paulinagrion 342 platystigma, Ceriagrion 306 Platystigma 456 platystyla, Amphicnemis 295 platystylus, Platylestes 128 pluotae, Elattoneura 427 pocomana, Argia 205 Podagrion 167 Podagrioninae 166 Podolestes 160 podolestoides podolestoides, Devadatta 5 - basilanensis, Devadatta 5 Podopteryx 161 pollutum, Enallagma 257 pollux, Perissolestes 174 polychromatica, Drepanosticta 410 polychromaticum, Enallagma 258 Polythore 96 Polythoridae 89 Polythorinae 93 polyxena, Philogenia 159 ponapensis, Teinobasis 354 pontica, subsp. of elegans, Ischnura 269 ponticum, Coenagrion 225 pontogenes, Pseudagrion 341 popoluca, Argia 205 porrectum, Leptagrion 310 posita posita, Ischnura 273 - acicularis, Ischnura 274 - azteca, Ischnura 274 possompsei, subsp. of cyathigerum, Enallagma 252 postnodalis, Argiolestes 145 poungyi, Coeliccia 377 pozuzina, Polythore 99 praecellens, Lestes 118

491 praeclava, subsp. of carnifex, Palaiargia 212 praeclavum, Pseudagrion 297 praemorsus praemorsus, Lestes 118 praemorsus andamanensis, Lestes 118 praemorsus decipiens, Lestes 119 - sikkimus, Lestes 119 praetermissum, Ceriagrion 306 praetextatum, Pseudagrion 335 praevarum, Enallagma 258 praevius, Lestes 119 pretiosa, Teinobasis 354 Prionocnemis 385 procera, Heliocharis 75 - Polythore 99 Prodasineura 437 prognata, Ischnura 274 Proischnura 248 prolixa, Melanesobasis 317 prolongata, Proneura 341 Proneura 451 pronoti, Mecistogaster 459 Prophaon 25 Proplatycnemis 392 proselytus, Protolestes 163 Protallagma 282 prothoracale, Papuagrin 322 prothoracalis, Argiolestes 145 prothoracica, Rhyacocnemis 385 - Teinobasis 354 prothoracicum, Leptagrion 310 Protolestes 162 Protoneura 451 Protoneuridae 417 Protoneurinae 443 Protosticta 413 protostictoides, Platycnemis 397 - Protoneura 453 proxima, Hetaerina 39 proximum, Microstigma 459 pruinescens, Idiocnemis 380 - Ischnura 274 pruinescens, Lestes 121 pruinosa, Elattoneura 427 pruinosa, Metacnemis 383 pruinosa pruinosa, Mnais 22 - costalis, Mnais 22 - costalis f. ogumi, Mnais 22 - nawai, Mnais 22 pruinosa, Mnesarete 43 pruinosum pruinosum, Pseudagrion 341 pruinosum fraseri, Pseudagrion 341 pruinosum ranauense, Pseudagrion 341 Psaironeura 454 Pseudagriinae 290 Pseudagrion 323

pseudalatipes, Platycnemis 397 pseudalatipes pallida, Platycnemis 397 pseudamericana, Hetaerina 34 Pseuderythromma 227 pseuderythromma, Pseudagrion 238 pseudexul, Nososticta 435 Pseudochalcolestes, subgen. of Lestes 110 Pseudocopera 392 Pseudolestes 175 Pseudolestidae 174 pseudoelongatum, Enallagma 254 pseudomassaicum, Pseudagrion 346 - doualae, Pseudagrion 346 - f. rusingae, Pseudagrion 346 Pseudophaea 84 pseudosjoestedti, subsp. of sjoestedti, Pseudagrion 344 Pseudostigma 461 Pseudostigmatidae 455 Pseudostigmatinae 455 Psilocnemis, subgen, of Platycnemis 388 Psolodesmus 26 psyche, Austrolestes 131 pterauratum, Pseudagrion 341 pudica, Mnesarete 44 Puella 109 puella, Umma 30 - puella, Coenagrion 225 - kocheri, Coenagrion 225 - sytiacum, Coenagrion 225 puellare, f. of najas, Erythromma 228 Puellidae 177 pulchella, Agrion 220 - , Sapho 97 pulchellum, Coanegrion 225 pulchellum ab. bilineatum, Agrion 226 pulcherrimus, Indölestes 135 pulchrum, Agrion 252 pulla, Argia pulverulans, race of incisa, Agriocnemis 181 pulverulans, Calicnemia 371 - , Nesolestes 155 pumilio, Ischnura 274 pumilio var. immaculata, Ischnura 275 - var. ornata, Ischnura 275 pumilio var. punctata, Ischnura 275 - f. stephanella, Ischnura 274 - var. striata, Ischnura 275 punctata, var. of pumilio, Ischnura 275 punctum, Pseudagrion 341 pupa, Agrion 225 pupilla, Agrion 268 purpurea, Umma 30 pusilla, subsp. of pagenstecheri, Rhinocypha 69 pusillissimus, Argiolestes 145

492 pusillus, Agriolestes 145 putridum, Agrion 203 pygmaea, Agriocnemis 184 pygmaea, var. of caerulescens, Coenagrion 220 pyriformis, Coeliccia 377 pyriformosa, subsp. of dispar, Chlorocypha 50 pyroprocta, Nososticta 435 Pyrrhosoma 231 pytho, Drepanosticta 410 quadrata, Drepanosticta 410 quadratum, Acanthagrion 236 quadricolor, Telagrion 358 quadricornis, Metaleptobasis 289 quadrigerum, Cercion 217 quadrimaculata, Aristocypha 47 quadrimaculata hemihyalina, Rhinocypha 47 quadrimaculata, Disparoneura 423 quadrioculatum, Pseudagrion 342 quadristigma, Prodasineura 441 quadristriatus, Lestes 119 quattuordecimmaculata, Rhinocypha 57 quercifolius, Lestes 119 quigyanensis, Anisopleura 79 quinta, Paraphlebia 157 radama, Nesolestes 155 radians, Chalcopteryx 93 radiatus, Lestes 119 radix, subsp. of glauca, Chlorcypha 51 raineyi, Telagrion 358 rajah, Teinobasis 355 ramajana, Disparoneura 423 ramburi, Caconeura 418 Coenagriocnemis 188 ramburi ramburi, Ischnura 275 - credura, Ischnura 275 ramosa, Lieftinckia 382 ranatra, Selysioneura 366 ranauense, subsp. of pruinosum, Pseudagrion 341 ranavalona, Nesolestes 155 ranee, Teinobasis 355 rangifera, Nososticta 435 - , Selysioneura 367 raphaella, Philogenia 159 rarum, Aciagrion 242 rasoherinae, Phaon 26 reclusa, Argia 205 rectangula, Argia 205 rectangulare, Papuagrion 322 rectangularis, Lestes 120 rectangulata, Calicnemia 371 recurva, Teinobasis 355 recurvatum, Enallagma 258

reductum, Papuagrion 322 reflexa, Risiocnemis 386 refulgens, Euphaea 86 regalis, Archilestes 109 regina, Mnesarete 44 regulatus, Lestes 120 reinholdi, Diphlebia 76 remiger, Amphicnemis 295 remissa, Psaironeura 454 remotus, Perissolestes 174 remus, Perissolestes 174 renaudi, Pseudagrion 342 renifera, Coeliccia 377 resecta, Coeliccia 377 resolutum, Coenagrion 226 retrograda, subsp. of tincta, Rhinocypha 72 reuniensis, Coenagriocnemis 188 reventazoni, Palamnema 402 rhapia, Selysioneura 367 Rhinagrion 163 Rhinocypha 65 Rhinoneura 73 Rhipidolestes 175 rhoadsi, Argia 206 Rhodischnura 283 rhodosoma, Ischnura 276 rhomboidalis, Agrion 302 Rhomboides 177 Rhyacocnemis 385 ribeiroi, Minagrion 317 - , Telagrion 358 riccii, Megalestes 107 ridleyi, Lestes 127 Rima 100 Rimanella 100 Rimanellidae 100 risi, Acanthagrion 237 risi, Caconeura 418 - , subsp. of orientalis, Calopteryx 14 - , Chorismagrion 106 risi, Disparoneura 443 risi Pinhey, Enallagma 259 risi Schmidt, Enallagma 258 - , Indolestes 135 - , Pseudagrion 342 Risiocnemis 385 Risiolestes 140 Risioneura 429 rita, Argia 203 robinsoni, Tatocnemis 170 robusta, Drepanosticta 410 - , Mesocnemis 383 - , Philoganga 6 - , Protosticta 415 robustior, Isosticta 364

493 robustum, Enallagma 258 robustus, Nesolestes 155 rogersi, Argia 206 rollinati, Mnesarete 44 romani, Heteragrion 152 romulus, Parissolestes 174 Roppaneura 455 rosea, Hetaerina 40 - , Metaleptobasis 290 rosea rosea, Nososticta 435 - crientata, Nososticta 435 roseonotata, Podopteryx 162 rossii, subsp. of nubilipennis, Chlorocnemis 422 rotschildi, Lestes 125 rotundatum, var. of cyathigerum, Enallagma 252 rotundatum rotundatum, Microstigma 460 - exustum, Microstigma 461 rotundipenne, Enallagma 258 rouxi, Argiolestes 144 rua, Enallagma 258 rubella, Agrion 307 rubellocerinum, Ceriagrion 306 rubellum, Agrion 274 rubens, Agrion 268 rubeola, subsp. of rubescens, Agriocnemis 187 ruberrima ruberrima, Agriocnemis 184 - albifrons, Agriocnemis 184 rubescens rubescens, Agriocnemis 186 - intermedia, Agriocnemis 186 - lunulata, Agriocnemis 186 - nigricans, Agriocnemis 186 - obscura, Agriocnemis 187 - rubeola, Agriocnemis 187 rubiae, Ceriagrion 306 rubicauda, Teinobasis 355 ribicaudum, race of allopterum, Anisagrion 296 rubiceps rubiceps, Pseudagrion 342 - ceylonicum, Pseudagrion 342 rubida, Chlorocypha 52 rubidum rubidum, Oxyagrion 281 - rufulum, Oxyagrion 281 rubilio, Ischnura 276 rubricauda, Agriocnemis 184 rubridorsum, subsp. of angolense, Pseudagrion 333 rubrifrons, Acanthagrion 236 rubripes, Risiocnemis 387 - , Copera 390 - , Rhipidolestes 176 rubristigma, Enallagma 259 - , Nesolestes 155 rubriventris, Chlorocypha 53 - , Neoneura 449 rubropictus, Micromerus 73 rubropunctata, Palaiargia 214

rubroviridis, subsp. of sudanicum, Pseudagrion 346 rudis, Hetaerina 40 rufescens, Agrion 268 rufescens, Libellago 61 - , Protolestes 163 ruficollis, Heterophaea 88 Teinobasis 355 rufipedum, Papuagrion 322 rufipes, Coenagriocnemis 188 - , Platycnemis 397 rufithorax, Neoneura 449 - , Teinobasis 355 rufitibia rufitibia, Platycypha 65 - lucalaensis, Platycypha 65 rufocinctum, Pseudagrion 342 rufostigma, Drepanosticta 411 rufostigma rufostigma, Ischnura 276 - annandalei, Ischnura 276 - mildredae, Ischnura 276 - , Protosticta 416 Pseudagrion 343 rufovittata, Ischnura 276 rufum, Leptagrion 310 rufulum, subsp. of rubidum, Oxyagrion 281 runtuni, Protallagma 282 rupamnensis, Hetaerina 41 rupinsulensis, Hetaerina 41 rusingae, f. of pseudmassaicum, Pseudagrion 346 rutilans rutilans, Chalcopteryx 93 - confluens, Chalcopteryx 93 ryukyuanum, subsp. of latericium, Ceriagrion 304 ryukyuensis ryukyuensis, Coeliccia 378 - amamii, Coeliccia 378 saalasi, Argia 204 sagitta, subsp. of tincta, Rhinocypha 72 sagittiferum, Austroallagma 246 saharensis, subsp. of elegans, Ischnura 269 sakejii, Ceriagrion 306 salicoti, Andinagrion 244 salisburyense, Pseudagrion 343 salisburyense f. chongwe, Pseudagrion 343 salomonensis, Lieftinckia 382 salomonis salomonis, Nososticta 435 - eburnea, Nososticta 435 salomonis geminata, Nososticta 435 salomonis, Agriocnemis 184 saltator, Argiolestes 145 saltum, Mnesarete 44 salutarius, Argiolestes 145 salva, Telebasis 361 samarcandica, Calopteryx 14

494 samaritis, Teinobasis 356 samfyae, subsp. of coeleste, Pseudagrion 328 samoense, Pseudagrion 343 sancta, Psaironeura 454 sanguinalis, Telebasis 361 sanguinea, Hetaerina 40 sanguineum, Agrion 231 sanguinipes, Heteropodagrion 153 - , Platycnemis 397 Psaironeura 454 sanguinolenta, Hetaerina 40 Rhinocypha 70 sanguinostigma, Ceriagrion 307 - , Ischnura 277 Protostigma 416 sania, Agriocnemis 184 santosi, Oxyagrion 281 saphirina, Umma 30 Sapho 27 sapphirina, Enallagma 259 sarasini, Caledopteryx 147 saskii, subsp. of foliacea, Platycnemis 394 safeties, Agrion 312 saucium, Amphiagrion 243 sauteri, Agrion 218 saundersii, Thore 99 scalaris, Lestes 120 schmidti, Chlorocypha 53 - , Enallagma 259 - , Philogenia 160 - , Schmidtiphaea 89 schmidtianum, Pseudagrion 343 Schmidtiphaea 89 schoana, Ischnura 260 schreiben, Neoneura 449 schultzei, Enallagma 255 scintilla, Protoneura 453 scintillans, Chalcopteryx 93 Teinobasis 356 Sciotropis 164 scitulum, Coenagrion 226 Sclerocypha 58 scutellum scutellum, Coeliccia 378 - hainanensis, Coeliccia 378 seabrai, Chalcopteryx 94 secundaris, Metacnemis 384 secundum, Dimeragrion 148 sedula, Argia 206 selenion, Mortonagrion 190 sellata, Lestes 125 selysi, Chlorocypha 53 - , Metaleptobasis 290 - , Nehalennia 230 Nesobasis 319 - , Nososticta 436

- , Orolestes 127 Podopteryx 162 - , Rhinocypha 70 - , Synlestes 105 Selysioneura 365 semicirculare, Enallagma 259 semicolon, Pseudagrion 329 semlikiensis, subsp. of nigripes, Chlorocnemis 421 semilimbata, Dyspaea 82 semiopaca, Cora 96 - , Libellago 61 - , Mnais 22 semitincta, subsp. of tincta, Rhinocypha 72 semperi, Euphaea 86 sempronia, Hetaerina 40 senegalensis, Ischnura 277 septentrionalis, Hetaerina 40 septima, Drepanosticta 411 serapica, subsp. of vittata, Copera 392 Serena serena, Teinobasis 356 - humeralis, Teinobasis 356 serracipoensis, Telagrion 317 serrata, Risiocnemis 387 serrulatum, Pseudagrion 343 serva, Argia 206 setigerum, Megapodagrion 168 sexlineatum, Cercion 218 Seychellibasis 348 seydeli, Chlorocypha 53 seyrigi, Pseudagrion 343 sharpae, Chlorocypha 53 sharpi, Drepanostica 411 siamense, Pseudagrion 344 sidonicus, Argiolestes 145 siebersi, Drepanosticta 411 sieboldi, Cercion 218 sigma, Lestes 120 signatum, Enallagma 259 signatus, Micromerus 60 sikassoensis, Platycnemis 397 sikkimus, subsp. of praemorsus, Lestes 119 sikorae, Pseudagrion 255 silaceum, Pseudagrion 344 silenta, Trichocnemis 375 silenus, Drepanosticta 411 silvarum, Heteragrion 152 - , Neosticta 365 Philogenia 160 silvaticus, Lestes 120 silviae, Argentagrion 246 silvicola, Nososticta 436 sincerus, subsp. of jocaste, Mecistogaster 457 sinense, Ceriagrion 307 singularis, Mesochemis 383

495 sinhalensis, Drepanosticta 411 Sinhalestes 128 Sinolestes 104 sinuatipennis, Arrhenocnemis 166 - , Tatocnemis 170 sinuatum, Enallagma 259 sinuatum f. fugax, Enallagma 259 simba, Disparoneura 426 simalurum, subsp. of pilidorsum, Pseudagrion 341 simile, race of civile, Enallagma 250 simile, Mortonagrion 190 - , Oxyagrion 281 - , Pseudagrion 344 simillima, Coeliccia 378 simillimum, Coenagrion 226 simmondsi, Melanesobasis 317 simonae, Pseudagrion 344 Tanymecocsticta 368 simonei, Protolestes 163 simplex, Argiolestes 146 - , Hetaerina 41 - , Isosticta 364 - , Lestes 120 simplicinervis, Protosticta 416 simulacrum, Helveciagrion 262 simulans, Lestes 120 simulatrix, Hylaeargia 210 - , Lestes 121 simulatum, Heteragrion 152 siqueirai, Leptagrion 310 sita, Prodasineura 441 sjoestedti, subsp. of anomalum, Microstigma 460 - sjoestedti, Pseudagrion 344 - jacksoni, Pseudagrion 344 - nigeriense, Pseudagrion 344 - pseudosjoestedti, Pseudagrion 344 - wittei, Pseudagrion 345 skaloni, morpha of uncatus, Lestes 113 Skiallagma 283 skinneri, Cora 96 smaragdina, Calopteryx 10 smaragdina, Mnesarete 44 - smaragdina, Vestalis 33 - velata, Vestalis 33 smaragdulus, Lestes 121 smedleyi, Amphicnemis 295 smithiana, Argia 206 snellemanni snellemanni, Melanocypha 62 - javana, Melanocphya 63 sobrina, Ischnura 277 sobrinum, Xanthocnemis 363 solida, Nososticta 436 solitaria, Argiocnemis 187

- , Nososticta 436 Solomocnemis 387 solutus, Perilestes 173 somalicum, Enallagma 260 somalicus, Lestes 121 sooretamae, Metaleptobasis 358 sophia, Coenagrion 226 sophia Selys, 1840, Agrion 230 - Selys, 1876, Agrion 230 sordida, Argia 206 soror, Austrosticta 363 souteri, Elattoneura 427 spathulifera, Drepanosticta 411 spatulae, subsp. of angolensis, Agriocnemis 179 spatulata, Epipleoneura 445 spatulus, Lestes 121 speciosa, Nehalennia 230 speciosum, Heteragrion 152 spectrum, Agrion 121 spegazzinii, Argia 206 spencei, Pseudagrion 345 spernatum spernatum, Pseudagrion 345 - gerstaeckeri, Pseudagrion 345 - natalense, Pseudagrion 345 spinicauda, Ischnura 277 spinicaudum, Papuagrion 322 spinifer, Rhinocypha 70 spinipes, Isosticta 364 spinithoracicum, Pseudagrion 345 splendens splendens, Calopteryx 14 - amasina, Calopteryx 15 splendens race ancilla, Calopteryx 15 splendens balcanica, Calopteryx 15 - caprai, Calopteryx 15 - cartvelica, Calopteryx 15 splendens var. cretensis, Calopteryx 15 splendens erevanensis, Calopteryx 15 splendens var. faivrei, Calopteryx 15 splendens johanseni, Calopteryx 15 - mingrelica, Calopteryx 16 - pfeifferi, Calopteryx 16 splendens, Euphae 87 splendida, Austrocnemis 187 splendida, Umma 29 splendidissima., Agriocnemis 185 sponsa, Lestes 121 sponsus, Agriolestes 146 spumarius, Lestes 122 spuria, Aristocypha 47 stahli, Pentaphlebia 5 starmuehlneri, Drepanosticta 412 starreanum, Pseudagrion 345 steeleae steeleae, Aciagrion 242 - abercornensis, Aciagrion 242 stellata, Palaiargia 214

496 stellatum, Pseudagrion 326 - f. major, Pseudagrion 326 Stenagrion 349 Stenobasis, subgen. of Telebasis 296 Stenocnemis 387 Stenocora 92 stenomantis, Selysioneura 367 Stenosticta 367 Stephanella, f. of pumilio, Ischnura 274 stephanodera, Asthenocnemis 369 sternalis, Lestes 122 stevensi, subsp. of annulata, Copera 389 stevensi, Protosticta 414 stictica, Libellago 62 Pericnemis 322 stigmatica, Argia 206 stigmaticus, Lestes 122 stillacruoris, Paramecocnemis 385 stigmatizans, Libellago 62 - , Teinobasis 356 straeleni, Chlorocypha 54 - bwamba, Chlorocypha 54 striata, var. of pumilio, Ischnura 275 striata, subsp. of paedisca, Sympecma 137 striatipes, Psilocnemis 390 striatus, Lestes 122 strigata, Mnais 22 strigosa, Teinobasis 356 strohmi, Acanthallagma 238 strouhali, Enallagma 260 strumidens, Idiocnemis 380 stuckenbergi, Pseudagrion 346 stueberi, Ischnura 277 - , Papuargia 214 stultus, Lestes 122 stygia, Rhinocypha 71 stygium, Mortonagrion 190 suave, Ceriagrion 307 suavis, Teinobasis 356 subaequistyla, Copera 388 subapicalis, Argia 206 - , Miocora 92 subannulata, Psilocnemis 389 subcostalis, Euphaea 87 subdilatata, Platycnemis 398 subfumata, Cora 96 subfurcatum, Enallagma 260 subgriseus, subsp. of griseus, Argiolestes 142 subhumeralis, Nesobasis 319 sublacteum, Pseudagrion 346 sublatus, Lestes 122 sublimata, var. of occisa, Hetaerina 38 submontana, Drepanosticta 412 subsp. of apicalis, Vestaiis 32 subnodalis, Euphaea 87

Isomecocnemis 429 subornatus, subsp. of ornatus, Argiolestes 144 subplatystyla, Anisopleura 79 subtile, Enallagma 260 subtropica, Drepanosticta 412 sucra, Philogenia 160 sudanicum, Enallagma 261 sudanicum sudanicum, Pseudagrion 346 - rubroviridis, Pseudagrion 346 sufficiens, subsp. of zebrina, Idiocnemis 381 - , Rhyacocnemis 385 sulcatum, Enallagma 261 sulinum, Qxyagrion 282 sumbana, Rhinocypha 71 sumatrana, Argiocnemis 187 sumatrana, Libellago 62 Sundacypha 73 sundana, Drepanosticta 412 superba, subsp. of marshalli, Chlorocnemis 421 - , Euphaea 87 superba, Sapho 27 superba, Teinobasis 356 superbum, Heteropodagrion 153 suberbum, Pseudagrion 343 Super lestes 108 superplatypes, Copera 390 suphuripes, Libellago 64 suensoni, Megalestes 107 sushmae, Archibasis 132 sutteri, Indolestes 135 svenhedini, Indocypha 58 svihleri, Coeliccia 378 sylvatica, Neoneura 449 Sylphis 9 symoensis, Pseudagrion 346 Sympecmatinae 128 Sympecma 135 Sympycna, subgen. of Agrion 135 Synlestidae 101 Synlestinae 101 Synlestes 105 syriaca, Calopteryx 16 syriaca, Platycnemis 393 syriacum, subsp. of puella, Coenagrion 225 - , Pseudagrion 346 taipokauensis, Protosticta 416 taitensis, Ischnura 277 talamanca, Argia 207 tamoyo, Argia 207 Tanymecocsticta 367 tanysiptera, Palaiargia 214 Taolestes 175 taracumbi, Nososticta 436 tarascana, Argia 207

497 tarry i, Lestes 118 tasmanica, var. of heterosticta, Ischnura 271 Tatocnemididae 166 Tatocnemis 169 taurica taurica, Calopteryx 16 - tshaldirica, Calopteryx 16 taurus, Drepanosticta 412 taxaense, Acanthagrion 236 t-coerulea, Indoneura 418 Teinobasis 349 Telagrion 357 Teleallagma 248 Telebasis Selys 1865, 358 Telebasis Selys 1877, 349 telegastrum, Nesobasis 319 temporale, Acanthagrion 236 - , Megapodagrion 168 temporalis, Lestes 123 tenax, Elattoneura 428 tenax, Leptobasis 358 tendipes, Risiocnemis 387 tenebricosa, Prodasineura 442 tenella, Archibasis 298 - , Drepanosticta 412 tenellum tenellum, Ceriagrion 307 - nielseni, Ceriagrion 307 tenuatus, Lestes 123 tenuis, race of griseus, Argiolestes 142 tenuis, Chlorocypha 53 - , Mnais 23 - , Protoneura 453 - , Teinobasis 357 tenuispinus, Argiolestes 146 tenussima, Psaironeura 454 tenuissimus, Indolestes 135 terira, Argia 207 terminale, Oxyagrion 282 terminalis race bogotensis, Cora 95 terminalis, Euthore 91 terminata, race of derivata, Polythore 98 terminata, Rhinocypha 71 terminatum, Anonisma 455 ternaria, Argia 207 terraba Philogenia 160 terue, Calopteryx 16 tessellatus, Chlorolestes 103 tetragena, Metaleptobasis 290 Tetraneura 4 tetrica, Elattoneura 428 texana, Hetaerina 35 tezpi, Argia 207 thailandica, Caliphaea 8 thalassina, Nososticta 436 thalia, Selysioneura 367 Thaumatagrion 388

Thaumatoneura 171 Thaumatoneurinae 171 theebawi, Prodasineura 442 thelmae, Ischnura 278 thenartum, Pseudagrion 347 theodori, Argia 207 Telebasis 361 Thermagrion 248 theryi, subsp. of caerulescens, Coenagrion 220 thespis, Argia 207 thisma, Argia 208 thoracicus, Lestes 123 subsp. of earnshawi, Mnais 21 Thore 96 tibetanum, Mesopodagrion 169 tibialis, Argia 208 Tigriagrion 283 tikalus, Lestes 123 tillyardi, Aciagrion 242 - , subsp. of lestoides, Diphlebia 77 - , Isosticta 364 tillyardi, var. of adytum, Megalagrion 311 tillyardi, Synlestes 105 timorana, subsp. of pagenstecheri, Rhinocypha 69 tincta, Polythore 99 tincta tincta, Rhinocypha 71 - adusta, Rhinocypha 71 - amanda, Rhinocypha 72 - dentiplaga, Rhinocypha 72 - retrograda, Rhinocypha 72 - sagitta, Rhinocypha 72 - semitincta, Rhinocypha 72 tinctipennis, Argia 208 - , Pseudagrion 347 - , Pyrrhosoma 232 tindalei, Austrolestes 131 tinlandica, Philogenia 160 tiomanensis, subsp. of argyroides, Devadatta 4 titia, Hetaerina 41 titicacae, Protallagma 283 tokyoensis, Copera 390 tolteca, Hetaerina 41 tonsillare, Aciagrion 243 tonto, Agria 208 Torrenticnemis 388 torrenticola, Caconeura 429 torresiana, Ischnura 272 torridum torridum, Pseudagrion 347 - hulae, Pseudagrion 347 - f. orientis, Pseudagrion 347 tractipennis, Argia 208 transcaspica, Calopteryx 16 translata, Argia 208

498 transvaalica, subsp. of falcifera, Agriocnemis 180 traviatum traviatum, Enallagma 261 - westfalli, Enallagma 261 trebbaui, Iridictyon 19 triangulare, Heteragrion 152 triangularis, Periocnemis 323 tricellulare, Heteragrion 152 Trichocnemis 371 Tricholestes 131 tricolor, Calopteryx 41 tricolor, Euphaea 87 Lestes 123 - , Rhinagrion 164 tricornis, Pseudagrion 347 tricrenaticeps, Ceriagrion 307 tridens, Lestes 123 trifaria Karsch, Libellago 53 trifaria Schoudeten, Libellago 52 trifasciata, Aristocypha 47 trifoliata, Argia 209 trigonale, Pseudagrion 347 trilineatum, Coenagrion 218 trilobata, Agriocnemis 185 - , Protosticta 416 trilobatum, Acanthagrion 237 trimaculata, Drepanosticta 412 - , Rhinocypha 72 trimaculatum, Cyanallagma 248 trina, Argia 209 trinervis, Neuragrion 156 Trineuragrion 164 trinitatis, Hypolestes 166 tripartita, subsp. of margarita, Echo 18 tristis, Argiolestes 146 tristis, Vestalis 11 tropica, Drepanosticta 412 tropicalis, Elattoneura 428 tropicanum, subsp. of hageni, Pseudagrion 333 tropicus, Synlestes 105 trotteri, Copera 391 truncatum, Acanthagrion 237 truncatidens, Rhipidolestes 176 truncatipennis, Anisagrion 296 truncatum, Enallagma 261 Xiphiagrion 284 truncatus, Sinolestes 104 tshaldirica, subsp. of taurica, Calopteryx 16 tsiamae, Elattoneura 428 tuanuii, Xanthocnemis 363 tuberculoid, Leptobasis 287 tuberculatum Selys, Agrion 275 - Charpentier, Agrion 268 tuberculatus, Archilestes 109 tuberculicornis, Nesolestes 155

tupi, Argia 209 turconii, Rhinocypha 73 udeana, Lestes 127 uenoi, Rhinocypha 73 ulmeca, Argia 209 ulmifolia, Platycnemis 397 ultima, Ischnura 278 umbargae, Argiocnemis 187 umbra, Dicterias 75 umbratilis, Selysioneura 167 umbratus, Chlorolestes 103 umbriaca, Argia 209 umbrinus, Lestes 124 umbrosa, Philogenia 160 Umma 28 umsingaziense, Pseudagrion 347 uncata, Protosticta 416 meatus, Lestes 113 meatus morpha skaloni, Lestes 113 uncifer, Lestes 124 underwoodi, Argia 209 unicolor, Calopteryx 17 - , Cyrano 54 - , subsp. of ochraceus, Lestes 117 uniformis, Echo 19 unimaculata, Paracypha 63 uniseries, Caledargiolestes 147 undulatus, Lestes 124 unguiculatus, Lestes 124 ungulatum, Pseudagrion 348 urnifera, Ischnura 260 urubamba, Lestes 124 ussurense, subsp. of hylas, Coenagrion 222 ustulata, Libellago 73 ustum, Pseudagrion 348 utahensis, Ischnura 265 uxor, Micromerus 60 vaalense, Pseudagrion 348 vacca, Coeliccia 378 vacillans vacillans, Leptobasis 287 - atrodorsum, Leptobasis 287 vacillans race ecuadorica, Leptobasis 287 - race guinae, Leptobasis 287 vagabundum, Megalagrion 315 vaginale vaginale, Enallagma 261 - longfieldi, Enallagma 261 vakoanae, Pseudagrion 348 valga, Cyanocharis 74 valida, Metacnemis 384 vallei, Anisopleura 79 vallisi, Oristicta 365 vansomereni, Enallagma 261 - , Pseudagrion 348

499 variata, Argia 209 variabilis, Argia 196 varians, Ceriagrion 307 variegata, Argia 209 variegata variegata, Euphae 87 - intermedia, Euphea 87 varralli, Mortonagrion 190 velaris, Agriocnemis 184 velata, subsp. of smaragdina, Vestalis 33 venale, Megapodagrion 168 vereninotata, Agrion 245 venezuelensis, Epipleoneura 445 venilia, Selysioneura 367 venusta, Sapho 28 vernale, Coenagrion 227 - , Enallagma 262 veronica, Indolestes 132 versicolor, Protosticta 416 - , Telebasis 362 verticalis, Ischnura 278 verticalis verticalis, Prodasineura 442 - andamensis, Prodasineura 442 - annandalei, Prodasineura 442 - burmanensis. Prodasineura 442 - delia, Prodasineura 442 - humeralis, Prodasineura 443 vesperum, Enallagma 262 vesta, Agrion 17 Vestalaria 30 Vestalinae 8 Vestalis 30 vestalis, subsp. of virens, Lestes 125 Vestinus 30 vetusta, Philoganga 6 vicentius, subsp. of jocaste, Mecistogaster 457 vicinum, subsp. of anomalum, Microstigma 460 victoria, Agriocnemis 185 - , Polythore 100 victoriae, Chlorocypha 54 viduata, subsp. of aurora, Ischnura 265 viduum, Acanthagrion 237 viduus, Lestes 124 vigilax, Lestes 125 villiersi, Ceriagrion 300 villiersi, Prodasineura 443 villosipes, Rhinoneura 73 vinsoni, Ischnura 278 viola, Archibasis 298 violacea, Argia 210 violacea, Coenagrion 221 virens virens, Lestes 125 - marikovskii, Lestes 125 - vestalis, Lestes 125 virens, Vestalis 34 virgatus, Lestes 125

virgatus, Mecistogaster 456 virginica, Calopteryx 10 virgo virgo, Calopteryx 17 - britannica, Calopteryx 17 - festica, Calopteryx 17 virgo ab.fusca, Calopteryx 17 virgo meridionalis, Calopteryx 17 virgula, Selysioneura 367 viridatum, Rhinagrion 164 viridescens, Acanthagrion 237 viridis viridis, Lestes 126 - parvidens, Lestes 126 - , Drepanosticta 413 - , Protoneura 454 viridissima, Pseudophaea 87 viridivittatum, Agrion 124 viridulum, Erythromma 228 viridulum f. latistrigii, Erythromma 228 - Orientale, Erythromma 228 viridulus, Lestes 126 vitiensis, Agriocnemis 185 - , Indolestes 135 vitrella, Rhinocypha 56 vitrinella, Heterocypha 58 vittata vittata, Copera 391 - acutimargo, Copera 391 - assamensis, Copera 391 - deccanensis, Copera 391 - javana, Copera 391 - palawana, Copera 392 - serapica, Copera 392 Indocypha 58 vittata, Lestes 124 vittata, Polythore 100 Prodasineura 443 vittigera, Onychargia 211 vivida, Argia 210 v-nigrum, Cercion 218 vulcanoae, Helveciagrion 262 vumbaense, Pseudagrion 348 vridaghi, Elattoneura 428 vriesianum, Leptagrion 311 vulnerata, Hetaerina 41 - , Telebasis 362 wahlbergi, Lestes 126 Wahnesia 139 wainaeanum, subsp. of amaurodytum, Megalagrion 311 walkeri, Enallagma 226 wallacei, Amphicnemis 295 - , Nososticta 436 - , Orolestes 127 - , Teinobasis 357 walli, Drepanosticta 413

500 Dysphaea 82 waltheri, Minagrion 317 Neoneura 449 waterstoni, Calopteryx 17 watsoni, Austroagrion 216 webbianum, Enallagma 262 weewa, Enallagma 262 weibezahni, Metaleptobasis 290 werneri, Coeliccia 378 westermanni, Phylloneura 419 westfalli, subsp. of traviatum, Enallagma 261 Hetaerina 41 Metaleptobasis 290 weyseri, Synlestes 105 wheeleri, race of fontinalis, Drepanosticta 407 whellani, Ceriagrion 308 whellani, Pseudagrion 334 whiteheadi, Rhinocypha 57 williamsi, Pseudagrion 348 williamsoni, Acanthagrion 237 williamsoni, Disparoneura 421 williamsoni, Epipleoneura 445 - , Megalagrion 315 Oxystigma 156 - , Polythore 100 Pseudagrion 348 willinki, Telebasis 362 wilsoni, Argia 210 wittei, Chlorocnemis 422 Chlorocypha 54 - , subsp. of sjoestedti, Pseudagrion 345 woytkowskii, Protoneura 454 wynaadensis, Indophaea 89 Xanthagrion 362 xanthe, Nososticta 436 Rhinocypha 73 Xanthocnemis 362

xanthocyana, Libellago 62 xanthocyane, Oreagrion 279 xanthomelaena, Caconeura 434 xanthomelas, Megalagrion 315 xanthopterum, Agrion 274 xanthopus, Platycnemis 398 Xanthostigma 456 xanthostoma, Calopteryx 17 - , Cora 96 Xerolestes 109 Xiphiagrion 284

yachowensis, Mesopodagrion 169 yakima, subsp. of aequabilis, Calopteryx 10 yakusimensis, subsp. of aculeatus, Rhipidolestes 175 yayeyamana, Euphaea 88 yezoense, subsp. of deserti, Enallagma 253 yoshikoae, Noguchiphaea 8 ypsilon, Enallagma 251 yungarum, Acanthagrion 237

zambiense, Aciagrion 243 zambesiensis, subsp. of glaucescens, Pseudagrion 333 zebrina zebrina, Idiocnemis 381 - sufficiens, Idiocnemsis 381 zelandicum, Xanthocnemis 381 zelandicum race antipodum, Xanthagrion 362 zerafica, Agriocnemis 185 zernyi, subsp. of lindeni, Cercion 217 zeteki, Philogenia 160 zoe, Paraphlebia 157 Zoniagrion 284 zumbense, Pseudagrion 332

As of Part 102 Das Tierreich is published in hard cover. Since 1958 the following have appeared Contribution 72

Author

Title

Year

No. of pages

H.Wermuth

REPTILIA: Helodermalidae

1958

16 pages, 9 figures

73

M. Beier

ORTHOPTERA: Tettigoniidae (Pseudophyllinae I)

1962

XII, 468 pages, 245 figures

74

M. Beier

ORTHOPTERA: Tettignoniidae (Pseudophyllinae II)

1960

396 pages, 241 figures

75

K.Odening

TREMATODA (Digenea): Plagiorchiidae III (Haematoloechinae) und Omphalometridae

1960

IV, 77 pages 60 figures

76

K. Sanft

AVES (Upupae): Bucerotidae

1960

IV, 176 pages 106 figures

77

R. Kilias

GASTROPODA

1962

63 pages, 47 figures

PROSOBRANCHIA:

Tonnidae 78

St. W. Gorham

GYMNOPHIONA, Liste der rezenten Amphibien und Reptilien

1962

X, 25 pages

79

R. Mertens

HELODERMATIDAE, VARANIDAE, LANTHANOTIDAE. Liste der rezenten Amphibien und Reptilien

1963

X, 26 pages

80

Η. Wermuth

GEKKONIDAE, PYGOPODIAE, XANTUSIIDAE. Liste der rezenten Amphibien und Reptilien

1965

XXII, 246 pages

81

J.A.Peters

COLUBRIDAE (Dipsadinae). Liste der rezenten Amphibien und Reptilien

1965

VIII, 18 pages

82

J. lilies

PLECOPTERA, Katalog

1966

XXX, 632 pages, 20 figures

83

R. Mertens

CHAMAELEONIDAE. Liste der rezenten Amphibien und Reptilien

1966

X, 37 pages

84

C. Gans

UROPELTIDAE. Liste der rezenten Amphibien und Reptilien

1966

29 pages

85

St. W. Gorham

ASCAPHIDAE, LEIOPELMATIDAE, PIPIDAE, DISCOGLOSSIDAE, PELOBATIDAE, LEPTODACTYLIDAE, RHINOPHRYNIDAE. Liste der rezenten Amphibien und Reptilien

1966

XVI, 222 pages

86

H.Wermuth

AGAMIDAE. Liste der rezenten Amphibien und Reptilien

1967

XIV, 127 pages

87

H.Wermuth

CORDYLIDAE (Cordylinae + Gerrhosaurinae). Liste der rezenten Amphibien und Reptilien

1968

X, 30 pages

88

F. Haas

BIVALVIA: Unionacea

1969

X, 663 pages, 5 figures

Contri- Author bution

Title

Year

No. of pages

89

A. F. Stimson

BOIDAE, (Boinae + Bolyeriinae + Loxoceminae + Pythoninae). Liste der rezenten Amphibien und Reptilien

1969

XI, 49 pages

90

H. Wermuth

ANGUIDAE, ANNIELLIDAE, XENOSAURIDAE. Liste der rezenten Amphibien und Reptilien

1969

XII, 41 pages

91

M. Fischer

HYMENOPTERA: Braconidae (Opiinae I)

1972

XII, 620 pages, 463 figures

92

R. Kilias

GASTROPODA PROSOBRANCHIA: Cymatiidae

1973

VIII, 235 pages, 149 figures

93

P. Banarescu & T.T. Nalbant

PISCES, TELEOSTEI: Cyprinidae (Gobioninae)

1973

VII, 304 pages, 154 figures, 19 maps

94

P. Zwick

PLECOPTERA, Phylogenetisches System und Katalog

1973

XXXII, 465 pages, 75 figures

95

E. Duellmann

HYLIDAE, CENTROLENIDAE, PSEUDIDAE. Liste der rezenten Amphibien und Reptilien

1976

XIX, 225 pages

96

M. Fischer

HYMENOPTERA: Braconidae (Optiinae Ii/Amerika)

1976

1038 pages, 890 figures

97

S.G. Kiriakoff

LEPIDOPTERA NOCTUIFORMES: Agaristidae I

1976

IX, 180 pages, 39 figures

98

S.G.Kiriakoff

LEPIDOPTERA NOCTUIFORMES: Agaristidae II

1976

VIII, 165 pages, 52 figures

99

S.G.Kiriakoff

LEPIDOPTERA NOCTUIFORMES: Agaristidae III

1976

VI, 86 pages 26 figures

100

H. Wermuth & tR.Mertens

TESTUDINES, CROCODYLIA, RHYNCHOCEPHALIA. Liste der rezenten Amphibien und Reptilien

1976

XXVII, 174 pages

101

D.E.Hahn

ANOMALEPIDIDAE, LEPTOTYPHLOPIDAE, TYPHLOPIDAE. Liste der rezenten Amphibien und Reptilien

1980

XII, 93 pages

102

H. Steinmann

DERMAPTERA Catadermaptera I

1986

XIV, 343 pages

103

A. Kaltenbach

SAGIN AE Saltatoria- Tettigoniidae

1986

VIII, 92 pages

104

M. Fischer

HYMENOPTERA Optiinae III

1987

XV, 734 pages

105

H. Steinmann

DERMAPTERA Catadermaptera II

1989

XIX, 504 pages

Contribution

Author

106

H. Steinmann

107

R.L. Hoffman

108

H. Steinmann

109

Title

DERMAPTERA Eudermaptera I MYRIAPODA 4 Polydesmida: Oxydesmidae

Year

No. of pages

1989

XX, 558 pages

1990

XVI, 512 pages

DERMAPTERA Eudermaptera II

1993

XXII, 711 pages

A.M. Bauer

GEKKONIDAE (Reptilia Sauria)

1994 XIV, 306 pages

110

H. Steinmann

1997 XXII, 500 pages

111

H. Steinmann

World Catalogue of Odonata Volume I Zygoptera World Catalogue of Odonata Volume II Anisoptera

1997

XIV, 636 pages